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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1039.64740" ID-GBIF-Dataset="8fe0041e-40a6-4fd0-a674-948f3f919703" ID-PMC="PMC8159918" ID-Pensoft-Pub="1313-2970-1039-1" ID-Pensoft-UUID="FA2622A854585C59A7E8182A4E66C6DF" ID-PubMed="34084064" ID-ZooBank="3487C3570FAC4907A5C0C0A0C801C6E3" ModsDocID="1313-2970-1039-1" checkinTime="1621541808324" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2021" docId="5ACDE1923ED251A58221A192D338A26A" docLanguage="en" docName="ZooKeys 1039: 1-108" docOrigin="ZooKeys 1039" docPubDate="2021-05-20" docSource="http://dx.doi.org/10.3897/zookeys.1039.64740" docTitle="Pyrrhalta ishiharai Kimoto 1976" docType="treatment" docVersion="4" id="FA2622A854585C59A7E8182A4E66C6DF" lastPageNumber="1" masterDocId="FA2622A854585C59A7E8182A4E66C6DF" masterDocTitle="Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei" masterLastPageNumber="108" masterPageNumber="1" pageNumber="1" updateTime="1668152740594" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:affiliation>Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4358-7211</mods:nameIdentifier>
<mods:affiliation>Mendel University in Brno, Department of Zoology, Fisheries, Hydrobiology and Apiculture, Zemĕdĕlska 1, 613 00, Brno, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1039</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>108</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1039.64740</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1039.64740</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1039-1</mods:identifier>
<mods:identifier type="ZooBank">3487C3570FAC4907A5C0C0A0C801C6E3</mods:identifier>
<mods:identifier type="Pensoft-UUID">FA2622A854585C59A7E8182A4E66C6DF</mods:identifier>
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<treatment ID-GBIF-Taxon="182248282" LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A" httpUri="http://treatment.plazi.org/id/5ACDE1923ED251A58221A192D338A26A" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="5ACDE192-3ED2-51A5-8221-A192D338A26A" authority="Kimoto, 1976" authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">Pyrrhalta ishiharai Kimoto, 1976</taxonomicName>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Field photographs of adults of Pyrrhalta species on host plants A P. tsoui Bezdek &amp; Lee B P. formosanensis sp. nov. C P. semifulva (Jacoby) D P. ishiharai Kimoto." figureDoi="10.3897/zookeys.1039.64740.figure26" httpUri="https://binary.pensoft.net/fig/545556" pageId="0" pageNumber="1">Figs 26D</figureCitation>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Habitus of Pyrrhalta ishiharai Kimoto and P. wulaiensis sp. nov. A P. ishiharai, female, typical form, dorsal view B ditto, ventral view C ditto, lateral view D P. wulaiensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1">, 34A-C</figureCitation>
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">, 35</figureCitation>
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Field photographs of Pyrrhalta wulaiensis sp. nov. on host plant A egg B third-instar larva C mature larva D pupa E adult F adults: P. wulaiensis sp. nov. (left) and P. ishiharai (right)." figureDoi="10.3897/zookeys.1039.64740.figure37" httpUri="https://binary.pensoft.net/fig/545567" pageId="0" pageNumber="1">, 37F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta aurata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aurata">Pyrrhalta aurata</taxonomicName>
: Kimoto, 1976: 4 (Taiwan). Misidentification (after
<bibRefCitation author="Kimoto, S" journalOrPublisher="The Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="1" pagination="191 - 192" refId="B36" refString="Kimoto, S, 1994. Description of a new galerucid species from Taiwan, China (Coleoptera: Chrysomelidae). The Transactions of the Shikoku Entomological Society 20: 191 - 192" title="Description of a new galerucid species from Taiwan, China (Coleoptera: Chrysomelidae)." volume="20" year="1994">Kimoto 1994</bibRefCitation>
)!
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">Pyrrhalta ishiharai</taxonomicName>
Kimoto, 1994: 191;
<bibRefCitation author="Kimoto, S" journalOrPublisher="Bulletin of the Institute of Comparative Studies of International Cultures and Societies" pageId="0" pageNumber="1" pagination="1 - 152" refId="B39" refString="Kimoto, S, Chu, Y-I, 1996. Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera). Bulletin of the Institute of Comparative Studies of International Cultures and Societies 16: 1 - 152" title="Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera)." volume="16" year="1996">Kimoto and Chu 1996</bibRefCitation>
: 56 (catalogue);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="0" pageNumber="1" refId="B42" refString="Kimoto, S, Takizawa, H, 1997. Leaf beetles (Chrysomelidae) of Taiwan. Tokai University Press, Tokyo" title="Leaf beetles (Chrysomelidae) of Taiwan." year="1997">Kimoto and Takizawa 1997</bibRefCitation>
: 300 (key), 373;
<bibRefCitation author="Beenen, R" editor="Loebl, I" journalOrPublisher="6). Chrysomeloidea. Apollo Books, Stenstrup" pageId="0" pageNumber="1" pagination="443 - 491" publicationUrl="https://www.cabdirect.org/cabdirect/abstract/20123189777" refId="B3" refString="Beenen, R, 2010. Galerucinae. In: Loebl, I, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera (Vol. 6). Chrysomeloidea. Apollo Books, Stenstrup: 443 - 491, https://www.cabdirect.org/cabdirect/abstract/20123189777" title="Galerucinae." url="https://www.cabdirect.org/cabdirect/abstract/20123189777" volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2010">Beenen 2010</bibRefCitation>
: 452 (catalogue);
<bibRefCitation author="Xue, HJ" journalOrPublisher="Junk, ' s-Gravenhage" pageId="0" pageNumber="1" publicationUrl="https://www.cabdirect.org/cabdirect/abstract/20103331643" refId="B93" refString="Xue, HJ, Yang, X-K, 2010. Species catalogue of Pyrrhalta Joannis (Coleoptera: Chrysomelidae: Galerucinae) of the world. Entomotaxonomia 32 (supplement): 119-136. https://www.cabdirect.org/cabdirect/abstract/20103331643" title="Species catalogue of Pyrrhalta Joannis (Coleoptera: Chrysomelidae: Galerucinae) of the world. Entomotaxonomia 32 (supplement): 119 - 136." url="https://www.cabdirect.org/cabdirect/abstract/20103331643" year="2010">Xue and Yang 2010</bibRefCitation>
: 124 (catalogue);
<bibRefCitation author="Yang, X" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B98" refString="Yang, X, Ge, S, Nie, R, Ruan, Y, Li, W, 2015. Chinese leaf beetles. Science Press, Beijing" title="Chinese leaf beetles." year="2015">Yang et al. 2015</bibRefCitation>
: 117 (catalogue).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♀ (EUMJ, by original designation): &quot;(TAIWAN) / Kueishan [龜山] / ~ Wulai [烏來] / Taipei Hsien / 5. VI, 1970 / Y. Hori leg. [p, w] //
<taxonomicName authorityName="Joannis" authorityYear="1865" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pyrrhalta</taxonomicName>
/
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">Pyrrhalta ishiharai</taxonomicName>
/ Kimoto, n. sp [h] / Det. S. Kimoto, 19[p]93 [h, w] // msp [h, w] // PHOTO [p, r] // HOLOTYPE [p, r]&quot;.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
1♀ (KMNH): &quot;NANSHANCHI [南山溪] / TAIWAN / 2. V. 1982 / F. KIMURA [p, y] //
<taxonomicName authorityName="Joannis" authorityYear="1865" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pyrrhalta</taxonomicName>
/
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">Pyrrhalta ishiharai</taxonomicName>
/ Kimoto, n. sp [h] / Det. S. Kimoto, 19[p]93 [h, w] // PARATYPE [p, b]&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2010-07-10" collectorName="M. - H. Tsou" country="Taiwan" location="Chienshih" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hsinchu">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
.
<collectingRegion country="Taiwan" name="Hsinchu">Hsinchu</collectingRegion>
:
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A:10BB13BA99A79AD1C731872B94B8264F" country="Taiwan" name="Chienshih" stateProvince="Hsinchu">Chienshih</location>
(尖石),
<collectingDate value="2010-07-10">10.VII.2010</collectingDate>
, leg.
<collectorName>M.-H. Tsou</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2012-08-05" collectorName="Y. - L. Lin" country="Taiwan" location="Chienshih" specimenCount="1" specimenCount-male="1" stateProvince="Hsinchu">
<specimenCount type="male">1♂</specimenCount>
(TARI), same locality,
<collectingDate value="2012-08-05">5.VIII.2012</collectingDate>
, leg.
<collectorName>Y.-L. Lin</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2003-01-07" collectingDateMax="2003-02-13" collectingDateMin="2003-01-07" collectorName="C. S. Lin, W. T. Yang" country="Taiwan" location="Chunyang" specimenCount="1" specimenCount-male="1" stateProvince="Nantou">
<collectingRegion country="Taiwan" name="Nantou">Nantou</collectingRegion>
:
<specimenCount type="male">1♂</specimenCount>
(NMNS),
<location LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A:C8A816CF1ED6646D7A168F960776A668" country="Taiwan" name="Chunyang" stateProvince="Nantou">Chunyang</location>
(春陽),
<collectingDate value="2003-01-07" valueMax="2003-02-13" valueMin="2003-01-07">7.I. -13.II.2003</collectingDate>
, leg.
<collectorName>C. S. Lin</collectorName>
&amp;
<collectorName>W. T. Yang</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2011-06-26" collectorName="M. - H. Tsou" country="Taiwan" location="Fushan" specimenCount="2" specimenCount-female="2" stateProvince="Taipei">
<collectingRegion country="Taiwan" name="Taipei">Taipei</collectingRegion>
:
<specimenCount type="female">2♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A:AC6B53F99888D8961C1FD76E0D2BC7E1" country="Taiwan" name="Fushan" stateProvince="Taipei">Fushan</location>
(福山),
<collectingDate value="2011-06-26">26.VI.2011</collectingDate>
, leg.
<collectorName>M.-H. Tsou</collectorName>
</materialsCitation>
;
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“8.VII.2011”">
&quot;
<date value="2011-07-08">8.VII.2011</date>
&quot;
</normalizedToken>
;
<specimenCount type="male">3♂</specimenCount>
,
<specimenCount type="female">3♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“21.VI.2015”">
&quot;
<date value="2015-06-21">21.VI.2015</date>
&quot;
</normalizedToken>
;
<materialsCitation collectingDate="2011-07-08" collectorName="M. - H. Tsou" country="Taiwan" location="Hsinhsien" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Taipei">
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A:80F89C09D784BC91B2008322D3CD44E2" country="Taiwan" name="Hsinhsien" stateProvince="Taipei">Hsinhsien</location>
(信賢),
<collectingDate value="2011-07-08">8.VII.2011</collectingDate>
, leg.
<collectorName>M.-H. Tsou</collectorName>
</materialsCitation>
;
<specimenCount type="male">4♂</specimenCount>
,
<specimenCount type="female">3♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“6.VII.2019”">
&quot;
<date value="2019-07-06">6.VII.2019</date>
&quot;
</normalizedToken>
;
<specimenCount type="male">4♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
(TARI), same but with
<date value="2019-07-07">7.VII.2019</date>
&quot;;
<specimenCount type="male">3♂</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“27.VI.2020”">
&quot;
<date value="2020-06-27">27.VI.2020</date>
&quot;
</normalizedToken>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“5.VII.2020”">
&quot;
<date value="2020-07-05">5.VII.2020</date>
&quot;
</normalizedToken>
;
<materialsCitation collectingDate="2011-07-08" collectorName="M. - H. Tsou" country="Taiwan" location="Wulai" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Taipei">
<specimenCount type="male">4♂</specimenCount>
,
<specimenCount type="female">3♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A:0B2A40E9A098546A4E5919CA4B6675E6" country="Taiwan" name="Wulai" stateProvince="Taipei">Wulai</location>
(烏來),
<collectingDate value="2011-07-08">8.VII.2011</collectingDate>
, leg.
<collectorName>M.-H. Tsou</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“17.VI.2018”">
&quot;
<date value="2018-06-17">17.VI.2018</date>
&quot;
</normalizedToken>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">7♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“27.VI.2020”">
&quot;
<date value="2020-06-27">27.VI.2020</date>
&quot;
</normalizedToken>
;
<materialsCitation collectingDate="2020-07-05" country="Taiwan" location="Taitung" specimenCount="13" specimenCount-female="8" specimenCount-male="5" stateProvince="Taitung">
<specimenCount type="male">5♂</specimenCount>
,
<specimenCount type="female">8♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“5.VII.2020”">
&quot;
<collectingDate value="2020-07-05">5.VII.2020</collectingDate>
&quot;
</normalizedToken>
;
<collectingRegion country="Taiwan" name="Taitung">Taitung</collectingRegion>
</materialsCitation>
:
<materialsCitation collectingDate="2013-06-19" collectorName="Y. - T. Chung" country="Taiwan" location="Liyuan" specimenCount="2" specimenCount-male="2" stateProvince="Taitung">
<specimenCount type="male">2♂</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:5ACDE1923ED251A58221A192D338A26A:AE99E86565391EBDBC81413139481DB9" country="Taiwan" name="Liyuan" stateProvince="Taitung">Liyuan</location>
(栗園),
<collectingDate value="2013-06-19">19.VI.2013</collectingDate>
, leg.
<collectorName>Y.-T. Chung</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="B. - X. Guo" country="Taiwan" specimenCount="2" specimenCount-male="2" stateProvince="Taitung">
<specimenCount type="male">2♂</specimenCount>
(TARI), same but with &quot;leg.
<collectorName>B.-X. Guo</collectorName>
&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Length 4.8-5.1 mm, width 2.3-2.5 mm. Body color (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Habitus of Pyrrhalta ishiharai Kimoto and P. wulaiensis sp. nov. A P. ishiharai, female, typical form, dorsal view B ditto, ventral view C ditto, lateral view D P. wulaiensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1">34A-C</figureCitation>
) yellowish brown; vertex with one dark spot at center; antennae dark brown, but four or five basal antennomeres basally paler; pronotum with three black spots, one elongate spot at center, one pair laterally; scutellum basally darker; four pairs of transverse dark spots on elytra, one pair near base and behind scutellum, three pairs at basal 2/5, 3/5, 4/5 respectively, intercepted by two pairs of longitudinal yellowish brown ridges, all dark spots poorly defined; meso- and metathoracic ventrites darker; apical 2/3 of tibiae and entire tarsi black except inner side of protibia. Eyes small, interocular space 2.35-2.38
<normalizedToken originalValue="×">x</normalizedToken>
diameter of eye. Antennae filiform in males (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35A</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.6: 0.9: 0.7: 0.7: 0.7: 0.6: 0.6: 0.6: 0.6: 0.9, length to width ratios of antennomeres I-XI 3.1: 2.1: 3.2: 2.5: 2.4: 2.1: 1.8: 1.8: 1.8: 1.6: 2.2; filiform in females (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.6: 0.7: 0.7: 0.6: 0.6: 0.6: 0.6: 0.5: 0.5: 0.8, length to width ratios of antennomeres I-XI 3.4: 2.5: 2.9: 2.8: 2.2: 1.7: 1.6: 1.6: 1.5: 1.4: 2.3. Pronotum and elytra convex. Pronotum 1.7-1.9
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, disc with reticulate microsculpture; coarse, extremely dense punctures, and extremely short pubescence; with median longitudinal and lateral depressions; lateral margins angular, widest at apical 1/3, apical and basal margins truncate; posterior setiferous punctures strongly erect. Elytra elongate, broad, parallel-sided, 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with reticulate microsculpture and coarse, extremely dense punctures and short pubescence; with two pairs of long longitudinal ridges near suture, apically abbreviated; several oblique ridges exterior to longitudinal ridges. Apical spur of middle tibia of middle small (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35E</figureCitation>
), tarsomere I with a small tooth at middle ventrally in males (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35H</figureCitation>
). Aedeagus (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35C, D</figureCitation>
) slender in dorsal view, 5.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, sides asymmetric, widest at middle, apex angular; strongly curved near base in lateral view, weakly recurved apically, apex acute; ostium longitudinal, not covered by membrane; two endophallic sclerites elongate, apex of primary endophallic sclerite with several teeth, 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, secondary sclerite much shorter, 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as primary sclerite, apex acute, with one additional tooth near apex. Only apices of gonocoxae (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35K</figureCitation>
) sclerotized and longitudinal, with dense, long setae along lateral and apical margins. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35F</figureCitation>
) narrow; disc with several long setae and dense short setae along apical margin; spiculum long. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35G</figureCitation>
) very swollen; pump short and strongly curved; sclerotized proximal spermathecal duct wide and short. Apical margin of abdominal ventrite V slightly concave, with deeply rounded depression at middle in males (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35J</figureCitation>
); slightly concave in females (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35I</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1" start="Figure 34" startId="F34">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 34.</emphasis>
Habitus of
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta ishiharai</emphasis>
</taxonomicName>
Kimoto and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. ishiharai" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">P. ishiharai</emphasis>
</taxonomicName>
, female, typical form, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
ditto, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
ditto, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov., female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
ditto, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
ditto, lateral view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1" start="Figure 35" startId="F35">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 35.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta ishiharai</emphasis>
</taxonomicName>
Kimoto
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna, male
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna, female
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
ditto, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
apex of tibia of middle leg, male
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
spermatheca
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
tarsi of middle leg, male
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
abdominal ventrite V, female
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
abdominal ventrite V, male
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
gonocoxae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adults of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. ishiharai" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">P. ishiharai</emphasis>
</taxonomicName>
Kimoto and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. are easily separated from other species within the species group by the longitudinal ridges on the elytra (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Habitus of Pyrrhalta ishiharai Kimoto and P. wulaiensis sp. nov. A P. ishiharai, female, typical form, dorsal view B ditto, ventral view C ditto, lateral view D P. wulaiensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1">34</figureCitation>
) and the angular apices of the aedeagi (Figs
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35C</figureCitation>
,
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36C</figureCitation>
).
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta ishiharai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. by the larger body size (Fig.
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Field photographs of Pyrrhalta wulaiensis sp. nov. on host plant A egg B third-instar larva C mature larva D pupa E adult F adults: P. wulaiensis sp. nov. (left) and P. ishiharai (right)." figureDoi="10.3897/zookeys.1039.64740.figure37" httpUri="https://binary.pensoft.net/fig/545567" pageId="0" pageNumber="1">37F</figureCitation>
), 4.8-5.1 mm long (3.3-3.7 mm long in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov.), dark spots present between the longitudinal ridges on the elytra (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Habitus of Pyrrhalta ishiharai Kimoto and P. wulaiensis sp. nov. A P. ishiharai, female, typical form, dorsal view B ditto, ventral view C ditto, lateral view D P. wulaiensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1">34A</figureCitation>
) (dark spots absent between longitudinal ridges on elytra in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Habitus of Pyrrhalta ishiharai Kimoto and P. wulaiensis sp. nov. A P. ishiharai, female, typical form, dorsal view B ditto, ventral view C ditto, lateral view D P. wulaiensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1">34D</figureCitation>
), apical spine present on tibia (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35E</figureCitation>
) and modified tarsomere I of middle leg (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35H</figureCitation>
) in males (lacking apical spine on tibia and normal tarsomere I of middle leg in males of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov.), longitudinal ostium and aedeagus recurved in apical 1/3 (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35C, D</figureCitation>
) (transverse ostium and aedeagus curved at middle in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36C, D</figureCitation>
)), longitudinally cylindrical gonocoxae with dense, long setae (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35K</figureCitation>
) (transversely rounded gonocoxae with scattered short setae in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36I</figureCitation>
)).
</paragraph>
<caption doi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2021" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta wulaiensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta wulaiensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna, male
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna, female
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
ditto, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
spermatheca
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
abdominal ventrite V, female
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
abdominal ventrite V, male
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
gonocoxae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="food plant">
<paragraph pageId="0" pageNumber="1">Food plant.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adults feed on flowers of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Meliosma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Meliosma rhoifolia" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="rhoifolia">
<emphasis italics="true" pageId="0" pageNumber="1">Meliosma rhoifolia</emphasis>
</taxonomicName>
Maxim. (
<taxonomicName class="Dicotyledoneae" family="Sabiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Sabiaceae</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Field photographs of adults of Pyrrhalta species on host plants A P. tsoui Bezdek &amp; Lee B P. formosanensis sp. nov. C P. semifulva (Jacoby) D P. ishiharai Kimoto." figureDoi="10.3897/zookeys.1039.64740.figure26" httpUri="https://binary.pensoft.net/fig/545556" pageId="0" pageNumber="1">26D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">The species is widespread at lowlands (0-1,500 m) in Taiwan.</paragraph>
</subSubSection>
</treatment>
</document>