treatments-xml/data/03/FE/87/03FE87F14C5C7F0B26D2BACB0432FDC4.xml

<document id="4D4F77FE8B58F16740A55AE8E1CF1FC3" ID-DOI="10.11646/zootaxa.5293.3.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="7961217" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1684825370997" checkinUser="plazi" docAuthor="Zhang, Yong-Qin &amp; Yin, Zi-Wei" docDate="2023" docId="03FE87F14C5C7F0B26D2BACB0432FDC4" docLanguage="en" docName="zootaxa.5293.3.11.pdf" docOrigin="Zootaxa 5293 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.3.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrodes tibialis Zhang &amp; Yin 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="599" masterDocId="FFC7FF894C5D7F082645BA700551FF9B" masterDocTitle="A new species of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from China" masterLastPageNumber="599" masterPageNumber="596" pageNumber="597" updateTime="1684825878875" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="EB2D4EBF9538CB807E783F651FFD6AF1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EFA141E270B6A2D1E3BC4A5A7668250D">
<mods:title id="63CBA89EEED0F805507C32C7195134B0">A new species of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from China</mods:title>
</mods:titleInfo>
<mods:name id="F0CE30BFCAE0BA3760DE12C176152606" type="personal">
<mods:role id="8C41BDDB3CCDC14C393F78BAE0B80DC8">
<mods:roleTerm id="3496E6EA1BD644922847E4CD6BEFDFEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1774CA55EA4F2DCFE5FBA0B74606C1C0">Zhang, Yong-Qin</mods:namePart>
<mods:nameIdentifier id="084127B428C35203472A4C6F1F555C67" type="ORCID">0009-0004-6112-6640</mods:nameIdentifier>
<mods:affiliation id="012D8DE1DD48AA8E14BB0C74510D19F3">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, Shanghai 200234, China &amp; 18772757263 @ 163. com; https: // orcid. org / 0009 - 0004 - 6112 - 6640</mods:affiliation>
</mods:name>
<mods:name id="859BF30951FCA28556B90254B599849B" type="personal">
<mods:role id="20B63BE5ABF26D50CB259AFB65DFB4C3">
<mods:roleTerm id="E7730E48221C24F770E849CFD912D7CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D97BCAA749D91FAE077A7AAC36F05EED">Yin, Zi-Wei</mods:namePart>
<mods:nameIdentifier id="3898AF4D254C857623F9E039068E5291" type="ORCID">0000-0001-6659-9448</mods:nameIdentifier>
<mods:affiliation id="72ADF7565D98AFFE04E9C739C61A9BE8">pselaphinae @ gmail. com; https: // orcid. org / 0000 - 0001 - 6659 - 9448</mods:affiliation>
<mods:nameIdentifier id="06613EB059A848099DC9C2C4622CD3AB" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="DEEA93BAB02C65076C21D892F58F4A5A">text</mods:typeOfResource>
<mods:relatedItem id="FFB86636C9CE2CA787E34FC2B49E8B57" type="host">
<mods:titleInfo id="FCE6F2AA66F89191091D67C769A191A6">
<mods:title id="A9A6E077228BD98C4D5A886271C2424C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9A1E8825F96684C3DF46EEDBBC7A8583">
<mods:date id="9264BEBB54B008E5C35E77DBE9AC853E">2023</mods:date>
<mods:detail id="C0891229132A945BB4C9E89244B02CB5" type="pubDate">
<mods:number id="ABF01A06940BB6CF6D9E0516051001FB">2023-05-23</mods:number>
</mods:detail>
<mods:detail id="1851DF3CD42E5DB56E3BD528EFC1D7CD" type="volume">
<mods:number id="B72C645A44840CA6D76BB3BE2E0BA31C">5293</mods:number>
</mods:detail>
<mods:detail id="668F07A71F229AA7D37E2909F173F508" type="issue">
<mods:number id="E872CF30A65906B3BA9C19D246573747">3</mods:number>
</mods:detail>
<mods:extent id="7207AE67E7AA273296E68A704915CE37" unit="page">
<mods:start id="2EBF7C1D52871120ED3B0E9FCBE0B107">596</mods:start>
<mods:end id="D6F19ED2B137150BB7A903065DA9DDC6">599</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C6536CB8D84E194A4EF1D8378CBC4C3A">
<mods:url id="92BCDBECAF82E9ED988E0A6B31EDA7CF">http://dx.doi.org/10.11646/zootaxa.5293.3.11</mods:url>
</mods:location>
<mods:classification id="FFDC66A60CE1D0632F3BDC558B87260C">journal article</mods:classification>
<mods:identifier id="50DDF67939A565CB63E28EBAF1494445" type="DOI">10.11646/zootaxa.5293.3.11</mods:identifier>
<mods:identifier id="652440E3DE301C6BE56199BCF5281492" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C19C3EB880E36DCE95AF5FFCD6A60732" type="Zenodo-Dep">7961217</mods:identifier>
</mods:mods>
<treatment id="03FE87F14C5C7F0B26D2BACB0432FDC4" ID-DOI="http://doi.org/10.5281/zenodo.7961204" ID-Zenodo-Dep="7961204" LSID="urn:lsid:plazi:treatment:03FE87F14C5C7F0B26D2BACB0432FDC4" httpUri="http://treatment.plazi.org/id/03FE87F14C5C7F0B26D2BACB0432FDC4" lastPageId="3" lastPageNumber="599" pageId="1" pageNumber="597">
<subSubSection id="C34D656C4C5C7F0926D2BACB04E5FF4E" box="[151,436,187,214]" pageId="1" pageNumber="597" type="nomenclature">
<paragraph id="8BE836E74C5C7F0926D2BACB04E5FF4E" blockId="1.[151,1437,151,2013]" box="[151,436,187,214]" pageId="1" pageNumber="597">
<heading id="D0A0818B4C5C7F0926D2BACB04E5FF4E" bold="true" box="[151,436,187,214]" fontSize="11" level="1" pageId="1" pageNumber="597" reason="1">
<emphasis id="B923EAF54C5C7F0926D2BACB04E5FF4E" bold="true" box="[151,436,187,214]" pageId="1" pageNumber="597">
<taxonomicName id="4C574D644C5C7F0926D2BACB0401FF4D" authority="Zhang &amp; Yin, 2023" authorityName="Zhang &amp; Yin" authorityYear="2023" box="[151,336,187,214]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B923EAF54C5C7F0926D2BACB0401FF4D" bold="true" box="[151,336,187,214]" italics="true" pageId="1" pageNumber="597">Tyrodes tibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210578E4C5C7F092713BACB04E5FF4E" box="[342,436,187,213]" pageId="1" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D656C4C5C7F0926D2BA91070AFE80" pageId="1" pageNumber="597" type="description">
<paragraph id="8BE836E74C5C7F0926D2BA9105B9FF62" blockId="1.[151,1437,151,2013]" box="[151,232,225,249]" pageId="1" pageNumber="597">
(
<figureCitation id="136C2A624C5C7F0926DABA91058CFF62" box="[159,221,225,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="1" pageNumber="597">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8BE836E74C5C7F0926D2BB75070AFE80" blockId="1.[151,1437,151,2013]" box="[151,603,261,285]" pageId="1" pageNumber="597">
Chinese common name: 
<emphasis id="B923EAF54C5C7F0927E5BB75070AFE80" bold="true" box="[416,603,261,283]" pageId="1" pageNumber="597">ẊṘṁLjaeẘffl</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34D656C4C5C7F092682BB59073BFE4A" pageId="1" pageNumber="597" type="materials_examined">
<paragraph id="8BE836E74C5C7F092682BB59073BFE4A" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">
<emphasis id="B923EAF54C5C7F092682BB590498FEDA" box="[199,457,297,321]" inLineHeading="true" pageId="1" pageNumber="597" reason="1">
<emphasis id="B923EAF54C5C7F092682BB590437FEDA" bold="true" box="[199,358,297,321]" pageId="1" pageNumber="597">Type material</emphasis>
(3 exx.).
</emphasis>
<materialsCitation id="3B3F3CBA4C5C7F092795BB5901B9FEFE" collectingDate="2021-04-23" collectionCode="UMAE, SNUC" collectorName="Huaping N. R. &amp; Yin &amp; Zhang &amp; Pan &amp; Shen" country="China" elevation="13560" latitude="25.562433" location="Guilin City" longLatPrecision="1" longitude="109.93808" pageId="1" pageNumber="597" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B923EAF54C5C7F092795BB590788FEDA" bold="true" box="[464,729,297,321]" pageId="1" pageNumber="597">
<typeStatus id="54EC88454C5C7F092795BB59073AFEDA" box="[464,619,297,321]" pageId="1" pageNumber="597">HOLOTYPE</typeStatus>
: 
<collectingCountry id="F34076774C5C7F092432BB590785FEDA" box="[631,724,297,321]" name="China" pageId="1" pageNumber="597">CHINA</collectingCountry>
:
</emphasis>
♁: ‘China: 
<collectingRegion id="4993F8054C5C7F09251EBB5906EBFEDA" box="[859,954,297,321]" country="China" name="Guangxi" pageId="1" pageNumber="597">Guangxi</collectingRegion>
, 
<location id="8E88603C4C5C7F092581BB59016EFEDA" LSID="urn:lsid:plazi:treatment:03FE87F14C5C7F0B26D2BACB0432FDC4:8E88603C4C5C7F092581BB59016EFEDA" box="[964,1087,297,321]" country="China" latitude="25.562433" longLatPrecision="1" longitude="109.93808" name="Guilin City" pageId="1" pageNumber="597" stateProvince="Guangxi">Guilin City</location>
, 
<collectorName id="26A253314C5C7F09220FBB5901B8FEDB" box="[1098,1257,297,321]" pageId="1" pageNumber="597">
Huaping 
<taxonomicNameLabel id="A210578E4C5C7F0922E8BB5901B8FEDB" box="[1197,1257,297,320]" pageId="1" pageNumber="597" rank="subSpecies">N. R.</taxonomicNameLabel>
</collectorName>
, 
<geoCoordinate id="EE6350204C5C7F0922B0BB5700C9FEDB" box="[1269,1432,295,320]" degrees="25" direction="north" minutes="33" orientation="latitude" pageId="1" pageNumber="597" precision="1" seconds="44.76" value="25.562433">25°33′44.76″N</geoCoordinate>
, 
<geoCoordinate id="EE6350204C5C7F0926D2BB3B0415FEFF" box="[151,324,331,356]" degrees="109" direction="east" minutes="56" orientation="longitude" pageId="1" pageNumber="597" precision="1" seconds="17.08" value="109.93808">109°56′17.08″E</geoCoordinate>
, 
<quantity id="4CAF9B024C5C7F092710BB3D04EAFEFF" box="[341,443,333,357]" metricMagnitude="4" metricUnit="m" metricValue="1.3559999999999999" pageId="1" pageNumber="597" unit="m" value="13560.0">
<elevation id="007AD1D44C5C7F092710BB3D04EAFEFF" box="[341,443,333,357]" metricMagnitude="4" metricUnit="m" metricValue="1.3559999999999999" pageId="1" pageNumber="597" unit="m" value="13560.0">13560 m</elevation>
</quantity>
, 
<date id="FFE910274C5C7F092789BB3D076EFEFE" box="[460,575,333,357]" pageId="1" pageNumber="597" value="2021-04-23">
<collectingDate id="EFADE9CF4C5C7F092789BB3D076EFEFE" box="[460,575,333,357]" pageId="1" pageNumber="597" value="2021-04-23">23.iv.2021</collectingDate>
</date>
, 
<collectorName id="26A253314C5C7F09240ABB3D0729FEFE" box="[591,632,333,357]" pageId="1" pageNumber="597">Yin</collectorName>
, 
<collectorName id="26A253314C5C7F0924CDBB3D079EFEFE" box="[648,719,333,357]" pageId="1" pageNumber="597">Zhang</collectorName>
, 
<collectorName id="26A253314C5C7F0924A5BB3D0659FEFF" box="[736,776,333,356]" pageId="1" pageNumber="597">Pan</collectorName>
&amp; 
<collectorName id="26A253314C5C7F092573BB3D063AFEFE" box="[822,875,333,357]" pageId="1" pageNumber="597">Shen</collectorName>
leg. [
<emphasis id="B923EAF54C5C7F0925F1BB3D013EFEF8" bold="true" box="[948,1135,333,355]" pageId="1" pageNumber="597">
<collectionCode id="ED46AE224C5C7F0925F1BB3D0142FEF8" box="[948,1043,333,355]" pageId="1" pageNumber="597">ŨMAE</collectionCode>
ḢỄḾő
</emphasis>
]’ (
<collectionCode id="ED46AE224C5C7F0922D6BB3D01B1FEFE" box="[1171,1248,333,357]" pageId="1" pageNumber="597">SNUC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B3F3CBA4C5C7F0922B2BB3D0737FE4A" collectingDate="2021-04-21" collectingDateMax="2021-04-24" collectingDateMin="2021-04-21" collectionCode="SNUC" collectorName="Tang &amp; Peng &amp; Cai &amp; Song" country="China" county="Huanjiang" elevation="450" latitude="25.199821" location="Qinshuitang" longLatPrecision="1" longitude="108.796265" municipality="Jiuwan Mt" pageId="1" pageNumber="597" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="B923EAF54C5C7F0922B2BB3D05A8FE12" bold="true" pageId="1" pageNumber="597">
<typeStatus id="54EC88454C5C7F0922B2BB3D00C6FEFE" box="[1271,1431,333,357]" pageId="1" pageNumber="597">PARATYPES</typeStatus>
: 
<collectingCountry id="F34076774C5C7F0926D2BB0105A5FE12" box="[151,244,369,393]" name="China" pageId="1" pageNumber="597">CHINA</collectingCountry>
:
</emphasis>
1 ♁, same collection data as for holotype, except ‘ 
<date id="FFE910274C5C7F092570BB01069EFE12" box="[821,975,369,393]" pageId="1" pageNumber="597" value="2021-04-21" valueMax="2021-04-22" valueMin="2021-04-21">
<collectingDate id="EFADE9CF4C5C7F092570BB01069EFE12" box="[821,975,369,393]" pageId="1" pageNumber="597" value="2021-04-21" valueMax="2021-04-22" valueMin="2021-04-21">21-22.iv.2021</collectingDate>
</date>
, FIT (
<collectingMethod id="52164EF04C5C7F092252BB0101A5FE12" box="[1047,1268,369,393]" pageId="1" pageNumber="597">flight intercept trap</collectingMethod>
)’; 1 ♁, ‘China: 
<collectingRegion id="4993F8054C5C7F0926D2BBE505A7FE36" box="[151,246,405,429]" country="China" name="Guangxi" pageId="1" pageNumber="597">Guangxi</collectingRegion>
, 
<collectingCounty id="62894E6B4C5C7F0926BABBE50420FE36" box="[255,369,405,429]" pageId="1" pageNumber="597">Huanjiang</collectingCounty>
, 
<collectingMunicipality id="6B8CAC9D4C5C7F09273FBBE504BCFE37" box="[378,493,405,429]" pageId="1" pageNumber="597">Jiuwan Mt</collectingMunicipality>
, 
<location id="8E88603C4C5C7F0927B2BBE5072DFE36" LSID="urn:lsid:plazi:treatment:03FE87F14C5C7F0B26D2BACB0432FDC4:8E88603C4C5C7F0927B2BBE5072DFE36" box="[503,636,405,429]" country="China" county="Huanjiang" latitude="25.199821" longLatPrecision="1" longitude="108.796265" municipality="Jiuwan Mt" name="Qinshuitang" pageId="1" pageNumber="597" stateProvince="Guangxi">Qinshuitang</location>
, 
<geoCoordinate id="EE6350204C5C7F0924C0BBE30676FE37" box="[645,807,403,428]" degrees="25" direction="north" minutes="11" orientation="latitude" pageId="1" pageNumber="597" precision="1" seconds="59.36" value="25.199821">25°11′59.36″N</geoCoordinate>
, 
<geoCoordinate id="EE6350204C5C7F092574BBE3068FFE37" box="[817,990,403,428]" degrees="108" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="597" precision="1" seconds="46.55" value="108.796265">108°47′46.55″E</geoCoordinate>
, 
<quantity id="4CAF9B024C5C7F0925A2BBE5017DFE37" box="[999,1068,405,429]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="597" unit="m" value="450.0">
<elevation id="007AD1D44C5C7F0925A2BBE5017DFE37" box="[999,1068,405,429]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="597" unit="m" value="450.0">450 m</elevation>
</quantity>
, 
<date id="FFE910274C5C7F092273BBE501F9FE36" box="[1078,1192,405,429]" pageId="1" pageNumber="597" value="2021-04-24">
<collectingDate id="EFADE9CF4C5C7F092273BBE501F9FE36" box="[1078,1192,405,429]" pageId="1" pageNumber="597" value="2021-04-24">24.iv.2021</collectingDate>
</date>
, 
<collectorName id="26A253314C5C7F0922F7BBE501BBFE36" box="[1202,1258,405,429]" pageId="1" pageNumber="597">Tang</collectorName>
, 
<collectorName id="26A253314C5C7F0922B6BBE5007AFE36" box="[1267,1323,405,429]" pageId="1" pageNumber="597">Peng</collectorName>
, 
<collectorName id="26A253314C5C7F092371BBE5000DFE36" box="[1332,1372,405,429]" pageId="1" pageNumber="597">Cai</collectorName>
, 
<collectorName id="26A253314C5C7F092320BBE500CDFE36" box="[1381,1436,405,429]" pageId="1" pageNumber="597">Song</collectorName>
leg. [
<emphasis id="B923EAF54C5C7F09268BBBC904A5FE54" bold="true" box="[206,500,441,463]" pageId="1" pageNumber="597">ŨMhvhƜNjứRffiḵae</emphasis>
]’ (
<collectionCode id="ED46AE224C5C7F092454BBC9070FFE4A" box="[529,606,441,465]" pageId="1" pageNumber="597">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D656C4C5C7F092682BBAD0187FD56" pageId="1" pageNumber="597" type="diagnosis">
<paragraph id="8BE836E74C5C7F092682BBAD0187FD56" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">
<emphasis id="B923EAF54C5C7F092682BBAD046DFE6E" bold="true" box="[199,316,477,501]" inLineHeading="true" pageId="1" pageNumber="597" reason="1">Diagnosis.</emphasis>
<emphasis id="B923EAF54C5C7F092703BBAD042CFE6E" box="[326,381,477,501]" italics="true" pageId="1" pageNumber="597">Male</emphasis>
. Body length approximately 
<quantity id="4CAF9B024C5C7F092486BBAD061BFE6F" box="[707,842,477,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="1" pageNumber="597" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
. color reddish-brown, dorsal surface of body covered with relatively long pubescence. Head truncate at base, with punctiform vertexal and frontal fovea. Antenna relatively elongate, antennomere 1 (scape) angularly expanded at base of lateral margin, distinct club formed by apical three enlarged antennomeres. Pronotum with punctiform median and small lateral antebasal foveae connected by transverse sulcus. Discal striae of elytra shallow and short. Mesal margin of metatibia with small, short preapical projection. Sternite 3 (V) with lateral portions broadly concaved. Aedeagus dorso-ventrally asymmetric, relatively elongate, median lobe with lateral expansion at apical 1/3, endophallus armature with three elongate sclerites. 
<emphasis id="B923EAF54C5C7F09224DB8C50109FD56" box="[1032,1112,693,717]" italics="true" pageId="1" pageNumber="597">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34D656C4C5C7F0B2682B8A90756FE68" lastPageId="3" lastPageNumber="599" pageId="1" pageNumber="597" type="description">
<paragraph id="8BE836E74C5C7F092682B8A90786FC8E" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">
<emphasis id="B923EAF54C5C7F092682B8A9041AFD6A" bold="true" box="[199,331,729,753]" pageId="1" pageNumber="597">Description</emphasis>
<emphasis id="B923EAF54C5C7F092717B8A904D8FD6A" box="[338,393,729,753]" italics="true" pageId="1" pageNumber="597">Male</emphasis>
. Body (
<figureCitation id="136C2A624C5C7F0927A5B8A90765FD6A" box="[480,564,729,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="1" pageNumber="597">Fig. 1A</figureCitation>
) length 
<quantity id="4CAF9B024C5C7F0924CBB8A9067CFD6B" box="[654,813,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" metricValueMax="2.08" metricValueMin="1.86" pageId="1" pageNumber="597" unit="mm" value="1.97" valueMax="2.08" valueMin="1.86">1.86–2.08 mm</quantity>
; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with dense, long pubescence.
</paragraph>
<paragraph id="8BE836E74C5C7F092682B9510457FB5E" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">
Head (
<figureCitation id="136C2A624C5C7F09274AB9510431FCA2" box="[271,352,801,825]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="1" pageNumber="597">Fig. 1B</figureCitation>
) truncate at base, approximately as long as wide, length 
<quantity id="4CAF9B024C5C7F092581B9510133FCA3" box="[964,1122,801,825]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="1" pageNumber="597" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.40–0.43 mm</quantity>
, width across eyes 
<quantity id="4CAF9B024C5C7F092377B9510595FCC7" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.3" metricValueMin="4.1" pageId="1" pageNumber="597" unit="mm" value="0.42" valueMax="0.43" valueMin="0.41">0.41–0.43 mm</quantity>
; vertex with small, asetose, and moderately separated foveae (dorsal tentorial pits); tempora short, rounded, densely setose; frons roughly punctate admesally, anteriorly forming distinct rostrum, with punctiform frontal fovea, antennal tubercles weakly convex, cavities broad; clypeus connecting with frons by distinct median ridge, with smooth surface, its anterior margin moderately carinate and moderately raised; ocular-mandibular carinae absent, ocular canthi moderately convex. Venter with smooth surface; tiny gular foveae (posterior tentorial pits) in shared transverse opening, lacking median carina or sulcus. Compound eyes prominent, each composed of 45 ommatidia. Maxillary palpus (
<figureCitation id="136C2A624C5C7F09234FB989000DFB8A" box="[1290,1372,1017,1041]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="1" pageNumber="597">Fig. 1C</figureCitation>
) with palpomere 1 small, 2 basally pedunculate and broadened apically, 3 pedunculate at base for short distance and then broadened toward apex, 4 largest, with short stem at base, suboval, with long palpal cone at apex. Antenna relatively elongate, length 
<quantity id="4CAF9B024C5C7F092702BE1504B5FBE7" box="[327,484,1125,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" metricValueMax="1.37" metricValueMin="1.29" pageId="1" pageNumber="597" unit="mm" value="1.33" valueMax="1.37" valueMin="1.29">1.29–1.37 mm</quantity>
, antennomere 1 (scape) with coarse surface, angularly expanded at base of lateral margin, 2–8 sub-moniliform, 9–11 each enlarged and forming distinct club, 11 much shorter than 9 and 10 combined (
<quantity id="4CAF9B024C5C7F092368BEF900DCFB3B" box="[1325,1421,1161,1185]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="597" unit="mm" value="0.28">0.28 mm</quantity>
: 
<quantity id="4CAF9B024C5C7F0926D2BEDD05AAFB5E" box="[151,251,1197,1221]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="597" unit="mm" value="0.38">0.38 mm</quantity>
).
</paragraph>
<paragraph id="8BE836E74C5C7F092682BEA10634FA06" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">
Pronotum (
<figureCitation id="136C2A624C5C7F09277ABEA104C1FB72" box="[319,400,1233,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="1" pageNumber="597">Fig. 1B</figureCitation>
) slightly broader than long, length 
<quantity id="4CAF9B024C5C7F09254CBEA106F7FB73" box="[777,934,1233,1257]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="1" pageNumber="597" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
, width 
<quantity id="4CAF9B024C5C7F0925B1BEA101C0FB73" box="[1012,1169,1233,1257]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="1" pageNumber="597" unit="mm" value="0.43" valueMax="0.45" valueMin="0.41">0.41–0.45 mm</quantity>
, widest approximately at middle; lateral margins subparallel in basal half, convergent anteriorly from middle toward anterior margin, both anterior and posterior margins slightly arched; disc moderately convex, finely punctate and densely setose, with small, asetose median and lateral antebasal foveae connected by thin but distinct transverse sulcus, lacking carina or spine. Prosternum with anterior part slightly shorter than coxal part, with distinct, setose and widely separated procoxal foveae; hypomera fused with sternum, smooth; margin of coxal cavity thinly carinate.
</paragraph>
<paragraph id="8BE836E74C5C7F092682BFD9014AF9B6" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">
Elytra with long pubescence, much wider than long, length 
<quantity id="4CAF9B024C5C7F09250FBFD906B9FA5B" box="[842,1000,1449,1473]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="6.0" metricValueMin="5.2" pageId="1" pageNumber="597" unit="mm" value="0.56" valueMax="0.6" valueMin="0.52">0.52–0.60 mm</quantity>
, width 
<quantity id="4CAF9B024C5C7F092272BFD90187FA5B" box="[1079,1238,1449,1473]" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="9.1" metricValueMin="8.0" pageId="1" pageNumber="597" unit="mm" value="0.855" valueMax="0.91" valueMin="0.8">0.80–0.91 mm</quantity>
; each elytron with two moderately large, asetose basal foveae; discal striae short and shallow, extending posteriorly from outer basal fovea to approximately 3.9/10 of elytral length; humeral denticle absent, humeri rounded, lacking subhumeral fovea or marginal stria; posterior margin with row of dense setae. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8BE836E74C5C7F092682BC490452F97A" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="597">Mesoventrite short, laterally fused with metaventrite; median mesoventral foveae moderately separated, originating from shared setose, transverse opening, large lateral mesoventral foveae broadly forked internally, with short, apically truncate mesoventral process. Metaventrite at middle with large, blunt projection before posterior margin, with large, setose lateral mesocoxal foveae and one median metaventral fovea, posterior metaventral process with small emargination at middle.</paragraph>
<paragraph id="8BE836E74C5C7F092682BC9D0051F89E" blockId="1.[151,1437,151,2013]" box="[199,1280,1773,1797]" pageId="1" pageNumber="597">
Legs moderately elongate, mesal margin of metatibia (
<figureCitation id="136C2A624C5C7F092548BC9D060AF89E" box="[781,859,1773,1797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="1" pageNumber="597">Fig. 1F</figureCitation>
) with small, short preapical projection.
</paragraph>
<paragraph id="8BE836E74C5C7F0A2682BD610468FF48" blockId="1.[151,1437,151,2013]" lastBlockId="2.[151,1436,151,355]" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597">
Abdomen widest at lateral margins of tergite 1 (IV), length 
<quantity id="4CAF9B024C5C7F092524BD610150F8B3" box="[865,1025,1809,1833]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" metricValueMax="7.1" metricValueMin="6.1" pageId="1" pageNumber="597" unit="mm" value="0.6599999999999999" valueMax="0.71" valueMin="0.61">0.61–0.71 mm</quantity>
, width 
<quantity id="4CAF9B024C5C7F092210BD6101A6F8B3" box="[1109,1271,1809,1833]" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="8.7" metricValueMin="7.4" pageId="1" pageNumber="597" unit="mm" value="0.8049999999999999" valueMax="0.87" valueMin="0.74">0.74–0.87 mm</quantity>
. Tergite 1 (IV) longest, 1.5× as long as 2 (V), with broad, setose basal sulcus and two pairs of basolateral foveae, lacking discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, 3 shorter than 2 and 4; tergite 5 (VIII) semicircular, transverse, posterior margin broadly and shallowly emarginate at middle. Sternite 2 (IV) at middle as long as 3–6 (V– VIII), approximately 3.5× as long as 3, with deep, setose basal impression and pair of basolateral sockets, lateral portions of sternite 3 broadly concaved, 4 short at middle, lacking foveae, 5 at middle longer than 4, with pair of small basolateral foveae, 6 transverse, roundly emarginate at middle, 7 (IX) (
<figureCitation id="136C2A624C5F7F0A255CBAE7063DFF2B" box="[793,876,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="2" pageNumber="598">Fig. 1G</figureCitation>
) semi-sclerotized, subtriangular, apical margin with scattered setae.
</paragraph>
<paragraph id="8BE836E74C5F7F0A2682BAAF0498FEF8" blockId="2.[151,1436,151,355]" pageId="2" pageNumber="598">
Aedeagus (
<figureCitation id="136C2A624C5F7F0A277BBAAF04F6FF63" box="[318,423,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="2" pageNumber="598">Fig. 1H, I</figureCitation>
) 
<quantity id="4CAF9B024C5F7F0A27F0BA900743FF6C" box="[437,530,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="598" unit="mm" value="0.38">0.38 mm</quantity>
long, dorso-ventrally asymmetric, moderately elongate; median lobe with large basal capsule and round diaphragm, narrowing from base toward apex, with large expansion at left side (orientation according to 
<figureCitation id="136C2A624C5F7F0A26F7BB5705A9FEDB" box="[178,248,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1933,1957]" captionTargetBox="[151,1436,374,1904]" captionTargetId="figure-90@2.[151,1436,374,1904]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes tibialis sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I." figureDoi="http://doi.org/10.5281/zenodo.7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="2" pageNumber="598">Fig. 1I</figureCitation>
), apex with several small spines and one larger hook, parameres short, roundly triangular, endophallus armature with three elongate sclerites.
</paragraph>
<caption id="DF28666F4C5F7F0A26D2BDFD06EFF876" ID-DOI="http://doi.org/10.5281/zenodo.7961221" ID-Zenodo-Dep="7961221" httpUri="https://zenodo.org/record/7961221/files/figure.png" pageId="2" pageNumber="598" startId="2.[151,250,1933,1957]" targetBox="[151,1436,374,1904]" targetPageId="2" targetType="figure">
<paragraph id="8BE836E74C5F7F0A26D2BDFD06EFF876" blockId="2.[151,1436,1933,2030]" pageId="2" pageNumber="598">
<emphasis id="B923EAF54C5F7F0A26D2BDFD045EF83E" bold="true" box="[151,271,1933,1957]" pageId="2" pageNumber="598">FIGURE 1</emphasis>
. Morphology of 
<taxonomicName id="4C574D644C5F7F0A2787BDFD070EF83E" authority="Zhang &amp; Yin, 2023" authorityName="Zhang &amp; Yin" authorityYear="2023" box="[450,607,1933,1957]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B923EAF54C5F7F0A2787BDFD070EF83E" box="[450,607,1933,1957]" italics="true" pageId="2" pageNumber="598">Tyrodes tibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210578E4C5F7F0A242DBDFE07EBF83E" box="[616,698,1934,1957]" pageId="2" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
, male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Maxillary palpus. D. Scape. E. Antennal club. F. Apical portion of metatibia. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and dorsal (I). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, F, G; 0.2 mm in H. I.
</paragraph>
</caption>
<paragraph id="8BE836E74C5E7F0B2682BAE704C4FF34" blockId="3.[151,1437,151,607]" box="[199,405,151,175]" pageId="3" pageNumber="599">
<emphasis id="B923EAF54C5E7F0B2682BAE7044EFF34" box="[199,287,151,175]" italics="true" pageId="3" pageNumber="599">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BE836E74C5E7F0B2682BACB066FFE30" blockId="3.[151,1437,151,607]" pageId="3" pageNumber="599">
<emphasis id="B923EAF54C5E7F0B2682BACB04F9FF4F" bold="true" box="[199,424,187,212]" inLineHeading="true" pageId="3" pageNumber="599" reason="1">Comparative notes.</emphasis>
This species is distinct within the genus in having modified metatibiae, which bear a small preapical projection on the mesal margin. The presence of a lateral expansion of the aedeagal median lobe is shared by 
<taxonomicName id="4C574D644C5E7F0B26D2BB74049BFE87" authority="(Schaufuss, L. W.)" baseAuthorityName="Schaufuss, L. W." box="[151,458,259,284]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="B923EAF54C5E7F0B26D2BB7405F7FE80" box="[151,166,260,283]" italics="true" pageId="3" pageNumber="599">T</emphasis>
. 
<emphasis id="B923EAF54C5E7F0B26F1BB7305ABFE80" box="[180,250,259,283]" italics="true" pageId="3" pageNumber="599">histrio</emphasis>
(Schaufuss, L. W.)
</taxonomicName>
from 
<collectingCountry id="F34076774C5E7F0B2448BB730726FE80" box="[525,631,259,283]" name="Sri Lanka" pageId="3" pageNumber="599">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFC64B164C5E7F0B24C3BB730649FE80" author="Jeannel, R." box="[646,792,259,283]" pageId="3" pageNumber="599" pagination="5 - 32" refId="ref2285" refString="Jeannel, R. (1957) Sur quelques Pselaphides du Tonkin recueillis par le Pere A. de Cooman. Revue Francaise d'Entomologie, 24 (1), 5 - 32." type="journal article" year="1957">Jeannel 1957</bibRefCitation>
: fig. 29). These two species can be readily separated by the larger body size (
<quantity id="4CAF9B024C5E7F0B2716BB5804E4FEA4" box="[339,437,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" pageId="3" pageNumber="599" unit="mm" value="2.08">2.08 mm</quantity>
vs. 
<quantity id="4CAF9B024C5E7F0B27A1BB57076BFEDB" box="[484,570,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="599" unit="mm" value="1.5">1.5 mm</quantity>
), relatively much more elongate male antennal clubs, much narrower elytra and abdomen in relation to head and pronotum, and a different structure of the endophallus armature of 
<taxonomicName id="4C574D644C5E7F0B228FBB3C0060FEF8" authorityName="Zhang &amp; Yin" authorityYear="2023" box="[1226,1329,331,355]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B923EAF54C5E7F0B228FBB3C0188FEF8" box="[1226,1241,332,355]" italics="true" pageId="3" pageNumber="599">T</emphasis>
. 
<emphasis id="B923EAF54C5E7F0B22A2BB3B0060FEF8" box="[1255,1329,331,355]" italics="true" pageId="3" pageNumber="599">tibialis</emphasis>
</taxonomicName>
. The new species clearly differs from the only other Chinese congener, 
<taxonomicName id="4C574D644C5E7F0B2559BB000682FE1C" authority="Yin &amp; Li" authorityName="Yin &amp; Li" box="[796,979,367,392]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="species" species="jenisi">
<emphasis id="B923EAF54C5E7F0B2559BB00067AFE1C" box="[796,811,368,391]" italics="true" pageId="3" pageNumber="599">T</emphasis>
. 
<emphasis id="B923EAF54C5E7F0B2572BB000620FE1C" box="[823,881,368,391]" italics="true" pageId="3" pageNumber="599">jenisi</emphasis>
Yin &amp; Li
</taxonomicName>
from Yunnan (
<bibRefCitation id="EFC64B164C5E7F0B222ABB1F005FFE13" author="Yin, Z-W &amp; Li, L. - Z." box="[1135,1294,367,392]" pageId="3" pageNumber="599" pagination="101 - 105" refId="ref2516" refString="Yin, Z-W &amp; Li, L. - Z. (2013) First record of Tyrodes Raffray (Coleoptera, Staphylinidae, Pselaphinae) in China, with description of T. jenisi sp. n. from Yunnan Province. ZooKeys, 301, 101 - 105. https: // doi. org / 10.3897 / zookeys. 301.4911" type="journal article" year="2013">Yin &amp; Li 2013</bibRefCitation>
), by the more elongate antennal clubs, as well as by the form of the aedeagus.
</paragraph>
<paragraph id="8BE836E74C5E7F0B2682BBC70756FE68" blockId="3.[151,1437,151,607]" pageId="3" pageNumber="599">
<emphasis id="B923EAF54C5E7F0B2682BBC70414FE54" bold="true" box="[199,325,439,463]" inLineHeading="true" pageId="3" pageNumber="599" reason="1">Bionomics.</emphasis>
Two individuals were found from sifted leaf litter samples in broad-leaved forests, while one was collected by a flight intercept trap.
</paragraph>
</subSubSection>
<subSubSection id="C34D656C4C5E7F0B2682BB8F07F8FD8C" box="[199,681,511,535]" pageId="3" pageNumber="599" type="distribution">
<paragraph id="8BE836E74C5E7F0B2682BB8F07F8FD8C" blockId="3.[151,1437,151,607]" box="[199,681,511,535]" pageId="3" pageNumber="599">
<emphasis id="B923EAF54C5E7F0B2682BB8F040BFD8C" bold="true" box="[199,346,511,535]" inLineHeading="true" pageId="3" pageNumber="599" reason="1">Distribution.</emphasis>
Southwestern 
<collectingCountry id="F34076774C5E7F0B27BDBB8F076DFD8C" box="[504,572,511,535]" name="China" pageId="3" pageNumber="599">China</collectingCountry>
: 
<collectingRegion id="4993F8054C5E7F0B2400BB8F07F5FD8C" box="[581,676,511,535]" country="China" name="Guangxi" pageId="3" pageNumber="599">Guangxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D656C4C5E7F0B2682B8540432FDC4" pageId="3" pageNumber="599" type="etymology">
<paragraph id="8BE836E74C5E7F0B2682B8540432FDC4" blockId="3.[151,1437,151,607]" pageId="3" pageNumber="599">
<emphasis id="B923EAF54C5E7F0B2682B8540419FDA7" bold="true" box="[199,328,548,572]" inLineHeading="true" pageId="3" pageNumber="599" reason="1">Etymology.</emphasis>
The new specific epithet, 
<taxonomicName id="4C574D644C5E7F0B2412B85307F0FDA0" authorityName="Zhang &amp; Yin" authorityYear="2023" box="[599,673,547,571]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B923EAF54C5E7F0B2412B85307F0FDA0" box="[599,673,547,571]" italics="true" pageId="3" pageNumber="599">tībiālis</emphasis>
</taxonomicName>
, is a Latin adjective, meaning, ‘tibial’, referring to the tibial modification of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>