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<document ID-DOI="http://doi.org/10.5281/zenodo.4756906" ID-GBIF-Dataset="87dcf476-dd02-4e6f-94f1-3f611d12ddd7" ID-GBIF-Taxon="182066746" ID-ISSN="1854-0392" ID-Zenodo-Dep="4756906" checkinTime="1620835414876" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2010" docId="2F298794934EFFF2FC47F9E51F9F558F" docLanguage="en" docName="Illiesia.6.24.303-314.pdf" docOrigin="Illiesia 6 (24)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Claassenia drukpa Stark &amp; Sivec 2010, sp. n." docType="treatment" docVersion="5" lastPageId="3" lastPageNumber="305" masterDocId="D310FFEC934EFFF1FF9AFFB71E315137" masterDocTitle="Systematic Notes On The Genus Claassenia Wu (Plecoptera: Perlidae), With Description Of A New Species" masterLastPageNumber="314" masterPageNumber="303" pageId="0" pageNumber="303" updateTime="1632015929349" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematic Notes On The Genus Claassenia Wu (Plecoptera: Perlidae), With Description Of A New Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>24</mods:number>
</mods:detail>
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<mods:start>303</mods:start>
<mods:end>314</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4753177</mods:identifier>
<mods:identifier type="GBIF-Dataset">87dcf476-dd02-4e6f-94f1-3f611d12ddd7</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4756906" ID-GBIF-Taxon="182066746" ID-Zenodo-Dep="4756906" LSID="urn:lsid:plazi:treatment:2F298794934EFFF2FC47F9E51F9F558F" httpUri="http://treatment.plazi.org/id/2F298794934EFFF2FC47F9E51F9F558F" lastPageId="3" lastPageNumber="305" pageId="0" pageNumber="303">
<subSubSection box="[989,1289,1618,1643]" pageId="0" pageNumber="303" type="nomenclature">
<paragraph blockId="0.[989,1289,1618,1643]" box="[989,1289,1618,1643]" pageId="0" pageNumber="303">
<heading bold="true" box="[989,1289,1618,1643]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="303" reason="2">
<emphasis bold="true" box="[989,1289,1618,1643]" pageId="0" pageNumber="303">
<taxonomicName authority="Stark &amp; Sivec, 2010" authorityName="Stark &amp; Sivec" authorityYear="2010" box="[989,1210,1618,1642]" class="Insecta" family="Perlidae" genus="Claassenia" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="drukpa" status="sp. n.">
<emphasis bold="true" box="[989,1210,1618,1642]" italics="true" pageId="0" pageNumber="303">Claassenia drukpa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1224,1289,1619,1643]" pageId="0" pageNumber="303" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1074,1204,1654,1679]" pageId="0" pageNumber="303" type="description">
<paragraph blockId="0.[1074,1204,1654,1679]" box="[1074,1204,1654,1679]" pageId="0" pageNumber="303">
(
<figureCitation box="[1083,1195,1654,1679]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="1.[189,241,1468,1492]" captionStartId-1="2.[189,241,1255,1279]" captionTargetBox-0="[306,1341,226,1447]" captionTargetBox-1="[257,1374,259,1165]" captionTargetId-0="figure-218@1.[287,1344,215,1453]" captionTargetId-1="figure-51@2.[249,1383,251,1175]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." captionText-1="Figs. 8-11. Claassenia drukpa egg. 8. Egg, lateral. 9. Collar end with anchor. 10. Collar end, apical aspect. 11. Chorionic detail." figureDoi-0="http://doi.org/10.5281/zenodo.4753183" figureDoi-1="http://doi.org/10.5281/zenodo.4753185" httpUri-0="https://zenodo.org/record/4753183/files/figure.png" httpUri-1="https://zenodo.org/record/4753185/files/figure.png" pageId="0" pageNumber="303">Figs. 1-11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="305" pageId="0" pageNumber="303" type="materials_examined">
<paragraph blockId="0.[835,1443,1725,1928]" pageId="0" pageNumber="303">
<emphasis bold="true" box="[835,1086,1726,1750]" pageId="0" pageNumber="303">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111164319" collectingDate="2006-04-24" collectionCode="PMSL" collectorName="H. Malicky" country="Bhutan" elevation="1370" latitude="27.483334" location="Dungkhar Rongchhu" longLatPrecision="1237" longitude="89.76667" pageId="0" pageNumber="303" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Punakha" typeStatus="holotype">
<typeStatus box="[1108,1220,1726,1750]" pageId="0" pageNumber="303">Holotype</typeStatus>
<specimenCount box="[1241,1261,1725,1750]" pageId="0" pageNumber="303" type="male"></specimenCount>
and
<specimenCount box="[1348,1382,1725,1750]" pageId="0" pageNumber="303" type="male">2♂</specimenCount>
,
<specimenCount box="[1410,1443,1725,1750]" pageId="0" pageNumber="303" type="female">1♀</specimenCount>
<typeStatus box="[835,951,1762,1786]" pageId="0" pageNumber="303">paratypes</typeStatus>
from
<collectingCountry box="[1065,1152,1762,1786]" name="Bhutan" pageId="0" pageNumber="303">Bhutan</collectingCountry>
,
<collectingRegion box="[1184,1290,1762,1786]" country="Bhutan" name="Punakha" pageId="0" pageNumber="303">Punakha</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2F298794934EFFF2FC47F9E51F9F558F:A25F6059934EFFF1FAB0F9551D8F562A" country="Bhutan" latitude="27.483334" longLatPrecision="1237" longitude="89.76667" name="Dungkhar Rongchhu" pageId="0" pageNumber="303" stateProvince="Punakha">Dungkhar Rongchhu</location>
,
<geoCoordinate box="[968,1074,1797,1821]" degrees="27" direction="north" minutes="29" orientation="latitude" pageId="0" pageNumber="303" precision="925" value="27.483334">27° 29 N</geoCoordinate>
,
<geoCoordinate box="[1084,1184,1797,1821]" degrees="89" direction="east" minutes="46" orientation="longitude" pageId="0" pageNumber="303" precision="925" value="89.76667">89° 46 E</geoCoordinate>
,
<quantity box="[1193,1277,1797,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="0" pageNumber="303" unit="m" value="1370.0">
<elevation box="[1193,1277,1797,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="0" pageNumber="303" unit="m" value="1370.0">1370 m</elevation>
</quantity>
,
<date box="[1286,1439,1797,1821]" pageId="0" pageNumber="303" value="2006-04-24">
<collectingDate box="[1286,1439,1797,1821]" pageId="0" pageNumber="303" value="2006-04-24">24 April 2006</collectingDate>
</date>
,
<collectorName box="[835,967,1833,1857]" pageId="0" pageNumber="303">H. Malicky</collectorName>
(
<collectionCode box="[981,1058,1833,1857]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="0" pageNumber="303">PMSL</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="0.[835,1443,1725,1928]" lastBlockId="1.[835,1443,1646,1919]" lastPageId="1" lastPageNumber="304" pageId="0" pageNumber="303">
<emphasis bold="true" box="[835,1018,1868,1892]" pageId="0" pageNumber="303">Adult habitus.</emphasis>
General color tan patterned with darker brown. Head with dark pigment in ocellar triangle invaded by pale pigment (
<figureCitation box="[616,683,1646,1670]" captionStart="Figs" captionStartId="1.[189,241,1468,1492]" captionTargetBox="[306,1341,226,1447]" captionTargetId="figure-218@1.[287,1344,215,1453]" captionTargetPageId="1" captionText="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="1" pageNumber="304">Fig. 1</figureCitation>
); lappets dark brown but most of head pale brown. Basal antennal segment brown, segments 2-3 dark brown and remainder of flagellum pale brown. Pronotum margined laterally, posteriorly, anteriorly and along median suture with dark brown pigment; disk pale brown. Femora mostly pale brown, tibiae dark at knee but pale brown over most of surface (
<figureCitation box="[714,783,1894,1918]" captionStart="Figs" captionStartId="1.[189,241,1468,1492]" captionTargetBox="[306,1341,226,1447]" captionTargetId="figure-218@1.[287,1344,215,1453]" captionTargetPageId="1" captionText="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="1" pageNumber="304">Fig. 5</figureCitation>
). Wings with pale brown tint.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4753183" ID-Zenodo-Dep="4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="1" pageNumber="304" startId="1.[189,241,1468,1492]" targetBox="[306,1341,226,1447]" targetPageId="1">
<paragraph blockId="1.[189,1444,1468,1563]" pageId="1" pageNumber="304">
Figs. 1-7.
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2010" box="[300,495,1468,1492]" class="Insecta" family="Perlidae" genus="Claassenia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="304" phylum="Arthropoda" rank="species" species="drukpa">
<emphasis box="[300,495,1468,1492]" italics="true" pageId="1" pageNumber="304">Claassenia drukpa</emphasis>
</taxonomicName>
structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca.
</paragraph>
</caption>
<paragraph blockId="1.[835,1443,1646,1919]" pageId="1" pageNumber="304">
<emphasis bold="true" box="[835,904,1681,1705]" pageId="1" pageNumber="304">Male.</emphasis>
Brachypterous, forewing length ca.
<quantity box="[1363,1439,1681,1705]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="304" unit="mm" value="5.0">5 mm</quantity>
. Hemitergal lobes short, finger-like, projecting upward in lateral aspect and slightly forward of median in dorsal aspect (
<figureCitation box="[1132,1235,1788,1812]" captionStart="Figs" captionStartId="1.[189,241,1468,1492]" captionTargetBox="[306,1341,226,1447]" captionTargetId="figure-218@1.[287,1344,215,1453]" captionTargetPageId="1" captionText="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="1" pageNumber="304">Figs. 2-4</figureCitation>
); apical third and dorsal margin of hemiterga dark brown. Tergum 9 bearing a median hump covered with a circular patch of small sensilla basiconica (
<figureCitation box="[1166,1234,1895,1919]" captionStart="Figs" captionStartId="1.[189,241,1468,1492]" captionTargetBox="[306,1341,226,1447]" captionTargetId="figure-218@1.[287,1344,215,1453]" captionTargetPageId="1" captionText="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="1" pageNumber="304">Fig. 2</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4753185" ID-Zenodo-Dep="4753185" httpUri="https://zenodo.org/record/4753185/files/figure.png" pageId="2" pageNumber="305" startId="2.[189,241,1255,1279]" targetBox="[257,1374,259,1165]" targetPageId="2">
<paragraph blockId="2.[189,1442,1255,1315]" pageId="2" pageNumber="305">
Figs. 8-11.
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2010" box="[315,511,1255,1279]" class="Insecta" family="Perlidae" genus="Claassenia" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="drukpa">
<emphasis box="[315,511,1255,1279]" italics="true" pageId="2" pageNumber="305">Claassenia drukpa</emphasis>
</taxonomicName>
egg. 8. Egg, lateral. 9. Collar end with anchor. 10. Collar end, apical aspect. 11. Chorionic detail.
</paragraph>
</caption>
<paragraph blockId="2.[189,797,1397,1920]" pageId="2" pageNumber="305">
<emphasis bold="true" box="[189,283,1398,1422]" pageId="2" pageNumber="305">Female.</emphasis>
Macropterous, forewing length ca.
<quantity box="[709,793,1397,1421]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="305" unit="mm" value="26.0">26 mm</quantity>
. Subgenital plate projects over base of sternum 9 as a short triangular process with shallow median notch on posterior margin (
<figureCitation box="[442,510,1504,1528]" captionStart="Figs" captionStartId="1.[189,241,1468,1492]" captionTargetBox="[306,1341,226,1447]" captionTargetId="figure-218@1.[287,1344,215,1453]" captionTargetPageId="1" captionText="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="2" pageNumber="305">Fig. 6</figureCitation>
). Vagina a membranous flattened bag with rectangular outline and short, partially fused accessory glands at anterior angles; anterior margin of vagina bearing a median sclerite and posterior half sparsely lined with fine setal spines (
<figureCitation box="[284,357,1682,1706]" captionStart="Figs" captionStartId="1.[189,241,1468,1492]" captionTargetBox="[306,1341,226,1447]" captionTargetId="figure-218@1.[287,1344,215,1453]" captionTargetPageId="1" captionText="Figs. 1-7. Claassenia drukpa structures. 1. Head and pronotum. 2. Male genitalia, dorsal. 3. Male hemitergum and cercal base, lateral. 4. Male hemitergum, lateral. 5. Foreleg. 6. Female abdominal sterna 8-9. 7. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4753183" httpUri="https://zenodo.org/record/4753183/files/figure.png" pageId="2" pageNumber="305">Fig. 7</figureCitation>
). Spermatheca swollen, balloon-like and attached to vagina by a short, slender stalk.
</paragraph>
<paragraph blockId="2.[189,797,1397,1920]" box="[189,396,1753,1777]" pageId="2" pageNumber="305">
<emphasis bold="true" box="[189,264,1753,1777]" pageId="2" pageNumber="305">Larva.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="2.[189,797,1397,1920]" lastBlockId="2.[835,1443,1397,1919]" pageId="2" pageNumber="305">
<emphasis bold="true" box="[189,241,1789,1813]" pageId="2" pageNumber="305">Egg.</emphasis>
Outline oval, length ca. 420 μm, equatorial width ca. 354 μm (
<figureCitation box="[403,469,1825,1849]" captionStart="Figs" captionStartId="2.[189,241,1255,1279]" captionTargetBox="[257,1374,259,1165]" captionTargetId="figure-51@2.[249,1383,251,1175]" captionTargetPageId="2" captionText="Figs. 8-11. Claassenia drukpa egg. 8. Egg, lateral. 9. Collar end with anchor. 10. Collar end, apical aspect. 11. Chorionic detail." figureDoi="http://doi.org/10.5281/zenodo.4753185" httpUri="https://zenodo.org/record/4753185/files/figure.png" pageId="2" pageNumber="305">Fig. 8</figureCitation>
). Collar button- like, set on a raised, disc-like platform ca. 100 μm in diameter; margins irregularly crenulate and bearing short, obscure vertical struts (
<figureCitation box="[1104,1215,1397,1422]" captionStart="Figs" captionStartId="2.[189,241,1255,1279]" captionTargetBox="[257,1374,259,1165]" captionTargetId="figure-51@2.[249,1383,251,1175]" captionTargetPageId="2" captionText="Figs. 8-11. Claassenia drukpa egg. 8. Egg, lateral. 9. Collar end with anchor. 10. Collar end, apical aspect. 11. Chorionic detail." figureDoi="http://doi.org/10.5281/zenodo.4753185" httpUri="https://zenodo.org/record/4753185/files/figure.png" pageId="2" pageNumber="305">Figs. 8-10</figureCitation>
); anchor a relatively plain membranous bell-shaped structure with short cylindrical pedicel (
<figureCitation box="[1100,1183,1468,1493]" captionStart="Figs" captionStartId="2.[189,241,1255,1279]" captionTargetBox="[257,1374,259,1165]" captionTargetId="figure-51@2.[249,1383,251,1175]" captionTargetPageId="2" captionText="Figs. 8-11. Claassenia drukpa egg. 8. Egg, lateral. 9. Collar end with anchor. 10. Collar end, apical aspect. 11. Chorionic detail." figureDoi="http://doi.org/10.5281/zenodo.4753185" httpUri="https://zenodo.org/record/4753185/files/figure.png" pageId="2" pageNumber="305">Fig. 9</figureCitation>
). Chorionic surface punctate throughout, but mesal band including mixed larger, regularly spaced punctations and smaller irregularly spaced ones (
<figureCitation box="[1213,1288,1575,1599]" captionStart="Figs" captionStartId="2.[189,241,1255,1279]" captionTargetBox="[257,1374,259,1165]" captionTargetId="figure-51@2.[249,1383,251,1175]" captionTargetPageId="2" captionText="Figs. 8-11. Claassenia drukpa egg. 8. Egg, lateral. 9. Collar end with anchor. 10. Collar end, apical aspect. 11. Chorionic detail." figureDoi="http://doi.org/10.5281/zenodo.4753185" httpUri="https://zenodo.org/record/4753185/files/figure.png" pageId="2" pageNumber="305">Fig. 11</figureCitation>
); punctations on anterior pole shallow, fine and barely discernable at 500X. Micropylar row subequatorial, set ca. 135 μm from anterior end.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="305" type="etymology">
<paragraph blockId="2.[835,1443,1397,1919]" pageId="2" pageNumber="305">
<emphasis bold="true" box="[835,973,1717,1741]" pageId="2" pageNumber="305">Etymology.</emphasis>
The species name, used as a noun in apposition, is based on the Bhutanese word for dragon people, and honors the people and history of
<collectingCountry box="[835,923,1824,1848]" name="Bhutan" pageId="2" pageNumber="305">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="306" pageId="2" pageNumber="305" type="diagnosis">
<paragraph blockId="2.[835,1443,1397,1919]" lastBlockId="3.[189,797,1077,1208]" lastPageId="3" lastPageNumber="306" pageId="2" pageNumber="305">
<emphasis bold="true" box="[835,964,1859,1883]" pageId="2" pageNumber="305">Diagnosis.</emphasis>
Males of this species are distinguished by the short, upwardly directed hemiterga and females by the triangular, projecting and apically notched subgenital plate. This species is the only known
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2010" box="[189,302,1148,1172]" class="Insecta" family="Perlidae" genus="Claassenia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis box="[189,302,1148,1172]" italics="true" pageId="3" pageNumber="306">Claassenia</emphasis>
</taxonomicName>
in which the egg form varies from the usual spindle shape.
</paragraph>
</subSubSection>
</treatment>
</document>