240 lines
37 KiB
XML
240 lines
37 KiB
XML
<document id="222222CA30BD107C64160A6F5C541E3E" ID-CLB-Dataset="3115" ID-DOI="10.1093/zoolinnean/zlaa184" ID-GBIF-Dataset="c0c8a1d4-59e5-4d8a-80bf-de2018ab0bf3" ID-ISSN="0024-4082" ID-Zenodo-Dep="5753254" ID-ZooBank="A8B9DE67-0C57-49D6-B865-388AD51AA50F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638521882613" checkinUser="plazi" docAuthor="Hsiao, Yun, Li, Yun, Ren, Dong & Pang, Hong" docDate="2021" docId="03E3640FFF88FFEBD3BD2B54AA668C7D" docLanguage="en" docName="zlaa184.pdf" docOrigin="Zoological Journal of the Linnean Society 193" docSource="https://doi.org/10.1093/zoolinnean/zlaa184" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Hukawngichthyurus maha Hsiao & Li & Ren & Pang 2021, SP. NOV." docType="treatment" docVersion="6" lastPageNumber="1285" masterDocId="FFDA1C77FF85FFE5D03B2D2DA9158A03" masterDocTitle="Morphological phylogenetics provide new insights into the classification and evolution of fossil soldier beetles from Mid-Cretaceous Burmese amber (Coleoptera: Cantharidae)" masterLastPageNumber="1293" masterPageNumber="1271" pageNumber="1284" updateTime="1699471872165" updateUser="ExternalLinkService">
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<mods:title id="BB6CC426BE9532D6A8F213E227A1EEF1">Morphological phylogenetics provide new insights into the classification and evolution of fossil soldier beetles from Mid-Cretaceous Burmese amber (Coleoptera: Cantharidae)</mods:title>
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<mods:namePart id="38E7814B83A5D8BA01381B5D0F616F7A">Hsiao, Yun</mods:namePart>
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<mods:namePart id="5A23304755FB00499BA842FEF0B647A1">Li, Yun</mods:namePart>
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<mods:namePart id="5BB5C2819687E79A0E596FC324F406E6">Ren, Dong</mods:namePart>
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<mods:roleTerm id="2F790ABE111F80400E767D467C898DFB">Author</mods:roleTerm>
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<mods:namePart id="1D2CCD68BC6CDFA77E0DAB1ABE3DEDB5">Pang, Hong</mods:namePart>
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<mods:title id="AD9229CC0A512C3730D61DD6FFF4A1F0">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="F73D663FDDD173A151CD1B3FB92AC766">
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<mods:date id="F741CF767B6367D03554CC2647C64876">2021</mods:date>
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<mods:number id="40EE483754B320CA19B3102AF82FF074">2021-02-03</mods:number>
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<mods:number id="F903BB80A48F41E911BB34C186D6B2A2">193</mods:number>
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<mods:url id="5B07FA10512F1751D936C4CFC5C798CC">https://doi.org/10.1093/zoolinnean/zlaa184</mods:url>
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<treatment id="03E3640FFF88FFEBD3BD2B54AA668C7D" ID-DOI="http://doi.org/10.5281/zenodo.5753268" ID-GBIF-Taxon="190877393" ID-Zenodo-Dep="5753268" LSID="urn:lsid:plazi:treatment:03E3640FFF88FFEBD3BD2B54AA668C7D" httpUri="http://treatment.plazi.org/id/03E3640FFF88FFEBD3BD2B54AA668C7D" lastPageId="14" lastPageNumber="1285" pageId="13" pageNumber="1284">
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<subSubSection id="C3508692FF88FFE8D3BD2B54AC4D8C92" box="[902,1368,1657,1681]" pageId="13" pageNumber="1284" type="nomenclature">
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<paragraph id="8BF5D519FF88FFE8D3BD2B54AC4D8C92" blockId="13.[902,1368,1657,1681]" box="[902,1368,1657,1681]" pageId="13" pageNumber="1284">
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<heading id="D0BD6275FF88FFE8D3BD2B54AC4D8C92" box="[902,1368,1657,1681]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="1284" reason="2">
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<emphasis id="B93E090BFF88FFE8D3BD2B54ADFB8C93" bold="true" box="[902,1262,1657,1681]" italics="true" pageId="13" pageNumber="1284">
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†
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<taxonomicName id="4C4AAE9AFF88FFE8D3A92B57ADFB8C93" authority="Hsiao & Li & Ren & Pang, 2021" authorityName="Hsiao & Li & Ren & Pang" authorityYear="2021" box="[914,1262,1658,1681]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1290" phylum="Arthropoda" rank="species" species="maha" status="sp. nov.">
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<emphasis id="B93E090BFF88FFE8D3A92B57ADB68C93" bold="true" box="[914,1187,1658,1681]" pageId="13" pageNumber="1284">HUKAWNGICHTHYURUS</emphasis>
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MAHA
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</emphasis>
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SP
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<emphasis id="B93E090BFF88FFE8D5292B54AC4D8C92" bold="true" box="[1298,1368,1657,1681]" pageId="13" pageNumber="1284">. NOV.</emphasis>
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<subSubSection id="C3508692FF88FFEBD4082B87AA668C7D" lastPageId="14" lastPageNumber="1285" pageId="13" pageNumber="1284" type="description">
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<paragraph id="8BF5D519FF88FFE8D4082B87ADBF8CC0" blockId="13.[1075,1194,1706,1731]" box="[1075,1194,1706,1731]" pageId="13" pageNumber="1284">
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(
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<figureCitation id="1371C99CFF88FFE8D4002B87ADB78CC0" box="[1083,1186,1706,1731]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStartId-0="14.[146,224,783,805]" captionStartId-1="15.[163,241,1245,1267]" captionTargetBox-0="[149,757,195,743]" captionTargetBox-1="[403,1203,195,1205]" captionTargetId-0="figure-687@14.[149,757,195,743]" captionTargetId-1="figure-356@15.[403,1203,195,1205]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Figure 7. Habitus of Hukawngichthyurus maha: A, dorsal view; B, ventral-lateral view. Scale bars: 0.5 mm." captionText-1="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi-0="http://doi.org/10.5281/zenodo.5753274" figureDoi-1="http://doi.org/10.5281/zenodo.5753276" httpUri-0="https://zenodo.org/record/5753274/files/figure.png" httpUri-1="https://zenodo.org/record/5753276/files/figure.png" pageId="13" pageNumber="1284">FIGS 7–8</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF5D519FF88FFE8D3002BF9AC888D0B" blockId="13.[827,1443,1747,1800]" pageId="13" pageNumber="1284">
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<emphasis id="B93E090BFF88FFE8D3002BF9AD928CEA" box="[827,1159,1747,1769]" italics="true" pageId="13" pageNumber="1284">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n: l s i d: z o o b a n k. org:act:
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3F289BBE-89FF-414D-965C-691C8E06EC
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</uuid>
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.
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<paragraph id="8BF5D519FF88FFEBD3002A0BA9CB89C1" blockId="13.[827,1442,1830,1882]" lastBlockId="14.[145,761,879,962]" lastPageId="14" lastPageNumber="1285" pageId="13" pageNumber="1284">
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<emphasis id="B93E090BFF88FFE8D3002A0BAAFF8D38" box="[827,1002,1830,1851]" italics="true" pageId="13" pageNumber="1284">Type material:</emphasis>
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<materialsCitation id="3B22DF44FF88FFEBD43B2A0BA9CB89C1" ID-GBIF-Occurrence="3414066303" country="Myanmar" county="Guangzhou" lastPageId="14" lastPageNumber="1285" location="Male" municipality="Sun Yat-sen University" pageId="13" pageNumber="1284" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F16BBBFF88FFE8D43B2A0BAD7B8D3F" box="[1024,1134,1830,1852]" pageId="13" pageNumber="1284" type="holotype">Holotype</typeStatus>
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:
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<location id="8E9583C2FF88FFE8D4402A0BADAC8D3F" LSID="urn:lsid:plazi:treatment:03E3640FFF88FFEBD3BD2B54AA668C7D:8E9583C2FF88FFE8D4402A0BADAC8D3F" box="[1147,1209,1830,1852]" country="Myanmar" county="Guangzhou" municipality="Sun Yat-sen University" name="Male" pageId="13" pageNumber="1284">Male</location>
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; no. YL-COL00001; earliest
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<location id="8E9583C2FF88FFE8D39E2A69AD518D59" LSID="urn:lsid:plazi:treatment:03E3640FFF88FFEBD3BD2B54AA668C7D:8E9583C2FF88FFE8D39E2A69AD518D59" box="[933,1092,1860,1882]" country="Myanmar" county="Guangzhou" municipality="Sun Yat-sen University" name="Cenomanian" pageId="13" pageNumber="1284">Cenomanian</location>
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,
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<location id="8E9583C2FF88FFE8D46E2A69AC318D59" LSID="urn:lsid:plazi:treatment:03E3640FFF88FFEBD3BD2B54AA668C7D:8E9583C2FF88FFE8D46E2A69AC318D59" box="[1109,1316,1860,1882]" country="Myanmar" county="Guangzhou" municipality="Sun Yat-sen University" name="Hukawng Valley" pageId="13" pageNumber="1284">Hukawng Valley</location>
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, northern
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<collectingCountry id="F35D9589FF8BFFEBD0AA2E42A8168987" box="[145,259,879,900]" name="Myanmar" pageId="14" pageNumber="1285">Myanmar</collectingCountry>
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; deposited in the
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<location id="8E9583C2FF8BFFEBD1F02E42ABB08986" LSID="urn:lsid:plazi:treatment:03E3640FFF88FFEBD3BD2B54AA668C7D:8E9583C2FF8BFFEBD1F02E42ABB08986" box="[459,677,879,901]" country="Myanmar" county="Guangzhou" municipality="Sun Yat-sen University" name="Museum of Biology" pageId="14" pageNumber="1285">Museum of Biology</location>
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,
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<location id="8E9583C2FF8BFFEBD2942E42A85289A7" LSID="urn:lsid:plazi:treatment:03E3640FFF88FFEBD3BD2B54AA668C7D:8E9583C2FF8BFFEBD2942E42A85289A7" country="Myanmar" county="Guangzhou" municipality="Sun Yat-sen University" name="School of Life Sciences" pageId="14" pageNumber="1285">School of Life Sciences</location>
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,
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<collectingMunicipality id="6B914F63FF8BFFEBD1682EA3AB7789A7" box="[339,610,910,932]" pageId="14" pageNumber="1285">Sun Yat-sen University</collectingMunicipality>
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,
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<collectingCounty id="6294AD95FF8BFFEBD2552EA3ABE089A7" box="[622,757,910,932]" pageId="14" pageNumber="1285">Guangzhou</collectingCounty>
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,
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<collectingCountry id="F35D9589FF8BFFEBD0AA2E81A9CF89C1" box="[145,218,940,962]" name="China" pageId="14" pageNumber="1285">China</collectingCountry>
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.
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</materialsCitation>
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<caption id="DF358591FF8BFFEBD0A92E22AB988940" ID-DOI="http://doi.org/10.5281/zenodo.5753274" ID-Zenodo-Dep="5753274" httpUri="https://zenodo.org/record/5753274/files/figure.png" pageId="14" pageNumber="1285" startId="14.[146,224,783,805]" targetBox="[149,757,195,743]" targetPageId="14">
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<paragraph id="8BF5D519FF8BFFEBD0A92E22AB988940" blockId="14.[145,761,783,835]" pageId="14" pageNumber="1285">
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<emphasis id="B93E090BFF8BFFEBD0A92E22A9EF8926" bold="true" box="[146,250,783,805]" pageId="14" pageNumber="1285">Figure 7.</emphasis>
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Habitus of
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<taxonomicName id="4C4AAE9AFF8BFFEBD1432E22AB848926" authorityName="Hsiao & Li & Ren & Pang" authorityYear="2021" box="[376,657,783,805]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="maha">
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||
<emphasis id="B93E090BFF8BFFEBD1432E22AB848926" box="[376,657,783,805]" italics="true" pageId="14" pageNumber="1285">Hukawngichthyurus maha</emphasis>
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</taxonomicName>
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||
: A, dorsal view; B, ventral-lateral view. Scale bars: 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="8BF5D519FF8BFFEBD0AA2EC1ABEF8EF5" blockId="14.[145,762,1003,1270]" pageId="14" pageNumber="1285">
|
||
<emphasis id="B93E090BFF8BFFEBD0AA2EC1A81F8E02" box="[145,266,1004,1025]" italics="true" pageId="14" pageNumber="1285">Diagnosis:</emphasis>
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It can be distinguished from its congener by the head with distinct setae (
|
||
<figureCitation id="1371C99CFF8BFFEBD22B2927AB918E1C" box="[528,644,1034,1056]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8A, B</figureCitation>
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||
) (without setae in
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<taxonomicName id="4C4AAE9AFF8BFFEBD0D42904A8D88E3E" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[239,461,1064,1086]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="kyawkhaingwini">
|
||
<emphasis id="B93E090BFF8BFFEBD0D42904A8D88E3E" box="[239,461,1064,1086]" italics="true" pageId="14" pageNumber="1285">H. kyawkhaingwini</emphasis>
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||
</taxonomicName>
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||
); thick scape and enlarged antennomere 3–6 (
|
||
<figureCitation id="1371C99CFF8BFFEBD14B296AA8D18E5F" box="[368,452,1095,1117]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8C</figureCitation>
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||
) (scape thin and elongate and antennomere 3–5 stout, only slightly enlarged in
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<taxonomicName id="4C4AAE9AFF8BFFEBD08829A8A8B18E9A" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[179,420,1156,1178]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="kyawkhaingwini">
|
||
<emphasis id="B93E090BFF8BFFEBD08829A8A8B18E9A" box="[179,420,1156,1178]" italics="true" pageId="14" pageNumber="1285">H. kyawkhaingwini</emphasis>
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</taxonomicName>
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); apical tergite moderately developed, with obtuse apex (
|
||
<figureCitation id="1371C99CFF8BFFEBD22F298EAB648EBB" box="[532,625,1187,1209]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8E</figureCitation>
|
||
) (strongly developed,with acute apex in
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD1EA29EFABB98ED4" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[465,684,1218,1239]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="kyawkhaingwini">
|
||
<emphasis id="B93E090BFF8BFFEBD1EA29EFABB98ED4" box="[465,684,1218,1239]" italics="true" pageId="14" pageNumber="1285">H.kyawkhaingwini</emphasis>
|
||
</taxonomicName>
|
||
);tibial spurs present (
|
||
<figureCitation id="1371C99CFF8BFFEBD10229CDA89F8EF6" box="[313,394,1248,1270]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8G</figureCitation>
|
||
) (absent in
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD23629CCABFE8EF6" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[525,747,1248,1270]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="kyawkhaingwini">
|
||
<emphasis id="B93E090BFF8BFFEBD23629CCABFE8EF6" box="[525,747,1248,1270]" italics="true" pageId="14" pageNumber="1285">H. kyawkhaingwini</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF5D519FF8BFFEBD0AA280DAA948E14" blockId="14.[145,761,1311,1885]" lastBlockId="14.[809,1426,197,1047]" pageId="14" pageNumber="1285">
|
||
<emphasis id="B93E090BFF8BFFEBD0AA280DA80B8F36" box="[145,286,1312,1333]" italics="true" pageId="14" pageNumber="1285">Description:</emphasis>
|
||
Male. Body 3.0 mm long, 1.0 mm wide, parallel-sided, densely and finely pubescent, with pubescence distinct in head. Body yellowish brown, with dark brown elytra (
|
||
<figureCitation id="1371C99CFF8BFFEBD1FF2856AB1C8F93" box="[452,521,1403,1425]" captionStart="Figure 7" captionStartId="14.[146,224,783,805]" captionTargetBox="[149,757,195,743]" captionTargetId="figure-687@14.[149,757,195,743]" captionTargetPageId="14" captionText="Figure 7. Habitus of Hukawngichthyurus maha: A, dorsal view; B, ventral-lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5753274" httpUri="https://zenodo.org/record/5753274/files/figure.png" pageId="14" pageNumber="1285">Fig. 7</figureCitation>
|
||
). Head (
|
||
<figureCitation id="1371C99CFF8BFFEBD24E2856ABE58F93" box="[629,752,1403,1425]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8A, B</figureCitation>
|
||
)
|
||
<quantity id="4CB278FCFF8BFFEBD0AA28B4A9F38FAC" box="[145,230,1433,1455]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="1285" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long,
|
||
<quantity id="4CB278FCFF8BFFEBD11128B4A8958FAC" box="[298,384,1433,1455]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="1285" unit="mm" value="0.5">0.5 mm</quantity>
|
||
wide. Vertex finely wrinkled and with setae, slightly convex, slightly narrowed behind the eyes. Eyes large, hemispherically prominent, with fine facets, interocular distance 2.2 times maximum eye diameter. Anterior margin of clypeus somewhat concave, frontoclypeal suture absent. Labrum subquadrate, membranous, exposed. Gular sutures close, nearly confluent. Mandibles elongate, strongly curved at apical fourth, with a tooth on inflexed outer margin. Apical segment of maxillary palpi flattened, elongate, securiform, with apex straight-truncated; apical labial palpomere triangular. Antennae (
|
||
<figureCitation id="1371C99CFF8BFFEBD29B2BC6ABE58D03" box="[672,752,1771,1793]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8C</figureCitation>
|
||
) filiform, with central antennomeres enlarged, 11-segmented, extending to half of elytral length; scape thick, widened apically; pedicel short, stout, moniliform; antennomere 3–6 stout, enlarged, subequal in length, longer than pedicel; antennomeres 7–11 filiform; relative lengths of antennomeres 3–11: 1.0:1.2:1.2:1.2:1.2:1.1:1.1:1.1:1.2. Pronotum (
|
||
<figureCitation id="1371C99CFF8BFFEBD5082C0CAC9D8B35" box="[1331,1416,289,311]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8D</figureCitation>
|
||
)
|
||
<quantity id="4CB278FCFF8BFFEBD3122C6DAA928B56" box="[809,903,320,341]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="1285" unit="mm" value="0.4">0.4 mm</quantity>
|
||
long,
|
||
<quantity id="4CB278FCFF8BFFEBD3E12C6DAD228B56" box="[986,1079,320,342]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="1285" unit="mm" value="0.6">0.6 mm</quantity>
|
||
wide, transverse. Anterior margin slightly arcuate, posterior margin slightly concave, lateral margins narrowed apically in anterior half; disc irregular, with a large depression anteriorly. Scutellum narrowed apically, truncate at apex. Elytra (
|
||
<figureCitation id="1371C99CFF8BFFEBD30A2CF4AA9F8BED" box="[817,906,473,495]" captionStart="Figure 7" captionStartId="14.[146,224,783,805]" captionTargetBox="[149,757,195,743]" captionTargetId="figure-687@14.[149,757,195,743]" captionTargetPageId="14" captionText="Figure 7. Habitus of Hukawngichthyurus maha: A, dorsal view; B, ventral-lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5753274" httpUri="https://zenodo.org/record/5753274/files/figure.png" pageId="14" pageNumber="1285">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation id="1371C99CFF8BFFEBD3AC2CF4AAAC8BED" box="[919,953,473,494]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">8D</figureCitation>
|
||
)
|
||
<quantity id="4CB278FCFF8BFFEBD3F22CF4AD0A8BED" box="[969,1055,473,495]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="1285" unit="mm" value="2.5">2.5 mm</quantity>
|
||
long,
|
||
<quantity id="4CB278FCFF8BFFEBD45E2CF4ADAE8BED" box="[1125,1211,473,494]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="1285" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide, subparallelsided, somewhat narrowed before the middle, rounded at apex; elytra smooth; hindwing well-developed, exposed beyond elytra. Abdomen (
|
||
<figureCitation id="1371C99CFF8BFFEBD4EB2F18AC3D8849" box="[1232,1320,565,587]" captionStart="Figure 7" captionStartId="14.[146,224,783,805]" captionTargetBox="[149,757,195,743]" captionTargetId="figure-687@14.[149,757,195,743]" captionTargetPageId="14" captionText="Figure 7. Habitus of Hukawngichthyurus maha: A, dorsal view; B, ventral-lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5753274" httpUri="https://zenodo.org/record/5753274/files/figure.png" pageId="14" pageNumber="1285">Fig. 7B</figureCitation>
|
||
) weakly sclerotized, with eight ventrites; first six segments broad, as wide as elytra; last two segments (
|
||
<figureCitation id="1371C99CFF8BFFEBD50D2F5FAA2C88A5" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8E, F</figureCitation>
|
||
) narrower, apical tergite (
|
||
<figureCitation id="1371C99CFF8BFFEBD45E2FBCADA288A5" box="[1125,1207,657,679]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8E</figureCitation>
|
||
) distinctly furcate, ventrite 7 (
|
||
<figureCitation id="1371C99CFF8BFFEBD3802F9DAD0688C6" box="[955,1043,688,710]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8F</figureCitation>
|
||
) with apical margin straight, ventrite 8 tapered apically, forming narrow, elongate process. Aedeagus (
|
||
<figureCitation id="1371C99CFF8BFFEBD43A2FC0AD5A8901" box="[1025,1103,749,771]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8F</figureCitation>
|
||
) partially exposed, with pair of slender ventral processes, curved outwards. Legs (
|
||
<figureCitation id="1371C99CFF8BFFEBD30A2E07AA93893C" box="[817,902,810,832]" captionStart="Figure 8" captionStartId="15.[163,241,1245,1267]" captionTargetBox="[403,1203,195,1205]" captionTargetId="figure-356@15.[403,1203,195,1205]" captionTargetPageId="15" captionText="Figure 8. Details of body parts of Hukawngichthyurus maha: A, head, ventral-lateral view; B, ditto, dorsal view; C, antenna indicated by a black arrow; D, pronotum and elytra; E, apical tergite; F, caudal segments, with the process of aedeagus indicated by a black arrow; G, tarsus of left mid leg, with the tibial spurs indicated by a black arrow. Scale bars: 0.2 mm (A, B, E, F, G); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5753276" httpUri="https://zenodo.org/record/5753276/files/figure.png" pageId="14" pageNumber="1285">Fig. 8G</figureCitation>
|
||
) short; profemur length
|
||
<quantity id="4CB278FCFF8BFFEBD4912E07AC168943" box="[1194,1283,810,832]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="1285" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, mesofemur
|
||
<quantity id="4CB278FCFF8BFFEBD3122E64AA95895D" box="[809,896,841,862]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="1285" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, and metafemur
|
||
<quantity id="4CB278FCFF8BFFEBD47A2E64AD82895D" box="[1089,1175,841,862]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="1285" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; tibiae slender, nearly straight, with pair of tibial spurs, protibia length
|
||
<quantity id="4CB278FCFF8BFFEBD3122EABAA90899F" box="[809,901,902,924]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="1285" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, mesotibia
|
||
<quantity id="4CB278FCFF8BFFEBD4292EABAD7B899F" box="[1042,1134,902,924]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="1285" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, and metatibia
|
||
<quantity id="4CB278FCFF8BFFEBD50B2EABAC9E899F" box="[1328,1419,902,924]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="1285" unit="mm" value="0.8">0.8 mm</quantity>
|
||
; tarsal formula 5-5-5; tarsomeres 1–3 simple, tarsomere 1 as long as combined length of tarsomere 2 and 3; tarsomere 4 bilobed; claws simple, with minute dent at base.
|
||
</paragraph>
|
||
<paragraph id="8BF5D519FF8BFFEBD312296DAC878EB2" blockId="14.[809,1426,1088,1201]" pageId="14" pageNumber="1285">
|
||
<emphasis id="B93E090BFF8BFFEBD312296DAAB28E56" box="[809,935,1088,1109]" italics="true" pageId="14" pageNumber="1285">Etymology:</emphasis>
|
||
The specific epithet is derived from
|
||
<emphasis id="B93E090BFF8BFFEBD56B296DAC738E56" box="[1360,1382,1088,1109]" italics="true" pageId="14" pageNumber="1285">m</emphasis>
|
||
ā
|
||
<emphasis id="B93E090BFF8BFFEBD54A296DAC958E56" box="[1393,1408,1088,1109]" italics="true" pageId="14" pageNumber="1285">h</emphasis>
|
||
ā, ‘great or noble’ in Sanskrit, referring to the discovery of this species being a great contribution to the palaeontology of
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD3D929B6AD648EB2" authorityName="IMHOFF" authorityYear="1856" box="[994,1137,1179,1201]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
|
||
. It is a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="8BF5D519FF8BFFEBD31229F6AA668C7D" blockId="14.[809,1425,1242,1662]" pageId="14" pageNumber="1285">
|
||
<emphasis id="B93E090BFF8BFFEBD31229F6AAB28EF3" box="[809,935,1242,1264]" italics="true" pageId="14" pageNumber="1285">Remarks:</emphasis>
|
||
This species is closely related to
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD31229D7AD138F0D" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[809,1030,1273,1295]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="kyawkhaingwini">
|
||
<emphasis id="B93E090BFF8BFFEBD31229D7AD138F0D" box="[809,1030,1273,1295]" italics="true" pageId="14" pageNumber="1285">H. kyawkhaingwini</emphasis>
|
||
</taxonomicName>
|
||
with which it shares the small body size, the head shape with the small rostrum, the apical maxillary palpomere with straight-truncated apex, the filiform antennae with the short, stout and moniliform pedicel and the stout central antennomeres, the transverse pronotum, with large depression on the disc anteriorly, the smooth and distinctly shortened elytra, with largely exposed caudal segments and hindwings and the furcate caudal segment. Considering that only four minute differences are identified between
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD3122B01AD058C43" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[809,1040,1579,1601]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="kyawkhaingwini">
|
||
<emphasis id="B93E090BFF8BFFEBD3122B01AD058C43" box="[809,1040,1579,1601]" italics="true" pageId="14" pageNumber="1285">H. kyawkhaingwini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD4762B01ADA18C43" authorityName="Hsiao & Li & Ren & Pang" authorityYear="2021" box="[1101,1204,1579,1601]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="species" species="maha">
|
||
<emphasis id="B93E090BFF8BFFEBD4762B01ADA18C43" box="[1101,1204,1579,1601]" italics="true" pageId="14" pageNumber="1285">H. maha</emphasis>
|
||
</taxonomicName>
|
||
, we place it in the genus
|
||
<taxonomicName id="4C4AAE9AFF8BFFEBD3492B67AD4E8C5C" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[882,1115,1610,1631]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93E090BFF8BFFEBD3492B67AD4E8C5C" box="[882,1115,1610,1631]" italics="true" pageId="14" pageNumber="1285">Hukawngichthyurus</emphasis>
|
||
</taxonomicName>
|
||
, rather than propose a new genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |