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90 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1007.60012" ID-GBIF-Dataset="3d2f1c17-e64f-4d30-a1e1-b3db979f72fa" ID-PMC="PMC7788075" ID-Pensoft-Pub="1313-2970-1007-85" ID-Pensoft-UUID="0A16B87A3549522BB729306C3DA5FF88" ID-PubMed="33505182" ID-ZooBank="408A5B35D6054D90A46884D14E78AC3D" ModsDocID="1313-2970-1007-85" checkinTime="1609377148083" checkinUser="pensoft" docAuthor="Maddison, David R." docDate="2020" docId="440E52208B6159159AA47B17BBE8AAC8" docLanguage="en" docName="ZooKeys 1007: 85-128" docOrigin="ZooKeys 1007" docPubDate="2020-12-30" docSource="http://dx.doi.org/10.3897/zookeys.1007.60012" docTitle="Bembidion erosum" docType="treatment" docVersion="4" id="0A16B87A3549522BB729306C3DA5FF88" lastPageNumber="85" masterDocId="0A16B87A3549522BB729306C3DA5FF88" masterDocTitle="Shards, sequences, and shorelines: two new species of Bembidion from North America (Coleoptera, Carabidae)" masterLastPageNumber="128" masterPageNumber="85" pageNumber="85" updateTime="1668149658784" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Shards, sequences, and shorelines: two new species of Bembidion from North America (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maddison, David R.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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<mods:nameIdentifier type="email">david.maddison@science.oregonstate.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-12-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1007</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>85</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1007.60012</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1007.60012</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1007-85</mods:identifier>
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<mods:identifier type="ZooBank">408A5B35D6054D90A46884D14E78AC3D</mods:identifier>
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<mods:identifier type="Pensoft-UUID">0A16B87A3549522BB729306C3DA5FF88</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="176349025" LSID="urn:lsid:plazi:treatment:440E52208B6159159AA47B17BBE8AAC8" httpUri="http://treatment.plazi.org/id/440E52208B6159159AA47B17BBE8AAC8" lastPageNumber="85" pageId="0" pageNumber="85">
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<subSubSection pageId="0" pageNumber="85" type="nomenclature">
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<paragraph pageId="0" pageNumber="85">
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<taxonomicName LSID="440E5220-8B61-5915-9AA4-7B17BBE8AAC8" authority="(Motschulsky, 1850)" baseAuthorityName="Motschulsky" baseAuthorityYear="1850" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion erosum" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erosum">Bembidion erosum (Motschulsky, 1850)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="reference_group">
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<paragraph pageId="0" pageNumber="85">
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<taxonomicName authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Peryphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peryphus erosus" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erosus">Peryphus erosus</taxonomicName>
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Motschulsky, 1850:10. Lectotype female, in ZMUM, examined, designated by
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<bibRefCitation DOI="https://doi.org/10.4039/entm125167fv" author="Bousquet, Y" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="85" pagination="1 - 397" refId="B3" refString="Bousquet, Y, Larochelle, A, 1993. Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America north of Mexico. Memoirs of the Entomological Society of Canada 167: 1 - 397, DOI: https://doi.org/10.4039/entm125167fv" title="Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America north of Mexico." url="https://doi.org/10.4039/entm125167fv" volume="167" year="1993">Bousquet and Larochelle (1993)</bibRefCitation>
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, labeled
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<normalizedToken originalValue="“type”">"type"</normalizedToken>
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[handwritten],
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<normalizedToken originalValue="“California”">"California"</normalizedToken>
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[handwritten on red paper], "
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<taxonomicName authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Peryphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peryphus erosus" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erosus">Peryphus erosus</taxonomicName>
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Mots California" [handwritten on green paper], [a rectangle of blank red paper], "LECTOTYPE
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<taxonomicName authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Peryphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peryphus erosus" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erosus">Peryphus erosus</taxonomicName>
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Motschulsky Des. by Y. Bousquet
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<normalizedToken originalValue="’91”">'91"</normalizedToken>
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[partly handwritten on red paper]. Type locality California.
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</paragraph>
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<paragraph pageId="0" pageNumber="85">
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<taxonomicName authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion marinicum" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="marinicum">Bembidion marinicum</taxonomicName>
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Casey, 1918:57. Holotype female in USNM (type number 36919), examined. Type locality Marin County, California.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="diagnosis">
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<paragraph pageId="0" pageNumber="85">Diagnosis and geographic distribution.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Most adults of this species are the darkest members of this group (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Adult males of Bembidion transversale subgroup members A B. transversale (voucher V 101454, Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542 ° N, 113.7556 ° W) B B. erosum (voucher V 101453, USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051 ° N, 124.1005 ° W). C B. corgenoma (voucher V 101452, from type locality) D B. perspicuum, light form (voucher V 101461, USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239 ° N, 110.1722 ° W) E B. perspicuum, dark form (neotype of Bembidium haplogonum Chaudoir, USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881 ° N, 122.5400 ° W) F B. sarpedon (voucher V 101459, USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346 ° N, 104.6114 ° W). Scale bar 1.0 mm." figureDoi="10.3897/zookeys.1007.60012.figure11" httpUri="https://binary.pensoft.net/fig/492375" pageId="0" pageNumber="85">11B</figureCitation>
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), with the second and third antennomeres infuscated (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Antennae of B. transversale subgroup A B. transversale (voucher V 101457, Canada: Ontario: Thunder Bay Dist., Rossport) B B. erosum (voucher V 101456, USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051 ° N, 124.1005 ° W) C B. corgenoma (voucher V 101455, USA: Oregon: Coos Co., Crooked Creek S of Bandon, 43.0814 ° N, 124.4335 ° W). Scale bar 0.1 mm." figureDoi="10.3897/zookeys.1007.60012.figure12" httpUri="https://binary.pensoft.net/fig/492376" pageId="0" pageNumber="85">12B</figureCitation>
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), at least apically, and with dark femora, although the more northern populations (e.g., from mainland British Columbia) have paler legs and paler ground color of the body. Prothorax moderately cordate, with a smooth basal region with few punctures. Tip of aedeagus abruptly curved downward (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Male genitalia of B. transversale subgroup A B. transversale (voucher DNA 4219, USA: Oregon: Harney County, Banks of Silver Creek, 1379 m, 43.7278 ° N, 119.6256 ° W) B B. transversale (voucher DNA 2161, Canada: Alberta: Lethbridge, Oldman River, 800 m, 49.7043 ° N, 112.866 ° W) C B. erosum (voucher DNA 4033, USA: Oregon: Curry Co., Floras Creek at route 124 SE Langlois, 21 m, 42.9132 ° N, 124.4251 ° W) D B. erosum (voucher DNA 3562, USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051 ° N, 124.1005 ° W) E B. corgenoma (voucher DNA 2180, USA: California: Sonoma Co., Russian River, 3 mi NE Healdsburg) F B. corgenoma (holotype, voucher DNA 5673, USA: Oregon: Benton Co., Corvallis, Willamette River, 60 m, 44.5491 ° N, 123.2449 ° W). Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1007.60012.figure14" httpUri="https://binary.pensoft.net/fig/492378" pageId="0" pageNumber="85">14C, D</figureCitation>
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); basal sclerotized lobe large (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Basal sclerotized lobe of internal sac of male Bembidion transversale group members A B. transversale (voucher DNA 4219, USA: Oregon: Harney County, Banks of Silver Creek, 1379 m, 43.7278 ° N, 119.6256 ° W) B B. erosum (voucher DNA 4033, USA: Oregon: Curry Co., Floras Creek at route 124 SE Langlois, 21 m, 42.9132 ° N, 124.4251 ° W) C B. corgenoma (voucher DNA 2180, USA: California: Sonoma Co., Russian River, 3 mi NE Healdsburg). Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1007.60012.figure15" httpUri="https://binary.pensoft.net/fig/492379" pageId="0" pageNumber="85">15B</figureCitation>
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); apex of flagellar sheath with long, thin dark line (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Central sclerite complex of B. transversale subgroup A, B B. transversale (vouchers V 101437 and V 101436, USA: Michigan: Port aux Pins, Bois Blanc Isl.) C, D B. erosum (vouchers V 101440 and V 101439, USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051 ° N, 124.1005 ° W) E, F B. corgenoma (vouchers V 101428 and V 101430, USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5475 ° N, 123.2428 ° W). Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1007.60012.figure16" httpUri="https://binary.pensoft.net/fig/492380" pageId="0" pageNumber="85">16C, D</figureCitation>
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). This species is coastal, occurring from Haida Gwaii in British Columbia south along the coast to Big Sur and neighboring areas of California, with only one record from east of the Cascades (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Geographic distribution of B. erosum." figureDoi="10.3897/zookeys.1007.60012.figure20" httpUri="https://binary.pensoft.net/fig/492384" pageId="0" pageNumber="85">20</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |