865 lines
86 KiB
XML
865 lines
86 KiB
XML
<document id="B2921DBF7ED55B30450822691302CF04" ID-DOI="10.3897/zookeys.1203.122758" ID-ZooBank="C6B34F96-DFE7-475D-A021-73E0B1D61F3D" ID-publisher-id="122758" URI-arpha="1812E618-828A-5AF2-9224-A4A203BF0908" URI-zoobank="http://zoobank.org/C6B34F96-DFE7-475D-A021-73E0B1D61F3D" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1717085984932" checkinUser="pensoft" docAuthor="Grismer, L. Lee, Aowphol, Anchalee, Grismer, Jesse L., Aksornneam, Akrachai, Quah, Evan S. H., Murdoch, Matthew L., Gregory, Jeren J., Nguyen, Eddie, Kaatz, Amanda, Bringsøe, Henrik & Rujirawan, Attapol" docDate="2024" docId="B98A591B02275EF398445EB55624C364" docLanguage="en" docName="ZooKeys 1203: 211-238" docOrigin="ZooKeys 1203" docSource="https://zookeys.pensoft.net/article/122758/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Cyrtodactylus phamiensis Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan 2024, sp. nov." docType="treatment" docUuid="DDADBD8A-5234-4183-89A7-C5A3C015456A" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="238" masterDocId="1812E618828A5AF29224A4A203BF0908" masterDocTitle="A new species of the Cyrtodactylus chauquangensis group (Squamata, Gekkonidae) from the borderlands of extreme northern Thailand" masterLastPageNumber="238" masterPageNumber="211" pageNumber="211" updateTime="1717105598636" updateUser="admin">
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<mods:title id="EFA3AB28C396E1AD44870B9BA096D170">A new species of the Cyrtodactylus chauquangensis group (Squamata, Gekkonidae) from the borderlands of extreme northern Thailand</mods:title>
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<mods:namePart id="20FF47477153E035A99824E3C844AA4E">Grismer, L. Lee</mods:namePart>
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<mods:affiliation id="14CBA58701328FF028BBCAA0EB78F868">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA & Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Malaysia</mods:affiliation>
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<mods:namePart id="DFB706295813B6A92185FABC2AF83FD6">Aowphol, Anchalee</mods:namePart>
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<mods:affiliation id="D033A0F1531B44A0E10945E33B256060">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:namePart id="0D536866735E1EA99D7120874C8FA68D">Grismer, Jesse L.</mods:namePart>
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<mods:affiliation id="5ABE46C934FF05C148A8A3680D0CDD9C">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA</mods:affiliation>
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<mods:namePart id="75CAAD371B1A6B37CA0E46D43EA19028">Aksornneam, Akrachai</mods:namePart>
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<mods:affiliation id="2FB0DE62B4A73106C4B3DE0B9ED0E187">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:affiliation id="54FF759C830491089502A638716BCDB2">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Malaysia & Biodiversity Center, Kasetsart University, Bangkok 10900, Thailand & Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377, Singapore</mods:affiliation>
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<mods:namePart id="24FF56586180C421EFB105E9A816D84B">Murdoch, Matthew L.</mods:namePart>
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<mods:affiliation id="4B81F6D223276B60F6124027558369FD">School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
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<mods:namePart id="6F01CEB093F567EF1C68B23CEA273323">Gregory, Jeren J.</mods:namePart>
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<mods:affiliation id="EFEA5E1F6105F80D19FC5E62DC8B71B4">School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
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<mods:namePart id="2E3EED93BA147E7C6C31A134C9976CC1">Nguyen, Eddie</mods:namePart>
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<mods:affiliation id="540137713D4A0879EE8C27D0F9D8001E">School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
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<mods:namePart id="5FD1B00AB6C690EE539B8ABD765AD5C8">Kaatz, Amanda</mods:namePart>
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<mods:affiliation id="B0897EFD39BEC3C97A9AA62BACE4701D">School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
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<mods:namePart id="5B36A8737B9D77FA4D548308E30A23F9">Bringsøe, Henrik</mods:namePart>
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<mods:affiliation id="6C7DF4DDD723F92E35156EE5D9BB0069">Irisvej 8, DK- 4600 Køge, Denmark</mods:affiliation>
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<mods:namePart id="D195A02030C74602F3F816CE4524DEC6">Rujirawan, Attapol</mods:namePart>
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<mods:affiliation id="FBF47D0CD34318730DD597D0D7C74A62">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:part id="7AEE541DEE49E940AC246CAE9BA88888">
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<mods:date id="1270BC217F45E1546E0791A7F6C5ED1D">2024</mods:date>
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<mods:number id="04D0834F4BCACE15E5E81053779EDF30">2024-05-30</mods:number>
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<mods:number id="16E2800FA8BF6D06CD6D49375CDDDAA6">1203</mods:number>
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<mods:identifier id="8CF39780C15D0585B91BDA5D8865B928" type="DOI">10.3897/zookeys.1203.122758</mods:identifier>
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<mods:identifier id="160B3DC08BDB2AE30FBD58EF4D63208F" type="ZooBank">C6B34F96-DFE7-475D-A021-73E0B1D61F3D</mods:identifier>
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<subSubSection id="2488AA450EE3747C6AB5E4D2C05A5159" type="nomenclature">
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<paragraph id="5E5DF4DF337F5DB3EF2EB9A6014013DD">
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<taxonomicName id="CD222BB2B17523B6392C1763D5DE5ABF" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
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<emphasis id="7F488F7C8F6792255CC6EFEDF029957B" italics="true">Cyrtodactylus phamiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2466EE80349AD704FA7B0BA77535A227" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="1D673E77D2F3DBCA742572C93B2A71ED" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selected individuals of the type series and referred specimen of Cyrtodactylus phamiensis sp. nov. from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand." figureDoi="10.3897/zookeys.1203.122758.figure4" httpUri="https://binary.pensoft.net/fig/1059372">Figs 4</figureCitation>
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,
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<figureCitation id="FC4274C89E507CFE4C7F5233C04946B5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Adult male holotype of Cyrtodactylus phamiensis sp. nov. A dorsal view B ventral view C dorsal view of head D gular region E thighs and precloacal region F ventral view of right manus G ventral view of left pes H subcaudal region I lateral view of left side of head. Photographs by Attapol Rujirawan." figureDoi="10.3897/zookeys.1203.122758.figure5" httpUri="https://binary.pensoft.net/fig/1059373">5</figureCitation>
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,
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<figureCitation id="805230BF16AB4DFA3DD496C1EB0A15E0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cyrtodactylus cf. phamiensis sp. nov. from 5 km north of the type locality in Tham Pha Chom. Photograph by Ada Chornelia." figureDoi="10.3897/zookeys.1203.122758.figure6" httpUri="https://binary.pensoft.net/fig/1059374">6</figureCitation>
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,
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<figureCitation id="FC18B7E182E05AF585359F9A237A09E9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus phamiensis sp. nov. from 50 meters west-southwest of the type locality, immediately east of Pha Mi Village. 17 December 2022. Photograph by Henrik Bringsøe." figureDoi="10.3897/zookeys.1203.122758.figure7" httpUri="https://binary.pensoft.net/fig/1059375">7</figureCitation>
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,
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<figureCitation id="3134F0B6F1C229057A55AAB24B96EB81" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Paratypes of Cyrtodactylus phamiensis sp. nov. from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand. Photographs by Attapol Rujirawan." figureDoi="10.3897/zookeys.1203.122758.figure8" httpUri="https://binary.pensoft.net/fig/1059376">8</figureCitation>
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</paragraph>
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<paragraph id="FBF898A967A992DC6F1D77446174E7FB">
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<heading id="CAB33674310110FDE85F6CFDB92F9729" reason="title">Type material.</heading>
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</paragraph>
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<paragraph id="54D06E55D47C20AF216B06D03B29544A">
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<materialsCitation id="EAACD9242A13FB0369416F5BFC8D6A0E" collectingDate="2023-03-26" collectionCode="ZMKU" collectorName="A. Aowphol & A. Rujirawan & A. Aksornneam & L. L. Grismer & J. L. Grismer & E. S. H. Quah & M. L. Murdoch" country="Thailand" county="Mae Sai District" elevation="517" latitude="20.40134" location="Pha Mi Village" longLatPrecision="1" longitude="99.85369" municipality="Wiang Phang Kham Subdistrict" specimenCode="R 01086" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Rai Province" typeStatus="holotype">
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<emphasis id="7C5D686222D0B778B3411259A2040C9B" bold="true">
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<typeStatus id="7F29B03658086D7CF7EC297D945D47F0" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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(
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<collectionCode id="20A1C7C395302A2951E577A99C16469A" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
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</abbrev>
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<specimenCode id="8BA580909705ECFF250E862AB73339B5" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile">R 01086</specimenCode>
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) collected from
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,
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,
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,
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,
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<collectingCountry id="32E20115BC2A63F72313886203969F95" name="Thailand">Thailand</collectingCountry>
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(
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<geoCoordinate id="C1F4206E07CAEBDDDE3617251205957D" degrees="20.40134" direction="north" orientation="latitude" precision="1" value="20.40134">20.40134 ° N</geoCoordinate>
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,
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<geoCoordinate id="62ECE0951BBECEB773248D06463AA385" degrees="99.85369" direction="east" orientation="longitude" precision="1" value="99.85369">99.85369 ° E</geoCoordinate>
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</named-content>
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;
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<elevation id="C3ADFC8E7CC72DFC818C117F5326AF46" metricMagnitude="2" metricUnit="m" metricValue="5.17" unit="m" value="517.0">
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elevation
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<quantity id="9D05844854EEACA3899521E60DADF74F" metricMagnitude="2" metricUnit="m" metricValue="5.17" unit="m" value="517.0">517 m</quantity>
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a. s. l.
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</elevation>
|
||
) on
|
||
<date id="737A469CF0A9D13B5D738E2FE223E1D8" value="2023-03-26">
|
||
<collectingDate id="CFC55441710ACCB819B23A1650FA7822" value="2023-03-26">26 March 2023</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="74DD0BB404BA4D3BBA725519894AFE7A">A. Aowphol</collectorName>
|
||
,
|
||
<collectorName id="58F7C5D9B08F78145DCAEB917283910B">A. Rujirawan</collectorName>
|
||
,
|
||
<collectorName id="621ABF93D094D5F0560ED18EE963E4A6">A. Aksornneam</collectorName>
|
||
,
|
||
<collectorName id="28205E7277922E4B0FFC41856AB8DB01">L. L. Grismer</collectorName>
|
||
,
|
||
<collectorName id="AD8E291FF726102428DB184DE1AC90FA">J. L. Grismer</collectorName>
|
||
,
|
||
<collectorName id="C29167DA06AC136A569A4C853E535CA4">E. S. H. Quah</collectorName>
|
||
, and
|
||
<collectorName id="A9E6D623693F465E31A9CB47FC0C8061">
|
||
<collectorName id="C80D78613950549F4D73D33F258067A7">M. L. Murdoch</collectorName>
|
||
.
|
||
</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="3D4C552D30D8E8BDBC35878EEB1A59BF">
|
||
<materialsCitation id="85C676683C65608DB152DB1677DE2325" collectingDate="2023-03-26" collectionCode="ZMKU" collectorName="A. Aowphol & A. Rujirawan & A. Aksornneam & L. L. Grismer & J. L. Grismer & E. S. H. Quah & M. L. Murdoch" country="Thailand" county="Mae Sai District" elevation="517" latitude="20.40134" location="Pha Mi Village" longLatPrecision="1" longitude="99.85369" municipality="Wiang Phang Kham Subdistrict" specimenCode="R 01085, R 01087, R 01084" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiang Rai Province" typeStatus="paratype">
|
||
<emphasis id="A76D2C123F4009DED4E45C8037CE6C3D" bold="true">
|
||
<emphasis id="4FC943D2EAE136EF3ABB9204CCC924E5" bold="true" italics="true">
|
||
<typeStatus id="FB7E648ED648E60E361FD551CE5CAADE" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="406E5F5960721A1D0503D7EA88C4971D" count="2" type="male">Two adult males</specimenCount>
|
||
(
|
||
<abbrev id="ABBRID0EJOAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="8DB78F189B814F33B76BF921B6F05092" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="0FF722BEE6164EC29F99286FA9A69952">R 01085</specimenCode>
|
||
,
|
||
<abbrev id="ABBRID0ENOAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="D79A6C4A0EF09730FD1BCA8249D75850" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="9F6846131421DABE6B25F5E9427CE6B4">R 01087</specimenCode>
|
||
) and
|
||
<specimenCount id="449FC73862828D282748A454E3746A4B" count="1" type="female">one adult female</specimenCount>
|
||
(
|
||
<abbrev id="ABBRID0EROAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="906C65CBD2C12F20906E66F88A56E651" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="4092C992FE655B3A812B473CEEFC516F">R 01084</specimenCode>
|
||
) bear the same collection data as the holotype
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F4AC71A7AB75CC171781DD9049352D90" collectingDate="2023-03-25" collectionCode="ZMKU" collectorName="A. Aowphol & A. Rujirawan & A. Aksornneam & L. L. Grismer & J. L. Grismer & E. S. H. Quah & M. L. Murdoch" country="Thailand" county="Mae Sai District" elevation="517" latitude="20.40134" location="Pha Mi Village" longLatPrecision="1" longitude="99.85369" municipality="Wiang Phang Kham Subdistrict" specimenCode="R 01073-01075, R 01078, R 01081" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Chiang Rai Province">
|
||
<specimenCount id="839F01A588D96A546473B83371C1B474" count="4" type="female">Four adult females</specimenCount>
|
||
(
|
||
<abbrev id="ABBRID0EVOAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="B7FC4DA271E904DCC7D7CD86CB5EACF3" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="83574F085403AA32EBFA78E20EF5F1B2">R 01073–01075</specimenCode>
|
||
,
|
||
<abbrev id="ABBRID0EZOAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="731760478E534F72307A0F3938B0E5F4" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="D647AD51A173384C57416DB334B3096F">R 01078</specimenCode>
|
||
) and
|
||
<specimenCount id="F8FE5012A8D2F1C129D86E3E97713146" count="1" type="male">one adult male</specimenCount>
|
||
(
|
||
<abbrev id="ABBRID0E4OAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="3741C2C33E9B1FA828F859D539A55151" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="EFB873E013469F42960EC0BFD7157216">R 01081</specimenCode>
|
||
) bear the same collection data as the holotype except collected on
|
||
<date id="EF8950A7CE47E287BE2999C06F3D1CD3" value="2023-03-25">
|
||
<collectingDate id="C4DA7CA1D77FEF07981DA858C2762798" value="2023-03-25">25 March 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EBPAG" type="Referred specimens">
|
||
<paragraph id="62DC5448ED117E50BD3ACEE8B70661AB">
|
||
<heading id="7CDCB03D33604B9125A5CA24CE1F46C6" reason="title">Referred specimens.</heading>
|
||
</paragraph>
|
||
<paragraph id="1D707129C2F9B2B9D1682DDBEEE057AE">
|
||
Six hatchlings.
|
||
<abbrev id="ABBRID0EHPAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="C23499838E006D55ECA56A68404FDC07" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="C6CF80B081E91EC2C5711F6B8FF10D12">R 01076</specimenCode>
|
||
–01077,
|
||
<abbrev id="ABBRID0ELPAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="88466A23D87AE20B5D3CEB0BDFF38463" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="7E44E53558923F64051AC8F9FDC4E0E9">R 01079</specimenCode>
|
||
–01080 bear the same collection data as the
|
||
<typeStatus id="D49A991A7B3349305D02937C3B44ECE6" type="holotype">holotype</typeStatus>
|
||
except were collected on
|
||
<date id="A7024476D5431FEA2F1B3A717DDEAFF4" value="2023-03-25">25 March 2023</date>
|
||
.
|
||
<abbrev id="ABBRID0EPPAG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="7E47856D0E9806768B8BD1502BD61E75" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="2B62107F939959E1167E2A49D2DCC761">R 01082</specimenCode>
|
||
–01083 bear the same collection data as the
|
||
<typeStatus id="06868943399061F73360CB9AA86F06A0" type="holotype">holotype</typeStatus>
|
||
except collected from
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWPAG" specific-use="{"type":"Point","coordinates":[99.854660,20.398000]}">
|
||
<geoCoordinate id="819AC5760D9F4A617B0A9EB85AE28A1F" degrees="20.39800" direction="north" orientation="latitude" precision="1" value="20.398">20.39800 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="662DE2910B1089A3C4525AD45CDAF0D2" degrees="99.85466" direction="east" orientation="longitude" precision="1" value="99.85466">99.85466 ° E</geoCoordinate>
|
||
</named-content>
|
||
; elevation
|
||
<quantity id="3EA129E6DA215C0B2307C0A55CDFD3E0" metricMagnitude="2" metricUnit="m" metricValue="5.05" unit="m" value="505.0">505 m</quantity>
|
||
a. s. l., on
|
||
<date id="2CA7829D48AE463C35C42FDB91E410DC" value="2023-03-25">25 March 2023</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2PAG" type="diagnosis">
|
||
<paragraph id="711E052FAE7500B076BCFE6E889D2435">
|
||
<heading id="38D45D4838A57C6180665F91A71336F6" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="D40F9514C877E6C36A2B1F7B8E9D5DE3">
|
||
<taxonomicName id="9DAF9CCC33B2B609C4FB020A8A9E094C" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="F8327380719DD079A266B555F89E631B" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="944B5127431DC525376E9064CC43BE02" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from all other species of the
|
||
<taxonomicName id="32046B1A0B0B57C7F48A36198365DB0E" authorityName="Quang, Orlov, Ananjeva, Johns, Ngoc Thao & Quang Vinh" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chauquangensis">
|
||
<emphasis id="824CC7CAA3135305FF8D16A6F348B8FC" italics="true">chauquangensis</emphasis>
|
||
</taxonomicName>
|
||
group by the combination of having a maximum
|
||
<abbrev id="ABBRID0EWQAG" xlink_title="snout-vent length">SVL</abbrev>
|
||
=
|
||
<quantity id="846042E9E178160D16D0A80CB805B12C" metricMagnitude="-2" metricUnit="m" metricValue="7.44" unit="mm" value="74.4">74.4 mm</quantity>
|
||
(female); 8–12 supralabials; 9–11 infralabials; 30–43 paravertebral tubercles; 19–25 rows of longitudinally arranged tubercles; 29–37 longitudinal rows of ventrals; 6–9 expanded subdigital lamellae on the fourth toe; 12–14 unmodified subdigital lamellae on the fourth toe; 19–22 total subdigital lamellae on the fourth toe; 19–28 total number of enlarged femoral scales; 9–14 total number of femoral pores in males (
|
||
<emphasis id="E36B9D84F855645BE82B70377714BCE9" italics="true">n</emphasis>
|
||
= 4); 6–11 enlarged precloacals; 4–6 precloacal pores in males (
|
||
<emphasis id="EE891E36A2C1300B847B5A4F106242BD" italics="true">n</emphasis>
|
||
= 4); two or three rows of large post-precloacal scales; enlarged femorals and enlarged precloacals continuous; proximal femorals usually smaller than distal femorals; femoral pores restricted to distal scales; body tubercles weakly keeled; small tubercles on forelimbs; tubercles extend beyond base of tail; medial subcaudals 2–3 times wider than long but not extending onto lateral surface of tail; nuchal loop often divided medially, bearing two posteriorly directed projections, no anterior azygous notch, projecting posterior margin; usually no triangular marking anterior to nuchal loop; dark-colored band on nape variably present; dark-colored dorsal bands lack paravertebral elements, have variably lightened centers, are edged with white tubercles, usually jagged in shape, and the same width or wider than interspaces; dark-colored markings in dorsal interspaces; no whitish ventrolateral fold; top of head in adults diffusely mottled, blotched; no light-colored reticulum on top of head; 4–6 dark-colored transverse body bands; 10–13 light-colored caudal bands on an original tail bearing dark-colored markings and not encircling tail (
|
||
<emphasis id="399BDDACB106266E33AB2D8AABAA750F" italics="true">n</emphasis>
|
||
= 7); 9–12 dark-colored caudal bands on an original tail and wider than light-colored caudal bands (
|
||
<emphasis id="35088FBDFFFF7259EFE33BF5D6D7DDA8" italics="true">n</emphasis>
|
||
= 7); and mature regenerated tail mottled (
|
||
<emphasis id="303F5F1B86D82FC016699BC419BE910D" italics="true">n</emphasis>
|
||
= 3) (Table 4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EERAG" type="description">
|
||
<paragraph id="2C715A4FD5EBD34D38A784C63CCB93A7">
|
||
<heading id="9FA8DA9B331D66B4D65EB7F5BCC2C4EE" reason="title">
|
||
Description of
|
||
<typeStatus id="8801702B9E5B28B40A7E4B93BCA417EB" type="holotype">holotype</typeStatus>
|
||
.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="43F83969DD051E539E30F3F42F6C179B">
|
||
(Figs
|
||
<figureCitation id="AC00BE75749B8F3678CECD275767FCF7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selected individuals of the type series and referred specimen of Cyrtodactylus phamiensis sp. nov. from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand." figureDoi="10.3897/zookeys.1203.122758.figure4" httpUri="https://binary.pensoft.net/fig/1059372">4 A</figureCitation>
|
||
,
|
||
<figureCitation id="3E004B6FA3AEA4A8B404AD05A1F19917" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Adult male holotype of Cyrtodactylus phamiensis sp. nov. A dorsal view B ventral view C dorsal view of head D gular region E thighs and precloacal region F ventral view of right manus G ventral view of left pes H subcaudal region I lateral view of left side of head. Photographs by Attapol Rujirawan." figureDoi="10.3897/zookeys.1203.122758.figure5" httpUri="https://binary.pensoft.net/fig/1059373">5</figureCitation>
|
||
, Suppl. material
|
||
<xref id="BE20D9C095AEA343455836EFA6C8441E" ref-type="supplementary-material" rid="S3">3</xref>
|
||
) Adult male
|
||
<abbrev id="ABBRID0EWRAG" xlink_title="snout-vent length">SVL</abbrev>
|
||
<quantity id="7E323B4D4881B087EB4431B340FCC177" metricMagnitude="-2" metricUnit="m" metricValue="6.85" unit="mm" value="68.5">68.5 mm</quantity>
|
||
; head moderate in length (
|
||
<abbrev id="ABBRID0E1RAG" xlink_title="head length">HL</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0E5RAG" xlink_title="snout-vent length">
|
||
<collectionCode id="E0D62E996BAFA7213FB0C486C4F0BE6A">SVL</collectionCode>
|
||
</abbrev>
|
||
0.28), width (
|
||
<abbrev id="ABBRID0ECSAG" xlink_title="head width">
|
||
<collectionCode id="BD989B6F5DFF300EAC00DD706758DD25">HW</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EGSAG" xlink_title="head length">HL</abbrev>
|
||
0.72), flattened (
|
||
<abbrev id="ABBRID0EKSAG" xlink_title="head depth">
|
||
<collectionCode id="BC5EDCB334A0D625DE74F363A0287A05">HD</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EOSAG" xlink_title="head length">HL</abbrev>
|
||
0.40), distinct from neck, triangular in dorsal profile; lores weakly concave anteriorly, weakly inflated posteriorly; prefrontal region concave; canthus rostralis rounded; snout elongate (
|
||
<abbrev id="ABBRID0ESSAG" xlink_title="snout length">
|
||
<collectionCode id="1A5D69B493511C520D0FC9242C13BB3D">ES</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EWSAG" xlink_title="head length">HL</abbrev>
|
||
0.40), flat, rounded in dorsal profile; eye large (
|
||
<abbrev id="ABBRID0E1SAG" xlink_title="eye diameter">
|
||
<collectionCode id="9E9D2C96E7BCF1FAC6F7F25B74F725D9">ED</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0E5SAG" xlink_title="head length">HL</abbrev>
|
||
0.31); ear opening elliptical, obliquely oriented, moderate in size; eye to ear distance slightly greater than diameter of eye; rostral rectangular, partially divided dorsally by inverted Y-shaped furrow, bordered posteriorly by large left and right supranasals, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two moderately sized postnasals, bordered ventrally by first supralabial; nine (R, L) rectangular supralabials tapering abruptly to below midpoint of eye, first – fifth supralabials largest; 11 (R, L) infralabials tapering smoothly to slightly past the termination of enlarged supralabials to corner of mouth; scales of rostrum and lores flat, larger than granular scales on top of head and occiput; scales of occiput intermixed with small, rounded, tubercles; superciliaries elongate, largest dorsally; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapezoidal postmentals contacting medially for ~ 65 % of their length posterior to mental; one row of enlarged, sublabials extending posteriorly to fifth infralabials (R, L); gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales.
|
||
</paragraph>
|
||
<caption id="ED5667A25F879AF37B9475BD585E0661" ID-DOI="10.3897/zookeys.1203.122758.figure5" ID-arpha="9D782E66-1DCB-58E5-870A-8CAF05F2F781" httpUri="https://binary.pensoft.net/fig/1059373" startId="F5">
|
||
<paragraph id="F720848DB649D132C12AA6DA7FACA1B2">
|
||
<label id="66B911A7ECCF65A160249EDA93FAE63A">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="912DB3C76DA6C73FB61FE952C7E2E6D7">
|
||
Adult male holotype of
|
||
<taxonomicName id="BCF2747A273D3EE4C82C22FB6633A711" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="C009A85C4DB7FC9EA77ED2F1F8DCBD5A" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FC958073FDE96B616618B86D229D0E56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="341C8E498B6E504E8688EB18A80A3287" bold="true">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="82C81A4D15623F6E8BFBB8132BE52F90" bold="true">B</emphasis>
|
||
ventral view
|
||
<emphasis id="20C73E8C5CB4A8BCCA1718AAEBA06FE2" bold="true">C</emphasis>
|
||
dorsal view of head
|
||
<emphasis id="A1934D6BFBE98C7727C95F3EF74F2702" bold="true">D</emphasis>
|
||
gular region
|
||
<emphasis id="367199E681A599DE8933A14DB12F8447" bold="true">E</emphasis>
|
||
thighs and precloacal region
|
||
<emphasis id="AFB97BCEB1A292F3F78EAE0737D27E48" bold="true">F</emphasis>
|
||
ventral view of right manus
|
||
<emphasis id="3B6FD0C42ADA993D1F677A52E40AC4AD" bold="true">G</emphasis>
|
||
ventral view of left pes
|
||
<emphasis id="85A980157BDA9607AB0ED3BFF95E3752" bold="true">H</emphasis>
|
||
subcaudal region
|
||
<emphasis id="B43813ABCF1836405AE23A5D7D4A4D8F" bold="true">I</emphasis>
|
||
lateral view of left side of head. Photographs by Attapol Rujirawan.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3311B7720C54990713AF48A8833E9CC1">
|
||
Body relatively long (
|
||
<abbrev id="ABBRID0EOUAG" xlink_title="axilla to groin length">
|
||
<collectionCode id="677269B5299C2A09577FC70212F89E92">AG</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0ESUAG" xlink_title="snout-vent length">
|
||
<collectionCode id="D63E5164D7E70732DDAB969FB0854424">SVL</collectionCode>
|
||
</abbrev>
|
||
0.46) with well-defined ventrolateral folds; dorsal scales small, granular, interspersed with moderately sized, smooth, rounded, semi-regularly arranged tubercles extending from occiput to slightly beyond base of tail; ~ 25 longitudinal rows of tubercles at midbody; ~ 33 paravertebral tubercles; 33 flat, imbricate, ventral scales much larger than dorsal scales; eight enlarged precloacal scales, six bearing pores; no deep precloacal groove or depression; and two rows of large post-precloacal scales on midline.
|
||
</paragraph>
|
||
<paragraph id="99F3B97C95A5C124989872942CC1D520">
|
||
Forelimbs moderate in length and stature (
|
||
<abbrev id="ABBRID0EYUAG" xlink_title="forearm length">
|
||
<collectionCode id="9044C1F53C58C4E07CCA90A4419D099B">FL</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0E3UAG" xlink_title="snout-vent length">
|
||
<collectionCode id="0B1F85AF384FB6ECF2284E5C4E546117">SVL</collectionCode>
|
||
</abbrev>
|
||
0.16); granular scales of forelimbs slightly larger than those on body, small rounded tubercles on dorsal surface of forearms; palmar scales flat, juxtaposed; digits well-developed, inflected at basal interphalangeal joints, slightly narrower distal to inflections; subdigital lamellae transversely expanded, those proximal to joint inflections much wider than nearly unmodified lamellae distal to inflections; claws well-developed, sheathed by a dorsal and ventral scale; hind limbs robust, wider and longer than forelimbs (
|
||
<abbrev id="ABBRID0EAVAG" xlink_title="tibia length">
|
||
<collectionCode id="DF66249323B921A23CA6B457DD355A8A">TBL</collectionCode>
|
||
</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EEVAG" xlink_title="snout-vent length">
|
||
<collectionCode id="859CEDB5C83C9CE170281D929B83404E">SVL</collectionCode>
|
||
</abbrev>
|
||
0.20), covered dorsally by granular scales interspersed with moderately sized tubercles, larger and flat scales anteriorly; ventral scales of thighs flat, imbricate, slightly larger than dorsals; subtibial scales small, flat, imbricate; one row of 10 (R) 11 (L) enlarged femoral scales terminating distally before knee, continuous with enlarged precloacal scales; proximal femorals nearly same size as distal femorals, all femorals forming an abrupt union with smaller, granular, ventral scales of posteroventral scales of thigh; femoral pores 4 (R) 5 (L) restricted to distalmost femorals; plantar scales flat, juxtaposed; digits well-developed, inflected at basal interphalangeal joints; claws well-developed, sheathed by a dorsal and ventral scale at base; seven (R, L) wide subdigital lamellae on fourth toe proximal to joint inflection, 12 (R, L) narrower lamellae distal to joint inflection, 19 total subdigital lamellae.
|
||
</paragraph>
|
||
<paragraph id="02D1FDDA767FE985809798D538F67FA5">
|
||
Tail regenerated, long (
|
||
<abbrev id="ABBRID0EKVAG" xlink_title="tail length">TL</abbrev>
|
||
/
|
||
<abbrev id="ABBRID0EOVAG" xlink_title="snout-vent length">
|
||
<collectionCode id="18800A3428F5426C96610FFB1CA600A1">SVL</collectionCode>
|
||
</abbrev>
|
||
1.14), thin,
|
||
<quantity id="1A84A25F07510780B4D735FD576DDD8A" metricMagnitude="-2" metricUnit="m" metricValue="7.81" unit="mm" value="78.1">78.1 mm</quantity>
|
||
in length,
|
||
<quantity id="712C07169031AA3D47217DC37F0F4013" metricMagnitude="-3" metricUnit="m" metricValue="6.9" unit="mm" value="6.9">6.9 mm</quantity>
|
||
wide at base, tapering to a point; dorsal caudal scales small, generally square, juxtaposed; median row of subcaudals significantly larger than dorsal caudals, transversely expanded, not extending dorsally onto lateral side of tail; body tubercles extending slightly beyond base of tail; faint hemipenal swellings at base of tail, two large postcloacal tubercles on both sides; and postcloacal scales flat, imbricate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ESVAG" type="Coloration prior to preservation">
|
||
<paragraph id="00EAED3E4476B7C81BA37C56AAD45F47">
|
||
<heading id="843958D90A1D75C9AA56C2F740EC2A18" reason="title">Coloration prior to preservation.</heading>
|
||
</paragraph>
|
||
<paragraph id="482EAE38EA7DDAF4A09032A233E35ED1">
|
||
(Figs
|
||
<figureCitation id="3322900564846930FD4A805F5FE3D1A2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selected individuals of the type series and referred specimen of Cyrtodactylus phamiensis sp. nov. from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand." figureDoi="10.3897/zookeys.1203.122758.figure4" httpUri="https://binary.pensoft.net/fig/1059372">4</figureCitation>
|
||
,
|
||
<figureCitation id="F89A0C425A06305609E6F8D536CCFD5A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Adult male holotype of Cyrtodactylus phamiensis sp. nov. A dorsal view B ventral view C dorsal view of head D gular region E thighs and precloacal region F ventral view of right manus G ventral view of left pes H subcaudal region I lateral view of left side of head. Photographs by Attapol Rujirawan." figureDoi="10.3897/zookeys.1203.122758.figure5" httpUri="https://binary.pensoft.net/fig/1059373">5</figureCitation>
|
||
) Ground color of top of head, limbs, and dorsum straw to pale brown; top of head bearing poorly defined, irregularly shaped, dark brown markings; dark brown, nuchal loop bearing two posterior projections extend between postorbital regions; well-defined, rectangular dark brown band on nape; six dark brown, immaculate, weakly jagged, dorsal body bands terminating above the ventrolateral folds extending from shoulders to groin, same width as straw-colored interspaces, not edged with white or bright-colored tubercles; one darkly colored sacral band; dorsal interspaces faintly mottled, each bearing a brown “ fuzzy-edged ” longitudinal vertebral marking; forelimbs faintly mottled; hind limbs more darkly mottled, accentuating light-colored tubercles; one post-sacral and five wide, dark brown caudal bands slightly wider whitish caudal bands markings; whitish caudal bands do not encircle tail, bear subcircular dark brown markings; iris reddish gold with thin black reticulations; venter beige with faint, dark shadowing on lateral edges of belly and limbs; and subcaudal region dark-brown, weakly mottled with pale-colored markings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EAWAG" type="etymology">
|
||
<paragraph id="1E29AEDFB8F2193175911EE07D20174F">
|
||
<heading id="5DDE0794FB405621703A01435BE0A41C" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="3F3000ACED23C73330E788B1128E09AC">
|
||
The species name
|
||
<emphasis id="946F4C84AAD20D0858BE5873FEFBA794" italics="true">phamiensis</emphasis>
|
||
is in reference to the
|
||
<typeStatus id="56A19EE7BD2298C58FA83DB876E9698D">type</typeStatus>
|
||
locality at Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District,
|
||
<collectingRegion id="920180FC7C7A4C01B77D64200240C8C4" country="Thailand" name="Chiang Rai">Chiang Rai Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="1FAA7B25E8742DEE620A4A732AA60A3D" name="Thailand">Thailand</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="B08D4A433E359201E56BF76F3C324AE6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of nominal species and unnamed populations of the Cyrtodactylus chauquangensis group. The star denotes the type locality of Cyrtodactylus phamiensis sp. nov." figureDoi="10.3897/zookeys.1203.122758.figure1" httpUri="https://binary.pensoft.net/fig/1059369">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUWAG" type="distribution">
|
||
<paragraph id="333042B3E65834862DC2DF4A6CC6BC22">
|
||
<heading id="7F35302A2C616D5E9FE4F996F22C8E7D" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="C37B72E31502D808C326D6EEF9E56174">
|
||
The type series of
|
||
<taxonomicName id="F9E4CF763809D47A4D6EA17DC01DFB4E" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="491D8F75BA0A5D168670F1D9B47EC400" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="27083634638B551BA2D30E155D2BE9ED" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus id="3B6FDF2543A0C565734C31FA60E5B03D">type</typeStatus>
|
||
locality at Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District,
|
||
<collectingRegion id="851D7A6ACCC84EFCE6EBA7D86CC762F8" country="Thailand" name="Chiang Rai">Chiang Rai Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="51E92B09E02D7E586013C2554866C8DA" name="Thailand">Thailand</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="1CB6A2BE48D13077FEB042FD29D5916B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of nominal species and unnamed populations of the Cyrtodactylus chauquangensis group. The star denotes the type locality of Cyrtodactylus phamiensis sp. nov." figureDoi="10.3897/zookeys.1203.122758.figure1" httpUri="https://binary.pensoft.net/fig/1059369">1</figureCitation>
|
||
). On
|
||
<date id="890FA30EE30B479880393E11788BCF42" value="2023-03-30">30 March 2023</date>
|
||
, a bat researcher from the University of
|
||
<collectingCountry id="137D2BF4D18FA2BA35B6F8EE3CEA2E92" name="Hong Kong">Hong Kong</collectingCountry>
|
||
, Ada Chornelia, informed us of this species’ potential presence in a two adjacent karst caves at monastery
|
||
<quantity id="FDAD8D6A3FCC7E2E87227A3D2A0CC02C" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
|
||
north of the
|
||
<typeStatus id="7BA80385EA7AFE477BE6EE64D9288CC1">type</typeStatus>
|
||
locality in Tham Pha Chom along the same line of large karst formations. Examination of a photograph (Fig.
|
||
<figureCitation id="879EFBD37AAB770E89DA04970CD54A5D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cyrtodactylus cf. phamiensis sp. nov. from 5 km north of the type locality in Tham Pha Chom. Photograph by Ada Chornelia." figureDoi="10.3897/zookeys.1203.122758.figure6" httpUri="https://binary.pensoft.net/fig/1059374">6</figureCitation>
|
||
) from this locality, tentatively confirms this observation. Furthermore, on
|
||
<date id="7E612118D84BB6795095DA31479B5BFA" value="2022-12-17">17 December 2022</date>
|
||
three
|
||
<taxonomicName id="1CBEE908EA15A31DFF46BB72624D7920" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="C9CADDEADFC90BF7B2881DFB53E0C150" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="62F262BD4705F28D14E44F6E5EB977FE" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were observed, one of which was photographed, only 50 meters west-southwest of the
|
||
<typeStatus id="3B7D6BAAEF2F65E6668C42046486585A">type</typeStatus>
|
||
locality immediately east of Pha Mi Village by H. Bringsøe (Fig.
|
||
<figureCitation id="F513ECD944BA7CB5ABC0B53AEBFB751E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus phamiensis sp. nov. from 50 meters west-southwest of the type locality, immediately east of Pha Mi Village. 17 December 2022. Photograph by Henrik Bringsøe." figureDoi="10.3897/zookeys.1203.122758.figure7" httpUri="https://binary.pensoft.net/fig/1059375">7</figureCitation>
|
||
) from the same karstic formations. It is likely that
|
||
<taxonomicName id="A88BAD2AE78FD54A8E4C249758E0363A" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="59EBEEBC377BC723F2C05D547D5EB0F1" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BAE2882E01F31BDBE19E23FC9675E532" rank="species">sp. nov.</taxonomicNameLabel>
|
||
ranges throughout the karstic landscapes of this region.
|
||
</paragraph>
|
||
<caption id="441F4A80A6814FD7786DFA3FEA4CBC18" ID-DOI="10.3897/zookeys.1203.122758.figure6" ID-arpha="0451D01F-EEFD-5921-B4C6-74E7251E9DC6" httpUri="https://binary.pensoft.net/fig/1059374" startId="F6">
|
||
<paragraph id="93D113D9B5B1267AD6196164926908E3">
|
||
<label id="209C62510579D08C55838531CA761E7D">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="62D60710C57942FD772AAB98F03C4B70">
|
||
<taxonomicName id="F859C7CB4418EB3740499EA447E5586C" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" isUncertain="true" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="0600A8DC3610E78A7679CDB25C7FE8D9" italics="true">Cyrtodactylus</emphasis>
|
||
cf.
|
||
<emphasis id="72785FB4878EBB6A2F30A94513C2251C" italics="true">phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="053CBA0935A7212FE40B9077190F5AC6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from 5 km north of the type locality in Tham Pha Chom. Photograph by Ada Chornelia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D315BD107757D5CD6CCABE1967460D70" ID-DOI="10.3897/zookeys.1203.122758.figure7" ID-arpha="8FF90C35-77B9-5E8F-A2F8-A11CF5147E41" httpUri="https://binary.pensoft.net/fig/1059375" startId="F7">
|
||
<paragraph id="0DBA8D1110DFEA57E3C8673664AA643F">
|
||
<label id="801070FCAFF80BB0D1D901420A22AB58">Figure 7.</label>
|
||
</paragraph>
|
||
<paragraph id="ED471458531B9F049835856A224D5750">
|
||
<taxonomicName id="699C2FBB59E3912166B17748B0C1EC6D" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="0EEB179DD7F4BC5E07D3C81945708EC4" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="74E4DF94628010DFF094B0BED8279CDD" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from 50 meters west-southwest of the type locality, immediately east of Pha Mi Village. 17 December 2022. Photograph by Henrik Bringsøe.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZZAG" type="variation">
|
||
<paragraph id="A24D3215C049ED04A02743D4822B3B51">
|
||
<heading id="F7D9C747D6DDB3A6DAEF01128DB5DC7C" reason="title">Variation.</heading>
|
||
</paragraph>
|
||
<paragraph id="B1F6C3A3A3552F7A32885CCB6DF9BCF7">
|
||
(Table 4) The
|
||
<typeStatus id="B2343B100696DF89FE8C8D15870A0227" type="paratype">paratypes</typeStatus>
|
||
closely approach the
|
||
<typeStatus id="F1D9059898BE97F79F017416708650AC" type="holotype">holotype</typeStatus>
|
||
in general coloration and pattern (Fig.
|
||
<figureCitation id="CA9441175DF34D058D189F3E2EDDEFA1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Paratypes of Cyrtodactylus phamiensis sp. nov. from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand. Photographs by Attapol Rujirawan." figureDoi="10.3897/zookeys.1203.122758.figure8" httpUri="https://binary.pensoft.net/fig/1059376">8</figureCitation>
|
||
). The most notable variation pertains to the shape of the nuchal loop, nape band, the caudal pattern and morphology. In
|
||
<abbrev id="ABBRID0ED1AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="9C999FBE7CEC637396026D05CB175510" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="06D57E7D9FBD0C754CD47C7CF8EF8696">R 01073</specimenCode>
|
||
–74,
|
||
<abbrev id="ABBRID0EH1AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="64A56EBA94420850122ED2451DB24FCE" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="87E837C9D76B0C78F0BEB4A9A47AF145">R 01081</specimenCode>
|
||
,
|
||
<abbrev id="ABBRID0EL1AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="31834A4E5AD73C0FD849944A61B4DF6F" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="A31C70DE92DAD1A887B900128BEEAEBD">R 01083</specimenCode>
|
||
–84 and
|
||
<abbrev id="ABBRID0EP1AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="1210BB197CB1CAE1326725A1041F994E" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="B8A8B235BF3E6D10F310576C683F5737">R 01087</specimenCode>
|
||
, the nuchal loop is medially bifurcated. That of
|
||
<abbrev id="ABBRID0ET1AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="6551472EE855218E78656FC9ACA28D29" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="B73E047FC3212774376B52ECC7BC5A70">R 01075</specimenCode>
|
||
is irregular and ill-defined. The
|
||
<typeStatus id="EB300F10BEEB2F3F2266ADCC6CA3056D" type="holotype">holotype</typeStatus>
|
||
,
|
||
<abbrev id="ABBRID0EX1AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="4911362D1C5849057C6ED3568FE1652F" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="F1BFE736C2AA182295A0A74B11FE647F">R 01086</specimenCode>
|
||
, is the only specimen with a complete rectangular nape band. Nape bands of the
|
||
<typeStatus id="36599A5B126237E475D3D16AA4494B9D" type="paratype">paratypes</typeStatus>
|
||
are either tripartite or nearly so.
|
||
<abbrev id="ABBRID0E21AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="89BF6EEA1F1712C0D30FC64A373BFDC6" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="92BBC2455E545103E964A88C4EFAA226">R 01074</specimenCode>
|
||
,
|
||
<abbrev id="ABBRID0E61AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="406763D999883344A3F37DA07919DCDC" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="C68D07B82E015C7812DBE98E4533E2A8">R 01084</specimenCode>
|
||
and
|
||
<abbrev id="ABBRID0ED2AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="AAD4BDCCE3BB45EB6520FB40090CF177" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="73E56F541D9A462282359AB5E2E04B3B">R 01087</specimenCode>
|
||
have complete tails. Tails of
|
||
<abbrev id="ABBRID0EH2AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="681660B40DEB1A57F0B16DDEAB166999" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="41151E71A80DBE5D95D8E8E83B2FCC48">R 01075</specimenCode>
|
||
and
|
||
<abbrev id="ABBRID0EL2AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="4F44F43F750D398895E115F328879325" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="D5BE50C4BFACF6ECB31CDDEA7603FAA9">R 01078</specimenCode>
|
||
are three-quarters to one-half regenerated, respectively.
|
||
<abbrev id="ABBRID0EP2AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="8E6F9C18C19CDAEA9CDD85ABBE6EEA96" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="93F659BB42BD178B23A2AA6610B0B0EB">R 01081</specimenCode>
|
||
and
|
||
<abbrev id="ABBRID0ET2AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="0CB674E59E5BA40B896072E5F012E27F" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="CC1CDEA5D3C0EDEAC36CDAD287B8214C">R 01085</specimenCode>
|
||
lack the majority of their tails. The body bands of
|
||
<abbrev id="ABBRID0EX2AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="AB1E43A268EF1A0E1104FE0FD7DDEBED" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="63C5A4D3356119FA8F6E030066E9A3A6">R 01074</specimenCode>
|
||
–75,
|
||
<abbrev id="ABBRID0E22AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="41C7761D1A473671D841EBD3E54A233D" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="DD9A3941BD2FC1FE6B0F559D5705393C">R 01085</specimenCode>
|
||
and
|
||
<abbrev id="ABBRID0E62AG" xlink_title="Zoological Museum, Kasetsart University, Bangkok, Thailand">
|
||
<collectionCode id="1F690C6576B2D28D6BF4A8AD9FC8F5AC" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" type="">ZMKU</collectionCode>
|
||
</abbrev>
|
||
<specimenCode id="FCE86B0C4F89D87F0BAA74B9A9612B59">R 01087</specimenCode>
|
||
have lightened centers. Morphometric, meristic, and categorical data of the type series and referred specimens are listed in Suppl. material
|
||
<xref id="F14CDA6BA5590678EA0AAFB000A6641A" ref-type="supplementary-material" rid="S3">3</xref>
|
||
.
|
||
</paragraph>
|
||
<caption id="8CE82769AE75B7B3A465CCD3DED84171" ID-DOI="10.3897/zookeys.1203.122758.figure8" ID-arpha="A024602B-DAB4-596D-A0E7-0C6F88BE6303" httpUri="https://binary.pensoft.net/fig/1059376" startId="F8">
|
||
<paragraph id="107376DC10191A6A96C2F0C271413F58">
|
||
<label id="468309B2B7B9CAD8B8F659C8CAC5F9CC">Figure 8.</label>
|
||
</paragraph>
|
||
<paragraph id="93A75492248868FCD8162B2BA368215C">
|
||
Paratypes of
|
||
<taxonomicName id="6240DFE308FB84857B7610AA6E6A703C" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="8C7131456E8FFD98490EBE5A5B7353F3" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="725EBFCF206C3333ED0D28FB98B9008F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand. Photographs by Attapol Rujirawan.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E63AG" type="comparisons">
|
||
<paragraph id="87AED88A7A1A52A434DB88CF6B5E87EA">
|
||
<heading id="E6C1B871EFD9BE87A061129260AB8C9A" reason="title">Comparisons.</heading>
|
||
</paragraph>
|
||
<paragraph id="D1E3CCAC1D0610194BC46B392EF0F5CB">
|
||
<taxonomicName id="6CC6F05605D1168878F9E63B34610FD0" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="29E02B43965164411A6339A59B9BBD70" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CCD5AD5422D1EFA304A2CCDE473B0A97" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is embedded in clade 2 and is the sister species to a clade composed of three lineages,
|
||
<taxonomicName id="FE97FBCD49A118637C6E77E9F0F86131" authorityName="Kunya, Panmongkol, Pauwels, Sumontha, Meewasana, Bunkhwamdi & Dangsri" authorityYear="2014" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="doisuthep">
|
||
<emphasis id="DCCC4810716C0FCFE173B77B5ED4E2BF" italics="true">C. doisuthep</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="85435167D107249D09E074ECC320A369" authorityName="Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya" authorityYear="2009" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="erythrops">
|
||
<emphasis id="EF9593698EB7A38CBDB83E7857B36DC6" italics="true">C. erythrops</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="DC8859E067EBD0F973B8FF501B10A7D2" italics="true">C.</emphasis>
|
||
sp. 6.
|
||
<taxonomicName id="DB0B6406568844A9A2711551390D2A38" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="35566F598645830F86BD5821317F1A56" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="805B8E33F2559899008A6EC3026677B6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from those three lineages by mean uncorrected pairwise sequence divergence of 13.5–14.5 % and the remaining species in the
|
||
<taxonomicName id="B6CF48B8A59215D18A04FA64E56CF0E9" authorityName="Quang, Orlov, Ananjeva, Johns, Ngoc Thao & Quang Vinh" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chauquangensis">
|
||
<emphasis id="564F857BC942571E074FB2FB3029A665" italics="true">chauquangensis</emphasis>
|
||
</taxonomicName>
|
||
group by 13.7–17.3 % (Suppl. material
|
||
<xref id="0560F2A1E81F681CD51D14A198F181B0" ref-type="supplementary-material" rid="S2">2</xref>
|
||
). It differs from
|
||
<taxonomicName id="BE53424DB0FDBB1E25D99FDEA9113CA2" authorityName="Kunya, Panmongkol, Pauwels, Sumontha, Meewasana, Bunkhwamdi & Dangsri" authorityYear="2014" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="doisuthep">
|
||
<emphasis id="829698859A667EA85E5562041DDFACBD" italics="true">C. doisuthep</emphasis>
|
||
</taxonomicName>
|
||
by having maximum
|
||
<abbrev id="ABBRID0ER6AG" xlink_title="snout-vent length">SVL</abbrev>
|
||
<quantity id="CA8766CE3732E2CC40970D37B5D5D2C0" metricMagnitude="-2" metricUnit="m" metricValue="7.44" unit="mm" value="74.4">74.4 mm</quantity>
|
||
(vs
|
||
<quantity id="5C79E080B17A3F5292EEB34C07AC7401" metricMagnitude="-2" metricUnit="m" metricValue="9.05" unit="mm" value="90.5">90.5 mm</quantity>
|
||
); 9–14 total number of femoral pores in males (vs absent); and lacking light-colored reticulum on the top of the head (vs present;
|
||
<bibRefCitation id="BD7A62F8FF14D64D9FF00DB248340EA5" DOI="10.11646/zootaxa.3811.2.6" author="Kunya" etAl="et al." firstAuthor="Kunya" issue="2" journalOrPublisher="Zootaxa" pagination="251-261" refId="B22" refString="Kunya K, Panmongkol A, Pauwels OSG, Sumontha M, Meewasana J, Bunkhwamdi W, Dangsri S (2014) A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Suthep, Chiang Mai Province, northern Thailand. Zootaxa 3811 (2): 251–261. https://doi.org/10.11646/zootaxa.3811.2.6" title="A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Suthep, Chiang Mai Province, northern Thailand." volume="3811" year="2014">Kunya et al. 2014</bibRefCitation>
|
||
: fig. 1).
|
||
<taxonomicName id="B667E78D0A692D02AA093B802608463D" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="BDCB5EA6DB2E567E31D0ACEC3E82CD79" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DD6A2D6340EB289593C11C0C1A23A71C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="0FF5B7D0E75CAC37F4C28B89A123F89E" authorityName="Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya" authorityYear="2009" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="erythrops">
|
||
<emphasis id="D65860ED7ADB00177FE592CD11301916" italics="true">C. erythrops</emphasis>
|
||
</taxonomicName>
|
||
by having 29–37 longitudinal rows of ventrals (vs 28 rows); 9–14 total number of femoral pores in males (vs 19 pores); 4–6 precloacal pores in males (vs 9 pores); and hatchlings with yellow-colored heads (vs tan colored;
|
||
<bibRefCitation id="029F5BE3A87AE9B8524F2B37D2268B6E" DOI="10.11646/zootaxa.2124.1.4" author="Bauer" etAl="et al." firstAuthor="Bauer" issue="1" journalOrPublisher="Zootaxa" pagination="51-62" refId="B2" refString="Bauer AM, Kunya K, Sumontha M, Niyomwan P, Panitvong N, Pauwels OSG, Chanhome L, Kunya T (2009) Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand. Zootaxa 2124 (1): 51–62. https://doi.org/10.11646/zootaxa.2124.1.4" title="Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand." volume="2124" year="2009">Bauer et al. 2009</bibRefCitation>
|
||
: fig. 5). Additional comparisons (meristics, morphometrics, and subcaudal scale morphology) between
|
||
<taxonomicName id="58853004507110ADE6A080AD80FF6E0D" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="0C0FB1E8F56C0FB63D1366B41EB04974" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0C0F654A240B2B2D6CA3F7BE49CD1F2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and the remaining species in the
|
||
<taxonomicName id="04033BEA13CEEF8C3AE94A8895B92E12" authorityName="Quang, Orlov, Ananjeva, Johns, Ngoc Thao & Quang Vinh" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chauquangensis">
|
||
<emphasis id="4D7A445DDCE2626C060BA61029BDC91F" italics="true">chauquangensis</emphasis>
|
||
</taxonomicName>
|
||
group are presented in Table 4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EIBBG" type="natural history">
|
||
<paragraph id="648520E35B24E6ADAED8EAAC3D70448A">
|
||
<heading id="17AF76227A6B2053626163B5AA75ED5A" reason="title">Natural history.</heading>
|
||
</paragraph>
|
||
<paragraph id="82EA551C78488221FACD3678B9D572FD">
|
||
(Fig.
|
||
<figureCitation id="A43F545D7EF1CD625F93DC37243509FE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Karst habitat of the type locality from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand. Photograph by Evan S. H. Quah." figureDoi="10.3897/zookeys.1203.122758.figure9" httpUri="https://binary.pensoft.net/fig/1059377">9</figureCitation>
|
||
) All specimens were collected during the evening between 19: 30 and 20: 50 hours on karst formations, at the entrance of a karst cave, within the cave, or on karst vegetation outside the cave at varying distances from the cave entrance.
|
||
<specimenCount id="45DF79AFF5F5E65CB8025A165D5B2158" count="1" type="generic">One specimen</specimenCount>
|
||
was observed during the day in a crack ~
|
||
<quantity id="108CDCC7AD5D4F7D11CAC57B7EBCAF4E" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</quantity>
|
||
above the cave floor ~
|
||
<quantity id="629AF9614E3451CA81D840F21636D721" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</quantity>
|
||
in from the entrance. Juveniles (
|
||
<abbrev id="ABBRID0ESBBG" xlink_title="snout-vent length">
|
||
<collectionCode id="A8D7DAF53FD6DEF1041DA09BB17A31C9">SVL</collectionCode>
|
||
</abbrev>
|
||
<
|
||
<quantity id="ED8441381AA5624A0A50889AA94F8370" metricMagnitude="-2" metricUnit="m" metricValue="4.0" unit="mm" value="40.0">40 mm</quantity>
|
||
) were found outside the cave less than
|
||
<quantity id="C3652F3EFFF86360C0C29FA2F38CD68C" metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">1 m</quantity>
|
||
above ground level on karst boulders or on the base of small trees. Most were found farther away (~
|
||
<quantity id="F8F0127C6112D8E118FDC5525575C0BC" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">20–40 m</quantity>
|
||
) from the cave entrance than adults. On 26 March, four or
|
||
<specimenCount id="792242CBD6561310CFEEC23BA061EDC3" count="5" type="juvenile">five juveniles</specimenCount>
|
||
(not collected) were also observed far from the cave entrance on karst boulders and on the base of trees. That same night, other juveniles were observed near ground level on small karst outcroppings along a shallow ravine ~
|
||
<quantity id="B7D618BF4E44C5A7E487B4CA4C5B6F55" metricMagnitude="-1" metricUnit="m" metricValue="3.0" unit="m" value="0.3">0.3 m</quantity>
|
||
southeast of the
|
||
<typeStatus id="4E90F608EC4EF3BB45ED49BA577167AE">type</typeStatus>
|
||
locality. We have noted similar behavior in juveniles of
|
||
<taxonomicName id="182BA99E995A3412867ABA396446C234" authority="Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah, 2018" authorityName="Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah" authorityYear="2018" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aunglini">
|
||
<emphasis id="9F1B9957B40CB62C668CF0E738767E3E" italics="true">Cyrtodactylus aunglini</emphasis>
|
||
<bibRefCitation id="3C7FD378F8395F3B287A3B46D5D157F7" DOI="10.11646/zootaxa.4527.2.1" author="Grismer LL & Wood Jr PL & Thura MK & Win NM & Grismer MS & Trueblood LA & Quah ESH" issue="2" journalOrPublisher="Zootaxa" pagination="151-185" refId="B14" refString="Grismer LL, Wood Jr PL, Thura MK, Win NM, Grismer MS, Trueblood LA, Quah ESH (2018 c) A re-description of Cyrtodactylus chrysopylos Bauer (Squamata: Gekkonidae) with comments on the adaptive significance of orange coloration in hatchlings and descriptions of two new species from eastern Myanmar (Burma). Zootaxa 4527 (2): 151 - 185. https: // doi. org / 10.11646 / zootaxa. 4527.2. 1" title="A re-description of Cyrtodactylus chrysopylos Bauer (Squamata: Gekkonidae) with comments on the adaptive significance of orange coloration in hatchlings and descriptions of two new species from eastern Myanmar (Burma)." volume="4527" year="2018">Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C3BEF9363B55AE487483CB2E215F89D9" authority="Grismer, Wood, Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018" authorityName="Grismer, Wood, Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw" authorityYear="2018" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bayinnyiensis">
|
||
<emphasis id="35761174C89C211926B7DBE7CA9E6702" italics="true">C. bayinnyiensis</emphasis>
|
||
<bibRefCitation id="6E33119D03F1608BE9B6CDF8EC212F10" DOI="10.1080/00222933.2018.1449911" author="Grismer LL & Wood Jr PL & Thura MK & Quah ESH & Murdoch ML & Grismer MS & Herr MW & Lin A & Kyaw H" refId="B13" refString="Grismer LL, Wood Jr PL, Thura MK, Quah ESH, Murdoch ML, Grismer MS, Herr MW, Lin A, Kyaw H (2018 b) Three more new species of Cyrtodactylus (Squamata: Gekkonidae) from the Salween Basin of eastern Myanmar underscore the urgent need for the conservation of karst habitats. Journal of Natural History 52 (19 - 20): 19 - 20, 1243 - 1294. https: // doi. org / 10.1080 / 00222933.2018. 1449911" year="2018">Grismer, Wood, Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="366F4350AF6CBBD8217C923FE71C8BE6" authority="Bauer, 2003" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chrysopylos">
|
||
<emphasis id="9C540496A0BE2EFEF1E276DF01B39A96" italics="true">C. chrysopylos</emphasis>
|
||
Bauer, 2003
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="1F333DF9DFC4565B39797B27313FD65A" authority="Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017" authorityName="Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin" authorityYear="2017" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="shwetaungorum">
|
||
<emphasis id="34EB61BACE52891E60536FE4F650F13C" italics="true">C. shwetaungorum</emphasis>
|
||
Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017
|
||
</taxonomicName>
|
||
, of unrelated species groups in
|
||
<collectingCountry id="53DFEAA908A6D86AA4C4B2BBE1F203F6" name="Myanmar">Myanmar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="CEB3F94A96C5DC30253C33E88897C743" DOI="10.1093/zoolinnean/zlx057" author="Grismer LL & Wood Jr PL & Thura MK & Zin T & Quah ESH & Murdoch ML & Grismer MS & Lin A & Kyaw H & Lwin N" issue="4" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="862-959" refId="B12" refString="Grismer LL, Wood Jr PL, Thura MK, Zin T, Quah ESH, Murdoch ML, Grismer MS, Lin A, Kyaw H, Lwin N (2018 a) Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society 182 (4): 862–959. https://doi.org/10.1093/zoolinnean/zlx057" title="Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism." volume="182">
|
||
Grismer et al. 2018
|
||
<bibRefCitation id="41331786012FA2A314AAE74A781D398E" DOI="10.1093/zoolinnean/zlx057" author="Grismer LL & Wood Jr PL & Thura MK & Zin T & Quah ESH & Murdoch ML & Grismer MS & Lin A & Kyaw H & Lwin N" firstAuthor="a" issue="4" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="862-959" refId="B12" refString="Grismer LL, Wood Jr PL, Thura MK, Zin T, Quah ESH, Murdoch ML, Grismer MS, Lin A, Kyaw H, Lwin N (2018 a) Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society 182 (4): 862–959. https://doi.org/10.1093/zoolinnean/zlx057" title="Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism." volume="182" year="2018">
|
||
a,
|
||
<bibRefCitation id="3850A5656C693B4558FF9E95D06F875D" DOI="10.1080/00222933.2018.1449911" author="Grismer LL & Wood Jr PL & Thura MK & Quah ESH & Murdoch ML & Grismer MS & Herr MW & Lin A & Kyaw H" refId="B13" refString="Grismer LL, Wood Jr PL, Thura MK, Quah ESH, Murdoch ML, Grismer MS, Herr MW, Lin A, Kyaw H (2018 b) Three more new species of Cyrtodactylus (Squamata: Gekkonidae) from the Salween Basin of eastern Myanmar underscore the urgent need for the conservation of karst habitats. Journal of Natural History 52 (19–20): 19–20, 1243–1294. https://doi.org/10.1080/00222933.2018.1449911">2018 b</bibRefCitation>
|
||
</bibRefCitation>
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="253D02BEC98395A4ACDF20DCB60F6BA9" DOI="10.11646/zootaxa.4527.2.1" author="Grismer LL & Wood Jr PL & Thura MK & Win NM & Grismer MS & Trueblood LA & Quah ESH" issue="2" journalOrPublisher="Zootaxa" pagination="151-185" refId="B14" refString="Grismer LL, Wood Jr PL, Thura MK, Win NM, Grismer MS, Trueblood LA, Quah ESH (2018 c) A re-description of Cyrtodactylus chrysopylos Bauer (Squamata: Gekkonidae) with comments on the adaptive significance of orange coloration in hatchlings and descriptions of two new species from eastern Myanmar (Burma). Zootaxa 4527 (2): 151–185. https://doi.org/10.11646/zootaxa.4527.2.1" title="A re-description of Cyrtodactylus chrysopylos Bauer (Squamata: Gekkonidae) with comments on the adaptive significance of orange coloration in hatchlings and descriptions of two new species from eastern Myanmar (Burma)." volume="4527">2018 c</bibRefCitation>
|
||
). All hatchlings of these were found on the ground far from the adults on karst formations. We suspect this may be a way to avoid predation by adults as well as a means to disperse to other karst habitats. The fact that several juveniles of
|
||
<taxonomicName id="7D1A6743496120446DD2BBAE6CA0759B" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="B71F9E1A619935660BA7C47C4D77FA79" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2452A825E28DA1F6F60ED14136A35CA0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and no gravid females were observed indicates the reproductive season must have terminated prior to March. The three individuals of
|
||
<taxonomicName id="3F5D173A68196B270B8A3C32E73B698B" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="EF179D6850FC00C84DF829E98A8903DA" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4FCDFE623228439B6E76945E3143489F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
which were found 50 meters from the
|
||
<typeStatus id="BA7294DFFC1EF89B94B0810CD0C3E799">type</typeStatus>
|
||
locality on
|
||
<date id="AFF90C44A13AB17D6406255F997F14A1" value="2022-12-17">17 December 2022</date>
|
||
were adults and were observed on the karst walls outside caves at night between 21: 30 and 22: 30 hours (Fig.
|
||
<figureCitation id="38203BA4C082F8CC0554EC0AD6302AA2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus phamiensis sp. nov. from 50 meters west-southwest of the type locality, immediately east of Pha Mi Village. 17 December 2022. Photograph by Henrik Bringsøe." figureDoi="10.3897/zookeys.1203.122758.figure7" httpUri="https://binary.pensoft.net/fig/1059375">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A9999A6D71ADDE5CDE8B8E0D473347F0" ID-DOI="10.3897/zookeys.1203.122758.figure9" ID-arpha="52DFF871-E889-5393-B045-017E793BEC6D" httpUri="https://binary.pensoft.net/fig/1059377" startId="F9">
|
||
<paragraph id="3C32D6BF15C70BA35481078BCDE3094B">
|
||
<label id="B5DAA8002D8E518C8999790CD24EF50C">Figure 9.</label>
|
||
</paragraph>
|
||
<paragraph id="58C499420172D47C516B281313D56ED5">Karst habitat of the type locality from Pha Mi Village, Wiang Phang Kham Subdistrict, Mae Sai District, Chiang Rai Province, Thailand. Photograph by Evan S. H. Quah.</paragraph>
|
||
</caption>
|
||
<paragraph id="BD102EF895C4322D29A4051A23F047DF">
|
||
Other species of herpetofauna observed in the vicinity during this period were two species of frogs,
|
||
<taxonomicName id="106DBDA2AA36943CC8C97D8297B18DEF" authority="(Blyth, 1856)" baseAuthorityName="Blyth" baseAuthorityYear="1856" class="Amphibia" family="Ranidae" genus="Sylvirana" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nigrovittata">
|
||
<emphasis id="4A90887E11A3DC04074412BA75E92881" italics="true">Sylvirana nigrovittata</emphasis>
|
||
(Blyth, 1856)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="34EAB2F0A050FF113FA0FEDD0B60F97F" authority="Hallowel, 1861" authorityName="Hallowel" authorityYear="1861" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="9EE35C5A58A2D2EAC10359E4443A577C" italics="true">Polypedates megacephalus</emphasis>
|
||
Hallowel, 1861
|
||
</taxonomicName>
|
||
; four other
|
||
<taxonomicName id="D3DFE3C1C4F6613BC2359776228A0E72" baseAuthorityName="Linnaeua" baseAuthorityYear="1758" class="Squamata" family="Gekkonidae" genus="Gekko" kingdom="Animalia" phylum="Chordata" rank="species" species="gecko">
|
||
<emphasis id="12CB533EB1F76FFBD3187489A91A06A9" italics="true">gecko</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName id="5FE8E75AD690D0BA70E08D00F8081505" authority="(Wiegmann, 1834)" baseAuthorityName="Wiegmann" baseAuthorityYear="1834" class="Squamata" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" phylum="Chordata" rank="species" species="mutilata">
|
||
<emphasis id="295980E8DF6A58699F7935F26CE68ACF" italics="true">Gehyra mutilata</emphasis>
|
||
(Wiegmann, 1834)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="90809C8CF68A0FF98F3199D10AE16644" authority="(Linnaeua, 1758)" baseAuthorityName="Linnaeua" baseAuthorityYear="1758" class="Squamata" family="Gekkonidae" genus="Gekko" kingdom="Animalia" phylum="Chordata" rank="species" species="gecko">
|
||
<emphasis id="5D0C9C19B7E7AA2BE472ABACE6F0CBDF" italics="true">Gekko gecko</emphasis>
|
||
(Linnaeua, 1758)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="793CCA852E849C4330B62D23857A9621" authority="Dumeril & Bibron, 1836" authorityName="Dumeril & Bibron" authorityYear="1836" class="Squamata" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="garnotii">
|
||
<emphasis id="09457DF3511694606913F8AD91F9F020" italics="true">Hemidactylus garnotii</emphasis>
|
||
Duméril & Bibron, 1836
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="0E84F3D85EE3742069C4E72172380646" authority="(Schneider, 1797)" baseAuthorityName="Schneider" baseAuthorityYear="1797" class="Squamata" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="platyurus">
|
||
<emphasis id="4B2A3727C885009DF7EF9EE9D6BA3BD0" italics="true">Hemidactylus platyurus</emphasis>
|
||
(Schneider, 1797)
|
||
</taxonomicName>
|
||
; and a pitviper
|
||
<taxonomicName id="7FC33F3B00774431710E69FAC27116D1" authority="Kramer, 1977" authorityName="Kramer" authorityYear="1977" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="F0770512C463350537BF1C3D598BD622" italics="true">Trimeresurus macrops</emphasis>
|
||
Kramer, 1977
|
||
</taxonomicName>
|
||
. We postulate that the high number of adult
|
||
<taxonomicName id="3DD6B673CEB56CD4538FDFF259E052EA" authority="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan, 2024" authorityName="Grismer & Aowphol & Grismer & Aksornneam & Quah & Murdoch & Gregory & Nguyen & Kaatz & Bringsøe & Rujirawan" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="phamiensis" status="sp. nov.">
|
||
<emphasis id="DE30B29061C8A0073DF3E755B00BA7BC" italics="true">Cyrtodactylus phamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C2D053B217E9C840BF3FD84B327B4430" rank="species">sp. nov.</taxonomicNameLabel>
|
||
that had missing or regenerated tails as well as their skittish nature and that they did not stray far from their shelters could have been due to predation pressures from the large
|
||
<taxonomicName id="D6A16CF362EE5D2DB6AB163A14448F42" baseAuthorityName="Linnaeua" baseAuthorityYear="1758" class="Squamata" family="Gekkonidae" genus="Gekko" kingdom="Animalia" phylum="Chordata" rank="species" species="gecko">
|
||
<emphasis id="280A93433E43BB0C25F6CB074D9C79D0" italics="true">G. gecko</emphasis>
|
||
</taxonomicName>
|
||
that were also found on the karst walls and the pitvipers that were observed coiled in ambush position on vegetation beside the karst.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |