199 lines
22 KiB
XML
199 lines
22 KiB
XML
<document ID-CLB-Dataset="291769" ID-DOI="http://dx.doi.org/10.3897/asp.82.e111148" ID-GBIF-Dataset="de602973-4f52-4de2-8014-342e7778d97b" ID-Pensoft-Pub="1864-8312-82-151" ID-Pensoft-UUID="8D187145BFCA55A1A73960CDE5BD124B" ID-ZooBank="43AC036E93CC4D79939A07DF54BE1A2D" ModsDocID="1864-8312-82-151" checkinTime="1711188240520" checkinUser="pensoft" docAuthor="Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent" docDate="2024" docId="FCC83238CF44597C9C8CB733D9D5D303" docLanguage="en" docName="ArthropodSystPhyl 82: 151-181" docOrigin="Arthropod Systematics & amp; Phylogeny 82" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/asp.82.e111148" docTitle="Genkyhag innebula Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024, sp. nov." docType="treatment" docUuid="407F2F7B-CA5D-467C-AFF5-11C7A2E9BCDA" docUuidSource="ZooBank" docVersion="4" id="8D187145BFCA55A1A73960CDE5BD124B" lastPageNumber="151" masterDocId="8D187145BFCA55A1A73960CDE5BD124B" masterDocTitle="Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities" masterLastPageNumber="181" masterPageNumber="151" pageNumber="151" updateTime="1732880532958" updateUser="ExternalLinkService">
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<mods:title id="3CE265803A7060F84604E7B99731E3B0">Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities</mods:title>
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<mods:namePart id="7B5E909531C703E6272E9168FB0D9F7E">Brazidec, Manuel</mods:namePart>
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<mods:nameIdentifier id="B6DA8534EA81F4AF7D933DB494C20D45" type="ORCID">https://orcid.org/0000-0002-0860-8972</mods:nameIdentifier>
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<mods:affiliation id="9BDFFD192EDFC46BABB668B75E9B4245">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="C34A2356D8369A37570CC12DC12D1174" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9FAC8CDB230920852A0E77125A514B54">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="44E62A10655BCFC1E30B6FD199C00FDF">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark</mods:affiliation>
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<mods:namePart id="C79E9041373E7C4AD7142FDB03702C99">Boudinot, Brendon E.</mods:namePart>
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<mods:affiliation id="577A482BEA448501C64DA6763990626F">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany</mods:affiliation>
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<mods:namePart id="EE3FF29C99E7D13C97B576C5B56ACC01">Richter, Adrian</mods:namePart>
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<mods:affiliation id="12E2277DCED99A516A46C51A8C9DB3E5">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan</mods:affiliation>
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<mods:namePart id="1E1691ECB825442E7BA277FD35F80C50">Hammel, Joerg U.</mods:namePart>
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<mods:affiliation id="F3E5240F0192FE2148FC4572817E5441">Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="186B7E01A4D46BD727B290DA71D9F7E6">Perkovsky, Evgeny E.</mods:namePart>
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<mods:nameIdentifier id="A28616521CDD142B185EAA3998FDB36E" type="ORCID">https://orcid.org/0000-0002-7959-4379</mods:nameIdentifier>
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<mods:affiliation id="40C57F29191431D3C9123AFC0A0D2F9A">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:namePart id="808E85102ACDD690B52DE0A67CC5BDEC">Fan, Yong</mods:namePart>
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<mods:affiliation id="899613768BDFFF3436A6F0A74834D276">Fushun Amber Institute, Fushun 113005, China</mods:affiliation>
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<mods:namePart id="D5AB591CEE3B4602C466C63E4D5824D1">Wang, Zhen</mods:namePart>
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<mods:affiliation id="ACA18595ECE440052DA9344AB16BFE8A">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="D7660780CDCD65B8F0282F5E90C1B57E">Wu, Qiong</mods:namePart>
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<mods:affiliation id="8D90F88CE206B90DD1AE20FA414EF8E5">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="CC5CBF706ADB5C1CCF82D5C53CEEBEB1">Wang, Bo</mods:namePart>
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<mods:affiliation id="B23770454CE922CB7D59492D270AE7F3">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="B33EE9A73FE82EB7CE878E5BE5D27AD9">Perrichot, Vincent</mods:namePart>
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<mods:nameIdentifier id="44F67EE1B2F2023360FB2158C2C4D5E5" type="ORCID">https://orcid.org/0000-0002-7973-0430</mods:nameIdentifier>
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<mods:affiliation id="6BC17376EF77689159D04BEB7448676C">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France</mods:affiliation>
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<mods:title id="795CE86242042892EEF9E1B89A6D7A04">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="B3C073EE38B1D7785836A94BF7B3E582">
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<mods:date id="408D48815EE8499810217812E58D496B">2024</mods:date>
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<mods:number id="C98B43F4C20E03CD55A452197061082F">2024-03-22</mods:number>
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<mods:number id="614603C0A713B5584BD7CB853AB6FD6A">82</mods:number>
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<treatment id="FCC83238CF44597C9C8CB733D9D5D303" ID-GBIF-Taxon="224057645" LSID="urn:lsid:zoobank.org:act:407F2F7B-CA5D-467C-AFF5-11C7A2E9BCDA" httpUri="http://treatment.plazi.org/id/FCC83238CF44597C9C8CB733D9D5D303" lastPageNumber="151" pageId="0" pageNumber="151" scope_family="Dinapsidae" scope_order="Hymenoptera" scope_superFamily="Megalyroidea">
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<subSubSection id="5A038242A2838AED9FD758C8CFD0B0D5" pageId="0" pageNumber="151" type="nomenclature">
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<paragraph id="26C33D3C094669418C307CA1D165E5B8" pageId="0" pageNumber="151">
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3.1.1.2.1. †
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<taxonomicName id="99E7C9DBEE12C19930DC84ACEBADBA29" LSID="https://zoobank.org/407F2F7B-CA5D-467C-AFF5-11C7A2E9BCDA" authority="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" class="Insecta" family="Megalyridae" genus="Genkyhag" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Genkyhag innebula" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="innebula" status="sp. nov.">Genkyhag innebula</taxonomicName>
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<taxonomicNameLabel id="FB25A7FF427CAAABB6FAF32000C97314" pageId="0" pageNumber="151">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="8ECDDF04DB71E4DD743DE5D0E9E3FF67" pageId="0" pageNumber="151" type="description">
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<paragraph id="68907F933AD9AD13475235ABDFF79670" pageId="0" pageNumber="151">
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<figureCitation id="EDE00A600CEDF3DD97A8085AAE3F09F4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H)." figureDoi="10.3897/asp.82.e111148.figure2" httpUri="https://binary.pensoft.net/fig/1007131" pageId="0" pageNumber="151">Figures 2C</figureCitation>
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<figureCitation id="6322B7E9860DA4B9C4961549D65DEACA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Genkyhag innebula gen. et sp. nov., mid-Cretaceous, Kachin amber. A holotype female NIGP 203546, habitus in dorsal view; B paratype male FAI-BI 11324 a, habitus in dorsal view; C habitus in frontal view; D paratype male FAI-BI 11324 b, habitus in lateral view; E paratype FAI-BI 11324 c, fore and hind wings, with hamuli indicated by arrows. Scale bars: 1 mm (A - E). Abbreviations: AG = inner axillar groove; MMS = median mesoscutal sulcus; OC = occipital carina." figureDoi="10.3897/asp.82.e111148.figure5" httpUri="https://binary.pensoft.net/fig/1007134" pageId="0" pageNumber="151">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8821ED97D40716BF2C32F4453FED8EA" pageId="0" pageNumber="151" type="etymology">
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<paragraph id="A5D726F8531C284E99CEA0A0104C1730" pageId="0" pageNumber="151">Etymology.</paragraph>
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<paragraph id="F771422A364B5C8F7975A7174577A145" pageId="0" pageNumber="151">
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From Latin 'nebula, In
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<normalizedToken id="A6001AAAE5DB22A6646279AA9101CB42" originalValue="nebula’">nebula'</normalizedToken>
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, meaning "in the fog", in reference to the amber pieces where the specimens are found, that contains numerous terrigenous inclusions, sometimes partially hiding the specimens. It is a noun in apposition.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A3663BB218BE4A06308BFBB393F4F0B1" pageId="0" pageNumber="151" type="materials_examined">
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<paragraph id="8C9763F06877EC3F60D1FE2DC9E49B7A" pageId="0" pageNumber="151">Material studied.</paragraph>
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<paragraph id="C2D5EA7128A585A5F21C4A5CE16E9ECB" pageId="0" pageNumber="151">
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<materialsCitation id="D770B4BE12390992367AF1BC9C79717F" ID-GBIF-Occurrence="4597783302" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="China" county="Nanjing" location="Nanjing Institute of Geology" municipality="Chinese Academy of Sciences" specimenCount="Three" specimenCount-male="3" typeStatus="Holotype">
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<emphasis id="67A4CB8039D6B90D23910564D101219C" bold="true" italics="true" pageId="0" pageNumber="151">
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<typeStatus id="F59442F9A7EF9EAD043E2F3D8456019D" type="holotype">Holotype</typeStatus>
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</emphasis>
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female NIGP203546; housed in the
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<location id="034906769DA84AF8959F5635170BCD81" LSID="urn:lsid:plazi:treatment:FCC83238CF44597C9C8CB733D9D5D303:034906769DA84AF8959F5635170BCD81" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="Nanjing Institute of Geology">Nanjing Institute of Geology</location>
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and
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<location id="823B5639074F59AADDDDDF8A64EA9595" LSID="urn:lsid:plazi:treatment:FCC83238CF44597C9C8CB733D9D5D303:823B5639074F59AADDDDDF8A64EA9595" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="Paleontology">Paleontology</location>
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(NIGP),
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<collectingMunicipality id="2F966C50544DF062C1FE8E59D8B8B563">Chinese Academy of Sciences</collectingMunicipality>
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,
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<collectingCounty id="22CB2BAE6757B804EE7518A08EE247E0">Nanjing</collectingCounty>
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,
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<collectingCountry id="71680D97E29A26FF75868729A329F3B5" name="China">China</collectingCountry>
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.
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Three
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male
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</specimenCount>
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||
FAI-BI 11324a, male FAI-BI 11324b, FAI-BI-11324c of unknown sex; housed in the Fushun Amber Institute.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B66B9FBDFAA73C68606029F2102F6DEE" pageId="0" pageNumber="151" type="type locality">
|
||
<paragraph id="EF39C0E8F608E0FC6250979E3B22CD2F" pageId="0" pageNumber="151">Type locality.</paragraph>
|
||
<paragraph id="205E96CBC5FFA46986E05AE116DBDCE7" pageId="0" pageNumber="151">Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CD1A337AFC57CB831645613158CFF899" pageId="0" pageNumber="151" type="age">
|
||
<paragraph id="67069CC86BC6ACCB40BBF351D1E43391" pageId="0" pageNumber="151">Age.</paragraph>
|
||
<paragraph id="8AA2F216516FCDBB908C29952EB066A6" pageId="0" pageNumber="151">Upper Albian to lower Cenomanian, mid-Cretaceous.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2021BF46E258975EED5908F3F849937" pageId="0" pageNumber="151" type="diagnosis">
|
||
<paragraph id="CFE60EA017030CCFB0766BBB67F116F4" pageId="0" pageNumber="151">Diagnosis.</paragraph>
|
||
<paragraph id="3DD01B79D74638F59CE76AFC4BD95D56" pageId="0" pageNumber="151">As for genus.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7EE68D9D7862B19A4D6B6D7CE47CCD8" pageId="0" pageNumber="151" type="description">
|
||
<paragraph id="C9E66AF8E0ADBCF20AD5EEDEFC94B357" pageId="0" pageNumber="151">Description.</paragraph>
|
||
<paragraph id="DFD423CB7B554EFB147214AC8E97D623" pageId="0" pageNumber="151">
|
||
Holotype female incompletely preserved (body length 3.12 mm), in a piece with multiple terrigenous inclusions. Paratypes preserved in a single piece: FAI-BI 11324a visible in dorsal and frontal views, FAI-BI 11324b in lateral view (body length ca. 3.45 mm), FAI-BI 11324c being one complete fore and hind wings and a partial fore wing. -
|
||
<emphasis id="C71403035E6E92B2F68365603ADE1BA7" bold="true" italics="true" pageId="0" pageNumber="151">Head</emphasis>
|
||
globular, apparently entirely glabrous (length for holotype ca. 0.47 mm); frons shagreened, convex, divided by a faintly impressed sulcus; compound eye oval, without postocular carina; vertex convex, shagreened; clypeus short; subantennal groove present; toruli inserted half way between clypeus and eyes; antenna short, barely reaching propodeum; scape globular; pedicel slightly longer; flagellomeres cylindrical, longer than wide; flagellomeres 7-11 shorter than flagellomeres 1-6; flagellomere 12 longest; occipital carina crenulate. -
|
||
<emphasis id="0F2ABF91AF5298B3CF3822CB0C507718" bold="true" italics="true" pageId="0" pageNumber="151">Mesosoma</emphasis>
|
||
slightly shorter than metasoma (length for holotype 1.12 mm; for paratype 11324a 1. 29 mm; for paratype 11324b 1.57 mm); pronotum not visible dorsally; mesoscutum shagreened, half as long as mesosoma, median mesoscutal sulcus crenulate (mesoscutum length for holotype 0.51 mm; for paratype 11324a 0.61 mm; width for holotype 0.65 mm; for paratype 11324a 0.71 mm); parapsidal lines present; axillae barely contiguous medially and inner grooves crenulate; mesoscutellum wider than long, shorter than mesoscutum; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; propodeum carinate, carinae delimitating row of small foveae in anterior region and larger foveae in posterior region. -
|
||
<emphasis id="AEDED96232D7CE593B7CEEEB4C1C01A1" bold="true" italics="true" pageId="0" pageNumber="151">Fore wing</emphasis>
|
||
hyaline, covered with microtrichiae (length for holotype 1.84 mm; for paratype 11324a 2.10 mm; for paratype 11324b 2.11 mm; for paratype 11324c 2.86 mm); venation almost complete, Sc+R, M+Cu, A, Rs+M and M fully pigmented; Cu spectral; marginal cell very narrow, closed in straight line; R1 barely extending beyond marginal cell; medial cell rectangular and relatively broad. -
|
||
<emphasis id="4AFAC35886149AAF1A7D9CB53555B167" bold="true" italics="true" pageId="0" pageNumber="151">Hind wing</emphasis>
|
||
hyaline; venation reduced to Sc+R and R1, Sc+R almost not diverging from margin, R1 pigmented almost to tip; at least seven hamuli, located distal to middle of hind wing. -
|
||
<emphasis id="69CD50C3088C873EE9FC06114F58B728" bold="true" italics="true" pageId="0" pageNumber="151">Legs</emphasis>
|
||
with one mesotibial and two metatibial spurs; metafemur swollen; ventral surface of metatibia and metabasitarsus with row of comb-like setae, metabasitarsus at least three times as long as following metatarsomeres. -
|
||
<emphasis id="39DE49A769D69BF9B9B9E0E88D5D1CE9" bold="true" italics="true" pageId="0" pageNumber="151">Metasoma</emphasis>
|
||
elongate, oval, with sparse short setae (length for holotype 1.43 mm); tergites smooth; ovipositor shorter than metasoma (length for holotype 1.34 mm; OL/BL ratio 0.43).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |