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<mods:title id="27E333457F917D315575B34D8BD794FA">Revision of the genus Odocnemis Allard, 1876 (Coleoptera: Tenebrionidae: Helopini) from Turkey, the Caucasus and Iran with observations on feeding habits</mods:title>
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<treatment id="03ADFF28F87F495FFF1EF92C3FF4F910" ID-DOI="http://doi.org/10.5281/zenodo.5681120" ID-GBIF-Taxon="124940356" ID-Zenodo-Dep="5681120" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910" httpUri="http://treatment.plazi.org/id/03ADFF28F87F495FFF1EF92C3FF4F910" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45">
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<paragraph id="8BBB4E3EF87F495DFF1EF92C3EE1F8DA" blockId="44.[151,747,1770,1830]" box="[151,747,1770,1797]" pageId="44" pageNumber="45">
<heading id="D0F3F952F87F495DFF1EF92C3EE1F8DA" bold="true" box="[151,747,1770,1797]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<taxonomicName id="4C0435BDF87F495DFF1EF92C3EE1F8DA" authority="Reitter in Bodemeyer, 1900" authorityName="Reitter in Bodemeyer" authorityYear="1900" box="[151,747,1770,1797]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="protina">
<emphasis id="B970922CF87F495DFF1EF92C3EE1F8DA" bold="true" box="[151,747,1770,1797]" pageId="44" pageNumber="45">
<emphasis id="B970922CF87F495DFF1EF92C3D77F8DB" bold="true" box="[151,381,1770,1796]" italics="true" pageId="44" pageNumber="45">Odocnemis protina</emphasis>
(Reitter in
<bibRefCitation id="EF9533CFF87F495DFD84F92D3EE9F8DA" author="Bodemeyer" box="[525,739,1771,1797]" pageId="44" pageNumber="45" refString="Bodemeyer, E. von (1900) Quer durch Klein-Asien in den Bulghar-Dagh. Eine naturwissenschaftliche Studien-Reise. Verlangs- Aktiengesseschaft vormals Dolter, Emmendingen, v + 169 pp." type="book" year="1900">Bodemeyer, 1900</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBB4E3EF87F495DFF1EF8CB3DCCF8F9" blockId="44.[151,747,1770,1830]" box="[151,454,1805,1830]" pageId="44" pageNumber="45">
<figureCitation id="133F52BBF87F495DFF1EF8CB3CF9F8F9" box="[151,243,1805,1830]" captionStart="FIGURE 24. O" captionStartId="45.[151,250,1888,1910]" captionTargetBox="[151,1435,201,1863]" captionTargetId="figure@45.[151,1436,193,1866]" captionTargetPageId="45" captionText="FIGURE 24. O. protina. A) Aedeagus, ventral view. B) Aedeagus, lateral view. C) Inner sternite VIII. D) Gastral spicula. E) Protibia. F) Mesotibia. G) Metatibia. H) Head. I), J) Pronotum. A I) Male. J) Female. Scale bars— 1 mm." httpUri="https://zenodo.org/record/192243/files/figure.png" pageId="44" pageNumber="45">
Figs.
<date id="FFBA68FEF87F495DFF5CF8C83CF9F8F9" box="[213,243,1806,1830]" pageId="44" pageNumber="45" value="1940-01-24">24</date>
</figureCitation>
,
<figureCitation id="133F52BBF87F495DFF76F8C83D17F8F9" box="[255,285,1806,1830]" captionStart="FIGURE 40" captionStartId="73.[151,250,1854,1876]" captionTargetBox="[152,1436,194,1818]" captionTargetId="figure@73.[151,1436,193,1833]" captionTargetPageId="73" captionText="FIGURE 40. Epipleura of Odocnemis. A), B) O. anatolica. C), D) O. amanosica. E) O. torosica torosica (Çamlıyayla). F) O. torosica torosica (Arslanköy). G), H) O. inornata. I), J) O. protina. K), L) O. euritopica. M), N) O. allardi. A), C), G), I), K), M) Ventral view. B). D), E), F), H), J), L) Dorsal view." httpUri="https://zenodo.org/record/192260/files/figure.png" pageId="44" pageNumber="45">40</figureCitation>
I, J, 46C, D, 49
</paragraph>
</subSubSection>
<subSubSection id="C31E1DB5F87F495DFF1EF8903EC0F839" pageId="44" pageNumber="45" type="reference_group">
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<taxonomicName id="4C0435BDF87F495DFF1EF8903E5AF8B4" authority="Reitter in Bodemeyer, 1900: 167" authorityName="Reitter in Bodemeyer" authorityPageNumber="167" authorityYear="1900" box="[151,592,1877,1899]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="protinus">
<emphasis id="B970922CF87F495DFF1EF8903CE1F8B4" box="[151,235,1878,1899]" italics="true" pageId="44" pageNumber="45">protinus</emphasis>
Reitter in
<treatmentCitation id="0AA5682FF87F495DFEEBF8933E5AF8B4" author="Bodemeyer" box="[354,592,1877,1899]" page="167" pageId="44" pageNumber="45" year="1900">
<bibRefCitation id="EF9533CFF87F495DFEEBF8933E16F8B4" author="Bodemeyer" box="[354,540,1877,1899]" pageId="44" pageNumber="45" refString="Bodemeyer, E. von (1900) Quer durch Klein-Asien in den Bulghar-Dagh. Eine naturwissenschaftliche Studien-Reise. Verlangs- Aktiengesseschaft vormals Dolter, Emmendingen, v + 169 pp." type="book" year="1900">Bodemeyer, 1900</bibRefCitation>
: 167
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</taxonomicName>
(
<taxonomicName id="4C0435BDF87F495DFDEAF8933F30F8B4" authority="(Odocnemis)" baseAuthorityName="Odocnemis" box="[611,826,1877,1899]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFDEAF8933EA1F8B4" box="[611,683,1877,1899]" italics="true" pageId="44" pageNumber="45">Helops</emphasis>
(
<emphasis id="B970922CF87F495DFD37F8933F3AF8B4" box="[702,816,1877,1899]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
)
</taxonomicName>
, description);
<treatmentCitation id="0AA5682FF87F495DFC5AF893389AF8B4" author="Gebien" box="[979,1168,1877,1899]" page="551" pageId="44" pageNumber="45" year="1911">
<bibRefCitation id="EF9533CFF87F495DFC5AF8933856F8B4" author="Gebien" box="[979,1116,1877,1899]" pageId="44" pageNumber="45" refString="Gebien, H. (1911) Pars. 28. Tenebrionidae. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus auspiciis et auxilio. Fol. 18. Junk, W., Berlin, pp. 355 - 585" type="journal article" year="1911">Gebien 1911</bibRefCitation>
: 551
</treatmentCitation>
(
<taxonomicName id="4C0435BDF87F495DFB2AF8933973F8B4" authority="(Odocnemis)" baseAuthorityName="Odocnemis" box="[1187,1401,1877,1899]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFB2AF89338E1F8B4" box="[1187,1259,1877,1899]" italics="true" pageId="44" pageNumber="45">Helops</emphasis>
(
<emphasis id="B970922CF87F495DFB74F8933965F8B4" box="[1277,1391,1877,1899]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
)
</taxonomicName>
, in catalogue);
<treatmentCitation id="0AA5682FF87F495DFEC0F8B23E18F855" author="Reitter" box="[329,530,1908,1930]" page="132" pageId="44" pageNumber="45" year="1922">
<bibRefCitation id="EF9533CFF87F495DFEC0F8B23DDCF855" author="Reitter" box="[329,470,1908,1930]" pageId="44" pageNumber="45" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44, 113 - 171. http: // dx. doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter 1922</bibRefCitation>
: 132
</treatmentCitation>
(
<taxonomicName id="4C0435BDF87F495DFDA4F8B23F58F855" authority="(Odocnemis)" baseAuthorityName="Odocnemis" box="[557,850,1908,1930]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFDA4F8B23EB7F855" box="[557,701,1908,1930]" italics="true" pageId="44" pageNumber="45">Cylindronotus</emphasis>
(
<emphasis id="B970922CF87F495DFD5EF8B23F43F855" box="[727,841,1908,1930]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
)
</taxonomicName>
in key);
<bibRefCitation id="EF9533CFF87F495DFC59F8B23862F855" author="Winkler" box="[976,1128,1908,1930]" pageId="44" pageNumber="45" refString="Winkler, A. (1928 - 1929) Pars. 8, 9: D. Heteromera. In: Winkler, A. (Ed.) Catalogus Coleopterorum regionis Palaearctica. Verlag von Albert Winkler, Wien, pp. 881 - 1136." type="book chapter" year="1928">
Winkler
<date id="FFBA68FEF87F495DFBBDF8B23862F855" box="[1076,1128,1908,1930]" pageId="44" pageNumber="45" value="1928" valueMax="1929">1928</date>
</bibRefCitation>
1929: 1018 (
<taxonomicName id="4C0435BDF87F495DFA85F8B23D41F877" authority="(Odocnemis)" baseAuthorityName="Odocnemis" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFA85F8B23996F855" box="[1292,1436,1908,1930]" italics="true" pageId="44" pageNumber="45">Cylindronotus</emphasis>
(
<emphasis id="B970922CF87F495DFF46F8543D4BF877" box="[207,321,1938,1960]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
)
</taxonomicName>
, in catalogue);
<treatmentCitation id="0AA5682FF87F495DFE7BF8543EBAF877" author="Gebien" box="[498,688,1938,1960]" page="427" pageId="44" pageNumber="45" year="1943">
<bibRefCitation id="EF9533CFF87F495DFE7BF8543E70F877" author="Gebien" box="[498,634,1938,1960]" pageId="44" pageNumber="45" refString="Gebien, H. (1943) Katalog der Tenebrioniden. Teil. 3. Mitteilungen der Munchner Entomologischen Gesellschaft, 33, 399 - 430 (778 - 809)." type="journal article" year="1943">Gebien 1943</bibRefCitation>
: 427
</treatmentCitation>
(806) (
<taxonomicName id="4C0435BDF87F495DFD7EF8543819F877" authority="(Odocnemis)" baseAuthorityName="Odocnemis" box="[759,1043,1938,1960]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFD7EF8543F8DF877" box="[759,903,1938,1960]" italics="true" pageId="44" pageNumber="45">Cylindronotus</emphasis>
(
<emphasis id="B970922CF87F495DFC10F8543801F877" box="[921,1035,1938,1960]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
)
</taxonomicName>
in catalogue); Nabozhenko 2008: 37 (
<taxonomicName id="4C0435BDF87F495DFF46F8773D4BF818" box="[207,321,1969,1991]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFF46F8773D4BF818" box="[207,321,1969,1991]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
</taxonomicName>
, in catalogue);
<treatmentCitation id="0AA5682FF87F495DFE6BF8773F28F818" author="Nabozhenko" box="[482,802,1969,1991]" page="244" pageId="44" pageNumber="45" year="2008">
<bibRefCitation id="EF9533CFF87F495DFE6BF8773EFBF818" author="Nabozhenko" box="[482,753,1969,1991]" pageId="44" pageNumber="45" refString="Nabozhenko, M. V. &amp; Lobl, I. (2008) Tribe Helopini. In: Lobl, I. &amp; Smetana, A. (Eds.) Catalogue of Palearctic Coleoptera. Fol. 5. Tenebrionoidea. Stenstrup, Apollo books, pp. 241 - 257." type="journal article" year="2008">Nabozhenko &amp; Löbl 2008</bibRefCitation>
: 244
</treatmentCitation>
(
<taxonomicName id="4C0435BDF87F495DFCB8F8773FA9F818" box="[817,931,1969,1991]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFCB8F8773FA9F818" box="[817,931,1969,1991]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
</taxonomicName>
, in catalogue);
<treatmentCitation id="0AA5682FF87F495DFBCDF8773985F818" author="Nabozhenko" box="[1092,1423,1969,1991]" page="70" pageId="44" pageNumber="45" year="2013">
<bibRefCitation id="EF9533CFF87F495DFBCDF8773966F818" author="Nabozhenko" box="[1092,1388,1969,1991]" pageId="44" pageNumber="45" refString="Nabozhenko, M. V., Keskin, B. &amp; Alpagut Keskin, N. (2013) Molecular-genetic research of darkling beetles of the genus Odocnemis (Coleoptera: Tenebrionidae). In: Molecular genetic approaches in taxonomy and ecology. Abstracts of the Scientific Conference (March 25 - 29 2013, Rostov-on-Don, Russia). Southern Scientific Centre Publishers, Rostov-on- Don, p. 60. [in Russian]" type="journal article" year="2013">Nabozhenko &amp; Keskin 2013</bibRefCitation>
: 70
</treatmentCitation>
71 (
<taxonomicName id="4C0435BDF87F495DFF66F8163D6BF839" box="[239,353,2000,2022]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF87F495DFF66F8163D6BF839" box="[239,353,2000,2022]" italics="true" pageId="44" pageNumber="45">Odocnemis</emphasis>
</taxonomicName>
, description of Iranian population).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF7B1EB6F87E495CFF1EF8A638C9F84B" httpUri="https://zenodo.org/record/192243/files/figure.png" pageId="45" pageNumber="46" targetBox="[151,1435,201,1863]" targetPageId="45">
<paragraph id="8BBB4E3EF87E495CFF1EF8A638C9F84B" blockId="45.[151,1435,1888,1941]" pageId="45" pageNumber="46">
<emphasis id="B970922CF87E495CFF1EF8A63D2BF8AA" bold="true" box="[151,289,1888,1910]" pageId="45" pageNumber="46">FIGURE 24.</emphasis>
<emphasis id="B970922CF87E495CFEA1F8A73D93F8A9" box="[296,409,1889,1910]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="4C0435BDF87E495CFEA1F8A73D9FF8A9" box="[296,405,1889,1910]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="protina">O. protina</taxonomicName>
.
</emphasis>
A) Aedeagus, ventral view. B) Aedeagus, lateral view. C) Inner sternite VIII. D) Gastral spicula. E) Protibia. F) Mesotibia. G) Metatibia. H) Head. I), J) Pronotum. AI) Male. J) Female. Scale bars—1 mm.
</paragraph>
</caption>
<subSubSection id="C31E1DB5F87D495FFF1EFF513916FD5B" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BBB4E3EF87D495FFF1EFF513DFCFE9F" blockId="46.[151,1437,151,1744]" pageId="46" pageNumber="47">
<materialsCitation id="3B6C4463F87D495FFF1EFF513D91FF27" ID-GBIF-Occurrence="1413110379" collectionCode="HNHM" country="Turkey" county="Typen" location="Biledjek" municipality="Bodemeyer" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B970922CF87D495FFF1EFF513E3EFF6F" bold="true" box="[151,564,151,176]" pageId="46" pageNumber="47">
<typeStatus id="54BFF09CF87D495FFF1EFF513CD8FF6F" box="[151,210,151,176]" pageId="46" pageNumber="47">Type</typeStatus>
material (studied,
<collectionCode id="ED15D6FBF87D495FFE59FF513E26FF6F" box="[464,556,151,176]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="46" pageNumber="47">HNHM</collectionCode>
)
</emphasis>
.
<typeStatus id="54BFF09CF87D495FFDC2FF5E3EB1FF6F" box="[587,699,152,176]" pageId="46" pageNumber="47" type="lectotype">Lectotype</typeStatus>
(
<specimenCount id="9D0285B7F87D495FFD5DFF5E3EE0FF6F" box="[724,746,152,176]" pageId="46" pageNumber="47" type="male"></specimenCount>
) designated here, with labels: “
<collectingCountry id="F3130EAEF87D495FFBF3FF51390FFF6F" box="[1146,1285,151,176]" name="Turkey" pageId="46" pageNumber="47">Asia minor</collectingCountry>
,
<location id="8EDB18E5F87D495FFA90FF513972FF6F" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFA90FF513972FF6F" box="[1305,1400,151,176]" country="Turkey" county="Typen" municipality="Bodemeyer" name="Biledjek" pageId="46" pageNumber="47">Biledjek</location>
v.
<collectingMunicipality id="6BDFD444F87D495FFF1EFF7A3D17FF0A" box="[151,285,188,213]" pageId="46" pageNumber="47">Bodemeyer</collectingMunicipality>
”, “Coll. Reitter”, “
<taxonomicName id="4C0435BDF87D495FFE76FF7B3E74FF0B" box="[511,638,189,212]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="protinus">
<emphasis id="B970922CF87D495FFE76FF7B3E74FF0B" box="[511,638,189,212]" italics="true" pageId="46" pageNumber="47">O. protinus</emphasis>
</taxonomicName>
m. 1900
<typeStatus id="54BFF09CF87D495FFD7EFF7A3F48FF0B" box="[759,834,188,212]" pageId="46" pageNumber="47">
<collectingCounty id="62DA36B2F87D495FFD7EFF7A3F48FF0B" box="[759,834,188,212]" pageId="46" pageNumber="47">Typen</collectingCounty>
</typeStatus>
”, “
<typeStatus id="54BFF09CF87D495FFCEFFF7A3FD7FF0A" box="[870,989,188,213]" pageId="46" pageNumber="47" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C0435BDF87D495FFC60FF7A3CDBFF27" authority="Reitter, 1900" authorityName="Reitter" authorityYear="1900" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="protinus" subGenus="Odocnemis">
<emphasis id="B970922CF87D495FFC60FF7A3832FF0A" box="[1001,1080,188,213]" italics="true" pageId="46" pageNumber="47">Helops</emphasis>
(
<emphasis id="B970922CF87D495FFBC7FF7A38C6FF0A" box="[1102,1228,188,213]" italics="true" pageId="46" pageNumber="47">Odocnemis</emphasis>
)
<emphasis id="B970922CF87D495FFB68FF7B3934FF0B" box="[1249,1342,189,212]" italics="true" pageId="46" pageNumber="47">protinus</emphasis>
Reitter, 1900
</taxonomicName>
” (curators lable)
</materialsCitation>
.
<materialsCitation id="3B6C4463F87D495FFE2FFF193992FF27" ID-GBIF-Occurrence="1413110382" box="[422,1432,223,248]" country="Turkey" location="Goek-Dagh" municipality="Bodemeyer" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<typeStatus id="54BFF09CF87D495FFE2FFF193E35FF27" box="[422,575,223,248]" pageId="46" pageNumber="47" type="paralectotype">Paralectotype</typeStatus>
(
<specimenCount id="9D0285B7F87D495FFDD8FF263E63FF27" box="[593,617,224,248]" pageId="46" pageNumber="47" type="male"></specimenCount>
): “
<collectingCountry id="F3130EAEF87D495FFD00FF193F03FF27" box="[649,777,223,248]" name="Turkey" pageId="46" pageNumber="47">Asia minor</collectingCountry>
<location id="8EDB18E5F87D495FFC98FF193F9EFF27" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFC98FF193F9EFF27" box="[785,916,223,248]" country="Turkey" municipality="Bodemeyer" name="Goek-Dagh" pageId="46" pageNumber="47">Goek-Dagh</location>
v.
<collectingMunicipality id="6BDFD444F87D495FFC3EFF193837FF27" box="[951,1085,223,248]" pageId="46" pageNumber="47">Bodemeyer</collectingMunicipality>
”, “
<taxonomicName id="4C0435BDF87D495FFBE9FF273984FF27" authority="Rtt., Coll. Reitter" authorityName="Rtt., Coll. Reitter" box="[1120,1422,223,248]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="protinus">
<emphasis id="B970922CF87D495FFBE9FF2738B6FF27" box="[1120,1212,225,248]" italics="true" pageId="46" pageNumber="47">protinus</emphasis>
Rtt., Coll. Reitter
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B6C4463F87D495FFF1EFEC23DFCFE9F" ID-GBIF-Occurrence="1413110376" country="Turkey" location="Sabandja" municipality="Bodemeyer" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="54BFF09CF87D495FFF1EFEC23D25FEC2" box="[151,303,260,285]" pageId="46" pageNumber="47" type="paralectotype">Paralectotype</typeStatus>
(
<specimenCount id="9D0285B7F87D495FFECBFEC23D5FFEC3" box="[322,341,260,284]" pageId="46" pageNumber="47" type="female"></specimenCount>
): “
<collectingCountry id="F3130EAEF87D495FFEFFFEC23DFCFEC2" box="[374,502,260,285]" name="Turkey" pageId="46" pageNumber="47">Asia minor</collectingCountry>
<location id="8EDB18E5F87D495FFE74FEC23E6EFEC2" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFE74FEC23E6EFEC2" box="[509,612,260,285]" country="Turkey" municipality="Bodemeyer" name="Sabandja" pageId="46" pageNumber="47">Sabandja</location>
v.
<collectingMunicipality id="6BDFD444F87D495FFD0DFEC23F00FEC2" box="[644,778,260,285]" pageId="46" pageNumber="47">Bodemeyer</collectingMunicipality>
”, “Coll. Reitter”, “
<typeStatus id="54BFF09CF87D495FFC53FEC23841FEC3" box="[986,1099,260,284]" pageId="46" pageNumber="47" type="paratype">Paratypus</typeStatus>
<taxonomicName id="4C0435BDF87D495FFBDBFEC23D20FE9F" authority="Reitter, 1900" authorityName="Reitter" authorityYear="1900" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="protinus" subGenus="Odocnemis">
<emphasis id="B970922CF87D495FFBDBFEC238ABFEC2" box="[1106,1185,260,285]" italics="true" pageId="46" pageNumber="47">Helops</emphasis>
(
<emphasis id="B970922CF87D495FFB38FEC23925FEC2" box="[1201,1327,260,285]" italics="true" pageId="46" pageNumber="47">Odocnemis</emphasis>
)
<emphasis id="B970922CF87D495FFAB6FEC33991FEC3" box="[1343,1435,261,284]" italics="true" pageId="46" pageNumber="47">protinus</emphasis>
Reitter, 1900
</taxonomicName>
” (curators label).
</materialsCitation>
</paragraph>
<paragraph id="8BBB4E3EF87D495FFF4EFE8A3938FEBB" blockId="46.[151,1437,151,1744]" box="[199,1330,332,357]" pageId="46" pageNumber="47">
One
<typeStatus id="54BFF09CF87D495FFF74FE8A3D9EFEBA" box="[253,404,332,357]" pageId="46" pageNumber="47" type="paralectotype">paralectotype</typeStatus>
is deposited in collection of Natural History Museum Stuttgart (
<bibRefCitation id="EF9533CFF87D495FFBEDFE8A392CFEBB" author="Schawaller" box="[1124,1318,332,357]" pageId="46" pageNumber="47" refString="Schawaller, W. (1994) Die Kafersammlung am Naturkundemuseum Stuttgart. Stuttgarter Beitrage zur Naturkunde Serie A (Biologie), 508, 1 - 40." type="journal article" year="1994">Schawaller 1994</bibRefCitation>
).
</paragraph>
<paragraph id="8BBB4E3EF87D495FFF4EFEA93916FD5B" blockId="46.[151,1437,151,1744]" pageId="46" pageNumber="47">
<materialsCitation id="3B6C4463F87D495FFF4EFEA93E58FE73" ID-GBIF-Occurrence="1413110380" collectingDate="1983-05-18" collectorName="Gasko" country="Turkey" county="Other" elevation="1200" location="Mts. Abant Dagi" municipality="Turcia" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bolu">
<emphasis id="B970922CF87D495FFF4EFEA93DEEFE57" bold="true" box="[199,484,367,392]" pageId="46" pageNumber="47">
<collectingCounty id="62DA36B2F87D495FFF4EFEA93D05FE57" box="[199,271,367,392]" pageId="46" pageNumber="47">Other</collectingCounty>
studied material.
</emphasis>
<collectingMunicipality id="6BDFD444F87D495FFE65FEA93E3DFE57" box="[492,567,367,392]" pageId="46" pageNumber="47">Turcia</collectingMunicipality>
, Mt.
<collectingRegion id="49C080DCF87D495FFDFCFEA93EA1FE57" box="[629,683,367,392]" country="Turkey" name="Bolu" pageId="46" pageNumber="47">Bolu</collectingRegion>
Dag Abant [
<collectingCountry id="F3130EAEF87D495FFCC8FEA93F9FFE57" box="[833,917,367,392]" name="Turkey" pageId="46" pageNumber="47">Turkey</collectingCountry>
,
<collectingRegion id="49C080DCF87D495FFC2BFEA93842FE57" box="[930,1096,367,392]" country="Turkey" name="Bolu" pageId="46" pageNumber="47">Bolu Province</collectingRegion>
,
<location id="8EDB18E5F87D495FFBDCFEB63910FE57" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFBDCFEB63910FE57" box="[1109,1306,367,392]" country="Turkey" county="Other" municipality="Turcia" name="Mts. Abant Dagi" pageId="46" pageNumber="47" stateProvince="Bolu">Mts. Abant Dağı</location>
], +
<quantity id="4CFCE3DBF87D495FFAB4FEB63992FE58" box="[1341,1432,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="46" pageNumber="47" unit="m" value="1200.0">
<elevation id="0029A90DF87D495FFAB4FEB63992FE58" box="[1341,1432,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="46" pageNumber="47" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="FFBA68FEF87D495FFF1EFE523D0DFE73" box="[151,263,404,428]" pageId="46" pageNumber="47" value="1983-05-18">
<collectingDate id="EFFE9116F87D495FFF1EFE523D0DFE73" box="[151,263,404,428]" pageId="46" pageNumber="47" value="1983-05-18">18.v.1983</collectingDate>
</date>
, leg.
<collectorName id="26F12BE8F87D495FFEDAFE523D97FE72" box="[339,413,404,429]" pageId="46" pageNumber="47">Gaskó</collectorName>
,
<specimenCount id="9D0285B7F87D495FFE39FE523DDEFE73" box="[432,468,404,428]" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFE61FE523E01FE73" box="[488,523,404,428]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
(CS)
</materialsCitation>
;
<materialsCitation id="3B6C4463F87D495FFDEFFE523F11FE2B" ID-GBIF-Occurrence="1413110372" collectingDate="2009-05-29" collectionCode="ZIN, ZDEU" collectorName="M. V. and S. V. Nabozhenko &amp; B. Keskin" country="Turkey" elevation="1260" latitude="31.25" location="Abant Dagi" municipality="Mudanya District" pageId="46" pageNumber="47" specimenCount="31" specimenCount-female="12" specimenCount-male="19" stateProvince="Bolu Province">
<collectingCountry id="F3130EAEF87D495FFDEFFE523EBDFE72" box="[614,695,404,429]" name="Turkey" pageId="46" pageNumber="47">Turkey</collectingCountry>
,
<collectingRegion id="49C080DCF87D495FFD40FE523F7FFE72" box="[713,885,404,429]" country="Turkey" name="Bolu" pageId="46" pageNumber="47">Bolu Province</collectingRegion>
,
<collectingMunicipality id="6BDFD444F87D495FFC01FE52385DFE72" box="[904,1111,404,429]" pageId="46" pageNumber="47">Mudanya District</collectingMunicipality>
,
<location id="8EDB18E5F87D495FFBE3FE5238FCFE72" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFBE3FE5238FCFE72" box="[1130,1270,404,429]" country="Turkey" latitude="31.25" municipality="Mudanya District" name="Abant Dagi" pageId="46" pageNumber="47" stateProvince="Bolu Province">Abant Dağı</location>
, 40°35135″
<geoCoordinate id="EE3028F9F87D495FFA0EFE523CD6FE0F" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="31.25">N 31°15</geoCoordinate>
894″E,
<quantity id="4CFCE3DBF87D495FFEB1FE713D9BFE10" box="[312,401,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="46" pageNumber="47" unit="m" value="1260.0">
<elevation id="0029A90DF87D495FFEB1FE713D9BFE10" box="[312,401,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="46" pageNumber="47" unit="m" value="1260.0">1260 m</elevation>
</quantity>
,
<date id="FFBA68FEF87D495FFE29FE7E3E1AFE0F" box="[416,528,440,464]" pageId="46" pageNumber="47" value="2009-05-29">
<collectingDate id="EFFE9116F87D495FFE29FE7E3E1AFE0F" box="[416,528,440,464]" pageId="46" pageNumber="47" value="2009-05-29">29.v.2009</collectingDate>
</date>
, leg.
<collectorName id="26F12BE8F87D495FFDD8FE7E3F84FE0F" box="[593,910,439,464]" pageId="46" pageNumber="47">M.V. and S.V. Nabozhenko</collectorName>
,
<collectorName id="26F12BE8F87D495FFC12FE7E381AFE0F" box="[923,1040,439,464]" pageId="46" pageNumber="47">B. Keskin</collectorName>
,
<specimenCount id="9D0285B7F87D495FFB97FE7E3853FE0F" box="[1054,1113,440,464]" pageId="46" pageNumber="47" type="male">8♂♂</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFBEEFE7E3895FE0F" box="[1127,1183,440,464]" pageId="46" pageNumber="47" type="female">4♀♀</specimenCount>
(
<collectionCode id="ED15D6FBF87D495FFB39FE7E38EAFE0F" box="[1200,1248,440,464]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="46" pageNumber="47">ZIN</collectionCode>
and CN),
<specimenCount id="9D0285B7F87D495FFAD4FE7E399DFE0F" box="[1373,1431,440,464]" pageId="46" pageNumber="47" type="male">4♂♂</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFF1EFE1A3CC5FE2B" box="[151,207,476,500]" pageId="46" pageNumber="47" type="female">2♀♀</specimenCount>
(dry material) and
<specimenCount id="9D0285B7F87D495FFE23FE1B3DEEFE2B" box="[426,484,477,500]" pageId="46" pageNumber="47" type="male">7♂♂</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFE79FE1A3E22FE2A" box="[496,552,476,501]" pageId="46" pageNumber="47" type="female">6♀♀</specimenCount>
(in ethanol) (
<collectionCode id="ED15D6FBF87D495FFD4BFE1A3F19FE2B" box="[706,787,476,500]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="46" pageNumber="47">ZDEU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6C4463F87D495FFCAEFE1A3EC8FDC7" ID-GBIF-Occurrence="1413110357" collectingDate="2009-05-02" collectionCode="CAL" collectorName="R. Krolik" country="Turkey" elevation="146" latitude="41.116665" location="Polonezkoy" longLatPrecision="1158" longitude="29.216667" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Istanbul Province">
<collectingCountry id="F3130EAEF87D495FFCAEFE1A3F72FE2A" box="[807,888,476,501]" name="Turkey" pageId="46" pageNumber="47">Turkey</collectingCountry>
,
<collectingRegion id="49C080DCF87D495FFC0AFE1A3846FE2A" box="[899,1100,476,501]" country="Turkey" name="Istanbul" pageId="46" pageNumber="47">Istanbul Province</collectingRegion>
,
<location id="8EDB18E5F87D495FFBD1FE1A38D7FE2A" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFBD1FE1A38D7FE2A" box="[1112,1245,476,501]" country="Turkey" latitude="41.116665" longLatPrecision="1158" longitude="29.216667" name="Polonezkoy" pageId="46" pageNumber="47" stateProvince="Istanbul Province">Polonezköy</location>
,
<quantity id="4CFCE3DBF87D495FFB60FE1A3938FE2B" box="[1257,1330,476,501]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="46" pageNumber="47" unit="m" value="146.0">
<elevation id="0029A90DF87D495FFB60FE1A3938FE2B" box="[1257,1330,476,501]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="46" pageNumber="47" unit="m" value="146.0">146 m</elevation>
</quantity>
,
<geoCoordinate id="EE3028F9F87D495FFAB6FE1A3996FE2A" box="[1343,1436,476,501]" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="41.116665">41°07N</geoCoordinate>
<geoCoordinate id="EE3028F9F87D495FFF1EFE393CF9FDC7" box="[151,243,511,536]" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="29.216667">29°13E</geoCoordinate>
,
<date id="FFBA68FEF87D495FFE89FDC63D68FDC7" box="[256,354,512,536]" pageId="46" pageNumber="47" value="2009-05-02">
<collectingDate id="EFFE9116F87D495FFE89FDC63D68FDC7" box="[256,354,512,536]" pageId="46" pageNumber="47" value="2009-05-02">2.v.2009</collectingDate>
</date>
, leg.
<collectorName id="26F12BE8F87D495FFE16FDC63E06FDC7" box="[415,524,511,536]" pageId="46" pageNumber="47">R. Królik</collectorName>
,
<specimenCount id="9D0285B7F87D495FFD91FDC63E36FDC7" box="[536,572,512,536]" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFDC0FDC63E66FDC7" box="[585,620,512,536]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
(
<collectionCode id="ED15D6FBF87D495FFDF5FDC63EB0FDC7" box="[636,698,512,536]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="46" pageNumber="47">CAL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6C4463F87D495FFD47FE393E44FDBF" ID-GBIF-Occurrence="1413110377" collectingDate="2014-04-10" collectionCode="ZIN, ZDEU" collectorName="M. V. and S. V. Nabozhenko &amp; B. Keskin" country="Turkey" elevation="160" latitude="41.08711" location="Omerli" longLatPrecision="1" longitude="29.369444" municipality="Alem Dag" pageId="46" pageNumber="47" specimenCount="23" specimenCount-female="12" specimenCount-male="11" stateProvince="Istanbul Province">
<collectingCountry id="F3130EAEF87D495FFD47FE393F15FDC7" box="[718,799,511,536]" name="Turkey" pageId="46" pageNumber="47">Turkey</collectingCountry>
,
<collectingRegion id="49C080DCF87D495FFCA3FE393FF9FDC7" box="[810,1011,511,536]" country="Turkey" name="Istanbul" pageId="46" pageNumber="47">Istanbul Province</collectingRegion>
,
<collectingMunicipality id="6BDFD444F87D495FFC77FE39387EFDC7" box="[1022,1140,511,536]" pageId="46" pageNumber="47">Alem Dağ</collectingMunicipality>
, near
<location id="8EDB18E5F87D495FFB31FE393900FDC7" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFB31FE393900FDC7" box="[1208,1290,511,536]" country="Turkey" latitude="41.08711" longLatPrecision="1" longitude="29.369444" municipality="Alem Dag" name="Omerli" pageId="46" pageNumber="47" stateProvince="Istanbul Province">Ömerli</location>
,
<quantity id="4CFCE3DBF87D495FFA9FFE39399DFDC8" box="[1302,1431,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="46" pageNumber="47" unit="m" value="160.0" valueMax="170.0" valueMin="150.0">
<elevation id="0029A90DF87D495FFA9FFE39399DFDC8" box="[1302,1431,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="46" pageNumber="47" unit="m" value="160.0" valueMax="170.0" valueMin="150.0">150170 m</elevation>
</quantity>
,
<geoCoordinate id="EE3028F9F87D495FFF1EFDE23D3BFDE2" box="[151,305,548,573]" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="41.08711">41°0513.6″N</geoCoordinate>
<geoCoordinate id="EE3028F9F87D495FFEB5FDE23DDCFDE2" box="[316,470,548,573]" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="29.369444">29°2210.0″E</geoCoordinate>
,
<date id="FFBA68FEF87D495FFE6CFDE23E57FDE2" box="[485,605,548,573]" pageId="46" pageNumber="47" value="2014-04-10">
<collectingDate id="EFFE9116F87D495FFE6CFDE23E57FDE2" box="[485,605,548,573]" pageId="46" pageNumber="47" value="2014-04-10">10.iv.2014</collectingDate>
</date>
, leg.
<collectorName id="26F12BE8F87D495FFD28FDE33FEAFDE2" box="[673,992,548,573]" pageId="46" pageNumber="47">M.V. and S.V. Nabozhenko</collectorName>
,
<collectorName id="26F12BE8F87D495FFC66FDE3386CFDE2" box="[1007,1126,548,573]" pageId="46" pageNumber="47">B. Keskin</collectorName>
,
<specimenCount id="9D0285B7F87D495FFBFCFDE238BAFDE3" box="[1141,1200,548,572]" pageId="46" pageNumber="47" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFB36FDE238FDFDE3" box="[1215,1271,548,572]" pageId="46" pageNumber="47" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED15D6FBF87D495FFA83FDE23937FDE3" box="[1290,1341,548,572]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="46" pageNumber="47">ZIN</collectionCode>
, CN),
<specimenCount id="9D0285B7F87D495FFA07FDE23CD1FDBF" pageId="46" pageNumber="47" type="male">8 males</specimenCount>
,
<specimenCount id="9D0285B7F87D495FFF60FD8E3D52FDBF" box="[233,344,583,608]" pageId="46" pageNumber="47" type="female">9 females</specimenCount>
(in ethanol) (
<collectionCode id="ED15D6FBF87D495FFE71FD8E3E4EFDBF" box="[504,580,584,608]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="46" pageNumber="47">ZDEU</collectionCode>
<emphasis id="B970922CF87D495FFDCCFD813E44FDBF" bold="true" box="[581,590,583,608]" pageId="46" pageNumber="47">)</emphasis>
</materialsCitation>
;
<materialsCitation id="3B6C4463F87D495FFDE9FD813912FD5B" ID-GBIF-Occurrence="1413110381" collectingDate="2014-04-11" collectionCode="ZDEU" collectorName="M. V. and S. V. Nabozhenko &amp; B. Keskin" country="Turkey" elevation="1180" latitude="40.84514" location="Pinus-Fagus forest" longLatPrecision="1" longitude="31.658306" pageId="46" pageNumber="47" specimenCount="5" specimenCount-male="5" stateProvince="Bolu Province">
<collectingCountry id="F3130EAEF87D495FFDE9FD813EBBFDBF" box="[608,689,583,608]" name="Turkey" pageId="46" pageNumber="47">Turkey</collectingCountry>
,
<collectingRegion id="49C080DCF87D495FFD36FD813F6CFDBF" box="[703,870,583,608]" country="Turkey" name="Bolu" pageId="46" pageNumber="47">Bolu Province</collectingRegion>
,
<location id="8EDB18E5F87D495FFCFCFD813846FDBF" LSID="urn:lsid:plazi:treatment:03ADFF28F87F495FFF1EF92C3FF4F910:8EDB18E5F87D495FFCFCFD813846FDBF" box="[885,1100,583,608]" country="Turkey" latitude="40.84514" longLatPrecision="1" longitude="31.658306" name="Pinus-Fagus forest" pageId="46" pageNumber="47" stateProvince="Bolu Province">Pinus-Fagus forest</location>
,
<geoCoordinate id="EE3028F9F87D495FFBD3FD8138FEFDBF" box="[1114,1268,583,608]" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="40.84514">40°5042.5″N</geoCoordinate>
<geoCoordinate id="EE3028F9F87D495FFB77FD81399DFDBF" box="[1278,1431,583,608]" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="31.658306">31°3929.9″E</geoCoordinate>
,
<quantity id="4CFCE3DBF87D495FFF1EFDAA3CE7FD5B" box="[151,237,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="46" pageNumber="47" unit="m" value="1180.0">
<elevation id="0029A90DF87D495FFF1EFDAA3CE7FD5B" box="[151,237,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="46" pageNumber="47" unit="m" value="1180.0">1180 m</elevation>
</quantity>
,
<date id="FFBA68FEF87D495FFF70FDAA3D7AFD5A" box="[249,368,620,645]" pageId="46" pageNumber="47" value="2014-04-11">
<collectingDate id="EFFE9116F87D495FFF70FDAA3D7AFD5A" box="[249,368,620,645]" pageId="46" pageNumber="47" value="2014-04-11">11.iv.2014</collectingDate>
</date>
, leg.
<collectorName id="26F12BE8F87D495FFE25FDAB3EEAFD5A" box="[428,736,620,645]" pageId="46" pageNumber="47">M.V. and S.V. Nabozhenko</collectorName>
,
<collectorName id="26F12BE8F87D495FFD63FDAB3F57FD5A" box="[746,861,620,645]" pageId="46" pageNumber="47">B. Keskin</collectorName>
,
<specimenCount id="9D0285B7F87D495FFCE0FDAA3FABFD5B" box="[873,929,620,644]" pageId="46" pageNumber="47" type="male">2♂♂</specimenCount>
(CN),
<specimenCount id="9D0285B7F87D495FFC66FDAA382DFD5B" box="[1007,1063,620,644]" pageId="46" pageNumber="47" type="male">3♂♂</specimenCount>
(in ethanol) (
<collectionCode id="ED15D6FBF87D495FFB36FDAA391AFD5B" box="[1215,1296,620,644]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="46" pageNumber="47">ZDEU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E1DB5F87D495FFF4EFD493FF4F910" pageId="46" pageNumber="47" type="description">
<paragraph id="8BBB4E3EF87D495FFF4EFD493E59FA07" blockId="46.[151,1437,151,1744]" pageId="46" pageNumber="47">
<emphasis id="B970922CF87D495FFF4EFD493DB6FD77" bold="true" box="[199,444,655,680]" pageId="46" pageNumber="47">Redescription, male.</emphasis>
Body length
<quantity id="4CFCE3DBF87D495FFDDBFD563EB5FD78" box="[594,703,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="46" pageNumber="47" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
, body width
<quantity id="4CFCE3DBF87D495FFCD9FD493FD3FD78" box="[848,985,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="46" pageNumber="47" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.83.5 mm</quantity>
.
<emphasis id="B970922CF87D495FFC6AFD493815FD77" bold="true" box="[995,1055,655,680]" italics="true" pageId="46" pageNumber="47">Body</emphasis>
slender, shining, black, legs darkbrown, rarely elytra dark-brown.
<emphasis id="B970922CF87D495FFD99FD753E44FD13" bold="true" box="[528,590,691,716]" italics="true" pageId="46" pageNumber="47">Head</emphasis>
widest at level of eyes. Eyes large, convex. Ratio of head width at eyes to distance between eyes: 1.6. Genae angular basally and straight in anterior 2/3. Lateral margin of head between gena and clypeus with very small weak emargination. Punctation of head coarse and dense: diameter of punctures about twice as wide as distance between punctures, punctures often connected on sides. Head pubescent with short recumbent yellowish hairs. Antennae strongly elongated (reaching almost 1/2 of elytral length; elytra 1.15 × as long as antennae), with 4 apical antennomeres extending beyond base of pronotum.
<emphasis id="B970922CF87D495FFB28FCA3391CFCA3" bold="true" box="[1185,1302,869,892]" italics="true" pageId="46" pageNumber="47">Pronotum</emphasis>
cordiform, transverse (1.151.2 × as wide as long), widest before middle, 1.4 × as wide as head. Lateral margins weakly rounded or almost straight, widely emarginated laterally. Anterior margin weakly rounded; base weakly rounded at middle and weakly widely sinuate laterally. Anterior angles acute or right, distinct on apex, projected; posterior angles right, distinct on apex. All margins narrowly beaded. Disc regularly strongly convex. Punctation of disc coarse and dense; diameter of punctures visibly more than distance between punctures; punctures round. Hypomera with coarse longitudinal wrinkles, not flattened on lateral margins.
<emphasis id="B970922CF87D495FFC21FBFF3FFAFB8D" bold="true" box="[936,1008,1081,1106]" italics="true" pageId="46" pageNumber="47">Elytra</emphasis>
convex, elongated (1.7 × as long as wide), widest at middle, 1.2 × as wide and 2.6 × as long as pronotum, 1.75 × as wide as head; width of elytral base subequal to width of pronotal base. Strial punctures merged in entire deep furrows. Elytral intervals weakly convex, with fine and sparse punctation (23 punctures in transverse distance of interval) and small distinct tubercles apically and laterally. Eighth intervals apically narrow and connected with margin of elytra and 2nd intervals. Epipleural carinae completely visible dorsally; epipleuron not reaching elytral apex.
<emphasis id="B970922CF87D495FFB34FB2D394CFADB" bold="true" box="[1213,1350,1259,1284]" italics="true" pageId="46" pageNumber="47">Ventral side</emphasis>
. Meso- and metaventrite with short pubescence. Abdominal ventrites with fine dense punctation, regularly covered with short yellowish hairs. Ventrite 1 with dense hair brush in middle; ventrite 5 without depression, completely beaded.
<emphasis id="B970922CF87D495FFF1EFA903CC4FAB2" bold="true" box="[151,206,1366,1389]" italics="true" pageId="46" pageNumber="47">Legs</emphasis>
slender. Inner side of femora pubescent basally and smooth and bare in apical 1/3. Protibiae straight, often weakly bisinuate on inner side, with 48 large and some small teeth; mesotibiae curved, with 38 small teeth; metatibiae straight, with small teeth or granules (rarely without teeth and granules in very small specimens). Trochanters with very dense hair brush.
</paragraph>
<paragraph id="8BBB4E3EF87D495FFF4EFA2538DBF9BA" blockId="46.[151,1437,151,1744]" pageId="46" pageNumber="47">
<emphasis id="B970922CF87D495FFF4EFA253DD9FA23" bold="true" box="[199,467,1507,1532]" pageId="46" pageNumber="47">Redescription, female.</emphasis>
Body length
<quantity id="4CFCE3DBF87D495FFDE6FA253F0BFA24" box="[623,769,1507,1532]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.2" metricValueMin="1.05" pageId="46" pageNumber="47" unit="mm" value="11.25" valueMax="12.0" valueMin="10.5">10.512 mm</quantity>
, body width
<quantity id="4CFCE3DBF87D495FFC1EFA253FF2FA24" box="[919,1016,1507,1532]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="46" pageNumber="47" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
. Pronotum more transverse (1.3 × as wide as long), widest at middle. Elytra wider (1.6 × as long as wide), 1.4 × as wide as pronotum. Intervals flat, wide (45 punctures in transverse distance of interval). Abdominal ventrite 5 without depression. Antennae shorter (reaching 1/4 of elytral length), only 3 apical antennomeres extending beyond base of pronotum.
</paragraph>
<paragraph id="8BBB4E3EF87D495FFF4EF9B73FF4F910" blockId="46.[151,1437,151,1744]" pageId="46" pageNumber="47">
<emphasis id="B970922CF87D495FFF4EF9B73D40F955" bold="true" box="[199,330,1649,1674]" pageId="46" pageNumber="47">Bionomics.</emphasis>
The species inhabits
<taxonomicName id="4C0435BDF87D495FFDDEF9B43FA3F956" authority="Abant Dag" authorityName="Abant Dag" box="[599,937,1649,1674]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="cerris">
<emphasis id="B970922CF87D495FFDDEF9B43F0FF956" box="[599,773,1650,1673]" italics="true" pageId="46" pageNumber="47">Quercus cerris</emphasis>
(Abant Dağ)
</taxonomicName>
,
<taxonomicName id="4C0435BDF87D495FFC35F9B43808F956" box="[956,1026,1650,1673]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="B970922CF87D495FFC35F9B43808F956" box="[956,1026,1650,1673]" italics="true" pageId="46" pageNumber="47">Fagus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0435BDF87D495FFBCFF9B43992F955" authority="Yedigoller" authorityName="Yedigoller" box="[1094,1432,1649,1674]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B970922CF87D495FFBCFF9B4390AF955" box="[1094,1280,1649,1674]" italics="true" pageId="46" pageNumber="47">Picea orientalis</emphasis>
(Yedigöller)
</taxonomicName>
,
<taxonomicName id="4C0435BDF87D495FFF1EF9533DD0F972" authority="Alem Dag" authorityName="Alem Dag" box="[151,474,1684,1709]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="robur">
<emphasis id="B970922CF87D495FFF1EF9533D4BF972" box="[151,321,1684,1709]" italics="true" pageId="46" pageNumber="47">Quercus robur</emphasis>
(Alem Dağ)
</taxonomicName>
at early night (20:0023:00). Iranian population inhabit relic forests of
<taxonomicName id="4C0435BDF87D495FFAB7F9533E8EF90F" authority="Nabozhenko &amp; Keskin 2013" authorityName="Nabozhenko &amp; Keskin" authorityYear="2013" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="castaneifolia">
<emphasis id="B970922CF87D495FFAB7F9533D22F90F" italics="true" pageId="46" pageNumber="47">Quercus castaneifolia</emphasis>
(
<bibRefCitation id="EF9533CFF87D495FFEBFF9713E76F90F" author="Nabozhenko" box="[310,636,1719,1744]" pageId="46" pageNumber="47" refString="Nabozhenko, M. V., Keskin, B. &amp; Alpagut Keskin, N. (2013) Molecular-genetic research of darkling beetles of the genus Odocnemis (Coleoptera: Tenebrionidae). In: Molecular genetic approaches in taxonomy and ecology. Abstracts of the Scientific Conference (March 25 - 29 2013, Rostov-on-Don, Russia). Southern Scientific Centre Publishers, Rostov-on- Don, p. 60. [in Russian]" type="journal article" year="2013">Nabozhenko &amp; Keskin 2013</bibRefCitation>
)
</taxonomicName>
. See also bionomics of
<taxonomicName id="4C0435BDF87D495FFC05F97E3FFDF90F" box="[908,1015,1720,1744]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="operta">
<emphasis id="B970922CF87D495FFC05F97E3FFDF90F" box="[908,1015,1720,1744]" italics="true" pageId="46" pageNumber="47">O. operta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>