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<document id="3234A306FF290BFB272E4661231CFC2E" ID-DOI="10.1080/00222933.2019.1606953" ID-GBIF-Dataset="d2615e10-8703-400b-b571-6c431b0ed9a7" ID-ISSN="1464-5262" ID-Zenodo-Dep="3675571" ID-ZooBank="09AA5C86-5138-42A7-9646-9FF2A7F46767" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582134181895" checkinUser="valdenar" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="9F138799E554B06E9BCB5A14D3EBFCA8" docLanguage="en" docName="JNATHIST.53.1112.725747.pdf.imf" docOrigin="Journal of Natural History 53 (11)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Aegidinus Arrow 1904" docType="treatment" docVersion="12" lastPageNumber="727" masterDocId="632AFFE1E556B06D9A1D5E52D024FFC2" masterDocTitle="Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature" masterLastPageNumber="747" masterPageNumber="725" pageNumber="726" updateTime="1698719875536" updateUser="plazi">
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<mods:title id="8832E5F72E44AD14065A0FA7DDD9F3F9">Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature</mods:title>
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<mods:namePart id="E2B496CB968FD8D8EC2FCF61FCF1198A">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="D86B6D3C156F60B7BAE22781081E5559">Akhmetova, Lilia A.</mods:namePart>
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<mods:namePart id="B9F144E7B73A375CE673397A6CB6D6E5">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="67F2CF05C0F59AB36741359E1DBC01F9">Journal of Natural History</mods:title>
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<paragraph id="1705368FE554B06F9BCB5A14D368FBA2" blockId="2.[470,844,1094,1120]" box="[470,844,1094,1120]" pageId="2" pageNumber="726">
<heading id="4C4D81E3E554B06F9BCB5A14D368FBA2" box="[470,844,1094,1120]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="726" reason="2">
Genus
<taxonomicName id="D0BA4D0CE554B06F98335A14D368FBA2" authority="Arrow, 1904" authorityName="Arrow" authorityYear="1904" box="[558,844,1094,1120]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="726" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE554B06F98335A14D288FBA2" bold="true" box="[558,684,1094,1120]" italics="true" pageId="2" pageNumber="726">Aegidinus</emphasis>
Arrow, 1904
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<subSubSection id="5FA06504E554B06F9ABD5A29D345FB74" pageId="2" pageNumber="726" type="materials_examined">
<paragraph id="1705368FE554B06F9ABD5A29D102FB51" blockId="2.[160,865,1147,1206]" box="[160,294,1147,1171]" pageId="2" pageNumber="726">
<emphasis id="25CEEA9DE554B06F9ABD5A29D102FB51" box="[160,294,1147,1171]" italics="true" pageId="2" pageNumber="726">
<typeStatus id="C801882DE554B06F9ABD5A29D0F6FB51" box="[160,210,1147,1171]" pageId="2" pageNumber="726">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="1705368FE554B06F9ABD5ACCD345FB74" blockId="2.[160,865,1147,1206]" box="[160,865,1182,1206]" pageId="2" pageNumber="726">
<taxonomicName id="D0BA4D0CE554B06F9ABD5ACCD22CFB74" authority="Westwood," authorityName="Westwood" authorityYear="1845" box="[160,520,1182,1206]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="726" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="25CEEA9DE554B06F9ABD5ACCD1A7FB74" box="[160,387,1182,1206]" italics="true" pageId="2" pageNumber="726">Aegidium guianensis</emphasis>
Westwood,
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designated by
<bibRefCitation id="732B4B7EE554B06F98A65ACCD379FB74" author="Paulian R" box="[699,861,1182,1206]" pageId="2" pageNumber="726" pagination="65 - 92" refId="ref9518" refString="Paulian R. 1984. Les Orphnidae Americains (Coleopteresi Scarabaeoidea). Annales de la Societe Entomologique de France (N S). 20: 65 - 92." type="journal article" year="1984">Paulian (1984)</bibRefCitation>
.
</paragraph>
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<subSubSection id="5FA06504E554B06E9ABD5AA0D3EBFCA8" lastPageId="3" lastPageNumber="727" pageId="2" pageNumber="726" type="diagnosis">
<paragraph id="1705368FE554B06F9ABD5AA0D130FAC9" blockId="2.[160,1156,1266,1707]" box="[160,276,1266,1291]" pageId="2" pageNumber="726">
<emphasis id="25CEEA9DE554B06F9ABD5AA0D130FAC9" bold="true" box="[160,276,1266,1291]" italics="true" pageId="2" pageNumber="726">Diagnosis</emphasis>
</paragraph>
<paragraph id="1705368FE554B06E9ABD5B44D227FF0B" blockId="2.[160,1156,1266,1707]" lastBlockId="3.[160,1156,142,444]" lastPageId="3" lastPageNumber="727" pageId="2" pageNumber="726">
Body small to mid-sized (length
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), reddish brown to dark brown. Mandibles subsymmetrical, with long processes on the outer sides. Clypeus with tubercle or horn on anterior margin medially in males, without horn in females. Pronotum variably excavated medially in males, convex to depressed medially in females; anterior margin of pronotum in males with a tubercle or horn medially. Propleura with carinae separating anterolateral areas from basal area. Scutellum narrowly rounded apically, about 1/15 length of elytra. Elytra convex, with marked humeral umbones and striae marked with elongated punctures. Wings fully developed. Metepisternon triangular, its posterior angle rounded to triangular and situated in distinct concavity of epipleuron. Protibiae with three strong outer teeth in both sexes and with a smaller medial tooth in majority of males. Each procoxa with two cavities. Phallobase tube shaped with strongly sclerotised ventral side but without differentiation of ventral and dorsal sclerites. Parameres relatively short, with complex shape, apices without setae. Gonocoxites separated into two more or less distinct parts.
</paragraph>
<paragraph id="1705368FE555B06E9ADD5E86D161FE7E" blockId="3.[160,1156,142,444]" pageId="3" pageNumber="727">
<taxonomicName id="D0BA4D0CE555B06E9ADD5E86D10EFF2E" authorityName="Arrow" authorityYear="1904" box="[192,298,212,236]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE555B06E9ADD5E86D10EFF2E" box="[192,298,212,236]" italics="true" pageId="3" pageNumber="727">Aegidinus</emphasis>
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is distinct among the orphnines in having the following unique characters: mandibles with processes on the outer sides; shape of the gonocoxites is diverse and species specific at least in majority of species; procoxae with two cavities each; parameres are asymmetric in two species; phallobase with protruding ventroapical plate in some species. From other Aegidiini,
<taxonomicName id="D0BA4D0CE555B06E98485F0CD29BFEB4" authorityName="Arrow" authorityYear="1904" box="[597,703,350,374]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE555B06E98485F0CD29BFEB4" box="[597,703,350,374]" italics="true" pageId="3" pageNumber="727">Aegidinus</emphasis>
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also differs in the absence of a hole connecting mesocoxal cavities, and in having more or less distinct transverse carina on metatibiae.
</paragraph>
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<emphasis id="25CEEA9DE555B06E9ABD5FBDD23BFDCA" bold="true" box="[160,543,495,520]" italics="true" pageId="3" pageNumber="727">Diagnostic characters of species</emphasis>
</paragraph>
<paragraph id="1705368FE555B06E9ABD5C41D435FD77" blockId="3.[160,1155,495,693]" pageId="3" pageNumber="727">
Most
<taxonomicName id="D0BA4D0CE555B06E9AC25C41D16DFDE9" authorityName="Arrow" authorityYear="1904" box="[223,329,531,555]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE555B06E9AC25C41D16DFDE9" box="[223,329,531,555]" italics="true" pageId="3" pageNumber="727">Aegidinus</emphasis>
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species are similar externally and, despite some interspecific variation of the body sculpture and head and protoracic armature in some species, reliable identification is only possible by examining the shape of the male genitalia. External female genitalia (gonocoxites) are also highly specific, although their usefulness in distinguishing closely related species requires further evaluation (see Discussion below).
</paragraph>
<paragraph id="1705368FE555B06E9ABD5CBBD27EFCC0" blockId="3.[160,1155,745,874]" box="[160,602,745,770]" pageId="3" pageNumber="727">
<emphasis id="25CEEA9DE555B06E9ABD5CBBD27EFCC0" bold="true" box="[160,602,745,770]" italics="true" pageId="3" pageNumber="727">Species composition and distribution</emphasis>
</paragraph>
<paragraph id="1705368FE555B06E9ABD5D5FD3EBFCA8" blockId="3.[160,1155,745,874]" pageId="3" pageNumber="727">
<taxonomicName id="D0BA4D0CE555B06E9ABD5D5FD12EFCE7" authorityName="Arrow" authorityYear="1904" box="[160,266,781,805]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="25CEEA9DE555B06E9ABD5D5FD12EFCE7" box="[160,266,781,805]" italics="true" pageId="3" pageNumber="727">Aegidinus</emphasis>
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comprises 14 species including two newly described in this work. Members of the genus occur in South America, mostly in the Amazon and Guianan moist forest regions to Yungas in the west, and on the island of
<collectingCountry id="6FAD761FE555B06E98EA5D00D377FCA8" box="[759,851,850,874]" name="Trinidad and Tobago" pageId="3" pageNumber="727">Trinidad</collectingCountry>
(
<figureCitation id="8F812A0AE555B06E997F5D00D3E7FCA8" box="[866,963,850,874]" captionStart="Figure 7" captionStartId="18.[160,225,1042,1065]" captionTargetBox="[166,1149,142,1011]" captionTargetId="figure@18.[166,1149,142,1011]" captionTargetPageId="18" captionText="Figure 7. Aegidinus spp. Locality records map." figureDoi="http://doi.org/10.5281/zenodo.3675585" httpUri="https://zenodo.org/record/3675585/files/figure.png" pageId="3" pageNumber="727">Figure 7</figureCitation>
).
</paragraph>
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