149 lines
16 KiB
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149 lines
16 KiB
XML
<document id="AEDE03F4B54A4620CD8EF840551CC994" ID-CLB-Dataset="35372" ID-DOI="http://dx.doi.org/10.3897/zookeys.374.6711" ID-GBIF-Dataset="0024d9ca-9daa-4899-8dfe-6eca9553faa5" ID-PMC="PMC3909809" ID-Pensoft-Pub="1313-2970-374-1" ID-PubMed="24493962" ID-ZooBank="B88BFB082A7B453DAFFB254337D93D85" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-374-1" ModsDocOrigin="ZooKeys 374" ModsDocTitle="The millipede genus Orthomorpha Bollman, 1893 in Laos (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species" checkinTime="1451246400552" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2014" docId="35BEB5B55B2B35AEDD4F351842D3FE2C" docLanguage="en" docName="ZooKeys 374: 1-22" docOrigin="ZooKeys 374" docSource="http://dx.doi.org/10.3897/zookeys.374.6711" docTitle="Orthomorpha cambodjana Attems 1953" docType="treatment" docVersion="5" lastPageNumber="7" masterDocId="E274FFEBB61AFB5D4B2B923FFFA1FFE6" masterDocTitle="The millipede genus Orthomorpha Bollman, 1893 in Laos (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species" masterLastPageNumber="22" masterPageNumber="1" pageNumber="6" updateTime="1701380550218" updateUser="ExternalLinkService">
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<mods:title id="DC600BCE45152460CEE1A9A43852E541">The millipede genus Orthomorpha Bollman, 1893 in Laos (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species</mods:title>
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<mods:namePart id="A650F08EB1F8597518322B5968224BAA">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="42914A540730C06DF25D309F521A0155">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="1F1F5EA0A8782FB821640FFC97640D5F">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="D163012672271B7EB9AA3DD93955FF4D">text</mods:typeOfResource>
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<mods:date id="4EEC211D1315D18E33EA17853BC366D4">2014</mods:date>
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<mods:detail id="673E5D7824880CB36812B6E24FC596AB" type="volume">
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<mods:number id="A2A0C92E1A1EA4E1448A3A4EF4B0D6C5">374</mods:number>
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<mods:url id="653AC03CA2AA69F3E06C174521569BC9">http://dx.doi.org/10.3897/zookeys.374.6711</mods:url>
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<mods:classification id="EA1C50228A6FB4A583D36634E34838EC">journal article</mods:classification>
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<mods:identifier id="7EC665AE15FEE279B8D752FC9CAB46FA" type="DOI">http://dx.doi.org/10.3897/zookeys.374.6711</mods:identifier>
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<treatment id="35BEB5B55B2B35AEDD4F351842D3FE2C" ID-GBIF-Taxon="152050708" LSID="urn:lsid:plazi:treatment:35BEB5B55B2B35AEDD4F351842D3FE2C" httpUri="http://treatment.plazi.org/id/35BEB5B55B2B35AEDD4F351842D3FE2C" lastPageId="7" lastPageNumber="7" pageId="5" pageNumber="6">
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<subSubSection id="F50842B5299F93E4A473C92D7CBD96C0" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="45CF61F39295B399BC8967154D311DD1" pageId="5" pageNumber="6">
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<taxonomicName id="2C6C59790DDC1ABE4BDF0ECA1CFA73B5" ID-CoL="6T8P5" LSID="http://species-id.net/wiki/Orthomorpha_cambodjana" authority="Attems, 1953" authorityName="Attems" authorityYear="1953" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cambodjana">
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<pageBreakToken id="E8BDE660857F62B058231229BE9E9A65" pageId="5" pageNumber="6" start="start">Orthomorpha</pageBreakToken>
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cambodjana (Attems, 1953)
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</taxonomicName>
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Figs 4-6
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</paragraph>
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</subSubSection>
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<subSubSection id="A8E65C45E5658C831C3A3DE900634C5A" pageId="5" pageNumber="6" type="reference_group">
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<paragraph id="02F98FFDDCC53C47168C063E64290F7A" pageId="5" pageNumber="6">
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<taxonomicName id="BDAC479A436A161E9DD313C25AD2220F" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus cambodjanus" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cambodjanus">Pratinus cambodjanus</taxonomicName>
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<bibRefCitation id="E7D69AF004F2C60DB16B77C6987B42E8" author="Attems, C" journalOrPublisher="Memoires du Museum national d' Histoire naturelle, serie A, Zoologie" pageId="20" pageNumber="21" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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: 168 (D).
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</paragraph>
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<paragraph id="5CAF1C55ECDAAB5D65E4CCAE3465649F" pageId="5" pageNumber="6">
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<taxonomicName id="EF06729632622EC707EFAF2DAE334DA8" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cambodjana">Orthomorpha cambodjana</taxonomicName>
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-
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<bibRefCitation id="012290B808683FB50E7C3BCF019AAE05" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="20" pageNumber="21" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M);
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<bibRefCitation id="1E5DA8951631E59B110D4DDE4D8C3780" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="20" pageNumber="21" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
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: 361 (M, D);
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<bibRefCitation id="6B6D3544FE2E125B37397C9CD3D48582" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="20" pageNumber="21" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
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: 56 (M);
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<bibRefCitation id="0268BA5B57B122CAD66C13C75621E17B" author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="20" pageNumber="21" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation id="8F741509C095BA3B32D08480158D30D8" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="20" pageNumber="21" pagination="35 - 46" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda Polydesmida), VI." volume="6" year="1998">Golovatch 1998</bibRefCitation>
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: 42 (M, D);
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<bibRefCitation id="C3337BF5EBDCE62B98710B76F992999D" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="20" pageNumber="21" pagination="1 - 161" title="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)." url="10.3897/zookeys.131.1921" volume="131" year="2011">Likhitrakarn et al. 2011</bibRefCitation>
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: 66 (D).
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</paragraph>
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</subSubSection>
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<subSubSection id="6FE08B06D240B6F641958972A378AA80" pageId="5" pageNumber="6" type="material examined">
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<paragraph id="6619F7EA69B9BAA7BF45CB31733013A4" pageId="5" pageNumber="6">Material examined.</paragraph>
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<paragraph id="666BCC8388F506D9D909BD8FC2AE3CE7" pageId="5" pageNumber="6">
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2 ♂ (CUMZ), Laos, Attapu Province, Xaysetha District, Ban Kasom, 116 m a.s.l.,
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<geoCoordinate id="E52DC2632335597348A8EFECD66F605E" direction="north" orientation="latitude" precision="15" value="15.1075">15°06'27"N</geoCoordinate>
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,
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<geoCoordinate id="0638ABFC12F0E8278906478A15DF2400" direction="east" orientation="longitude" precision="15" value="106.85389">106°51'14"E</geoCoordinate>
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, 16.10.2013, leg. C. Sutcharit and W. Siriwut.
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</paragraph>
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</subSubSection>
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<subSubSection id="592CA8670D8841D77861DFB473DA42A7" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="descriptive notes">
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<paragraph id="7FCCCB888E6B71B5E417D01E9FADB03C" pageId="5" pageNumber="6">Descriptive notes.</paragraph>
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<paragraph id="5CDBFB9898C6B9868A102118A3608BC3" pageId="5" pageNumber="6">Length 32-35 mm (♂), width of midbody pro- and metazona 2.6-2.7 and 3.8-3.9 mm (♂), respectively.</paragraph>
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<paragraph id="19DE8D4E5B52C6AE28A06C64049E7B60" pageId="5" pageNumber="6">
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Coloration in alcohol, after one month of preservation, blackish (Fig. 4
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<normalizedToken id="37BEBD43D8EF8833C3834033048F3842" originalValue="A–F">A-F</normalizedToken>
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), paraterga and epiproct light yellow to whitish yellow; venter, legs and antennae whitish to pale brown distally (Fig. 4
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<normalizedToken id="DC7085A8C15F8C20D031E0CF8448ADB5" originalValue="A–I">A-I</normalizedToken>
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).
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</paragraph>
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<caption id="44205A4F6D5C1E13D254DFE5B689E7DD" pageId="5" pageNumber="6">
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<paragraph id="9B29D061F1E5FB85DC2194145E380E00" pageId="5" pageNumber="6">
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Figure 4.
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<taxonomicName id="9BDC6E740EC22844EABDB6D5F5AC5574" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cambodjana">Orthomorpha cambodjana</taxonomicName>
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(Attems, 1953), ♂. A, B anterior part of body, dorsal and lateral views, respectively; C segments 10-11, dorsal view D segments 9-11, lateral view
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<normalizedToken id="9709D832B92AB80583B14C2BF1EFC648" originalValue="E–G">E-G</normalizedToken>
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posterior part of body, lateral, dorsal and ventral views, respectively; H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<paragraph id="5D54CC749CA8B5955437479752377A4E" pageId="6" pageNumber="7">
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<pageBreakToken id="94708AFA3529C86C4AB861CA5357561C" pageId="6" pageNumber="7" start="start">Antennae</pageBreakToken>
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reaching segment 3 when stretched dorsally. Paraterga very strongly developed (Fig. 4
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<normalizedToken id="4D5B8A527F272300AAD47338283EBD97" originalValue="A–G">A-G</normalizedToken>
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), anterolateral edge clearly convex. Calluses on paraterga 2 and 3 delimited by a sulcus only dorsally, following paraterga by a sulcus both dorsally and ventrally, strongly bordered. Pleurosternal carinae complete crests only in segment 2 (Fig. 4B, D, E), a small, sharp, caudal tooth in segments 3-7, a very faint tubercle until segment 15. Legs long and slender, midbody ones ca 1.1-1.3 times as long as body height, prefemora without modifications, tarsal brushes present until ♂ leg 7.
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</paragraph>
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<paragraph id="D310C7CF03AEE327858252ABBF065590" pageId="6" pageNumber="7">Gonopod solenophore with a tridentate tip (Figs 5, 6), dorsalmost prong being subacuminate, longer than both others, whereas middle denticle shortest and pointed.</paragraph>
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<caption id="580DF439FE5F5A665D19F765107CE7B9" pageId="6" pageNumber="7">
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<paragraph id="04DE2A2C6BFEE1E98FAB2080BE6873D5" pageId="6" pageNumber="7">
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Figure 5.
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<taxonomicName id="4DF85D43DBBC5E9122A4F7A8BD99A2B5" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cambodjana">Orthomorpha cambodjana</taxonomicName>
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(Attems, 1953), ♂, right gonopod. A, B mesal and lateral views, respectively
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<normalizedToken id="00833A9DCBFE1777E899EFBEC3DFA617" originalValue="C–F">C-F</normalizedToken>
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distal part of left gonopod, sublateral, submesal, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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<caption id="3ADB0491F3FC6A691675A4CB451F08B4" pageId="6" pageNumber="7">
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<paragraph id="CAD71A45BD0D37C9E199AF8718E07D42" pageId="6" pageNumber="7">
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Figure 6.
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<taxonomicName id="53878E254CD6C08EE7C0D9E9DF9BAD05" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cambodjana">Orthomorpha cambodjana</taxonomicName>
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(Attems, 1953), ♂. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="ADFB87DBF87B6E60623D71884966B38C" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="remarks">
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<paragraph id="4796161170EDA9136E9853EE0B9EC998" pageId="6" pageNumber="7">Remarks.</paragraph>
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<paragraph id="6BA8276E6F77F8193864235AF74854F8" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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This species has hitherto been known only from a ♂ lectotype and a ♀ paralectotype, currently both rather strongly faded, taken at Sre-Umbell, Kampot
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<pageBreakToken id="5AB1EA55D73CA403733B68CBAEECE6F5" pageId="7" pageNumber="8" start="start">Province</pageBreakToken>
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, southern Cambodia (
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<bibRefCitation id="0462F08C70A6052299FAA071C0DAF65B" author="Attems, C" journalOrPublisher="Memoires du Museum national d' Histoire naturelle, serie A, Zoologie" pageId="20" pageNumber="21" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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,
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<bibRefCitation id="7948B3A4A3CAEC2FA8266D5F2AF0620E" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="20" pageNumber="21" pagination="1 - 161" title="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)." url="10.3897/zookeys.131.1921" volume="131" year="2011">Likhitrakarn et al. 2011</bibRefCitation>
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). The above fresh samples show only slight variations, mainly a darker coloration, as opposed to the types which have recently been redescribed in due detail (
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<bibRefCitation id="69365743D6C5D20E428F719F8CCB0937" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="20" pageNumber="21" pagination="1 - 161" title="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)." url="10.3897/zookeys.131.1921" volume="131" year="2011">Likhitrakarn et al. 2011</bibRefCitation>
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). This species is thus new to the fauna of Laos, suggesting a far wider distribution in Indochina.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |