168 lines
20 KiB
XML
168 lines
20 KiB
XML
<document id="CB80008ACD450B1F54D020CABEA1B886" ID-CLB-Dataset="308416" ID-DOI="10.11646/zootaxa.5590.4.4" ID-GBIF-Dataset="5dcdcbb9-5269-4e93-b06c-ecc53d8e5aff" ID-ISSN="1175-5326" ID-Zenodo-Dep="14953515" ID-ZooBank="2AC35A62-6004-44C5-9D4D-33F6E410C087" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1740851019834" checkinUser="plazi" docAuthor="Belokobylskij, Sergey A. & Manukyan, Andranik R." docDate="2025" docId="03FE8795FF93FFC3FF29FC7F67A960E8" docLanguage="en" docName="zootaxa.5590.4.4.pdf" docOrigin="Zootaxa 5590 (4)" docSource="https://doi.org/10.11646/zootaxa.5590.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aulosaphes Muesebeck 1935" docType="treatment" docVersion="6" lastPageNumber="539" masterDocId="FFC7FFEDFF94FFCBFFBEFFDB65366161" masterDocTitle="New fossil species from two genera of the subfamily Hormiinae (Hymenoptera: Braconidae) from Eocene Baltic amber" masterLastPageNumber="544" masterPageNumber="531" pageNumber="538" updateTime="1741175249156" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B54E6045F9F2D660DEBA6B6240F1AE8F">New fossil species from two genera of the subfamily Hormiinae (Hymenoptera: Braconidae) from Eocene Baltic amber</mods:title>
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<mods:namePart id="2FC59DCDE9AB510A3B021585489F3D78">Belokobylskij, Sergey A.</mods:namePart>
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<mods:affiliation id="58758DA2C09B5452A3B860D8FBBC1FB9">Zoological Institute of the Russian Academy of Sciences, St Petersburg 199034, Russia</mods:affiliation>
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<mods:namePart id="2AA8BBBE45DF589FDC49F0C68A6E1BD2">Manukyan, Andranik R.</mods:namePart>
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<mods:affiliation id="2FDF266FF129FDCC33CF00EFAF61E383">Kaliningrad Amber Museum, Marshal Vasilevskiy Square, 1, Kaliningrad 236016, Russia</mods:affiliation>
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<mods:date id="1BCC7F72DFFF5B42A23B29428FBEACA4">2025</mods:date>
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<treatment id="03FE8795FF93FFC3FF29FC7F67A960E8" ID-DOI="http://doi.org/10.5281/zenodo.14973952" ID-GBIF-Taxon="255165495" ID-Zenodo-Dep="14973952" LSID="urn:lsid:plazi:treatment:03FE8795FF93FFC3FF29FC7F67A960E8" httpUri="http://treatment.plazi.org/id/03FE8795FF93FFC3FF29FC7F67A960E8" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538" scope_class="Insecta" scope_family="Braconidae" scope_order="Hymenoptera" scope_subFamily="Hormiinae" scope_tribe="Lysitermini">
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<emphasis id="B923EA91FF93FFCCFF29FC7F676E62DF" bold="true" box="[151,600,931,958]" pageId="7" pageNumber="538">
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Genus
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<taxonomicName id="4C574D00FF93FFCCFF50FC78676E62DF" ID-CoL="35T3" authority="Muesebeck, 1935" authorityName="Muesebeck" authorityYear="1935" box="[238,600,931,958]" class="Insecta" family="Braconidae" genus="Aulosaphes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EA91FF93FFCCFF50FC78644C62DF" bold="true" box="[238,378,931,958]" italics="true" pageId="7" pageNumber="538">Aulosaphes</emphasis>
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<bibRefCitation id="EFC64B72FF93FFCCFE3CFC7F676E62DF" author="Muesebeck, C. F. W." box="[386,600,932,958]" pageId="7" pageNumber="538" pagination="241 - 250" refId="ref7475" refString="Muesebeck, C. F. W. (1935) On the genus Oncophanes Foerster, with descriptions of two new related genera (Hymenoptera: Braconidae). Annals of the Entomological Society of America, 28, 241-250. https://doi.org/10.1093/aesa/28.2.241" type="journal article" year="1935">Muesebeck, 1935</bibRefCitation>
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</taxonomicName>
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</emphasis>
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<subSubSection id="C34D6508FF93FFCCFF29FC3364B9657E" pageId="7" pageNumber="538" type="reference_group">
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<treatmentCitation id="0AF61092FF93FFCCFF29FC3364CB6560" author="Muesebeck, C. F. W." box="[151,509,1000,1025]" page="248" pageId="7" pageNumber="538" year="1935">
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<taxonomicName id="4C574D00FF93FFCCFF29FC3364CB6560" ID-CoL="35T3" authority="Muesebeck, 1935: 248" authorityName="Muesebeck" authorityPageNumber="248" authorityYear="1935" box="[151,509,1000,1025]" class="Insecta" family="Braconidae" genus="Aulosaphes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EA91FF93FFCCFF29FC33643A6561" box="[151,268,1000,1024]" italics="true" pageId="7" pageNumber="538">Aulosaphes</emphasis>
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<bibRefCitation id="EFC64B72FF93FFCCFEADFC3364CB6560" author="Muesebeck, C. F. W." box="[275,509,1000,1025]" pageId="7" pageNumber="538" pagination="241 - 250" refId="ref7475" refString="Muesebeck, C. F. W. (1935) On the genus Oncophanes Foerster, with descriptions of two new related genera (Hymenoptera: Braconidae). Annals of the Entomological Society of America, 28, 241-250. https://doi.org/10.1093/aesa/28.2.241" type="journal article" year="1935">Muesebeck, 1935: 248</bibRefCitation>
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;
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<treatmentCitation id="0AF61092FF93FFCCFDB6FC3367D36560" author="Shenefelt, R. D." box="[520,741,1000,1025]" page="1140" pageId="7" pageNumber="538" year="1975">
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<bibRefCitation id="EFC64B72FF93FFCCFDB6FC3367D36560" author="Shenefelt, R. D." box="[520,741,1000,1025]" pageId="7" pageNumber="538" pagination="1115 - 1262" refId="ref7575" refString="Shenefelt, R. D. (1975) Braconidae 8. Exothecinae, Rogadinae. Hymenopterorum Catalogus, Nova Editio, Pars 12, 1115-1262" type="journal article" year="1975">Shenefelt 1975: 1140</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF61092FF93FFCCFD51FC3266D96560" author="van Achterberg, C." box="[751,1007,1000,1025]" page="80" pageId="7" pageNumber="538" year="1995">
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<bibRefCitation id="EFC64B72FF93FFCCFD51FC3266D96560" author="van Achterberg, C." box="[751,1007,1000,1025]" pageId="7" pageNumber="538" pagination="3 - 242" refId="ref7634" refString="van Achterberg, C. (1995) Generic revision of the subfamily Betylobraconinae (Hymenoptera: Braconidae) and other groups with modified fore tarsus. Zoologische Verhandelingen, 298, 3-242." type="journal article" year="1995">van Achterberg 1995: 80</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF61092FF93FFCCFC44FC3361CF6560" author="Belokobylskij, S. A." box="[1018,1273,1000,1025]" page="134" pageId="7" pageNumber="538" year="1998">
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<bibRefCitation id="EFC64B72FF93FFCCFC44FC3361CF6560" author="Belokobylskij, S. A." box="[1018,1273,1000,1025]" pageId="7" pageNumber="538" pagination="111 - 159" refId="ref6807" refString="Belokobylskij, S. A. (1998) Subfamily Exothecinae. In: Lehr, P. A. (Ed.), Key to the insects of the Russian Far East. Vol. 4. Neuropteroidea, Mecoptera, Hymenoptera. Pt 3. Dal'nauka, Vladivostok, pp. 111-159. [in Russian]" type="book chapter" year="1998">Belokobylskij 1998: 134</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF61092FF93FFCCFABAFC3265C6657E" author="He, J. H. & Chen, X. X. & Ma, Y." page="675" pageId="7" pageNumber="538" year="2000">
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<bibRefCitation id="EFC64B72FF93FFCCFABAFC3265C6657E" author="He, J. H. & Chen, X. X. & Ma, Y." pageId="7" pageNumber="538" refId="ref7240" refString="He, J. H., Chen, X. X. & Ma, Y. (2000) Hymenoptera, Braconidae. Fauna Sinica. Insecta. Vol. 18. Science Press, Beijing, 757 pp. [in Chinese with English summary]" type="book" year="2000">
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He
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||
<emphasis id="B923EA91FF93FFCCFA96FC32606A6561" box="[1320,1372,1000,1024]" italics="true" pageId="7" pageNumber="538">et al.</emphasis>
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2000: 675
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</treatmentCitation>
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;
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||
<bibRefCitation id="EFC64B72FF93FFCCFF44FBD364BD657E" author="Yu, D. S. & van Achterberg, C. & Horstmann, K." box="[250,395,1031,1055]" pageId="7" pageNumber="538" refId="ref7670" refString="Yu, D. S., van Achterberg, C. & Horstmann, K. (2016) Taxapad 2016. Ichneumonoidea 2015. Nepean, Ottawa, Ontario. [database on flash-drive]" type="book" year="2016">
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||
Yu
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||
<emphasis id="B923EA91FF93FFCCFEA2FBD36479657E" box="[284,335,1031,1055]" italics="true" pageId="7" pageNumber="538">et al.</emphasis>
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||
2016
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||
</bibRefCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34D6508FF93FFCCFF29FB9E67B76438" pageId="7" pageNumber="538" type="type_taxon">
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<paragraph id="8BE83683FF93FFCCFF29FB9E67A4653D" blockId="7.[151,658,1092,1117]" box="[151,658,1092,1117]" pageId="7" pageNumber="538">
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||
<emphasis id="B923EA91FF93FFCCFF29FB9E6410653D" bold="true" box="[151,294,1092,1117]" pageId="7" pageNumber="538">
|
||
<typeStatus id="54EC8821FF93FFCCFF29FB9E65FA653C" box="[151,204,1093,1117]" pageId="7" pageNumber="538">Type</typeStatus>
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species:
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</emphasis>
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||
<taxonomicName id="4C574D00FF93FFCCFE92FB9F67A4653D" authority="Ashmead, 1905" authorityName="Ashmead" authorityYear="1905" box="[300,658,1092,1116]" class="Insecta" family="Braconidae" genus="Rhyssalus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="unicolor">
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||
<emphasis id="B923EA91FF93FFCCFE92FB9F64D8653D" box="[300,494,1092,1116]" italics="true" pageId="7" pageNumber="538">Rhyssalus unicolor</emphasis>
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Ashmead, 1905
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</taxonomicName>
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||
</paragraph>
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||
<paragraph id="8BE83683FF93FFCCFF29FB50616D658C" blockId="7.[151,1437,1163,2017]" pageId="7" pageNumber="538">
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||
<emphasis id="B923EA91FF93FFCCFF29FB5065EC65C4" bold="true" box="[151,218,1163,1189]" pageId="7" pageNumber="538">Notes</emphasis>
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||
.
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<taxonomicName id="4C574D00FF93FFCCFF59FB5064D965C4" ID-CoL="35T3" authority="Muesebeck" authorityName="Muesebeck" authorityYear="1935" box="[231,495,1163,1189]" class="Insecta" family="Braconidae" genus="Aulosaphes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B923EA91FF93FFCCFF59FB50645E65C4" box="[231,360,1163,1189]" italics="true" pageId="7" pageNumber="538">Aulosaphes</emphasis>
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Muesebeck
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</taxonomicName>
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is relatively small extant genus with only ten known tropical and subtropical species from the hormiine tribe
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<taxonomicName id="4C574D00FF93FFCCFE1CFB7467EA65A8" authority="(Yu et al. 2016)" baseAuthorityName="Yu" baseAuthorityYear="2016" box="[418,732,1199,1225]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="tribe" tribe="Lysitermini">
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Lysitermini (
|
||
<bibRefCitation id="EFC64B72FF93FFCCFD8FFB7467E265A8" author="Yu, D. S. & van Achterberg, C. & Horstmann, K." box="[561,724,1199,1225]" pageId="7" pageNumber="538" refId="ref7670" refString="Yu, D. S., van Achterberg, C. & Horstmann, K. (2016) Taxapad 2016. Ichneumonoidea 2015. Nepean, Ottawa, Ontario. [database on flash-drive]" type="book" year="2016">
|
||
Yu
|
||
<emphasis id="B923EA91FF93FFCCFDE7FB7467A765A8" box="[601,657,1199,1225]" italics="true" pageId="7" pageNumber="538">et al.</emphasis>
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||
2016
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||
</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
. The members of this tribe have never been recorded as fossils, so our find as a Eocene amber inclusion is the first record of a fossil from this genus..
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||
</paragraph>
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<paragraph id="8BE83683FF93FFCCFF79FB2C67B76438" blockId="7.[151,1437,1163,2017]" pageId="7" pageNumber="538">
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||
The present amber specimen has a number of morphological differences from the
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<typeStatus id="54EC8821FF93FFCCFBC5FB2C619D6470" box="[1147,1195,1271,1297]" pageId="7" pageNumber="538">type</typeStatus>
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||
species of the genus
|
||
<emphasis id="B923EA91FF93FFCCFF29FAC064296454" box="[151,287,1307,1333]" italics="true" pageId="7" pageNumber="538">
|
||
<taxonomicName id="4C574D00FF93FFCCFF29FAC0642D6454" ID-CoL="35T3" authorityName="Muesebeck" authorityYear="1935" box="[151,283,1307,1333]" class="Insecta" family="Braconidae" genus="Aulosaphes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">Aulosaphes</taxonomicName>
|
||
,
|
||
</emphasis>
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||
<taxonomicName id="4C574D00FF93FFCCFE96FAC0678B6454" authority="Ashmead, 1905" authorityName="Ashmead" authorityYear="1905" box="[296,701,1306,1333]" class="Insecta" family="Braconidae" genus="Rhyssalus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B923EA91FF93FFCCFE96FAC067366454" box="[296,512,1307,1333]" italics="true" pageId="7" pageNumber="538">Rhyssalus unicolor</emphasis>
|
||
Ashmead, 1905
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||
</taxonomicName>
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||
, so the new species described here is placed in a new subgenus,
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||
<emphasis id="B923EA91FF93FFCCFF29FAE4640A6438" box="[151,316,1343,1369]" italics="true" pageId="7" pageNumber="538">Antiquasaphes</emphasis>
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||
<taxonomicName id="4C574D00FF93FFCCFEFDFAE464D56438" authorityName="Belokobylskij & Manukyan" authorityYear="2025" box="[323,483,1343,1369]" pageId="7" pageNumber="538" rank="subGenus" status="subgen. nov." subGenus="Belokobylskij">Belokobylskij</taxonomicName>
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||
,
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||
<emphasis id="B923EA91FF93FFCCFE50FAE467B76438" bold="true" box="[494,641,1343,1369]" pageId="7" pageNumber="538">
|
||
<taxonomicNameLabel id="A21057EAFF93FFCCFE50FAE467B76438" box="[494,641,1343,1369]" pageId="7" pageNumber="538" rank="subGenus">subgen. nov.</taxonomicNameLabel>
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||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D6508FF93FFC3FF79FAB967A960E8" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538" type="diagnosis">
|
||
<paragraph id="8BE83683FF93FFCCFF79FAB967DC6750" blockId="7.[151,1437,1163,2017]" pageId="7" pageNumber="538">
|
||
<emphasis id="B923EA91FF93FFCCFF79FAB964EB641C" bold="true" box="[199,477,1378,1405]" pageId="7" pageNumber="538">Diagnosis of the genus</emphasis>
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||
. Head transverse, rather strongly narrowed posteriorly (dorsal view). Ocelli often arranged in almost equilateral triangle. Eyes glabrous. Hypoclypeal depression large and circular or suboval. Malar suture usually distinct. Occipital carina complete, fused ventrally with hypostomal carina upper base of mandible. Maxillary palpi 6-segmented, labial palpi 4-segmented. Mandible bidentate, but second tooth sometimes small. Antenna relatively slender, filiform or weakly setiform. Pedicel enlarged. First flagellomere not shorter than second flagellomere; apical flagellomere acuminate apically.
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</paragraph>
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<paragraph id="8BE83683FF93FFCCFF79F9E067866784" blockId="7.[151,1437,1163,2017]" pageId="7" pageNumber="538">Mesosoma not or only weakly depressed dorso-ventrally. Pronotum anteriorly with flange. Mesonotum highly and curvedly elevated above pronotum. Notauli complete, deep, crenulate. Medial lobe of mesoscutum sometimes with medioposterior longitudinal sulcus. Precoxal sulcus deep at least medially, medial length, usually straight and at least partly crenulate. Prepectal carina distinct and complete. Propodeum without lateral tubercles, with areas distinctly delineated by carinae, areola variable.</paragraph>
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<paragraph id="8BE83683FF93FFCCFF79F93467286680" blockId="7.[151,1437,1163,2017]" pageId="7" pageNumber="538">Wings. Pterostigma rather wide. Radial (marginal) cell of fore wing not shortened. Radial vein (r) arising from, weakly before or weakly behind middle of pterostigma. Both radiomedial veins (2-SR and r-m) present. Recurrent vein (m-cu) distinctly postfurcal to first radiomedial vein (2-SR). Nervulus (cu-a) postfurcal. Parallel vein (CU1a) arising from about middle of distal side (3-CU1) of brachial (subdiscal) cell. Brachial (subdiscal) cell closed distally. Transverse anal veins (2A and a) absent. In hind wing, first abscissa of mediocubital vein (M+CU) weakly shorter than or almost equal to second abscissa (1-M). Submedial (subbasal) cell rather large. Recurrent vein (m-cu) present, curved, faintly pigmented.</paragraph>
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<paragraph id="8BE83683FF9CFFC3FF79FF4C663061B4" blockId="8.[151,1436,151,393]" pageId="8" pageNumber="539">Legs. Hind femur relatively slender or weakly thickened, elongate oval. Basitarsus of hind leg rather long, about 0.8 × as long as second-fifth segments combined.</paragraph>
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<paragraph id="8BE83683FF9CFFC3FF79FF0467A960E8" blockId="8.[151,1436,151,393]" pageId="8" pageNumber="539">Metasoma. First tergite wide and short, with distinct dorsope, laterope absent. Second and third tergites enlarged, strongly sclerotized, covered proceeding soft segments, with completely separated laterotergites, their spiracles situated near margin of dorsal part of tergites. Second suture deep, narrow, evenly curved. Third tergite on posterior margin with short lamella (border), usually distinctly serrate. Ovipositor medium size, distinctly shorter than metasoma, without any teeth or serration.</paragraph>
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</subSubSection>
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</treatment>
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</document> |