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<document ID-DOI="http://dx.doi.org/10.3897/fr.26.e86784" ID-Pensoft-Pub="2193-0074-1-27" ID-Pensoft-UUID="303E295022035210A8C508424A99393D" ID-ZooBank="6A3587A79E5740ACB9FD753EDDBA73C5" ModsDocID="2193-0074-26-1-27" checkinTime="1673667838350" checkinUser="pensoft" docAuthor="Yamada, Kazutaka, Yamamoto, Shuhei &amp; Takahashi, Yui" docDate="2023" docId="44ED1E37F5025D65B67B7D2852F1D492" docLanguage="en" docName="FossilRecord 26(1): 27-38" docOrigin="Fossil Record 26 (1)" docPubDate="2023-01-13" docSource="http://dx.doi.org/10.3897/fr.26.e86784" docTitle="Ecpaglocoris Yamada &amp; Yamamoto 2023, gen. nov." docType="treatment" docUuid="4846A233-F2BF-4083-A2DD-0DEEA05247FB" docUuidSource="ZooBank" docVersion="1" id="303E295022035210A8C508424A99393D" lastPageNumber="27" masterDocId="303E295022035210A8C508424A99393D" masterDocTitle="A new remarkable cimicoid genus and species (Hemiptera, Heteroptera, Cimicomorpha) from mid-Cretaceous Burmese amber, with implications for its aberrant male genitalia" masterLastPageNumber="38" masterPageNumber="27" pageNumber="27" updateTime="1673667838350" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new remarkable cimicoid genus and species (Hemiptera, Heteroptera, Cimicomorpha) from mid-Cretaceous Burmese amber, with implications for its aberrant male genitalia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yamada, Kazutaka</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4210-6693</mods:nameIdentifier>
<mods:affiliation>Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6 - chome, Sanda-shi, Hyogo 669 - 1546, Japan &amp; Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6 - chome, Sanda-shi, Hyogo 669 - 1546, Japan</mods:affiliation>
<mods:nameIdentifier type="email">yamada.kaz@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yamamoto, Shuhei</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4162-8457</mods:nameIdentifier>
<mods:affiliation>The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Takahashi, Yui</mods:namePart>
<mods:affiliation>Keio Yochisha Elementary School, Ebisu 2 - 35 - 1, Shibuya, Tokyo 150 - 0013, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Fossil Record</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>26</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>38</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/fr.26.e86784</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.26.e86784</mods:identifier>
<mods:identifier type="Pensoft-Pub">2193-0074-1-27</mods:identifier>
<mods:identifier type="ZooBank">6A3587A79E5740ACB9FD753EDDBA73C5</mods:identifier>
<mods:identifier type="Pensoft-UUID">303E295022035210A8C508424A99393D</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:4846A233-F2BF-4083-A2DD-0DEEA05247FB" httpUri="http://treatment.plazi.org/id/44ED1E37F5025D65B67B7D2852F1D492" lastPageNumber="27" pageId="0" pageNumber="27">
<subSubSection pageId="0" pageNumber="27" type="nomenclature">
<paragraph pageId="0" pageNumber="27">
Genus
<taxonomicName LSID="https://zoobank.org/4846A233-F2BF-4083-A2DD-0DEEA05247FB" authority="Yamada &amp; Yamamoto" authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ecpaglocoris" order="Hemiptera" pageId="0" pageNumber="27" rank="genus" status="gen. nov.">Ecpaglocoris Yamada &amp; Yamamoto</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="27">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="type species">
<paragraph pageId="0" pageNumber="27">Type species.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" kingdom="Animalia" lsidName="Ecpaglocoris ditomeus" order="Hemiptera" pageId="0" pageNumber="27" rank="species" species="ditomeus">
<emphasis italics="true" pageId="0" pageNumber="27">Ecpaglocoris ditomeus</emphasis>
</taxonomicName>
Yamada &amp; Yamamoto, sp. nov., by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="etymology">
<paragraph pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="27">
The genus name is a combination of the Greek ekpaglos (= wondrous, astounding) and koris (= bug), referring to this new fossil bug possessing unique male genitalia amongst
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="27" rank="superFamily" superFamily="Cimicoidea">Cimicoidea</taxonomicName>
; gender masculine.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
Body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1</figureCitation>
) elongated, dorsoventrally flattened; head (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3A</figureCitation>
) porrect; labium (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1D</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2B, C</figureCitation>
) reaching middle of mesosternum; pronotum (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1A, C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3A</figureCitation>
) with pair of long erect setae near antero-lateral and postero-lateral angles and a pair of similarly long setae behind anterior margin; pronotal callus flat, with longitudinal shallow groove on the mid-line; hemelytral membrane (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3D</figureCitation>
) with 10 or more long, slightly curved veins; a very long trichobothrium (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2E</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3D, E</figureCitation>
) present on middle of corium-membrane boundary; ostiolar peritreme (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2F</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3F</figureCitation>
) slightly curved forward at apex, distinctly continued to a fine carina which reaches anterior margin of metapleura; fore femur (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2G</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3G</figureCitation>
) extremely enlarged; fore tibia (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2G</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3G</figureCitation>
) strongly expanded towards apex, bearing 4-5 long, stout spines and 4-5 small teeth on ventral side and with greatly developed fossula spongiosa at apex; middle and hind tibiae (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1B, D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3H, I</figureCitation>
) with several long, stout spines, lacking fossula spongiosa; dorsal laterotergites (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2D</figureCitation>
) not fused with mediotergites on abdominal segments I to VIII; pygophore (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A, B. Apex of abdomen, dorsal (A) and left laterodorsal (B) views; C. Pygophore with parameres, dorsal view; D. Right paramere, right lateral view. Abbreviations: lpm, left paramere; pt, proctiger; py, pygophore; rpm, right paramere; tg 7 - 8, tergite 7 - 8. Scale bars: 0.1 mm." figureDoi="10.3897/fr.26.e86784.figure4" httpUri="https://binary.pensoft.net/fig/795263" pageId="0" pageNumber="27">4A-C</figureCitation>
) symmetrical, longer than combined length of abdominal sterna VII and VIII in ventral view, very broadly connected to abdominal segment VIII; proctiger well-developed; parameres (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A, B. Apex of abdomen, dorsal (A) and left laterodorsal (B) views; C. Pygophore with parameres, dorsal view; D. Right paramere, right lateral view. Abbreviations: lpm, left paramere; pt, proctiger; py, pygophore; rpm, right paramere; tg 7 - 8, tergite 7 - 8. Scale bars: 0.1 mm." figureDoi="10.3897/fr.26.e86784.figure4" httpUri="https://binary.pensoft.net/fig/795263" pageId="0" pageNumber="27">4</figureCitation>
) symmetrical, orientated anteriorly, very slender and long, moderately curved, acute towards apex, with groove running throughout the paramere.
</paragraph>
<caption doi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 1.</emphasis>
General habitus of
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" kingdom="Animalia" lsidName="Ecpaglocoris ditomeus" order="Hemiptera" pageId="0" pageNumber="27" rank="species" species="ditomeus">
<emphasis italics="true" pageId="0" pageNumber="27">Ecpaglocoris ditomeus</emphasis>
</taxonomicName>
gen. et sp. nov., male, holotype (AMNH Bu-SY33).
<emphasis bold="true" pageId="0" pageNumber="27">A.</emphasis>
Dorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">B.</emphasis>
Left laterodorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">C.</emphasis>
Ventral view;
<emphasis bold="true" pageId="0" pageNumber="27">D.</emphasis>
Right laterodorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="differential diagnosis">
<paragraph pageId="0" pageNumber="27">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
The new genus
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" kingdom="Animalia" lsidName="Ecpaglocoris" order="Hemiptera" pageId="0" pageNumber="27" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Ecpaglocoris</emphasis>
</taxonomicName>
differs from the genus
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
Tang, Wang &amp; Yao, 2022 by the head shorter than pronotum (in
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
, slightly longer than pronotum); vertex wider than twice the width of an eye in dorsal view (in
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
, narrower than the width of an eye); lateral margin of pronotum not membranous, only carinated at antero-lateral angle (in
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
, membranous, flattened and widely carinated); hemelytra parallel-sided (in
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
, strongly curved); PCu and R+M absent on hemelytra (in
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
, present); and fore tibia strongly expanded towards apex (in
<taxonomicName class="Insecta" family="Vetanthocoridae" genus="Pubivetanthocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pubivetanthocoris" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Pubivetanthocoris</emphasis>
</taxonomicName>
, slender, not expanded).
</paragraph>
<caption doi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 2.</emphasis>
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" kingdom="Animalia" lsidName="Ecpaglocoris ditomeus" order="Hemiptera" pageId="0" pageNumber="27" rank="species" species="ditomeus">
<emphasis italics="true" pageId="0" pageNumber="27">Ecpaglocoris ditomeus</emphasis>
</taxonomicName>
gen. et sp. nov., male, holotype (AMNH Bu-SY33).
<emphasis bold="true" pageId="0" pageNumber="27">A.</emphasis>
Right antenna, laterodorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">B, C.</emphasis>
Labium, ventral (
<emphasis bold="true" pageId="0" pageNumber="27">B</emphasis>
) and right lateral (
<emphasis bold="true" pageId="0" pageNumber="27">C</emphasis>
) views;
<emphasis bold="true" pageId="0" pageNumber="27">D.</emphasis>
Thorax and abdomen, right laterodorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">E.</emphasis>
Trichobothrium on corium-membrane boundary of right hemelytron;
<emphasis bold="true" pageId="0" pageNumber="27">F.</emphasis>
Ostiolar peritreme and evaporatorium, left lateroventral view;
<emphasis bold="true" pageId="0" pageNumber="27">G.</emphasis>
Right fore-leg, outer view. Abbreviations: a1-4, antennal segment 1-4; dlt1-6, dorsal laterotergite 1-6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls1-4. labial segment 1-4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="27">D</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="27">A-C, E-G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="description">
<paragraph pageId="0" pageNumber="27">Description.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Male.</emphasis>
Body elongated, dorsoventrally flattened.
</paragraph>
<paragraph pageId="0" pageNumber="27">
Head (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3A</figureCitation>
) porrect, slightly shorter than width across eyes; three pairs of long, erect trichobothria (cephalic macrosetae) on each side of anterior clypeus, near front margin of each eye and each side of vertex between eye and ocellus; anteocular region slightly longer than length of eye in dorsal view; eyes prominent, exceeding level of dorsal and ventral surfaces of head in lateral view; ocelli situated between eyes in front of an imaginary line that passes through posterior margin of eyes; vertex wider than twice the width of an eye in dorsal view; postocular region constricted, demarcated by transverse shallow furrow; neck long, smooth, highly polished. Antennal segment I (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3A</figureCitation>
) stout, exceeding apex of head, with a few short setae; prepedicellite present between segments I and II; segment II (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3B</figureCitation>
) stout, slightly thickened towards apex, about as long as head width across eyes, covered with suberect setae that are much shorter than width of the segment, intermixed with long setae that are longer than width of the segment; segments III and IV (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3B</figureCitation>
) filiform, much narrower than maximum width of segment II, equal in length, about 0.7 times as long as segment II, sparsely covered with long erect setae intermixed with short procumbent setae, longest setae much longer than twice the width of the respective segment. Labrum short, rounded at apex, not wholly covering labial segment I. Labium (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1D</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2B, C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3C</figureCitation>
) long, reaching middle of mesosternum, weakly curving; segment I visible, much shorter than the other segments; segment II stout, basally narrowed, approximately eye length long; segment III extremely long, slightly thickened near base and gradually narrowed towards apex, approximately 3.6 times as long as segment II; segment IV much slender, half as long as segment III.
</paragraph>
<caption doi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 3.</emphasis>
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" kingdom="Animalia" lsidName="Ecpaglocoris ditomeus" order="Hemiptera" pageId="0" pageNumber="27" rank="species" species="ditomeus">
<emphasis italics="true" pageId="0" pageNumber="27">Ecpaglocoris ditomeus</emphasis>
</taxonomicName>
gen. et sp. nov., male, holotype (AMNH Bu-SY33).
<emphasis bold="true" pageId="0" pageNumber="27">A.</emphasis>
Head and pronotum, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">B.</emphasis>
Right antenna, laterodorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">C.</emphasis>
Labium, lateral view;
<emphasis bold="true" pageId="0" pageNumber="27">D.</emphasis>
Left hemelytron, as seen in situ, not flattened;
<emphasis bold="true" pageId="0" pageNumber="27">E.</emphasis>
Trichobothrium on corium-membrane boundary of right hemelytron;
<emphasis bold="true" pageId="0" pageNumber="27">F.</emphasis>
Ostiolar peritreme and evaporatorium, left lateroventral view;
<emphasis bold="true" pageId="0" pageNumber="27">G.</emphasis>
Right fore-leg, outer view;
<emphasis bold="true" pageId="0" pageNumber="27">H.</emphasis>
Right middle leg, outer view;
<emphasis bold="true" pageId="0" pageNumber="27">I.</emphasis>
Right hind leg, inner view. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="27">D</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="27">A-C, F-I</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="27">
Pronotum (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1A, C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3A</figureCitation>
) nearly trapezoidal, shallowly depressed postero-medially, sparsely covered with tiny punctures, with pair of long erect setae near antero-lateral and postero-lateral angles and with pair of similarly long setae behind anterior margin; anterior margin slightly concave; lateral margin strongly angulate antero-laterally in dorsal view, densely covered with short setae; lateral carinae strongly expanded at antero-lateral angle; collar absent; callus flat, with longitudinal shallow groove on the mid-line; posterior margin deeply concave. Scutellum (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1A, B</figureCitation>
) large, sub-equilateral, mesal length longer than basal width, weakly depressed through middle, sparsely covered with short procumbent setae, with a pair of long erect setae near lateral margin base. Hemelytra (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3D</figureCitation>
) parallel-sided, surpassing apex of abdomen, overall covered with short procumbent setae; claval suture, medial fracture and costal fracture clearly visible; distal end of medial fracture not contiguous with costal fracture; costal margin slightly sinuate. Membrane (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3D</figureCitation>
) with a cross vein running along corium-membrane boundary and 10 or more long, slightly curved veins radiating posteriorly from it; stub (processus corial) present at distal end of a cross vein; a very long trichobothrium (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2E</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3D, E</figureCitation>
) present on middle of corium-membrane boundary, the length much longer than three times that of other setae on hemelytron. Mesosternum wide, coarse, mesally with longitudinal carina. Metasternum extremely swollen, obtuse at apex. Metepisternum wide, overall occupied by evaporatorium surrounding the metathoracic scent gland. Metathoracic scent gland (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2F</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3F</figureCitation>
) with a wide ostiolar peritreme; ostiolar peritreme curved slightly forward at apex, distinctly continued to a fine carina that reaches anterior margin of metapleura while gently curving; median furrow running throughout the ostiolar peritreme. Fore femur (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2G</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3G</figureCitation>
) extremely enlarged, unarmed, much thicker than width of middle femur; fore tibia (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2G</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3G</figureCitation>
) strongly expanded towards apex, bearing 4-5 long, stout spines and 4-5 small teeth on ventral side, with well-developed fossula spongiosa at apex; middle coxae widely separated from each other; middle tibia (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3H</figureCitation>
) expanded towards apex, covered with several long, stout spines on apical two-thirds and with tibial comb at apex, lacking fossula spongiosa; hind coxae proximate with each other; hind femur thickened, slightly narrower than width of fore femur, equal to hind tibia; hind tibia (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1B-D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Head and pronotum, dorsal view; B. Right antenna, laterodorsal view; C. Labium, lateral view; D. Left hemelytron, as seen in situ, not flattened; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view; H. Right middle leg, outer view; I. Right hind leg, inner view. Scale bars: 0.5 mm (D); 0.2 mm (A-C, F-I)." figureDoi="10.3897/fr.26.e86784.figure3" httpUri="https://binary.pensoft.net/fig/795262" pageId="0" pageNumber="27">3I</figureCitation>
) nearly cylindrical, densely covered with long, stout spines on apical three-fourths, with small tibial comb at apex, lacking fossula spongiosa. Tarsus three-segmented. Pretarsus of each leg with long, slender, symmetrical claws.
</paragraph>
<paragraph pageId="0" pageNumber="27">
Abdomen (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Dorsal view; B. Left laterodorsal view; C. Ventral view; D. Right laterodorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/fr.26.e86784.figure1" httpUri="https://binary.pensoft.net/fig/795260" pageId="0" pageNumber="27">1C, D</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2D</figureCitation>
) lateroventrally covered with dense short setae; lateral margins of sterna VI-VIII densely covered with short, suberect setae intermixed with thickly setae; dorsal laterotergites (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A. Right antenna, laterodorsal view; B, C. Labium, ventral (B) and right lateral (C) views; D. Thorax and abdomen, right laterodorsal view; E. Trichobothrium on corium-membrane boundary of right hemelytron; F. Ostiolar peritreme and evaporatorium, left lateroventral view; G. Right fore-leg, outer view. Abbreviations: a 1 - 4, antennal segment 1 - 4; dlt 1 - 6, dorsal laterotergite 1 - 6; ev, evaporatorium; fs, fossula spongiosa; lr, Labrum; ls 1 - 4. labial segment 1 - 4; op, ostiolar peritreme; tr, trichobothrium. Scale bars: 0.5 mm (D); 0.2 mm (A-C, E-G)." figureDoi="10.3897/fr.26.e86784.figure2" httpUri="https://binary.pensoft.net/fig/795261" pageId="0" pageNumber="27">2D</figureCitation>
) not fused with mediotergites on abdominal segments I to VIII; sterna II and III broad, and VIII very narrow.
</paragraph>
<paragraph pageId="0" pageNumber="27">
Male genitalia (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A, B. Apex of abdomen, dorsal (A) and left laterodorsal (B) views; C. Pygophore with parameres, dorsal view; D. Right paramere, right lateral view. Abbreviations: lpm, left paramere; pt, proctiger; py, pygophore; rpm, right paramere; tg 7 - 8, tergite 7 - 8. Scale bars: 0.1 mm." figureDoi="10.3897/fr.26.e86784.figure4" httpUri="https://binary.pensoft.net/fig/795263" pageId="0" pageNumber="27">4</figureCitation>
): Pygophore (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A, B. Apex of abdomen, dorsal (A) and left laterodorsal (B) views; C. Pygophore with parameres, dorsal view; D. Right paramere, right lateral view. Abbreviations: lpm, left paramere; pt, proctiger; py, pygophore; rpm, right paramere; tg 7 - 8, tergite 7 - 8. Scale bars: 0.1 mm." figureDoi="10.3897/fr.26.e86784.figure4" httpUri="https://binary.pensoft.net/fig/795263" pageId="0" pageNumber="27">4A-C</figureCitation>
) symmetrical, large, longer than combined length of abdominal sterna VII and VIII in ventral view, very broadly connected to abdominal segment VIII, dorsally depressed, posteriorly narrowed and rounded in dorsal view, densely covered with short, erect setae along outer margin and on posteroventral surface, lacking long trichobothria; proctiger well-developed; parameres (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ecpaglocoris ditomeus gen. et sp. nov., male, holotype (AMNH Bu-SY 33). A, B. Apex of abdomen, dorsal (A) and left laterodorsal (B) views; C. Pygophore with parameres, dorsal view; D. Right paramere, right lateral view. Abbreviations: lpm, left paramere; pt, proctiger; py, pygophore; rpm, right paramere; tg 7 - 8, tergite 7 - 8. Scale bars: 0.1 mm." figureDoi="10.3897/fr.26.e86784.figure4" httpUri="https://binary.pensoft.net/fig/795263" pageId="0" pageNumber="27">4</figureCitation>
) symmetrical, arising from near mid-line of dorsocaudal part of pygophore, orientated anteriorly, very slender and long, moderately curved, acute towards apex, with groove running throughout paramere.
</paragraph>
<caption doi="10.3897/fr.26.e86784.figure4" httpUri="https://binary.pensoft.net/fig/795263" pageId="0" pageNumber="27" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 4.</emphasis>
<taxonomicName authorityName="Yamada &amp; Yamamoto" authorityYear="2023" family="Vetanthocoridae" genus="Ecpaglocoris" kingdom="Animalia" lsidName="Ecpaglocoris ditomeus" order="Hemiptera" pageId="0" pageNumber="27" rank="species" species="ditomeus">
<emphasis italics="true" pageId="0" pageNumber="27">Ecpaglocoris ditomeus</emphasis>
</taxonomicName>
gen. et sp. nov., male, holotype (AMNH Bu-SY33).
<emphasis bold="true" pageId="0" pageNumber="27">A, B.</emphasis>
Apex of abdomen, dorsal (
<emphasis bold="true" pageId="0" pageNumber="27">A</emphasis>
) and left laterodorsal (
<emphasis bold="true" pageId="0" pageNumber="27">B</emphasis>
) views;
<emphasis bold="true" pageId="0" pageNumber="27">C.</emphasis>
Pygophore with parameres, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="27">D.</emphasis>
Right paramere, right lateral view. Abbreviations: lpm, left paramere; pt, proctiger; py, pygophore; rpm, right paramere; tg7-8, tergite 7-8. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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