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<document id="AC1E523B619522A0D09521AC0AF303AC" ID-DOI="10.5281/zenodo.174510" ID-GBIF-Dataset="d3d75e14-dbe7-4783-bb91-070a4a411e7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="174510" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459783530056" checkinUser="plazi" docAuthor="M, E L E N A V., Va, I K H A L J O &amp; Lim, Kil-Young" docDate="2006" docId="9F06B848FFEE3463FEC4F934C807FD0C" docLanguage="en" docName="zt01350p053.pdf" docOrigin="Zootaxa 1350" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Epanerchodus bacilliferus M, Va &amp; Lim, 2006, sp.n." docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="633FC030FFED3466FFCCFFB0CA4CFF92" masterDocTitle="The millipede genus Epanerchodus Attems, 1901 in the Korean Peninsula, with a description of a new species (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="53" masterPageNumber="45" pageNumber="48" updateTime="1698218205287" updateUser="plazi">
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<mods:title id="1D88519A06D18E3C6F98993295533EBE">The millipede genus Epanerchodus Attems, 1901 in the Korean Peninsula, with a description of a new species (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:namePart id="8B506002739B7668808042B1DCCB020C">M, E L E N A V.</mods:namePart>
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<mods:namePart id="9621F578EED2D7DEC933DB6E6859A735">Va, I K H A L J O</mods:namePart>
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<mods:namePart id="04D5D47CBE6FE3FC538837184C99E7A5">Lim, Kil-Young</mods:namePart>
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<mods:date id="35A039BFCDE216E1957B4BAD34CC4F75">2006</mods:date>
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<treatment id="9F06B848FFEE3463FEC4F934C807FD0C" ID-DOI="http://doi.org/10.5281/zenodo.6255355" ID-GBIF-Taxon="119348623" ID-Zenodo-Dep="6255355" LSID="urn:lsid:plazi:treatment:9F06B848FFEE3463FEC4F934C807FD0C" httpUri="http://treatment.plazi.org/id/9F06B848FFEE3463FEC4F934C807FD0C" lastPageId="5" lastPageNumber="50" pageId="3" pageNumber="48">
<subSubSection id="5FB55AD5FFEE3465FEC4F934CB39F953" pageId="3" pageNumber="48" type="nomenclature">
<paragraph id="1710095EFFEE3465FEC4F934C8C6F90C" blockId="3.[264,650,1668,1729]" box="[264,650,1668,1694]" pageId="3" pageNumber="48">
<heading id="4C58BE32FFEE3465FEC4F934C8C6F90C" bold="true" box="[264,650,1668,1694]" fontSize="11" level="1" pageId="3" pageNumber="48" reason="1">
<emphasis id="25DBD54CFFEE3465FEC4F934C8C6F90C" bold="true" box="[264,650,1668,1694]" pageId="3" pageNumber="48">
<taxonomicName id="D0AF72DDFFEE3465FEC4F934C804F90C" ID-CoL="6FMS6" box="[264,584,1668,1694]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="bacilliferus" status="sp. nov.">
<emphasis id="25DBD54CFFEE3465FEC4F934C804F90C" bold="true" box="[264,584,1668,1694]" italics="true" pageId="3" pageNumber="48">Epanerchodus bacilliferus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EE86837FFEE3465FD83F934C8C6F90C" box="[591,650,1668,1694]" pageId="3" pageNumber="48" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="1710095EFFEE3465FEC4F917CB39F953" blockId="3.[264,650,1668,1729]" box="[264,373,1703,1729]" pageId="3" pageNumber="48">
<figureCitation id="8F9415DBFFEE3465FEC4F917CB3DF953" box="[264,369,1703,1729]" captionStart="FIGURES 1 3" captionStartId="5.[264,382,1780,1804]" captionTargetBox="[292,1281,741,1749]" captionTargetId="figure@5.[263,1324,695,1772]" captionTargetPageId="5" captionText="FIGURES 1 3. Epanerchodus bacilliferus sp. n. (male holotype). 1, distal part of antenna; 2, gonopod (lateral view); 3, distal part of gonopod (mesal view). Scales in mm." httpUri="https://zenodo.org/record/174511/files/figure.png" pageId="3" pageNumber="48">Figs 13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5FB55AD5FFEE3465FEC4F947C89CF8AB" pageId="3" pageNumber="48" type="materials_examined">
<paragraph id="1710095EFFEE3465FEC4F947C89CF8AB" blockId="3.[264,1323,1783,1889]" pageId="3" pageNumber="48">
<emphasis id="25DBD54CFFEE3465FEC4F947C824F882" box="[264,616,1783,1808]" italics="true" pageId="3" pageNumber="48">
Material examined:
<typeStatus id="C814B7FCFFEE3465FE31F947C824F882" box="[509,616,1783,1808]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="01A9C2D7FFEE3465FDB0F947C881F883" box="[636,717,1783,1809]" pageId="3" pageNumber="48" type="male">1 male</specimenCount>
(ChNU) from Jinan, Jeonllabuk-do,
<collectingCountry id="6FB849CEFFEE3465FB41F947CF6BF883" box="[1165,1319,1783,1809]" name="South Korea" pageId="3" pageNumber="48">South Korea</collectingCountry>
, collected
<date id="63112F9EFFEE3465FEB5F8AFC86AF8AB" box="[377,550,1823,1849]" pageId="3" pageNumber="48" value="1993-08-07">7 August 1993</date>
by K.-Y. Lim.
</paragraph>
</subSubSection>
<subSubSection id="5FB55AD5FFEE3462FEF4F8F7CF50FE44" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48" type="diagnosis">
<paragraph id="1710095EFFEE3462FEF4F8F7CF50FE44" blockId="3.[264,1323,1783,1889]" lastBlockId="4.[264,1323,284,1870]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
<emphasis id="25DBD54CFFEE3465FEF4F8F7CBFAF8F2" box="[312,438,1863,1888]" italics="true" pageId="3" pageNumber="48">Diagnosis.</emphasis>
The species seems to be particularly closely related to
<taxonomicName id="D0AF72DDFFEE3462FB4EF8F7C8D7FEA4" authority="Mikhaljova &amp; Kim, 1993" authorityName="Mikhaljova &amp; Kim" authorityYear="1993" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" lastPageId="4" lastPageNumber="49" order="Polydesmida" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="25DBD54CFFEE3462FB4EF8F7CB1FFEA7" italics="true" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">Epanerchodus beroni</emphasis>
<bibRefCitation id="733E74AFFFE93462FEACFEACC8D7FEA4" author="Mikhaljova" box="[352,667,284,310]" pageId="4" pageNumber="53" refString="Mikhaljova, E. V. &amp; Kim, J. - P. (1993) Contribution to the millipede fauna of Korea (Diplopoda). Korean Arachnology, 9 (1 - 2), 31 - 42." type="journal article" year="1993">Mikhaljova &amp; Kim, 1993</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="6FB849CEFFE93462FD21FEACC9C4FEA4" box="[749,904,284,310]" name="North Korea" pageId="4" pageNumber="49">North Korea</collectingCountry>
, but differs in the low and broad postfemoral process of the gonopods, bearing apical teeth (or by the low and broad caudal distofemoral process of gonopods following a more recently proposed terminology [
<bibRefCitation id="733E74AFFFE93462FEC3FE24CB94FE3C" author="Golovatch" box="[271,472,404,430]" pageId="4" pageNumber="52" refString="Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159." type="journal article" year="1991">Golovatch, 1991</bibRefCitation>
;
<bibRefCitation id="733E74AFFFE93462FE2FFE24C8AEFE3C" author="Djursvoll" box="[483,738,404,430]" pageId="4" pageNumber="52" refString="Djursvoll, P., Golovatch, S. I., Johanson, K. A. &amp; Meidell, B. (2001 (for 2000 )) Phylogenetic relationships within Polydesmus sensu lato (Diplopoda: Polydesmidae). Wytwer, J. &amp; Golovatch, S. I. (Eds.). Progress in studies on Myriapoda and Onychophora. Fragmenta Faunistica, 43 (Suppl.), 37 - 59." type="journal article" year="2001">
Djursvoll
<emphasis id="25DBD54CFFE93462FD95FE24C8C0FE3F" box="[601,652,404,429]" italics="true" pageId="4" pageNumber="49">et al</emphasis>
., 2001
</bibRefCitation>
]), the greater number of bacilli in the distal part of gonopods and the absence of ventral outgrowths on the postfemora and tibiae of the legs.
</paragraph>
</subSubSection>
<subSubSection id="5FB55AD5FFE93462FEF4FE54C964FDDC" pageId="4" pageNumber="49" type="etymology">
<paragraph id="1710095EFFE93462FEF4FE54C964FDDC" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="49">
<emphasis id="25DBD54CFFE93462FEF4FE54CBF7FE6F" box="[312,443,484,509]" italics="true" pageId="4" pageNumber="49">Etymology.</emphasis>
The specific epithet refers to the presence the group of bacilli on the gonopod telopodite (or on the cephalic distofemoral process of gonopods in terms of
<bibRefCitation id="733E74AFFFE93462FEC4FD84CB94FDDC" author="Golovatch" box="[264,472,564,590]" pageId="4" pageNumber="52" refString="Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159." type="journal article" year="1991">Golovatch (1991)</bibRefCitation>
and
<bibRefCitation id="733E74AFFFE93462FDDCFD84C95CFDDC" author="Djursvoll" box="[528,784,564,590]" pageId="4" pageNumber="52" refString="Djursvoll, P., Golovatch, S. I., Johanson, K. A. &amp; Meidell, B. (2001 (for 2000 )) Phylogenetic relationships within Polydesmus sensu lato (Diplopoda: Polydesmidae). Wytwer, J. &amp; Golovatch, S. I. (Eds.). Progress in studies on Myriapoda and Onychophora. Fragmenta Faunistica, 43 (Suppl.), 37 - 59." type="journal article" year="2001">
Djursvoll
<emphasis id="25DBD54CFFE93462FD4BFD84C8F6FDDF" box="[647,698,564,589]" italics="true" pageId="4" pageNumber="49">et al</emphasis>
. (2001
</bibRefCitation>
)).
</paragraph>
</subSubSection>
<subSubSection id="5FB55AD5FFE93462FEF4FDECC846F944" pageId="4" pageNumber="49" type="description">
<paragraph id="1710095EFFE93462FEF4FDECCEEAFBEC" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="49">
<emphasis id="25DBD54CFFE93462FEF4FDECCB85FDE7" box="[312,457,604,629]" italics="true" pageId="4" pageNumber="49">Description.</emphasis>
Male. Length
<quantity id="D057A4BBFFE93462FDA3FDECC888FDE4" box="[623,708,604,630]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="49" unit="mm" value="27.0">27 mm</quantity>
, width 3.5 mm. Coloration pink, antennae and distal parts of legs light pink. Head covered with minute pubescence in anterior part. Vertigial setae absent (or all broken off). Genae rectangular in dorsal view. Antennae long and slender, clavate, in situ reaching to segment 4. Antennomere 7 with two small knobs, one bare, the other with short conical setae; antennomere 6 distally with group of short dense setae (
<figureCitation id="8F9415DBFFE93462FE9EFC94CBDBFCAC" box="[338,407,804,830]" captionStart="FIGURES 1 3" captionStartId="5.[264,382,1780,1804]" captionTargetBox="[292,1281,741,1749]" captionTargetId="figure@5.[263,1324,695,1772]" captionTargetPageId="5" captionText="FIGURES 1 3. Epanerchodus bacilliferus sp. n. (male holotype). 1, distal part of antenna; 2, gonopod (lateral view); 3, distal part of gonopod (mesal view). Scales in mm." httpUri="https://zenodo.org/record/174511/files/figure.png" pageId="4" pageNumber="49">Fig. 1</figureCitation>
). Length ratios of antennomeres 27 as 2.5:3.7:2.8:2.8:2.3:1.0, width ratios as 0.9:1.0:1.0:1.0:1.2:1.0, respectively. Collum elliptical, somewhat narrower than head; posterior corners slightly angular. Segments 24 somewhat shorter and narrower than others. Body parallel-sided on somites 615, further on gradually tapering toward telson. Metatergal polygonal sculpture as usual, three transverse rows of bosses, second and third rows being developed stronger than first one. Tergal setae short, apically blunt, almost all broken off. Paraterga well-developed, their caudolateral corners beak-shaped, pointed on segments 818. Ozopores evident, dorsal, ozopore formula normal. Epiproct produced into a caudal process, or tail, medium-sized, conical, rounded at tip, with setae.
</paragraph>
<paragraph id="1710095EFFE93462FEF4FB3CCEA8FB64" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="49">Legs long and slender; postfemora, tibiae and tarsi with sphaerotrichs on ventral side. Leg pairs 1 and 2 reduced in size as compared to subsequent pairs; leg pair 1 about half the size of leg pairs of midbody. Legs without any outgrowths on postfemora and tibiae.</paragraph>
<paragraph id="1710095EFFE93462FEF4FAB4C932FA74" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="49">
Following traditional terminology, gonopods (
<figureCitation id="8F9415DBFFE93462FCACFAB4C987FA8C" box="[864,971,1284,1310]" captionStart="FIGURES 1 3" captionStartId="5.[264,382,1780,1804]" captionTargetBox="[292,1281,741,1749]" captionTargetId="figure@5.[263,1324,695,1772]" captionTargetPageId="5" captionText="FIGURES 1 3. Epanerchodus bacilliferus sp. n. (male holotype). 1, distal part of antenna; 2, gonopod (lateral view); 3, distal part of gonopod (mesal view). Scales in mm." httpUri="https://zenodo.org/record/174511/files/figure.png" pageId="4" pageNumber="49">Figs 23</figureCitation>
) with a usual, heavily setose prefemoral portion. Femur well-developed, without outer horn. Clivus relatively broad, with a lamelliform margin. Seminal (= prostatic) groove making a characteristic loop, ending at bottom of femoral cavity. Postfemoral process low and broad, with apical teeth. Telopodite flattened, at about distal third strongly curved, caudally with hook-shaped processes, subapically with group of styliform bacilli.
</paragraph>
<paragraph id="1710095EFFE93462FEF4FA44C86FF93C" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="49">
Following a recently proposed terminology (
<bibRefCitation id="733E74AFFFE93462FC86FA44CE59F99C" author="Golovatch" box="[842,1045,1524,1550]" pageId="4" pageNumber="52" refString="Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159." type="journal article" year="1991">Golovatch, 1991</bibRefCitation>
;
<bibRefCitation id="733E74AFFFE93462FBEEFA44CF6EF99C" author="Djursvoll" box="[1058,1314,1524,1550]" pageId="4" pageNumber="52" refString="Djursvoll, P., Golovatch, S. I., Johanson, K. A. &amp; Meidell, B. (2001 (for 2000 )) Phylogenetic relationships within Polydesmus sensu lato (Diplopoda: Polydesmidae). Wytwer, J. &amp; Golovatch, S. I. (Eds.). Progress in studies on Myriapoda and Onychophora. Fragmenta Faunistica, 43 (Suppl.), 37 - 59." type="journal article" year="2001">
Djursvoll
<emphasis id="25DBD54CFFE93462FB56FA44CE82F99F" box="[1178,1230,1524,1549]" italics="true" pageId="4" pageNumber="49">et al</emphasis>
., 2001
</bibRefCitation>
) gonopods (
<figureCitation id="8F9415DBFFE93462FE45F9ACCBBCF9A4" box="[393,496,1564,1590]" captionStart="FIGURES 1 3" captionStartId="5.[264,382,1780,1804]" captionTargetBox="[292,1281,741,1749]" captionTargetId="figure@5.[263,1324,695,1772]" captionTargetPageId="5" captionText="FIGURES 1 3. Epanerchodus bacilliferus sp. n. (male holotype). 1, distal part of antenna; 2, gonopod (lateral view); 3, distal part of gonopod (mesal view). Scales in mm." httpUri="https://zenodo.org/record/174511/files/figure.png" pageId="4" pageNumber="49">Figs 23</figureCitation>
) without exomerite (=outer horn) but with two distofemoral processes, of which cephalic one the strongest, flattened, bearing a group of styliform bacilli subapically and hook-shaped processes caudally. Caudal distofemoral process low and broad, with apical teeth.
</paragraph>
<paragraph id="1710095EFFE93462FEF4F90CC846F944" blockId="4.[264,1323,284,1870]" box="[312,522,1724,1750]" pageId="4" pageNumber="49">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="5FB55AD5FFE93463FEF4F954C807FD0C" lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49" type="discussion">
<paragraph id="1710095EFFE93463FEF4F954C9C1FE6C" blockId="4.[264,1323,284,1870]" lastBlockId="5.[264,1323,284,670]" lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49">
<emphasis id="25DBD54CFFE93462FEF4F954CBEAF96F" box="[312,422,1764,1789]" italics="true" pageId="4" pageNumber="49">Remarks.</emphasis>
<bibRefCitation id="733E74AFFFE93462FE7DF954C8C9F96C" author="Golovatch" box="[433,645,1764,1790]" pageId="4" pageNumber="52" refString="Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159." type="journal article" year="1991">Golovatch (1991)</bibRefCitation>
in his review of the family
<taxonomicName id="D0AF72DDFFE93462FC2AF954CEC7F96C" box="[998,1163,1764,1790]" class="Diplopoda" family="Polydesmidae" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="49" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
in Southeast Asia proposed new terminology for several portions of polydesmid gonopods. He acknowledges that the traditional denomination femorite, part lying distad of the densely setose, basal prefemoral portion, does not at all reflect primary leg segmentation. Hence, gonopod femorite of
<taxonomicName id="D0AF72DDFFE83463FDC1FEF4C8F9FECF" box="[525,693,324,349]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="25DBD54CFFE83463FDC1FEF4C8F9FECF" box="[525,693,324,349]" italics="true" pageId="5" pageNumber="50">Epanerchodus</emphasis>
</taxonomicName>
species can be supplied with endomerite, currently referred to as exomerite (
<bibRefCitation id="733E74AFFFE83463FDF9FEDCC974FE14" author="Djursvoll" box="[565,824,364,390]" pageId="5" pageNumber="52" refString="Djursvoll, P., Golovatch, S. I., Johanson, K. A. &amp; Meidell, B. (2001 (for 2000 )) Phylogenetic relationships within Polydesmus sensu lato (Diplopoda: Polydesmidae). Wytwer, J. &amp; Golovatch, S. I. (Eds.). Progress in studies on Myriapoda and Onychophora. Fragmenta Faunistica, 43 (Suppl.), 37 - 59." type="journal article" year="2001">
Djursvoll
<emphasis id="25DBD54CFFE83463FD63FEDCC8BEFE17" box="[687,754,364,389]" italics="true" pageId="5" pageNumber="50">et al.,</emphasis>
2001
</bibRefCitation>
) (following traditional terminology, outer horn) and distofemoral outgrowths: cephalic (following traditional terminology, telopodite), caudal (following traditional terminology, postfemoral process) and middle (following traditional terminology, additional process).
</paragraph>
<paragraph id="1710095EFFE83463FEF4FDBCC807FD0C" blockId="5.[264,1323,284,670]" pageId="5" pageNumber="50">
The resemblance of gonopod illustrations of new species and Paiks (1960)
<taxonomicName id="D0AF72DDFFE83463FEC4FD84CBFCFDDF" box="[264,432,564,589]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="25DBD54CFFE83463FEC4FD84CBFCFDDF" box="[264,432,564,589]" italics="true" pageId="5" pageNumber="50">Epanerchodus</emphasis>
</taxonomicName>
sp. from Mt. Jiri,
<collectingCountry id="6FB849CEFFE83463FD45FD84C950FDDC" box="[649,796,564,590]" name="South Korea" pageId="5" pageNumber="50">South Korea</collectingCountry>
permits us to suppose that the latter belongs to
<taxonomicName id="D0AF72DDFFE83463FEE5FDECCB9CFDE7" box="[297,464,604,629]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="bacilliferus" status="sp. nov.">
<emphasis id="25DBD54CFFE83463FEE5FDECCB9CFDE7" box="[297,464,604,629]" italics="true" pageId="5" pageNumber="50">E. bacilliferus</emphasis>
</taxonomicName>
<emphasis id="25DBD54CFFE83463FE16FDEBC859FDE7" bold="true" box="[474,533,603,629]" pageId="5" pageNumber="50">
<taxonomicNameLabel id="3EE86837FFE83463FE16FDEBC859FDE7" box="[474,533,603,629]" pageId="5" pageNumber="50" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
Also, Lims (1988)
<taxonomicName id="D0AF72DDFFE83463FCC0FDECC9F8FDE7" box="[780,948,604,629]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="25DBD54CFFE83463FCC0FDECC9F8FDE7" box="[780,948,604,629]" italics="true" pageId="5" pageNumber="50">Epanerchodus</emphasis>
</taxonomicName>
sp. 1 from
<collectingCountry id="6FB849CEFFE83463FB88FDECCE96FDE4" box="[1092,1242,604,630]" name="South Korea" pageId="5" pageNumber="50">South Korea</collectingCountry>
seems closely related to
<taxonomicName id="D0AF72DDFFE83463FE14FD34C80FFD0F" box="[472,579,644,669]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="25DBD54CFFE83463FE14FD34C80FFD0F" box="[472,579,644,669]" italics="true" pageId="5" pageNumber="50">E. beroni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>