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<mods:title id="E4BE263116D6BE31150BBA2C8E6BBDBF">First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species</mods:title>
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<paragraph id="AD3C368BFF96A17FFF2FFF13FE10C6DF" blockId="1.[151,439,221,247]" box="[151,439,221,247]" pageId="1" pageNumber="53">
<heading id="F67481E7FF96A17FFF2FFF13FE10C6DF" bold="true" box="[151,439,221,247]" fontSize="11" level="1" pageId="1" pageNumber="53" reason="1">
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Genus
<taxonomicName id="6A834D08FF96A17FFF56FF13FE10C6DF" ID-CoL="62X23" authority="Walker" authorityName="Walker" box="[238,439,221,247]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF96A17FFF56FF13FEF2C6DF" bold="true" box="[238,341,221,247]" italics="true" pageId="1" pageNumber="53">Leusaba</emphasis>
Walker
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<subSubSection id="E5996500FF96A17FFF2FFEEBFD04C714" box="[151,675,293,316]" pageId="1" pageNumber="53" type="reference_group">
<paragraph id="AD3C368BFF96A17FFF2FFEEBFD04C714" blockId="1.[151,982,293,348]" box="[151,675,293,316]" pageId="1" pageNumber="53">
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<taxonomicName id="6A834D08FF96A17FFF2FFEEBFE18C714" ID-CoL="62X23" authority="Walker, 1857: 144" authorityName="Walker" authorityPageNumber="144" authorityYear="1857" box="[151,447,293,316]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF96A17FFF2FFEEBFF56C714" box="[151,241,293,316]" italics="true" pageId="1" pageNumber="53">Leusaba</emphasis>
<treatmentCitation id="2C22109AFF96A17FFF40FEEBFE18C714" author="Walker" box="[248,447,293,316]" page="144" pageId="1" pageNumber="53" year="1857">
<bibRefCitation id="C9124B7AFF96A17FFF40FEEBFE2EC714" author="Walker" box="[248,393,293,316]" pageId="1" pageNumber="60" refString="Walker, F. (1857) Catalogue of the Homopterous Insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with Descriptions of New Species. Journal of the Proceedings of the Linnean Society. London, 1, 141 - 175." type="journal article" year="1857">Walker, 1857</bibRefCitation>
: 144
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</taxonomicName>
;
<treatmentCitation id="2C22109AFF96A17FFE72FEEBFD38C714" author="Melichar" box="[458,671,293,316]" page="105" pageId="1" pageNumber="53" year="1914">
<bibRefCitation id="C9124B7AFF96A17FFE72FEEBFDCDC714" author="Melichar" box="[458,618,293,316]" pageId="1" pageNumber="60" refString="Melichar, L. (1914) Monographie der Tropiduchinen (Homoptera). Verhandlungen des Naturforschenden Vereins in Brunn, 53, 1 - 145." type="journal article" year="1914">Melichar 1914</bibRefCitation>
: 105
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="E5996500FF96A17FFF2FFE8BFC71C774" box="[151,982,325,348]" pageId="1" pageNumber="53" type="materials_examined">
<paragraph id="AD3C368BFF96A17FFF2FFE8BFC71C774" blockId="1.[151,982,293,348]" box="[151,982,325,348]" pageId="1" pageNumber="53">
<typeStatus id="72388829FF96A17FFF2FFE8BFF6BC774" box="[151,204,325,348]" pageId="1" pageNumber="53">Type</typeStatus>
species:
<taxonomicName id="6A834D08FF96A17FFE95FE8BFD69C774" authority="Walker, 1857: 144" authorityName="Walker" authorityPageNumber="144" authorityYear="1857" box="[301,718,325,348]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="9FF7EA99FF96A17FFE95FE8BFDA7C774" box="[301,512,325,348]" italics="true" pageId="1" pageNumber="53">Leusaba marginalis</emphasis>
<bibRefCitation id="C9124B7AFF96A17FFDB0FE8BFD3EC774" author="Walker" box="[520,665,325,348]" pageId="1" pageNumber="60" refString="Walker, F. (1857) Catalogue of the Homopterous Insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with Descriptions of New Species. Journal of the Proceedings of the Linnean Society. London, 1, 141 - 175." type="journal article" year="1857">Walker, 1857</bibRefCitation>
: 144
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, by original designation.
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<subSubSection id="E5996500FF96A17AFF2FFE44FC60C42A" lastPageId="4" lastPageNumber="56" pageId="1" pageNumber="53" type="description">
<paragraph id="AD3C368BFF96A17FFF2FFE44FE6BC7E2" blockId="1.[151,1437,394,2026]" pageId="1" pageNumber="53">
<emphasis id="9FF7EA99FF96A17FFF2FFE44FE95C78C" bold="true" box="[151,306,394,420]" pageId="1" pageNumber="53">Description.</emphasis>
Relatively large and stout tropiduchids (
<figureCitation id="35B82A0EFF96A17FFC98FE44FC32C78C" box="[800,917,394,420]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2. L" captionStart-2="FIGURE 3. L" captionStart-3="FIGURE 4" captionStartId-0="2.[151,255,1923,1947]" captionStartId-1="3.[151,255,1472,1496]" captionStartId-2="5.[151,258,1547,1571]" captionStartId-3="7.[151,255,1872,1896]" captionTargetBox-0="[223,1366,464,1859]" captionTargetBox-1="[169,1384,210,1461]" captionTargetBox-2="[159,1414,231,1506]" captionTargetBox-3="[166,1412,475,1871]" captionTargetId-0="figure@2.[188,1398,423,1899]" captionTargetId-1="figure@3.[151,1436,139,1476]" captionTargetId-2="figure@5.[151,1436,194,1525]" captionTargetId-3="figure@7.[151,1436,409,1875]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="7" captionText-0="FIGURE 1. Dorsal habitus and face respectively of Leusaba species. (A B) L. philippina (Philippines); (C D) Leusaba sp. indet. 1 (Sabah); (E F) Leusaba sp. indet. 2 (W. Malaysia); (G H) L. rufitarsis (Sri Lanka)." captionText-1="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." captionText-2="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." captionText-3="FIGURE 4. Leusaba indet sp. 1 (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) right fore wing; (E) right hind wing; (F) female genitalia, lateral view; (G) female genitalia, dorsal view; (H) female genitalia, ventral view; (I) gonopophyses IX, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F I = 0.5 mm. Abbreviations: bc, bursa copulatrix; oc, common oviduct; spe, spermatheca; va, anterior vagina; vp, posterior vagina." httpUri-0="https://zenodo.org/record/275554/files/figure.png" httpUri-1="https://zenodo.org/record/275555/files/figure.png" httpUri-2="https://zenodo.org/record/275556/files/figure.png" httpUri-3="https://zenodo.org/record/275557/files/figure.png" pageId="1" pageNumber="53">Figs. 14</figureCitation>
), length (from apex of vertex to tip of anal tube) 8.0
<quantity id="6A7B9B6EFF96A17FFEB6FE7EFED1C7E2" box="[270,374,432,458]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="1" pageNumber="53" unit="mm" value="10.6">10.6 mm</quantity>
(N=7).
</paragraph>
<paragraph id="AD3C368BFF96A17FFF7CFE1BFE23C482" blockId="1.[151,1437,394,2026]" pageId="1" pageNumber="53">Colour. General color green to ochraceous; frons with anterior and lateral margins and medial line suffused with orange to reddish stripes (black medial line may be present), apex of rostrum, eyes and antennae with pedicel suffused reddish to brownish; vertex with posterior marginal carinae, pronotum with median disc and lateral portion all suffused yellowish to reddish; fore wings hyaline, veins and stigma brown; fore tibia and mid tibia with apices, hind tibia with bases, lateral spines with apices, long stripes on hind femora all suffused with black.</paragraph>
<paragraph id="AD3C368BFF96A17FFF7CFD7BFD70C2F2" blockId="1.[151,1437,394,2026]" pageId="1" pageNumber="53">
<emphasis id="9FF7EA99FF96A17FFF7CFD7BFE3CC4E7" bold="true" box="[196,411,693,719]" pageId="1" pageNumber="53">Head and thorax</emphasis>
. Head (
<figureCitation id="35B82A0EFF96A17FFE42FD7BFDEEC4E7" box="[506,585,693,719]" captionStart="FIGURE 1" captionStartId="2.[151,255,1923,1947]" captionTargetBox="[223,1366,464,1859]" captionTargetId="figure@2.[188,1398,423,1899]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus and face respectively of Leusaba species. (A B) L. philippina (Philippines); (C D) Leusaba sp. indet. 1 (Sabah); (E F) Leusaba sp. indet. 2 (W. Malaysia); (G H) L. rufitarsis (Sri Lanka)." httpUri="https://zenodo.org/record/275554/files/figure.png" pageId="1" pageNumber="53">Figs. 1</figureCitation>
A, 1C, 1E, 1G) including eyes slightly narrower than pronotum, vertex approximately twice as broad as long, projecting before eyes for less than half length of eye; apex broadly convex in dorsal view, evenly and broadly rounded to base of frons; vertex and frons not separated, lateral carinae continuous. Eyes oval. Face with frons laterally carinate, carinae attaining frontoclypeal suture, with or without median ridge; slightly longer medially than greatest width and distinctly longer than width at anterior margin; ventrally depressed each side of midline (
<figureCitation id="35B82A0EFF96A17FFCFCFCBEFC35C5A2" box="[836,914,880,906]" captionStart="FIGURE 1" captionStartId="2.[151,255,1923,1947]" captionTargetBox="[223,1366,464,1859]" captionTargetId="figure@2.[188,1398,423,1899]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus and face respectively of Leusaba species. (A B) L. philippina (Philippines); (C D) Leusaba sp. indet. 1 (Sabah); (E F) Leusaba sp. indet. 2 (W. Malaysia); (G H) L. rufitarsis (Sri Lanka)." httpUri="https://zenodo.org/record/275554/files/figure.png" pageId="1" pageNumber="53">Figs. 1</figureCitation>
B, 2C) or depressed lateroventrally, forming a v-shaped central elevated area (
<figureCitation id="35B82A0EFF96A17FFD8AFC5BFDD8C587" box="[562,639,917,943]" captionStart="FIGURE 1" captionStartId="2.[151,255,1923,1947]" captionTargetBox="[223,1366,464,1859]" captionTargetId="figure@2.[188,1398,423,1899]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus and face respectively of Leusaba species. (A B) L. philippina (Philippines); (C D) Leusaba sp. indet. 1 (Sabah); (E F) Leusaba sp. indet. 2 (W. Malaysia); (G H) L. rufitarsis (Sri Lanka)." httpUri="https://zenodo.org/record/275554/files/figure.png" pageId="1" pageNumber="53">Figs. 1</figureCitation>
D, 3C); disc smooth or covered with microsetae; clypeus (
<figureCitation id="35B82A0EFF96A17FFA88FC5BFAD9C587" box="[1328,1406,917,943]" captionStart="FIGURE 1" captionStartId="2.[151,255,1923,1947]" captionTargetBox="[223,1366,464,1859]" captionTargetId="figure@2.[188,1398,423,1899]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus and face respectively of Leusaba species. (A B) L. philippina (Philippines); (C D) Leusaba sp. indet. 1 (Sabah); (E F) Leusaba sp. indet. 2 (W. Malaysia); (G H) L. rufitarsis (Sri Lanka)." httpUri="https://zenodo.org/record/275554/files/figure.png" pageId="1" pageNumber="53">Figs. 1</figureCitation>
B, 1D, 1F, 1H) with distinct median carina, without lateral carinae; rostrum short, attaining mesotrochanters, apical segment short, longer than broad; ocelli distinct; antennae (
<figureCitation id="35B82A0EFF96A17FFC02FC2EFBB7C5D2" box="[954,1040,992,1018]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
B, 2C, 4B, 4C) with scape short, ring-like; pedicel cylindrical, covered with many microsetae extending to base of pedicel. Vertex (
<figureCitation id="35B82A0EFF96A17FFA94FBCBFAD8C237" box="[1324,1407,1029,1055]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
A, 4A) subquadrate, distinctly shorter than greatest width, slightly longer than pronotum at midline; lateral margins ridged and subparallel; posterior margin carinate and arched anteriorly; anterior part of vertex smoothly swollen, uniting with base of frons to form evenly rounded surface; posterior part of vertex distinctly depressed, lateral margins of depression incurving anteriorly and meeting medially to form nearly semicircular arc; with a very faint median carina.
</paragraph>
<paragraph id="AD3C368BFF96A17FFF7CFB2BFAFBC122" blockId="1.[151,1437,394,2026]" pageId="1" pageNumber="53">
Pronotum (
<figureCitation id="35B82A0EFF96A17FFEF6FB2BFE03C2D7" box="[334,420,1253,1279]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
A, 4A) distinctly shorter than mesonotum in midline; with two contiguous median carinae and oblique and separate lateral carinae; anterior margin slightly carinate, nearly straight; disc projected anteriorly, deeply depressed between median and lateral carinae, with impression on each side; posterior margin obtusely angled excavate. Mesonotum (
<figureCitation id="35B82A0EFF96A17FFCF2FA9BFC39C347" box="[842,926,1365,1391]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
A, 4A) tricarinate, median carina straight, reaching to transverse suture; pronotum and mesonotum together medially 4.85.9 times as long as median length of vertex. Fore wings (
<figureCitation id="35B82A0EFF96A17FFE4EFA6EFDE3C392" box="[502,580,1440,1466]" captionStart="FIGURE 1" captionStartId="2.[151,255,1923,1947]" captionTargetBox="[223,1366,464,1859]" captionTargetId="figure@2.[188,1398,423,1899]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus and face respectively of Leusaba species. (A B) L. philippina (Philippines); (C D) Leusaba sp. indet. 1 (Sabah); (E F) Leusaba sp. indet. 2 (W. Malaysia); (G H) L. rufitarsis (Sri Lanka)." httpUri="https://zenodo.org/record/275554/files/figure.png" pageId="1" pageNumber="53">Figs. 1</figureCitation>
A, 1C, 1E, 1G) hyaline, surpassing extremity of abdomen by about 0.5 0.7 times length of body, widening towards tips, 2.72.8 times as long as maximum breadth; corium without granulation; veins prominent, thickly covered with short setae dorsally and ventrally (veins in basal area of corium not covered with setae dorsally); costal cell without cross veins; Sc+R and M forking at nodal line; Cu forking after level of junction of claval veins, the latter forked at basal 2/5 of clavus; nodal line forming a zigzag transverse line, subapical line slightly oblique, with 6 subapical and 1720 apical cells, subapical cells a little longer than latter.
<collectingCountry id="D594761BFF96A17FFE7BF94EFE59C0B2" box="[451,510,1664,1690]" name="India" pageId="1" pageNumber="53">Hind</collectingCountry>
wings (
<figureCitation id="35B82A0EFF96A17FFDE6F94EFD17C0B2" box="[606,688,1664,1690]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
E, 4E) hyaline, veins prominent and thickly covered with short setae dorsally and ventrally, posterior margin strongly sinuate, with three disconnected, oblique transverse veins before apical area, venation as in
<figureCitation id="35B82A0EFF96A17FFDD2F904FD1CC0CC" box="[618,699,1738,1764]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
E, 4E. Legs elongate; hind tibiae each with 34 distinct lateral spines, abnormally with 5 (often on one leg only, basal one very fine); spinal formula of hind leg 662.
</paragraph>
<paragraph id="AD3C368BFF96A17CFF7CF8DBFCACC7BA" blockId="1.[151,1437,394,2026]" lastBlockId="2.[151,1437,152,402]" lastPageId="2" lastPageNumber="54" pageId="1" pageNumber="53">
<emphasis id="9FF7EA99FF96A17FFF7CF8DBFE24C107" bold="true" box="[196,387,1813,1839]" pageId="1" pageNumber="53">Male genitalia.</emphasis>
Pygofer (
<figureCitation id="35B82A0EFF96A17FFE44F8DBFDF7C107" box="[508,592,1813,1839]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
F, 2H,
<date id="D93D104BFF96A17FFD1CF8DBFD1AC107" box="[676,701,1813,1839]" pageId="1" pageNumber="53">2I</date>
) symmetrical, short, in lateral view forming a perpendicular irregular rectangle, wider dorsally than ventrally, with dorsal posterior margin angularly produced posteriorly; dorsal margin deeply excavated to accommodate anal tube (
<figureCitation id="35B82A0EFF96A17FFCC4F8AEFC60C152" box="[892,967,1888,1914]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Fig. 2</figureCitation>
H); ventral margin strongly excavated anteriorly to accommodate gonostyli (
<figureCitation id="35B82A0EFF96A17FFDD6F84BFD1CC1B7" box="[622,699,1925,1951]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">
Fig.
<date id="D93D104BFF96A17FFD11F84BFD1CC1B7" box="[681,699,1925,1951]" pageId="1" pageNumber="53">2</date>
</figureCitation>
I). Xth segment (
<figureCitation id="35B82A0EFF96A17FFC33F84BFC46C1B7" box="[907,993,1925,1951]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
F, 2H) very long and slender, apical margin distinctly concave in dorsal view, lateroventral margin with two pairs of distinct long spines (
<figureCitation id="35B82A0EFF96A17FFA8CF864FADCC1EC" box="[1332,1403,1962,1988]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Fig. 2</figureCitation>
F); anal style short and small. Gonostyli (
<figureCitation id="35B82A0EFF96A17FFDDAF81EFD11C1C2" box="[610,694,2000,2026]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="1" pageNumber="53">Figs. 2</figureCitation>
F, 2G,
<date id="D93D104BFF96A17FFCBDF81EFCB8C1C2" box="[773,799,2000,2026]" pageId="1" pageNumber="53">2I</date>
) bilaterally asymmetrical, membranously fused with pygofer at base, left gonostylus forming irregular ellipse, more than 3 times as long as maximum breadth in lateral view, tapering to apex in distal 1/6, dorsal edge with two processes (see
<figureCitation id="35B82A0EFF95A17CFBDCFF73FB0BC6FF" box="[1124,1196,189,215]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="2" pageNumber="54">Fig. 2</figureCitation>
F), right gonostylus forming irregular rectangle, relatively large and broad, more than 3 times as long as maximum breadth in lateral view, apex relatively broad, basal dorsal edge with a process (see
<figureCitation id="35B82A0EFF95A17CFC50FEC6FB89C70A" box="[1000,1070,264,290]" captionStart="FIGURE 2. L" captionStartId="3.[151,255,1472,1496]" captionTargetBox="[169,1384,210,1461]" captionTargetId="figure@3.[151,1436,139,1476]" captionTargetPageId="3" captionText="FIGURE 2. L. philippina (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F G = 0.5 mm; H = 1 mm; I = 0.5 mm." httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="2" pageNumber="54">Fig. 2</figureCitation>
G). Periandrium (
<figureCitation id="35B82A0EFF95A17CFB43FEC6FAECC70A" box="[1275,1355,264,290]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="2" pageNumber="54">Figs. 3</figureCitation>
A, 3B) dorsally connected with ventrobasal margin of anal-tube, membranously fused with pygofer at ventral side, irregularly ring-like, distinctly sclerotized, surrounding aedeagus at basal part and visible in lateral view, apical margin with a branched process at ventral side.
</paragraph>
<caption id="F9FC6603FF95A17CFF2FF84DFB2AC193" httpUri="https://zenodo.org/record/275554/files/figure.png" pageId="2" pageNumber="54" targetBox="[223,1366,464,1859]" targetPageId="2">
<paragraph id="AD3C368BFF95A17CFF2FF84DFB2AC193" blockId="2.[151,1436,1923,1979]" pageId="2" pageNumber="54">
<emphasis id="9FF7EA99FF95A17CFF2FF84DFEBEC1B3" bold="true" box="[151,281,1923,1947]" pageId="2" pageNumber="54">FIGURE 1.</emphasis>
Dorsal habitus and face respectively of
<taxonomicName id="6A834D08FF95A17CFD7FF84AFC86C1B3" box="[711,801,1924,1947]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF95A17CFD7FF84AFC86C1B3" box="[711,801,1924,1947]" italics="true" pageId="2" pageNumber="54">Leusaba</emphasis>
</taxonomicName>
species. (AB)
<taxonomicName id="6A834D08FF95A17CFC76F84AFB4CC1B3" authority="(Philippines)" baseAuthorityName="Philippines" box="[974,1259,1924,1947]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="9FF7EA99FF95A17CFC76F84AFBFFC1B3" box="[974,1112,1924,1947]" italics="true" pageId="2" pageNumber="54">L. philippina</emphasis>
(Philippines)
</taxonomicName>
; (CD)
<taxonomicName id="6A834D08FF95A17CFAFAF84AFA3BC1B3" box="[1346,1436,1924,1947]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF95A17CFAFAF84AFA3BC1B3" box="[1346,1436,1924,1947]" italics="true" pageId="2" pageNumber="54">Leusaba</emphasis>
</taxonomicName>
sp. indet. 1 (Sabah); (EF)
<taxonomicName id="6A834D08FF95A17CFE00F86AFDB5C193" box="[440,530,1956,1979]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF95A17CFE00F86AFDB5C193" box="[440,530,1956,1979]" italics="true" pageId="2" pageNumber="54">Leusaba</emphasis>
</taxonomicName>
sp. indet. 2 (W. Malaysia); (GH)
<taxonomicName id="6A834D08FF95A17CFC3FF86AFBA2C193" box="[903,1029,1956,1979]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="9FF7EA99FF95A17CFC3FF86AFBA2C193" box="[903,1029,1956,1979]" italics="true" pageId="2" pageNumber="54">L. rufitarsis</emphasis>
</taxonomicName>
(Sri Lanka).
</paragraph>
</caption>
<caption id="F9FC6603FF94A17DFF2FFA0EFCFCC01F" httpUri="https://zenodo.org/record/275555/files/figure.png" pageId="3" pageNumber="55" targetBox="[169,1384,210,1461]" targetPageId="3">
<paragraph id="AD3C368BFF94A17DFF2FFA0EFCFCC01F" blockId="3.[151,1436,1472,1591]" pageId="3" pageNumber="55">
<emphasis id="9FF7EA99FF94A17DFF2FFA0EFEBEC3F0" bold="true" box="[151,281,1472,1496]" pageId="3" pageNumber="55">FIGURE 2.</emphasis>
<taxonomicName id="6A834D08FF94A17DFE98FA0EFE0DC3FF" box="[288,426,1472,1495]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="9FF7EA99FF94A17DFE98FA0EFE0DC3FF" box="[288,426,1472,1495]" italics="true" pageId="3" pageNumber="55">L. philippina</emphasis>
</taxonomicName>
(A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) left fore wing; (E) left hind wing; (F) male genitalia without aedeagus, left view; (G) gonostylus, right view; (H) anal segment and pygofer, dorsal view; (I) genital capsule, ventral view. Scale bars: Figs. A C = 0.5 mm; DE = 1 mm; FG = 0.5 mm; H = 1 mm; I =0.5 mm.
</paragraph>
</caption>
<paragraph id="AD3C368BFF94A17DFF7CF9AEFCC0C127" blockId="3.[151,1436,1632,2031]" pageId="3" pageNumber="55">
Aedeagus (
<figureCitation id="35B82A0EFF94A17DFEF5F9AEFE06C052" box="[333,417,1632,1658]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="3" pageNumber="55">Figs. 3</figureCitation>
A, 3B) asymmetrical, elongate, robust and mostly sclerotized, with a long, sinuous process at basal part, basal 1/4 to 1/2 part of aedeagus expanded into ellipsoid, connected with membranous endosoma apically, with three projections at left side and a prominence at right side; shaft of aedeagus sclerotized, embraced by endosoma, apical part strongly curved and directed posteroventrally, phallotrema hole-like, exposed apically; endosoma with three projections.
</paragraph>
<paragraph id="AD3C368BFF94A17AFF7CF8D4FC60C42A" blockId="3.[151,1436,1632,2031]" lastBlockId="4.[151,1437,152,1074]" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55">
<emphasis id="9FF7EA99FF94A17DFF7CF8D4FE35C11C" bold="true" box="[196,402,1818,1844]" pageId="3" pageNumber="55">Female genitalia</emphasis>
. Anal tube (
<figureCitation id="35B82A0EFF94A17DFD9DF8D5FDD2C11D" box="[549,629,1819,1845]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="3" pageNumber="55">Figs. 3</figureCitation>
C, 3D, 4F, 4G) relatively short, apical margin convex in dorsal view, anal styles reaching beyond apical ventral margin of anal tube. Gonopophyses VIII (first valvulae) (
<figureCitation id="35B82A0EFF94A17DFA88F88EFA27C172" box="[1328,1408,1856,1882]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="3" pageNumber="55">Figs. 3</figureCitation>
C, 3F, 4F, 4H) strongly sclerotized with 5 blunt teeth on dorsal margin, 3 blunt apical teeth on ventral margin, and several minute teeth in two angle-like rows laterobasad of middle. Gonopophyses VIII nearly symmetrical or asymmetrical at base in ventral view (
<figureCitation id="35B82A0EFF94A17DFDCAF87EFD60C1E2" box="[626,711,1968,1994]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="3" pageNumber="55">Figs. 3</figureCitation>
F, 4H). Gonopophyses IX (second valvulae) (
<figureCitation id="35B82A0EFF94A17DFB46F87EFAF2C1E2" box="[1278,1365,1968,1994]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="3" pageNumber="55">Figs. 3</figureCitation>
E,
<date id="D93D104BFF94A17DFAC0F87EFA33C1E2" box="[1400,1428,1968,1994]" pageId="3" pageNumber="55">4I</date>
) triangular, fused together on inner-lateral margin and strongly reduced but well sclerotized, apical ends confluent or not; gonospiculum (
<figureCitation id="35B82A0EFF93A17AFD9FFF56FDDDC69A" box="[551,634,152,178]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
E,
<date id="D93D104BFF93A17AFD26FF56FD1FC69A" box="[670,696,152,178]" pageId="4" pageNumber="56">4I</date>
) shorter than median length of triangular part (0.750.95:1). Gonoplacs (third valvulae) (
<figureCitation id="35B82A0EFF93A17AFE4CFF73FDEEC6FF" box="[500,585,189,215]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
C, 3D, 3F, 4F, 4G) with 34 teeth at dorsal margin, dentition simple, forming one line, continue without interruption along ventral margin and involving a further 13 teeth (see
<figureCitation id="35B82A0EFF93A17AFF2FFEC6FF40C70A" box="[151,231,264,290]" captionStart="FIGURE 4" captionStartId="7.[151,255,1872,1896]" captionTargetBox="[166,1412,475,1871]" captionTargetId="figure@7.[151,1436,409,1875]" captionTargetPageId="7" captionText="FIGURE 4. Leusaba indet sp. 1 (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) right fore wing; (E) right hind wing; (F) female genitalia, lateral view; (G) female genitalia, dorsal view; (H) female genitalia, ventral view; (I) gonopophyses IX, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F I = 0.5 mm. Abbreviations: bc, bursa copulatrix; oc, common oviduct; spe, spermatheca; va, anterior vagina; vp, posterior vagina." httpUri="https://zenodo.org/record/275557/files/figure.png" pageId="4" pageNumber="56">Figs. 4</figureCitation>
F, 4G); or dentition complex, not forming one line, with 3 outer teeth at apical part, then continue with 1213 teeth from apical to ventral margin (see
<figureCitation id="35B82A0EFF93A17AFD64FEE3FC97C76F" box="[732,816,301,327]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
C, 3D, 3F). Gonocoxae VIII (
<figureCitation id="35B82A0EFF93A17AFB26FEE3FB53C76F" box="[1182,1268,301,327]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
C, 4F) almost rectangular sclerotized plates with a distinct black spot on caudadorsal edge. Genital opening (gonoporus) singular (monotrysian) occurring between gonopophyses VIII and IX (
<figureCitation id="35B82A0EFF93A17AFC69FEB6FB87C7BA" box="[977,1056,376,402]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
F, 4H). Sternite VII with median part sinuous or broad concave at ventral side. Posterior vagina strikingly sclerotized in its ventral part (
<figureCitation id="35B82A0EFF93A17AFAD9FE53FF03C7F4" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
F, 4H). Dorsally, a large bursa copulatrix (
<figureCitation id="35B82A0EFF93A17AFD2EFE0CFD40C7F4" box="[662,743,450,476]" captionStart="FIGURE 3. L" captionStartId="5.[151,258,1547,1571]" captionTargetBox="[159,1414,231,1506]" captionTargetId="figure@5.[151,1436,194,1525]" captionTargetPageId="5" captionText="FIGURE 3. L. philippina (A) aedeagus, left view; (B) aedeagus, right view; (C) female genitalia, lateral view; (D) female genitalia, dorsal view; (E) gonopophyses IX, ventral view; (F) female genitalia, ventral view. Scale bars = 0.5 mm. Abbreviations: ape, apical production of endosoma; at, anal tube; bc, bursa copulatrix; bpa, basal process of aedeagus; en, endosoma; fa, anterior fibula; fp, posterior fibula; gl, gonospiculum; gnp, gonoporus; gp, gonoplac; gxp, endogonocoxal process; gxVIII, gonocoxae VIII; gyVIII, gonopophyses VIII; gyIX, gonopophyses IX; lpaI, left production 1 of aedeagus; lpaII, left production 2 of aedeagus; lpaIII, left production 3 of aedeagus; Per, periandrium; pp, process of periandrium; pph, phallotrema; rpa, right prominence of aedeagus; rpe, right production of endosoma; sa, shaft of aedeagus; stVII, Sternite VII; vg, vagina; vp, posterior vagina; vpe, ventral production of endosoma." httpUri="https://zenodo.org/record/275556/files/figure.png" pageId="4" pageNumber="56">Figs. 3</figureCitation>
C, 4F) opens directly into vagina and an enlarged common oviduct and spermatheca open apically into anterior vagina (
<figureCitation id="35B82A0EFF93A17AFCE3FE26FC05C42A" box="[859,930,488,514]" captionStart="FIGURE 4" captionStartId="7.[151,255,1872,1896]" captionTargetBox="[166,1412,475,1871]" captionTargetId="figure@7.[151,1436,409,1875]" captionTargetPageId="7" captionText="FIGURE 4. Leusaba indet sp. 1 (A) head, pronotum and mesonotum, dorsal view; (B) head, pronotum and mesonotum, lateral view; (C) head, ventral view; (D) right fore wing; (E) right hind wing; (F) female genitalia, lateral view; (G) female genitalia, dorsal view; (H) female genitalia, ventral view; (I) gonopophyses IX, ventral view. Scale bars: Figs. A C = 0.5 mm; D E = 1 mm; F I = 0.5 mm. Abbreviations: bc, bursa copulatrix; oc, common oviduct; spe, spermatheca; va, anterior vagina; vp, posterior vagina." httpUri="https://zenodo.org/record/275557/files/figure.png" pageId="4" pageNumber="56">Fig. 4</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="E5996500FF93A17AFF7CFDC3FF4FC464" pageId="4" pageNumber="56" type="distribution">
<paragraph id="AD3C368BFF93A17AFF7CFDC3FF4FC464" blockId="4.[151,1437,152,1074]" pageId="4" pageNumber="56">
<emphasis id="9FF7EA99FF93A17AFF7CFDC3FECAC40F" bold="true" box="[196,365,525,551]" pageId="4" pageNumber="56">Distribution.</emphasis>
Species of
<taxonomicName id="6A834D08FF93A17AFE47FDC3FDC0C40E" box="[511,615,525,550]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFE47FDC3FDC0C40E" box="[511,615,525,550]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
are currently known from
<collectingCountry id="D594761BFF93A17AFC0EFDC3FB80C40F" box="[950,1063,525,551]" name="Malaysia" pageId="4" pageNumber="56">Malaysia</collectingCountry>
(Borneo),
<collectingCountry id="D594761BFF93A17AFB08FDC3FA9CC40F" box="[1200,1339,525,551]" name="Philippines" pageId="4" pageNumber="56">Philippines</collectingCountry>
and
<collectingCountry id="D594761BFF93A17AFAC0FDC3FF43C464" name="Sri Lanka" pageId="4" pageNumber="56">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E5996500FF93A17AFF7CFD99FEC7C21A" pageId="4" pageNumber="56" type="discussion">
<paragraph id="AD3C368BFF93A17AFF7CFD99FD3BC5B4" blockId="4.[151,1437,152,1074]" pageId="4" pageNumber="56">
<emphasis id="9FF7EA99FF93A17AFF7CFD99FEE8C459" bold="true" box="[196,335,599,625]" pageId="4" pageNumber="56">Discussion.</emphasis>
<taxonomicName id="6A834D08FF93A17AFEEFFD96FE1DC459" box="[343,442,600,625]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFEEFFD96FE1DC459" box="[343,442,600,625]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
belongs in the tribe Paricanini Melichar, which was redefined by
<bibRefCitation id="C9124B7AFF93A17AFB7FFD96FADFC45A" author="Fennah" box="[1223,1400,600,626]" pageId="4" pageNumber="60" refString="Fennah, R. G. (1982) A tribal classification of the Tropiduchidae (Homoptera: Fulgoroidea), with the description of a new species on tea in Malaysia. Bulletin of Entomological Research, 72, 631 - 643." type="journal article" year="1982">Fennah (1982)</bibRefCitation>
. In addition to
<taxonomicName id="6A834D08FF93A17AFEA7FDB3FE24C4BE" box="[287,387,637,662]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFEA7FDB3FE24C4BE" box="[287,387,637,662]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
, the tribe includes
<taxonomicName id="6A834D08FF93A17AFDD2FDB3FC26C4BF" authority="Walker, 1857" authorityName="Walker" authorityYear="1857" box="[618,897,637,663]" class="Insecta" family="Tropiduchidae" genus="Paricana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFDD2FDB3FD7FC4BE" box="[618,728,637,662]" italics="true" pageId="4" pageNumber="56">Paricana</emphasis>
<bibRefCitation id="C9124B7AFF93A17AFD58FDB3FC26C4BF" author="Walker" box="[736,897,637,663]" pageId="4" pageNumber="60" refString="Walker, F. (1857) Catalogue of the Homopterous Insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with Descriptions of New Species. Journal of the Proceedings of the Linnean Society. London, 1, 141 - 175." type="journal article" year="1857">Walker, 1857</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="6A834D08FF93A17AFC36FDB3FB60C4BF" authority="Liang, 2003" authorityName="Liang" authorityYear="2003" box="[910,1223,637,663]" class="Insecta" family="Tropiduchidae" genus="Paricanoides" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFC36FDB3FB8BC4BE" box="[910,1068,637,662]" italics="true" pageId="4" pageNumber="56">Paricanoides</emphasis>
<bibRefCitation id="C9124B7AFF93A17AFB8CFDB3FB60C4BF" author="Liang" box="[1076,1223,637,663]" pageId="4" pageNumber="60" refString="Liang, A. P. (2003) A new genus of Tropiduchidae (Hemiptera: Fulgoroidea) from China and Vietnam, with description of eggs. Florida Entomologist, 86 (3), 361 - 369." type="journal article" year="2003">Liang, 2003</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="6A834D08FF93A17AFABFFDB3FF72C494" authority="Stal, 1859" authorityName="Stal" authorityYear="1859" class="Insecta" family="Tropiduchidae" genus="Stacota" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFABFFDB3FAF8C4BE" box="[1287,1375,637,662]" italics="true" pageId="4" pageNumber="56">Stacota</emphasis>
<bibRefCitation id="C9124B7AFF93A17AFADFFDB3FF72C494" author="Stal" pageId="4" pageNumber="60" refString="Stal. C. (1859) Novae quaedem Fulgorinorum formae speciesque insigniores. Berliner Entomologische Zeitschrift, 3, 313 - 327." type="journal article" year="1859">Stål, 1859</bibRefCitation>
</taxonomicName>
. Species of
<taxonomicName id="6A834D08FF93A17AFEDBFD6CFE60C493" box="[355,455,674,699]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFEDBFD6CFE60C493" box="[355,455,674,699]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
are externally similar to those of
<taxonomicName id="6A834D08FF93A17AFCEFFD6CFC63C493" box="[855,964,674,699]" class="Insecta" family="Tropiduchidae" genus="Paricana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFCEFFD6CFC63C493" box="[855,964,674,699]" italics="true" pageId="4" pageNumber="56">Paricana</emphasis>
</taxonomicName>
but can be distinguished from the latter by the laterally carinate vertex with a median carina and fore wings with six subapical and 1820 apical cells, with subapical cells a little longer than the latter.
<taxonomicName id="6A834D08FF93A17AFD69FD23FC93C52E" box="[721,820,749,774]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFD69FD23FC93C52E" box="[721,820,749,774]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="6A834D08FF93A17AFBD6FD23FAADC52E" box="[1134,1290,749,774]" class="Insecta" family="Tropiduchidae" genus="Paricanoides" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFBD6FD23FAADC52E" box="[1134,1290,749,774]" italics="true" pageId="4" pageNumber="56">Paricanoides</emphasis>
</taxonomicName>
by the frons and postclypeus with a median carina and more numerous apical and subapical cells in the forewing and more numerous apical cells in the hindwing (see
<bibRefCitation id="C9124B7AFF93A17AFD2BFCF6FC82C57A" author="Liang" box="[659,805,824,850]" pageId="4" pageNumber="60" refString="Liang, A. P. (2003) A new genus of Tropiduchidae (Hemiptera: Fulgoroidea) from China and Vietnam, with description of eggs. Florida Entomologist, 86 (3), 361 - 369." type="journal article" year="2003">Liang, 2003</bibRefCitation>
). It can be separated from
<taxonomicName id="6A834D08FF93A17AFBE4FCF6FB14C579" box="[1116,1203,824,849]" class="Insecta" family="Tropiduchidae" genus="Stacota" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFBE4FCF6FB14C579" box="[1116,1203,824,849]" italics="true" pageId="4" pageNumber="56">Stacota</emphasis>
</taxonomicName>
by the vertex being distinctly shorter than greatest width (slightly longer medially than greatest width in
<taxonomicName id="6A834D08FF93A17AFBC5FC93FB73C55E" box="[1149,1236,861,886]" class="Insecta" family="Tropiduchidae" genus="Stacota" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFBC5FC93FB73C55E" box="[1149,1236,861,886]" italics="true" pageId="4" pageNumber="56">Stacota</emphasis>
</taxonomicName>
), and fore wings lacking transverse veinlets in the costal cell.
</paragraph>
<paragraph id="AD3C368BFF93A17AFF7EFC66FEC7C21A" blockId="4.[151,1437,152,1074]" pageId="4" pageNumber="56">
In addition to
<taxonomicName id="6A834D08FF93A17AFED3FC66FDA6C5E9" box="[363,513,936,962]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="9FF7EA99FF93A17AFED3FC66FEDCC5E9" box="[363,379,936,961]" italics="true" pageId="4" pageNumber="56">L</emphasis>
.
<emphasis id="9FF7EA99FF93A17AFE31FC66FDA6C5E9" box="[393,513,936,961]" italics="true" pageId="4" pageNumber="56">philippina</emphasis>
</taxonomicName>
the following other species belong in
<taxonomicName id="6A834D08FF93A17AFC78FC66FB84C5E9" box="[960,1059,936,961]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFC78FC66FB84C5E9" box="[960,1059,936,961]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
. Two undescribed females from
<collectingCountry id="D594761BFF93A17AFF2FFC03FEA4C5CF" box="[151,259,973,999]" name="Malaysia" pageId="4" pageNumber="56">Malaysia</collectingCountry>
are considered to represent two additional species of
<taxonomicName id="6A834D08FF93A17AFCCAFC03FC71C5CE" box="[882,982,973,998]" class="Insecta" family="Tropiduchidae" genus="Leusaba" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="9FF7EA99FF93A17AFCCAFC03FC71C5CE" box="[882,982,973,998]" italics="true" pageId="4" pageNumber="56">Leusaba</emphasis>
</taxonomicName>
but are not formally named because of the lack of male specimens. They indicate that the Malaysian fauna is more widespread and diverse than currently known.
</paragraph>
</subSubSection>
</treatment>
</document>