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<document id="12C7E0AB0092F33CE389F8FE634E7121" ID-CLB-Dataset="4642" ID-DOI="10.5281/zenodo.5320906" ID-GBIF-Dataset="f33526c4-a22f-4c32-9fd9-34a1255f44eb" ID-ISSN="0374-1036" ID-Zenodo-Dep="5320906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630136926086" checkinUser="felipe" docAuthor="Ito, Noboru" docDate="2009" docId="352E879FFF94FFC9FE85FB5CFF7DFE53" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.599-606.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Trichotichnus (Pseudotrichotichnus) tenuitibialis Ito 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="606" masterDocId="C917FFE7FF91FFCEFF99FF9AFFBBFFC7" masterDocTitle="Note on the subgenus Pseudotrichotichnus of the genus Trichotichnus from Southeast Asia (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="606" masterPageNumber="599" pageNumber="604" updateTime="1699168252378" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="AD9BD9F637BA48C46C35FC365E03BCA8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A2C6834CAA4EA88F0A77E1EDF9A5C4F1">
<mods:title id="46563FF5FF41E3FE72F600B3A10D39A3">Note on the subgenus Pseudotrichotichnus of the genus Trichotichnus from Southeast Asia (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
<mods:name id="C3017FA99F6572760DDAE1A3D9F914BD" type="personal">
<mods:role id="4158791FB2A5C9A193883F11EE460D76">
<mods:roleTerm id="8AA747675238A353793CEC751E69E435">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B80BF92B5F4A629FA02A3078134F4BCF">Ito, Noboru</mods:namePart>
</mods:name>
<mods:typeOfResource id="4AA3C443E4B0BCD239B438A014B2DEE5">text</mods:typeOfResource>
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<mods:titleInfo id="7C304F5208D2FA0F2821DE3D021C19B5">
<mods:title id="707F37741D7202ACDBD20F7FA6F2D69E">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="523A056288CEA66B7AFBA80CFA428ACD">
<mods:date id="D76203009C3206968D3CF23D1DDBFF14">2009</mods:date>
<mods:detail id="AB5A325C33C70D028DF5E78231CD805F" type="pubDate">
<mods:number id="FB6FB9162D7B32C0C607B09E9BEB57DD">2009-12-15</mods:number>
</mods:detail>
<mods:detail id="B15B9CADC44671C51C7BA516B21C974E" type="volume">
<mods:number id="67EE2063177F6E9833F533A581E8D4B6">49</mods:number>
</mods:detail>
<mods:detail id="B361B9B0956BC5394BFAC5D792EBF2C0" type="issue">
<mods:number id="4C137C6A1730CACD66732101CD139D87">2</mods:number>
</mods:detail>
<mods:extent id="0797141F3BCC03D42301FF461EE4B58F" unit="page">
<mods:start id="5160DC53F6D604DE9AFE9B661B107D2E">599</mods:start>
<mods:end id="017B6D865708720C4A3BB272882AFEA3">606</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="051201483FE657DAB9E90EB1C84CCBA5">journal article</mods:classification>
<mods:identifier id="49B763BB311025F83432075611DAE929" type="CLB-Dataset">4642</mods:identifier>
<mods:identifier id="C1A8C5463CFB2B32406980C4ADD87493" type="DOI">10.5281/zenodo.5320906</mods:identifier>
<mods:identifier id="2491A35800258FA408ECE183D37451B5" type="GBIF-Dataset">f33526c4-a22f-4c32-9fd9-34a1255f44eb</mods:identifier>
<mods:identifier id="457E2B7FC11070C41501203043ADFB36" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="73F3F03BE084F101F26287017B476146" type="Zenodo-Dep">5320906</mods:identifier>
</mods:mods>
<treatment id="352E879FFF94FFC9FE85FB5CFF7DFE53" ID-DOI="http://doi.org/10.5281/zenodo.5342764" ID-GBIF-Taxon="185202415" ID-Zenodo-Dep="5342764" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53" httpUri="http://treatment.plazi.org/id/352E879FFF94FFC9FE85FB5CFF7DFE53" lastPageId="7" lastPageNumber="606" pageId="5" pageNumber="604">
<subSubSection id="F59D6502FF94FFCBFE85FB5CFC2CFB18" box="[284,919,1222,1247]" pageId="5" pageNumber="604" type="nomenclature">
<paragraph id="BD383689FF94FFCBFE85FB5CFC2CFB18" blockId="5.[284,919,1222,1247]" box="[284,919,1222,1247]" pageId="5" pageNumber="604">
<heading id="E67081E5FF94FFCBFE85FB5CFC2CFB18" bold="true" box="[284,919,1222,1247]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="604" reason="3">
<emphasis id="8FF3EA9BFF94FFCBFE85FB5CFC2CFB18" bold="true" box="[284,919,1222,1247]" pageId="5" pageNumber="604">
<taxonomicName id="7A874D0AFF94FFCBFE85FB5CFC81FB18" ID-CoL="58LJ7" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[284,826,1222,1247]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="tenuitibialis" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF94FFCBFE85FB5CFE0CFB18" bold="true" box="[284,439,1222,1247]" italics="true" pageId="5" pageNumber="604">Trichotichnus</emphasis>
(
<emphasis id="8FF3EA9BFF94FFCBFE5FFB5CFD12FB18" bold="true" box="[454,681,1222,1247]" italics="true" pageId="5" pageNumber="604">Pseudotrichotichnus</emphasis>
)
<emphasis id="8FF3EA9BFF94FFCBFD21FB5CFC81FB18" bold="true" box="[696,826,1222,1247]" italics="true" pageId="5" pageNumber="604">tenuitibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF94FFCBFCD8FB5DFC2CFB18" box="[833,919,1223,1247]" pageId="5" pageNumber="604" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF94FFCBFDB3FB7FFDDCFADD" pageId="5" pageNumber="604" type="description">
<paragraph id="BD383689FF94FFCBFDB3FB7FFD33FB3F" blockId="5.[113,1089,1253,1509]" box="[554,648,1253,1272]" pageId="5" pageNumber="604">
(
<figureCitation id="25BC2A0CFF94FFCBFDA9FB7FFDD6FB3F" box="[560,621,1253,1272]" captionStart="Figs" captionStartId="2.[153,192,1499,1518]" captionTargetBox="[168,1126,250,1426]" captionTargetId="figure-10@2.[151,1131,219,1426]" captionTargetPageId="2" captionText="Figs. 1–5. 1–2 – Trichotichnus (Pseudtrichotichnus) jelineki sp. nov., holotype (1 – habitus; 2 – male genitalia); 3 – T. (P.) vietnamensis sp. nov.; 4 – T. (P.) tenuitibialis sp. nov.; 5 – T. (P.) niger Louwerens, 1951. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320908" httpUri="https://zenodo.org/record/5320908/files/figure.png" pageId="5" pageNumber="604">Figs. 4</figureCitation>
,
<figureCitation id="25BC2A0CFF94FFCBFDEFFB7FFD3AFB3F" box="[630,641,1253,1272]" captionStart="Figs" captionStartId="3.[111,150,1072,1091]" captionTargetBox="[139,1068,228,1054]" captionTargetId="figure-14@3.[131,1070,219,1069]" captionTargetPageId="3" captionText="Figs. 6–9. Male genitalia. 6 – Trichotichnus (Pseudotrichotichnus) vietnamensis sp. nov.; 7 – T. (P.) tenuitibialis sp. nov.; 8 – T. (P.) niger Louwerens, 1951; 9 – T. (P.) miyakei Habu, 1980. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="5" pageNumber="604">7</figureCitation>
)
</paragraph>
<paragraph id="BD383689FF94FFCBFFE8FA92FDDCFADD" blockId="5.[113,1089,1253,1509]" box="[113,615,1287,1307]" pageId="5" pageNumber="604">
<taxonomicName id="7A874D0AFF94FFCBFFE8FA92FE2DFADC" authority=": ITO (1991)" authorityName=": ITO" authorityYear="1991" box="[113,406,1288,1307]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF94FFCBFFE8FA92FE95FADD" box="[113,302,1288,1306]" italics="true" pageId="5" pageNumber="604">Trichotichnus miyakei</emphasis>
: ITO (1991)
</taxonomicName>
: 159 (misidentification).
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF94FFCBFFE8FAABFECBFA23" pageId="5" pageNumber="604" type="materials_examined">
<paragraph id="BD383689FF94FFCBFFE8FAABFECBFA23" blockId="5.[113,1089,1253,1509]" pageId="5" pageNumber="604">
<emphasis id="8FF3EA9BFF94FFCBFFE8FAABFF4EFA83" bold="true" box="[113,245,1328,1348]" pageId="5" pageNumber="604">Type material.</emphasis>
<materialsCitation id="0DEF3CD4FF94FFCBFF63FAA8FE7AFA98" ID-GBIF-Occurrence="3339964318" collectingDate="1995-03-21" collectingDateMax="1995-03-30" collectingDateMin="1995-03-21" collectionCode="HNHM" collectorName="O. Merkl" country="Malaysia" location="Tanah Rata" municipality="Cameron Highlands" pageId="5" pageNumber="604" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<typeStatus id="623C882BFF94FFCBFF63FAA8FEF6FA84" box="[250,333,1330,1349]" pageId="5" pageNumber="604" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<location id="B8586052FF94FFCBFEEDFAABFE61FA82" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53:B8586052FF94FFCBFEEDFAABFE61FA82" box="[372,474,1329,1349]" country="Malaysia" municipality="Cameron Highlands" name="Tanah Rata" pageId="5" pageNumber="604" stateProvince="Pahang">Tanah Rata</location>
, singled in and around the town,
<collectingMunicipality id="5D5CACF3FF94FFCBFD69FAABFC26FA83" box="[752,925,1329,1348]" pageId="5" pageNumber="604">Cameron Highlands</collectingMunicipality>
,
<collectingRegion id="7F43F86BFF94FFCBFC3DFAABFC5EFA83" box="[932,997,1329,1348]" country="Malaysia" name="Pahang" pageId="5" pageNumber="604">Pahang</collectingRegion>
,
<collectingCountry id="C5907619FF94FFCBFC74FAABFB86FA83" box="[1005,1085,1329,1348]" name="Malaysia" pageId="5" pageNumber="604">Malaysia</collectingCountry>
,
<date id="C9391049FF94FFCBFFE8FAD6FF49FA98" box="[113,242,1356,1375]" pageId="5" pageNumber="604" value="1995-03-21" valueMax="1995-03-30" valueMin="1995-03-21">
<collectingDate id="D97DE9A1FF94FFCBFFE8FAD6FF49FA98" box="[113,242,1356,1375]" pageId="5" pageNumber="604" value="1995-03-21" valueMax="1995-03-30" valueMin="1995-03-21">21.–30.iii.1995</collectingDate>
</date>
,
<collectorName id="1072535FFF94FFCBFF60FAD6FEFFFA98" box="[249,324,1356,1375]" pageId="5" pageNumber="604">O.Merkl</collectorName>
leg.’ (
<collectionCode id="DB96AE4CFF94FFCBFEEFFAD6FE00FA98" box="[374,443,1356,1375]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="604">HNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DEF3CD4FF94FFCBFE5EFAD6FF2DFABE" ID-GBIF-Occurrence="3339964313" collectingDate="1974-04-07" collectorName="Y. Kiyoyama" country="Malaysia" location="Cameron Hi" municipality="Tanah Rata" pageId="5" pageNumber="604" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="623C882BFF94FFCBFE5EFAD6FDA6FA99" box="[455,541,1356,1375]" pageId="5" pageNumber="604" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="AB81FD00FF94FFCBFDBFFAD6FDF8FA98" box="[550,579,1350,1375]" pageId="5" pageNumber="604" type="female">1♀</specimenCount>
, ‘
<location id="B8586052FF94FFCBFDC9FAD6FD07FA98" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53:B8586052FF94FFCBFDC9FAD6FD07FA98" box="[592,700,1356,1375]" country="Malaysia" municipality="Tanah Rata" name="Cameron Hi" pageId="5" pageNumber="604">Cameron Hi</location>
,
<collectingMunicipality id="5D5CACF3FF94FFCBFD5BFAD6FC99FA98" box="[706,802,1356,1375]" pageId="5" pageNumber="604">Tanah Rata</collectingMunicipality>
,
<collectingCountry id="C5907619FF94FFCBFCB1FAD6FCCCFA98" box="[808,887,1356,1375]" name="Malaysia" pageId="5" pageNumber="604">Malaysia</collectingCountry>
,
<date id="C9391049FF94FFCBFCE7FAD6FC75FA98" box="[894,974,1356,1375]" pageId="5" pageNumber="604" value="1974-04-07">
<collectingDate id="D97DE9A1FF94FFCBFCE7FAD6FC75FA98" box="[894,974,1356,1375]" pageId="5" pageNumber="604" value="1974-04-07">7.iv.1974</collectingDate>
</date>
,
<collectorName id="1072535FFF94FFCBFC4DFAD6FBFAFA98" box="[980,1089,1356,1375]" pageId="5" pageNumber="604">Y.Kiyoyama</collectorName>
leg.
</materialsCitation>
’;
<materialsCitation id="0DEF3CD4FF94FFCBFF3DFAFDFD01FABE" ID-GBIF-Occurrence="3339964319" box="[164,698,1377,1402]" collectingDate="1974-03-05" collectorName="Y. Kiyoyama" country="Malaysia" location="Maxwell's Hill" pageId="5" pageNumber="604" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="AB81FD00FF94FFCBFF3DFAFDFF7EFABD" box="[164,197,1377,1402]" pageId="5" pageNumber="604" type="female">1 ♀</specimenCount>
, ‘
<location id="B8586052FF94FFCBFF4AFAFCFEE1FABE" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53:B8586052FF94FFCBFF4AFAFCFEE1FABE" box="[211,346,1382,1401]" country="Malaysia" name="Maxwell's Hill" pageId="5" pageNumber="604">Maxwell’s Hill</location>
,
<collectingCountry id="C5907619FF94FFCBFEFDFAFCFE0FFABE" box="[356,436,1382,1401]" name="Malaysia" pageId="5" pageNumber="604">Malaysia</collectingCountry>
,
<date id="C9391049FF94FFCBFE27FAFCFDAEFABE" box="[446,533,1382,1401]" pageId="5" pageNumber="604" value="1974-03-05">
<collectingDate id="D97DE9A1FF94FFCBFE27FAFCFDAEFABE" box="[446,533,1382,1401]" pageId="5" pageNumber="604" value="1974-03-05">5.III.1974</collectingDate>
</date>
,
<collectorName id="1072535FFF94FFCBFD87FAFDFD34FABE" box="[542,655,1382,1402]" pageId="5" pageNumber="604">Y. Kiyoyama</collectorName>
leg.
</materialsCitation>
’;
<materialsCitation id="0DEF3CD4FF94FFCBFD5EFAFDFEE9FA53" ID-GBIF-Occurrence="3339964320" collectingDate="1975-05-29" collectionCode="NIKJ" collectorName="Y. Kiyoyama" country="Malaysia" location="Tanah Rata" pageId="5" pageNumber="604" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
1 J
<specimenCount id="AB81FD00FF94FFCBFD77FAFDFCB4FABD" box="[750,783,1377,1402]" pageId="5" pageNumber="604" type="female">1 ♀</specimenCount>
, ‘
<location id="B8586052FF94FFCBFC84FAFCFC3FFABD" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53:B8586052FF94FFCBFC84FAFCFC3FFABD" box="[797,900,1382,1402]" country="Malaysia" name="Tanah Rata" pageId="5" pageNumber="604">Tanah Rata</location>
,
<collectingCountry id="C5907619FF94FFCBFC17FAFCFC65FABE" box="[910,990,1382,1401]" name="Malaysia" pageId="5" pageNumber="604">Malaysia</collectingCountry>
,
<date id="C9391049FF94FFCBFC7EFAFCFB86FABE" box="[999,1085,1382,1401]" pageId="5" pageNumber="604" value="1975-05-29">
<collectingDate id="D97DE9A1FF94FFCBFC7EFAFCFB86FABE" box="[999,1085,1382,1401]" pageId="5" pageNumber="604" value="1975-05-29">29.v.1975</collectingDate>
</date>
,
<collectorName id="1072535FFF94FFCBFFE8FA18FF59FA53" box="[113,226,1409,1429]" pageId="5" pageNumber="604">Y. Kiyoyama</collectorName>
leg.’ (
<collectionCode id="DB96AE4CFF94FFCBFE83FA1BFEF6FA53" box="[282,333,1409,1428]" pageId="5" pageNumber="604">NIKJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF94FFCBFEC5FA1BFD5FFA68" ID-GBIF-Occurrence="3339964312" collectingDate="1995-03-29" collectorName="O. Merkl &amp; I. Szikossy" country="Malaysia" location="Cameron Highlands" pageId="5" pageNumber="604" specimenCount="4" specimenCount-female="4" stateProvince="Pahang" typeStatus="paratype">
1 J
<specimenCount id="AB81FD00FF94FFCBFE1BFA1BFE09FA53" box="[386,434,1403,1428]" pageId="5" pageNumber="604" type="female">4 ♀♀</specimenCount>
, ‘
<locationDeviation id="AC8A2FABFF94FFCBFE5BFA1BFD4FFA53" box="[450,756,1409,1429]" location="Tanah Rata on Tapah Road" pageId="5" pageNumber="604">
<quantity id="7A7F9B6CFF94FFCBFE5BFA1BFE4AFA53" box="[450,497,1409,1428]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="604" unit="km" value="2.0">2 km</quantity>
S Tanah Rata on Tapah Road
</locationDeviation>
, montane rainforest, at light,
<location id="B8586052FF94FFCBFC6BFA1BFF70FA68" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53:B8586052FF94FFCBFC6BFA1BFF70FA68" country="Malaysia" name="Cameron Highlands" pageId="5" pageNumber="604" stateProvince="Pahang">Cameron Highlands</location>
,
<collectingRegion id="7F43F86BFF94FFCBFF4FFA06FEACFA68" box="[214,279,1436,1455]" country="Malaysia" name="Pahang" pageId="5" pageNumber="604">Pahang</collectingRegion>
,
<collectingCountry id="C5907619FF94FFCBFEBBFA06FEC9FA68" box="[290,370,1436,1455]" name="Malaysia" pageId="5" pageNumber="604">Malaysia</collectingCountry>
,
<date id="C9391049FF94FFCBFEE4FA06FE67FA68" box="[381,476,1436,1455]" pageId="5" pageNumber="604" value="1995-03-29">
<collectingDate id="D97DE9A1FF94FFCBFEE4FA06FE67FA68" box="[381,476,1436,1455]" pageId="5" pageNumber="604" value="1995-03-29">29.iii.1995</collectingDate>
</date>
,
<collectorName id="1072535FFF94FFCBFE7EFA06FD8CFA68" box="[487,567,1436,1455]" pageId="5" pageNumber="604">O. Merkl</collectorName>
&amp;
<collectorName id="1072535FFF94FFCBFDCEFA06FD03FA68" box="[599,696,1436,1455]" pageId="5" pageNumber="604">I. Szikossy</collectorName>
leg.
</materialsCitation>
’;
<materialsCitation id="0DEF3CD4FF94FFCBFD6AFA06FCCCFA0E" ID-GBIF-Occurrence="3339964317" collectingDate="1995-03-21" collectingDateMax="1995-04-02" collectingDateMin="1995-03-21" collectorName="O. Merkl" country="Malaysia" location="Tanah Rata" municipality="Cameron Highlands" pageId="5" pageNumber="604" specimenCount="5" specimenCount-female="5" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="AB81FD00FF94FFCBFD6AFA06FC9EFA68" box="[755,805,1430,1455]" pageId="5" pageNumber="604" type="female">5 ♀♀</specimenCount>
, ‘
<location id="B8586052FF94FFCBFCACFA06FC25FA68" LSID="urn:lsid:plazi:treatment:352E879FFF94FFC9FE85FB5CFF7DFE53:B8586052FF94FFCBFCACFA06FC25FA68" box="[821,926,1436,1455]" country="Malaysia" municipality="Cameron Highlands" name="Tanah Rata" pageId="5" pageNumber="604" stateProvince="Pahang">Tanah Rata</location>
, edge of degraded rainforest, at light,
<collectingMunicipality id="5D5CACF3FF94FFCBFE8AFA2DFE7BFA0E" box="[275,448,1462,1482]" pageId="5" pageNumber="604">Cameron Highlands</collectingMunicipality>
,
<collectingRegion id="7F43F86BFF94FFCBFE51FA2CFDB2FA0E" box="[456,521,1462,1481]" country="Malaysia" name="Pahang" pageId="5" pageNumber="604">Pahang</collectingRegion>
,
<collectingCountry id="C5907619FF94FFCBFD88FA2CFDDAFA0E" box="[529,609,1462,1481]" name="Malaysia" pageId="5" pageNumber="604">Malaysia</collectingCountry>
,
<date id="C9391049FF94FFCBFDF3FA2CFD4CFA0E" box="[618,759,1462,1481]" pageId="5" pageNumber="604" value="1995-03-21" valueMax="1995-04-02" valueMin="1995-03-21">
<collectingDate id="D97DE9A1FF94FFCBFDF3FA2CFD4CFA0E" box="[618,759,1462,1481]" pageId="5" pageNumber="604" value="1995-03-21" valueMax="1995-04-02" valueMin="1995-03-21">21.iii.–2.iv.1995</collectingDate>
</date>
,
<collectorName id="1072535FFF94FFCBFD66FA2DFCF6FA0E" box="[767,845,1462,1482]" pageId="5" pageNumber="604">O. Merkl</collectorName>
leg.
</materialsCitation>
’;
<materialsCitation id="0DEF3CD4FF94FFCBFC1DFA2DFED7FA23" ID-GBIF-Occurrence="3339964306" collectingDate="1995-03-21" collectingDateMax="1995-04-02" collectingDateMin="1995-03-21" collectionCode="HNHM" collectorName="O. Merkl" country="Malaysia" location="Tanah Rata" municipality="Cameron Highlands" pageId="5" pageNumber="604" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="AB81FD00FF94FFCBFC1DFA2DFC19FA0D" box="[900,930,1457,1482]" pageId="5" pageNumber="604" type="female">1 ♀</specimenCount>
, the same locality,
<collectingMethod id="64C64E9EFF94FFCBFFE8FA4BFEAEFA22" box="[113,277,1489,1509]" pageId="5" pageNumber="604">flight intercept trap</collectingMethod>
(
<collectionCode id="DB96AE4CFF94FFCBFEB9FA4BFEDDFA23" box="[288,358,1489,1508]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="604">HNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF94FFC8FFE8FA66FDC6FA34" lastPageId="6" lastPageNumber="605" pageId="5" pageNumber="604" type="description">
<paragraph id="BD383689FF94FFCBFFE8FA66FC11F992" blockId="5.[113,1089,1532,1621]" pageId="5" pageNumber="604">
<emphasis id="8FF3EA9BFF94FFCBFFE8FA66FF47F9D3" bold="true" box="[113,252,1532,1556]" pageId="5" pageNumber="604">Description.</emphasis>
Body similar in shape to
<taxonomicName id="7A874D0AFF94FFCBFD95FA64FDC0F9D2" authorityName=": ITO" authorityYear="1991" box="[524,635,1533,1557]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF94FFCBFD95FA64FDA0F9D2" box="[524,539,1534,1557]" italics="true" pageId="5" pageNumber="604">T</emphasis>
.
<emphasis id="8FF3EA9BFF94FFCBFDB1FA64FDC0F9D2" box="[552,635,1534,1557]" italics="true" pageId="5" pageNumber="604">miyakei</emphasis>
</taxonomicName>
, pitchy black though a little lighter than in
<taxonomicName id="7A874D0AFF94FFCBFFE8F984FF5AF9F2" authorityName=": ITO" authorityYear="1991" box="[113,225,1565,1589]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF94FFCBFFE8F984FF3BF9F2" box="[113,128,1566,1589]" italics="true" pageId="5" pageNumber="604">T</emphasis>
.
<emphasis id="8FF3EA9BFF94FFCBFF17F984FF5AF9F2" box="[142,225,1566,1589]" italics="true" pageId="5" pageNumber="604">miyakei</emphasis>
</taxonomicName>
, iridescent on elytra; buccal parts, antennae and tarsi light brown, labrum, tibiae and femora reddish brown to light reddish brown, mandibles blackish brown.
</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFF41FE1EFDF4" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">Head more or less large, 0.69–0.71 times as wide as pronotum, very sparsely and microscopically punctate, moderately raised; labrum subquadrate, with apex almost straight or shallowly emarginate; clypeus weakly protruding at apical angles, sometimes slightly swollen behind anterior margin; clypeal suture deep, slightly sinuate in middle; frontal impressions also deepened throughout, straight, oblique; eyes not prominent; temples abruptly declined, one-third the eye length; buccal fissure widely separated from genuine ventral margin of eyes; antennae short, not reaching pronotal base, submoniliform, segment 3 as long as segment 4 and twice as long as segment 2; palpi slender, segment 3 of labial palpus 1.1 times as long as segment 2; suture of mentum not distinct only near lateral ends of submentum, median tooth narrow and pointed at apex, epilobes narrow and parallel-sided; microsculpture invisible under 80× magnification.</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFDA6FBB6FCB4" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">
Pronotum gently convex, transversely subquadrate, widest at anterior third, 1.5 times as wide as long, at base 1.25 times as wide as on anterior margin, sides moderately arcuate in anterior half and straight in posterior half; surface mostly impunctate, punctures in basal foveae more sparse and finer than in
<taxonomicName id="7A874D0AFF97FFC8FE5CFD07FD8BFD73" authorityName=": ITO" authorityYear="1991" box="[453,560,668,692]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="605" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF97FFC8FE5CFD07FE6FFD73" box="[453,468,669,692]" italics="true" pageId="6" pageNumber="605">T</emphasis>
.
<emphasis id="8FF3EA9BFF97FFC8FE46FD07FD8BFD73" box="[479,560,669,692]" italics="true" pageId="6" pageNumber="605">miyakei</emphasis>
</taxonomicName>
; anterior margin slightly emarginate, without border in middle; basal margin bordered only at sides or with broken border throughout; anterior angles weakly produced, clearly arcuate; basal angles angulate, slightly obtuse; lateral furrows wholly impressed in a line; basal foveae shallow or only flattened, very obscurely grooved at inner sides; front transverse impression very shallow, hind one more vague; median line distinct to more or less vague, not reaching both anterior and basal margin; microsculpture obscure, consisting of transverse meshes and lines on disc and of isodiametric meshes near basal punctures.
</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFCE1FE78FBB4" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">Elytra oblong, suboval, 1.25 times as wide as pronotum and 1.50–1.53 times as long as wide, uniformly convex; sides gently rounded behind humeri, subarcuate near middle and rounded in apical third; apices widely rounded, tips narrowly separated from each other; base almost straight, rounded at humeral angles; striae wide, deep and clearly crenulate, scutellar striole long; intervals almost flat or weakly elevated on disc, fairly convex apically and basally, interval 3 with discal pore near apical two-fifths; marginal series widely spaced, consisting of (8–9) + (9–11) umbilicate pores; microsculpture sparsely visible as transverse thin lines. Hind wings well developed.</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFBE1FCBEFB14" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">Ventral surface almost smooth; metepisterna convergent behind, 1.6 times as long as wide; apical margin of abdominal sternite 6 truncate or very slightly emarginate in male and narrowly rounded in female, with four setae in both sexes.</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFB46FC84FAB4" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">Legs slender; fore tibiae weakly dilated apicad, not sulcate in apical fifth, with three spines along external margin near apex; hind tarsi short, 0.78 times as long as the width of head in male and 0.64–0.66 times as long in female, segment 1 0.79 times as long as segments 2 and 3 taken together, segment 2 1.25 times as long as segment 3 and twice as long as segment 4, all claws with two setae on both inner and outer ventral margin.</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFAE1FED0FA14" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">
Aedeagus (
<figureCitation id="25BC2A0CFF97FFC8FEB6FAE1FECBFA53" box="[303,368,1403,1428]" captionStart="Figs" captionStartId="3.[111,150,1072,1091]" captionTargetBox="[139,1068,228,1054]" captionTargetId="figure-14@3.[131,1070,219,1069]" captionTargetPageId="3" captionText="Figs. 6–9. Male genitalia. 6 – Trichotichnus (Pseudotrichotichnus) vietnamensis sp. nov.; 7 – T. (P.) tenuitibialis sp. nov.; 8 – T. (P.) niger Louwerens, 1951; 9 – T. (P.) miyakei Habu, 1980. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="6" pageNumber="605">Fig. 7</figureCitation>
) large, stout, dorsal surface straight in middle in lateral aspect; apical orifice not wide, opening oblique, placed laterally, inner sac without sclerite; apical lobe wide, subtriangular.
</paragraph>
<paragraph id="BD383689FF97FFC8FF2CFA41FDC6FA34" blockId="6.[151,1127,219,1619]" box="[181,637,1499,1524]" pageId="6" pageNumber="605">
Length: 7.8–10.0 mm; width:
<quantity id="7A7F9B6CFF97FFC8FE6CFA46FDC2FA34" box="[501,633,1500,1524]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="6" pageNumber="605" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.7–4.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF97FFC8FF0EFA61FE5BF994" pageId="6" pageNumber="605" type="diagnosis">
<paragraph id="BD383689FF97FFC8FF0EFA61FE5BF994" blockId="6.[151,1127,219,1619]" pageId="6" pageNumber="605">
<emphasis id="8FF3EA9BFF97FFC8FF0EFA61FE28F9D4" bold="true" box="[151,403,1531,1555]" pageId="6" pageNumber="605">Differential diagnosis.</emphasis>
This new species is allied to
<taxonomicName id="7A874D0AFF97FFC8FD4EFA67FCC8F9D3" authorityName=": ITO" authorityYear="1991" box="[727,883,1532,1556]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="605" phylum="Arthropoda" rank="species" species="miyakei" subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF97FFC8FD4EFA67FD50F9D3" box="[727,747,1533,1556]" italics="true" pageId="6" pageNumber="605">T.</emphasis>
(
<emphasis id="8FF3EA9BFF97FFC8FD62FA67FCABF9D3" box="[763,784,1533,1556]" italics="true" pageId="6" pageNumber="605">P.</emphasis>
)
<emphasis id="8FF3EA9BFF97FFC8FCB9FA67FCC8F9D3" box="[800,883,1533,1556]" italics="true" pageId="6" pageNumber="605">miyakei</emphasis>
</taxonomicName>
, but in addition to the characters mentioned in the description, the body is smaller and the aedeagus is not arcuate dorsally but straight in middle.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF96FFC9FFE8FF46FDFDFED3" pageId="7" pageNumber="606" type="etymology">
<paragraph id="BD383689FF96FFC9FFE8FF46FDFDFED3" blockId="7.[113,1089,220,404]" pageId="7" pageNumber="606">
<emphasis id="8FF3EA9BFF96FFC9FFE8FF46FF4AFF33" bold="true" box="[113,241,220,244]" pageId="7" pageNumber="606">Etymology.</emphasis>
The species name is composed from the Latin
<emphasis id="8FF3EA9BFF96FFC9FD4BFF47FCAAFF33" box="[722,785,221,244]" italics="true" pageId="7" pageNumber="606">tenuis</emphasis>
(= slim, slender) and
<emphasis id="8FF3EA9BFF96FFC9FC68FF47FB81FF33" box="[1009,1082,221,244]" italics="true" pageId="7" pageNumber="606">tibialis</emphasis>
, referring to the slender tibiae of the species.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF96FFC9FFE8FE81FCD7FEF3" box="[113,876,283,308]" pageId="7" pageNumber="606" type="distribution">
<paragraph id="BD383689FF96FFC9FFE8FE81FCD7FEF3" blockId="7.[113,1089,220,404]" box="[113,876,283,308]" pageId="7" pageNumber="606">
<emphasis id="8FF3EA9BFF96FFC9FFE8FE81FEBFFEF4" bold="true" box="[113,260,283,307]" pageId="7" pageNumber="606">Distribution.</emphasis>
Cameron Highlands (Peninsular
<collectingCountry id="C5907619FF96FFC9FDFEFE86FD70FEF3" box="[615,715,284,308]" name="Malaysia" pageId="7" pageNumber="606">Malaysia</collectingCountry>
,
<collectingRegion id="7F43F86BFF96FFC9FD4FFE86FCDBFEF3" box="[726,864,284,308]" country="Malaysia" name="Pahang" pageId="7" pageNumber="606">Pahang state</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF96FFC9FFE8FEA6FF7DFE53" pageId="7" pageNumber="606" type="discussion">
<paragraph id="BD383689FF96FFC9FFE8FEA6FF7DFE53" blockId="7.[113,1089,220,404]" pageId="7" pageNumber="606">
<emphasis id="8FF3EA9BFF96FFC9FFE8FEA6FF6DFE93" bold="true" box="[113,214,316,340]" pageId="7" pageNumber="606">Remark.</emphasis>
I treated the Malaysian specimens as
<taxonomicName id="7A874D0AFF96FFC9FDEFFEA7FCD9FE93" authority="(ITO 1991)" baseAuthorityName="ITO" baseAuthorityYear="1991" box="[630,866,316,340]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="606" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF96FFC9FDEFFEA7FD3EFE93" box="[630,645,317,340]" italics="true" pageId="7" pageNumber="606">T</emphasis>
.
<emphasis id="8FF3EA9BFF96FFC9FD0DFEA7FD5CFE93" box="[660,743,317,340]" italics="true" pageId="7" pageNumber="606">miyakei</emphasis>
(ITO 1991)
</taxonomicName>
, but a more detailed examination including the study of the aedeagus revealed that they belong to a separate species.
</paragraph>
</subSubSection>
</treatment>
</document>
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