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<mods:title id="AE56D6396C3C464524E6FDB20C0886CF">Contributions To The Study Of The Early Jurassic Petrified Forest Of Holbav And Cristian Areas (Brașov Region, South Carpathians, Romania), 2 Part.</mods:title>
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<mods:namePart id="DDF77C43D04D01E3F7A08D2C07325EF4">Iamandei, Stanila</mods:namePart>
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<mods:affiliation id="7AA75D6A73F7A6C61920DA69D1AC402B">Geological Institute of Romania, 1 st, Caransebeş Street, 012271 - Bucharest, Romania. iamandei @ gmail. com</mods:affiliation>
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<mods:namePart id="B54CD4E9BE9E509DA74088C1839F123A">Grădinaru, Eugen</mods:namePart>
<mods:affiliation id="D0FD2CE64B345FAA69A387900021789C">University of Bucharest, Department of Geology, 1, Bălcescu Nicolae Blvd., 010041, Bucharest, Romania. egradinaru @ gmail. com</mods:affiliation>
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<taxonomicName id="7BB14D101D44FFB1972BE35DFB695903" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D44FFB1972BE35DFBA65922" box="[850,1138,820,844]" italics="true" pageId="8" pageNumber="33">Brachyoxylon cristianicum</emphasis>
<bibRefCitation id="D8204B621D44FFB19000E35DFB695903" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="F4AB65181D44FFB19056E31FFB4A59E0" box="[1071,1182,886,910]" pageId="8" pageNumber="33" type="description">
<paragraph id="BC0E36931D44FFB19056E31FFB4A59E0" blockId="8.[850,1402,820,910]" box="[1071,1182,886,910]" pageId="8" pageNumber="33">
<figureCitation id="248A2A161D44FFB19056E31FFBBB59E0" box="[1071,1135,886,910]" captionStart="Fig" captionStartId="9.[178,213,1935,1956]" captionTargetBox="[178,1433,149,1926]" captionTargetId="figure-19@9.[178,1433,149,1926]" captionTargetPageId="9" captionText="Fig. 4 Brachyoxylon cristianicum Iamandei, Iamandei and Grădinaru, 2018 - inventory no. 27690(field no. 1113b). a-c: Cross section - thick-walled tracheids, polygonal, with rounded corners and curled walls; dispersed parenchyma cells; linear rays; d-f: Tangential section - uniseriate rays with polygonal ray-cells and biseriate storeys; unpitted tracheids; parenchyma with end-walls smooth or nodular and resin content inside; g-i: Radial section - radial pitting on tracheids of mixed type; cross-fields with 2-3 cupressoid or araucarioid in a single row or, when more numerous, in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520859" httpUri="https://zenodo.org/record/10520859/files/figure.png" pageId="8" pageNumber="33">Fig. 4</figureCitation>
, a-i.
</paragraph>
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<paragraph id="BC0E36931D44FFB19749E3D1FC4759BE" blockId="8.[816,1436,952,1406]" box="[816,915,952,976]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E3D1FC4759BE" bold="true" box="[816,915,952,976]" pageId="8" pageNumber="33">Material</emphasis>
</paragraph>
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<materialsCitation id="0CD93CCE1D44FFB19749E3B0FA825F10" location="National Museum of Geology" pageId="8" pageNumber="33" specimenCount="1" stateProvince="Bucuresti">
The studied material is represented by 13 samples of silicified wood collected from the Holbav locality, on Maiului brook. All the studied samples are fragments of trunk or thick-branch fragments, with centimetric to decimetric size, dark color and, by hand lens or even naked eye, the regular fibrous structure without vessels is visible, evidence of a conifer wood. The specimens, with the following field numbers: 1071, 1072, 1073, 1075, 1076, 1080, 1081, 1101, 1103, 1108, 1110, 1112, 1113b belong to “Grădinaru Collection” and are stored at the
<location id="B96E60481D44FFB19138E56AFBCC5F52" LSID="urn:lsid:plazi:treatment:341887851D44FFB3972BE35DFB3A5B35:B96E60481D44FFB19138E56AFBCC5F52" name="National Museum of Geology" pageId="8" pageNumber="33" stateProvince="Bucuresti">National Museum of Geology</location>
, in
<collectingRegion id="7E75F8711D44FFB1903EE54DFB605F52" box="[1095,1204,1316,1340]" country="Romania" name="Bucuresti" pageId="8" pageNumber="33">Bucharest</collectingRegion>
, under the inventory numbers 27678, 276789, 27680, 27681, 27682, 27684, 27685, 27686, 27717, 27687, 27688, 27689, 27690
</materialsCitation>
.
</paragraph>
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<paragraph id="BC0E36931D44FFB19749E5C0FB955FAF" blockId="8.[816,1437,1449,2068]" box="[816,1089,1449,1473]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E5C0FB955FAF" bold="true" box="[816,1089,1449,1473]" pageId="8" pageNumber="33">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D44FFB19749E5A3FAD55C2B" blockId="8.[816,1437,1449,2068]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E5A3FC115F8C" box="[816,965,1482,1506]" italics="true" pageId="8" pageNumber="33">Growth rings</emphasis>
in cross-section show a tracheidoxylic structure without resin canals, usually with abrupt transition from early- to late-wood, and with rather distinct growth-ring boundary. Resin canals absent.
</paragraph>
<paragraph id="BC0E36931D44FFB39749E627FE8A5B77" blockId="8.[816,1437,1449,2068]" lastBlockId="10.[151,772,158,1175]" lastPageId="10" lastPageNumber="35" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E627FC4F5C08" box="[816,923,1614,1638]" italics="true" pageId="8" pageNumber="33">Tracheids</emphasis>
with polygonal (quadrangular) cross-section with rounded corners, unequal in size, diameters of 20- 35-(45-65) μm and with thick walls, of 5-8(9) μm the double wall, usually curled by compression. The late-wood is represented by 2-3 rows of thick-walled and radially flattened cells (radial diameter of 5-10 μm). Intercellular spaces are present. The interradial bundles have 1-10 radial regular rows of tracheids. Density is of 992- 1518 tracheids per mm
<superScript id="4BC49BDB1D44FFB19053E73CFBE75D0D" attach="right" box="[1066,1075,1877,1891]" fontSize="6" pageId="8" pageNumber="33">2</superScript>
. Tangential pitting is absent. Radial pitting is usually uniseriate or biseriate of mixed
<typeStatus id="630A88311D44FFB19749E7F3FCB55DDC" box="[816,865,1946,1970]" pageId="8" pageNumber="33">type</typeStatus>
. When biseriate, the pits appear alternate or opposite and contiguous on short rows often continued by uniseriate portions and do not occupy all the length of the wall. The pits have round to hexagonal borders, diameters of 18-22 μm, and are variably flattened when contiguous (d/D=0.65-0.88), bearing round or tilted elliptic apertures of 7.5-10 / 2-3 μm. Sometimes crassulae or helical thickenings can appear.
</paragraph>
<caption id="E8CE661B1D45FFB094CBE7E6FBA35275" ID-DOI="http://doi.org/10.5281/zenodo.10520859" ID-Zenodo-Dep="10520859" httpUri="https://zenodo.org/record/10520859/files/figure.png" pageId="9" pageNumber="34" startId="9.[178,213,1935,1956]" targetBox="[178,1433,149,1926]" targetPageId="9" targetType="figure">
<paragraph id="BC0E36931D45FFB094CBE7E6FAD05DCA" blockId="9.[178,1434,1934,2075]" box="[178,1284,1934,1956]" pageId="9" pageNumber="34">
<emphasis id="8EC5EA811D45FFB094CBE7E6FF3F5DCA" bold="true" box="[178,235,1935,1956]" pageId="9" pageNumber="34">Fig. 4</emphasis>
<taxonomicName id="7BB14D101D45FFB0948BE7E7FC515DCA" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" box="[242,901,1934,1956]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D45FFB0948BE7E7FE215DCA" box="[242,501,1934,1956]" italics="true" pageId="9" pageNumber="34">Brachyoxylon cristianicum</emphasis>
<bibRefCitation id="D8204B621D45FFB09582E7E7FC515DCA" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[507,901,1934,1956]" pageId="9" pageNumber="34" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
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- inventory no. 27690(field no. 1113b).
</paragraph>
<paragraph id="BC0E36931D45FFB094CBE7C5FBA35275" blockId="9.[178,1434,1934,2075]" pageId="9" pageNumber="34">
<emphasis id="8EC5EA811D45FFB094CBE7C5FF055DAF" bold="true" box="[178,209,1964,1985]" pageId="9" pageNumber="34">a-c</emphasis>
: Cross section - thick-walled tracheids, polygonal, with rounded corners and curled walls; dispersed parenchyma cells; linear rays;
<emphasis id="8EC5EA811D45FFB09491E7A3FED15DB1" bold="true" box="[232,261,1994,2015]" pageId="9" pageNumber="34">d-f</emphasis>
: Tangential section - uniseriate rays with polygonal ray-cells and biseriate storeys; unpitted tracheids; parenchyma with end-walls smooth or nodular and resin content inside;
<emphasis id="8EC5EA811D45FFB096B0E781FD305D93" bold="true" box="[713,740,2024,2045]" pageId="9" pageNumber="34">g-i</emphasis>
: Radial section - radial pitting on tracheids of mixed type; cross-fields with 2-3 cupressoid or araucarioid in a single row or, when more numerous, in 2-3 superposed rows.
</paragraph>
</caption>
<paragraph id="BC0E36931D46FFB394EEE14BFE825B8E" blockId="10.[151,772,158,1175]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE14BFEB65B54" box="[151,354,290,314]" italics="true" pageId="10" pageNumber="35">Axial parenchyma</emphasis>
in cross-section appear few, dispersed, often difficult to identify among the tracheids. Vertically, sometimes they can show simple pits. The horizontal walls are thin and smooth, or slightly nodular. Sometimes, a resin content is present inside the cells, as plugs or granules.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE183FE745E3A" blockId="10.[151,772,158,1175]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE183FF1F586F" box="[151,203,490,513]" italics="true" pageId="10" pageNumber="35">Rays</emphasis>
in cross-section appear usually uniseriate and linear. In tangential section, the rays show 1-20(-25) cells in height (i.e. 30-570(-650) μm). The taller uniseriate rays have biseriations, which give sometimes a fusiform aspect. The ray-cells have polygonal rounded to oval section, marginal spaces and sometimes have dark content. The ray density is of 5-11 rays per horizontal millimeter. The rays appear homogeneous, with cells all procumbent, of 20-30 μm tall, relatively thick walled, of 4-5(7) μm the double wall. Within the marginal rows, but not only, the ray cells are taller, of 30-40 μm, and have slightly corrugated outer walls. The cross-fields are of araucarioid
<typeStatus id="630A88311D46FFB394EEE31EFF1C59E1" box="[151,200,887,911]" pageId="10" pageNumber="35">type</typeStatus>
, have 1-6(-8) pits, hexagonal or slightly rounded to oval, of cupressoid
<typeStatus id="630A88311D46FFB39514E3F1FE4959DE" box="[365,413,920,944]" pageId="10" pageNumber="35">type</typeStatus>
, tending to podocarpoid or araucarioid. They have 15-19/8-12 μm in diameters, and show circular or short elliptic tilted apertures of 10-14/5-7 μm. The pits are arranged in 1-2(3) rows per cross field, within ray body fields, or in the marginal fields and are alternate to slightly irregular.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE437FE785E1B" blockId="10.[151,772,158,1175]" box="[151,428,1117,1141]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE437FECB5E1B" box="[151,287,1117,1141]" italics="true" pageId="10" pageNumber="35">Resin canals</emphasis>
are absent.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE416FE395EF9" blockId="10.[151,772,158,1175]" box="[151,493,1151,1175]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE416FE8B5EF9" box="[151,351,1151,1175]" italics="true" pageId="10" pageNumber="35">Mineral inclusions</emphasis>
are absent.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D46FFB394EEE4A8FB3A5B35" pageId="10" pageNumber="35" type="discussion">
<paragraph id="BC0E36931D46FFB394EEE4A8FE785EB7" blockId="10.[151,771,1217,2035]" box="[151,428,1217,1241]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE4A8FE785EB7" bold="true" box="[151,428,1217,1241]" pageId="10" pageNumber="35">Affinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE48AFE1C5F8C" blockId="10.[151,771,1217,2035]" pageId="10" pageNumber="35">
From the microscopic study of all the standard oriented thin sections, we selected
<specimenCount id="AAB7FD1A1D46FFB395BBE56AFD8C5F75" box="[450,600,1283,1307]" count="13" pageId="10" pageNumber="35" type="generic">13 specimens</specimenCount>
having similar generic characters, represented by tracheidoxylic structure, radial pitting of mixed
<typeStatus id="630A88311D46FFB395ADE52FFDD65F33" box="[468,514,1350,1373]" pageId="10" pageNumber="35">type</typeStatus>
on tracheids and araucarioid cross-fields, which are consistent with the structure of
<taxonomicName id="7BB14D101D46FFB3948EE5EEFE5F5FF1" box="[247,395,1415,1439]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D46FFB3948EE5EEFE5F5FF1" box="[247,395,1415,1439]" italics="true" pageId="10" pageNumber="35">Brachyoxylon</emphasis>
</taxonomicName>
wood-type (see
<bibRefCitation id="D8204B621D46FFB3962EE5EEFD2A5FF1" author="Philippe, M." box="[599,766,1415,1439]" pageId="10" pageNumber="35" pagination="74 - 80" refId="ref16155" refString="Philippe, M., 1993. Nomenclature generique des tracheidoxyles mesozoiques a champs araucarioides. Taxon, 42: 74 - 80." type="journal article" year="1993">Philippe, 1993</bibRefCitation>
; Philipe and Bamford, 2008), but a slightly different from the above-described species.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE582FE755CC6" blockId="10.[151,771,1217,2035]" pageId="10" pageNumber="35">
We do not repeat the discussion from above on the genus creation and about the previously described species, but, consulting the comparative Table 1, from
<bibRefCitation id="D8204B621D46FFB39618E644FF355C08" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="10" pageNumber="35" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al. (2018)</bibRefCitation>
, we observed beside the generic characters some other specific details that are different of those of the above-described species.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE6D8FD575D43" blockId="10.[151,771,1217,2035]" pageId="10" pageNumber="35">
Thus, we observed that the radial tracheidal pitting of mixed
<typeStatus id="630A88311D46FFB39498E6BAFEDB5C85" box="[225,271,1747,1771]" pageId="10" pageNumber="35">type</typeStatus>
are commonly discontinuous, the axial parenchyma is present, as dispersed cells with slightly nodular horizontal walls and with resin content inside.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE75FFC635AB9" blockId="10.[151,771,1217,2035]" lastBlockId="10.[816,1436,158,645]" pageId="10" pageNumber="35">
Other features are: the relatively tall uniseriate rays with biseriations, up to fusiform aspect, homogeneous and with cross-fields with 1-6(-8) hexagonal to rounded cupressoid oculipores, alternately arranged (araucarioid) to slightly irregular. All these details almost identical to those of the species
<taxonomicName id="7BB14D101D46FFB395F1E7B2FD7A5D9D" box="[392,686,2011,2035]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="35" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D46FFB395F1E7B2FD7A5D9D" box="[392,686,2011,2035]" italics="true" pageId="10" pageNumber="35">Brachyoxylon cristianicum</emphasis>
</taxonomicName>
, which was previously described from the same area (
<bibRefCitation id="D8204B621D46FFB3914EE0F7FC785AB9" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="10" pageNumber="35" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al., 2018</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D46FFB39749E088FB3A5B35" blockId="10.[816,1436,158,645]" pageId="10" pageNumber="35">
From these characteristics, we attribute all the 13 here studied specimens coming from Holbav and Cristian to the above specified form:
<taxonomicName id="7BB14D101D46FFB3900BE14BFB3F5B35" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="35" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D46FFB3900BE14BFA485B54" box="[1138,1436,290,314]" italics="true" pageId="10" pageNumber="35">Brachyoxylon cristianicum</emphasis>
<bibRefCitation id="D8204B621D46FFB39749E12AFB3F5B35" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[816,1259,323,347]" pageId="10" pageNumber="35" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>