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<document id="A7F16E03463D248839277D4C5CB6B47A" ID-CLB-Dataset="7408" ID-DOI="10.5252/adansonia2021v43a8" ID-GBIF-Dataset="49f3bd37-fa27-45a4-90b2-1e80c7f027ac" ID-ISSN="1639-4798" ID-Zenodo-Dep="4680762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1618231216314" checkinUser="carolina" docAuthor="Van de Beek, Abraham &amp; Widrlechner, Mark P." docDate="2021" docId="D5365613DD105D54498AFF5EFD5809CC" docLanguage="en" docName="Adansonia.43.8.67-98.pdf.imf" docOrigin="Adansonia (3) 43 (8)" docStyle="DocumentStyle:2FE19AC740987067EC2A6C89A85F9648.8:Adansonia.2018-.journal_article" docStyleId="2FE19AC740987067EC2A6C89A85F9648" docStyleName="Adansonia.2018-.journal_article" docStyleVersion="8" docTitle="Rubus floridus Tratt." docType="treatment" docVersion="11" lastPageNumber="88" masterDocId="290F2E6BDD075D4F4A75FF89FFCC085D" masterDocTitle="North American species of Rubus L. (Rosaceae) described from European botanical gardens (1789 - 1823)" masterLastPageNumber="1823" masterPageNumber="1789" pageNumber="88" updateTime="1698936340201" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DBDED1582B764CB88FDE01FD0885460D">North American species of Rubus L. (Rosaceae) described from European botanical gardens (1789 - 1823)</mods:title>
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<mods:namePart id="16E7690C62993904C4A67AE9A6F971E3">Van de Beek, Abraham</mods:namePart>
<mods:affiliation id="327E7606F073AC7C9BD6F6087DD1BFA0">Petenbos 8 3904 BN Veenendaal (The Netherlands)</mods:affiliation>
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<mods:namePart id="B5275CEA76817AF7CE49EFF94AC7DAAD">Widrlechner, Mark P.</mods:namePart>
<mods:affiliation id="DA016979323DF5C3F0C069C78538C82B">Iowa State University Departments of Horticulture and of Ecology, Evolution, &amp; Organismal Biology 360 Bessey Hall, 2200 Osborn Drive, Ames, Iowa 50011 (United States)</mods:affiliation>
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<mods:title id="031FDB1E324A8389F6A13C48C32C04E7">Adansonia</mods:title>
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<mods:date id="58478B46339938B348155FB477B7257F">2021</mods:date>
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<mods:start id="97446BF6EA0286038A8DB759E26026EF">1789</mods:start>
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<treatment id="D5365613DD105D54498AFF5EFD5809CC" ID-DOI="http://doi.org/10.5281/zenodo.4684291" ID-GBIF-Taxon="180999573" ID-Zenodo-Dep="4684291" LSID="urn:lsid:plazi:treatment:D5365613DD105D54498AFF5EFD5809CC" httpUri="http://treatment.plazi.org/id/D5365613DD105D54498AFF5EFD5809CC" lastPageId="27" lastPageNumber="88" pageId="23" pageNumber="88">
<subSubSection id="1585B48EDD105D58498AFF5EFB1208AF" box="[1023,1246,215,242]" pageId="23" pageNumber="88" type="nomenclature">
<paragraph id="5D20E705DD105D58498AFF5EFB1208AF" blockId="23.[1023,1246,215,274]" box="[1023,1246,215,242]" pageId="23" pageNumber="88">
<heading id="06685069DD105D58498AFF5EFB1208AF" box="[1023,1246,215,242]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="88" reason="6">
<taxonomicName id="9A9F9C86DD105D58498AFF5EFB1208AF" ID-CoL="799MK" authority="Tratt." authorityName="Tratt." box="[1023,1246,215,242]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="floridus">
<emphasis id="6FEB3B17DD105D58498AFF5EFB5008AF" bold="true" box="[1023,1180,215,242]" italics="true" pageId="23" pageNumber="88">Rubus floridus</emphasis>
Tratt.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1585B48EDD105D584E4AFF7EFB51094F" box="[1087,1181,247,274]" pageId="23" pageNumber="88" type="description">
<paragraph id="5D20E705DD105D584E4AFF7EFB51094F" blockId="23.[1023,1246,215,274]" box="[1087,1181,247,274]" pageId="23" pageNumber="88">
(
<figureCitation id="C5A4FB80DD105D584E3DFF7EFB58094C" box="[1096,1172,247,273]" captionStart="FIG" captionStartId="26.[559,570,2008,2025]" captionTargetBox="[245,1342,215,1915]" captionTargetId="figure-23@26.[245,1342,215,1915]" captionTargetPageId="26" captionText="FIG. 11. — Rubus floridus Tratt. holotype, W(W0079959)." figureDoi="http://doi.org/10.5281/zenodo.4680808" httpUri="https://zenodo.org/record/4680808/files/figure.png" pageId="23" pageNumber="88">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1585B48EDD105D584936FEB8FB3F09CF" pageId="23" pageNumber="88" type="reference_group">
<paragraph id="5D20E705DD105D584936FEB8FA870915" blockId="23.[813,1457,304,476]" box="[835,1355,304,328]" pageId="23" pageNumber="88">
In
<treatmentCitation id="DC3EC114DD105D58492AFEB8FA870915" author="Trattinnick L." box="[863,1355,304,328]" page="73" pageId="23" pageNumber="88" year="1823">
<emphasis id="6FEB3B17DD105D58492AFEB8FB830915" box="[863,1103,304,328]" italics="true" pageId="23" pageNumber="88">Rosacearum monographia</emphasis>
3: 73 (
<bibRefCitation id="390E9AF4DD105D584EE3FEB9FAF10915" author="TRATTINNICK L." box="[1174,1341,304,328]" pageId="23" pageNumber="88" refId="ref19304" refString="TRATTINNICK L. 1823. - Rosacearum monographia 3. Heubner, Vindobonae, xvi + 171 p." type="book" year="1823">Trattinnick 1823</bibRefCitation>
).
</treatmentCitation>
</paragraph>
<paragraph id="5D20E705DD105D584958FEE9FB3F09CF" blockId="23.[813,1457,304,476]" pageId="23" pageNumber="88">
<treatmentCitation id="DC3EC114DD105D584958FEE9FB3F09CF" author="Bailey L. H." page="370, 385" pageId="23" pageNumber="88" year="1898">
<taxonomicName id="9A9F9C86DD105D584958FEE9FADE0925" ID-CoL="799MK" authority="(Tratt.) L. H. Bailey, Sketch" authorityName="L. H. Bailey, Sketch" baseAuthorityName="Tratt." box="[813,1298,352,376]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="variety" species="argutus" variety="floridus">
<emphasis id="6FEB3B17DD105D584958FEE9FC42092A" box="[813,910,352,375]" italics="true" pageId="23" pageNumber="88">R. argutus</emphasis>
var.
<emphasis id="6FEB3B17DD105D5849CAFEE9FBC90925" box="[959,1029,352,376]" italics="true" pageId="23" pageNumber="88">floridus</emphasis>
(Tratt.) L.H. Bailey,
<emphasis id="6FEB3B17DD105D584EA2FEE9FADE0925" box="[1239,1298,352,376]" italics="true" pageId="23" pageNumber="88">Sketch</emphasis>
</taxonomicName>
<emphasis id="6FEB3B17DD105D584F6FFEE9FC3109CF" italics="true" pageId="23" pageNumber="88">of the Evolution of our Native Fruits</emphasis>
: 370, 385 (
<bibRefCitation id="390E9AF4DD105D584E05FEF3FB2909CF" author="BAILEY L. H." box="[1136,1253,378,402]" pageId="23" pageNumber="88" refId="ref16166" refString="BAILEY L. H. 1898. - Sketch of the Evolution of our Native Fruits. MacMillan, New York, 465 p. https: // doi. org / 10.5962 / bhl. title. 19869" type="book" year="1898">Bailey 1898</bibRefCitation>
).
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="1585B48EDD105D584958FE23FB810981" pageId="23" pageNumber="88" type="materials_examined">
<paragraph id="5D20E705DD105D584958FE23FB810981" blockId="23.[813,1457,304,476]" pageId="23" pageNumber="88">
<materialsCitation id="EDF7ED58DD105D584958FE23FB810981" ID-GBIF-Occurrence="3071146315" collectionCode="W" municipality="Enslen" pageId="23" pageNumber="88" specimenCode="W0079959" specimenCount="1" typeStatus="holotype">
<typeStatus id="822459A7DD105D584958FE23FC5B099D" box="[813,919,426,449]" pageId="23" pageNumber="88" type="holotype">HOLOTYPE</typeStatus>
. —
<emphasis id="6FEB3B17DD105D5849B0FE23FBCE099F" box="[965,1026,426,450]" italics="true" pageId="23" pageNumber="88">
<collectingMunicipality id="BD447D7FDD105D5849B0FE23FBCE099F" box="[965,1026,426,450]" pageId="23" pageNumber="88">Enslen</collectingMunicipality>
</emphasis>
,
<date id="2921C1C5DD105D584E66FE23FB95099C" box="[1043,1113,426,450]" pageId="23" pageNumber="88" value="1822-01">1822 I</date>
,
<taxonomicName id="9A9F9C86DD105D584E10FE23FB38099F" box="[1125,1268,426,450]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="floribundus">R. floribundus</taxonomicName>
[sic!] Tratt., Amer. bor. Enslen,
<collectionCode id="3B8E7FC0DD105D5849D9FE4DFC090981" box="[940,965,452,476]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="23" pageNumber="88" type="Herbarium">W</collectionCode>
(
<specimenCode id="0D394F7EDD105D5849B9FE4DFB8E0981" box="[972,1090,452,476]" pageId="23" pageNumber="88">W0079959</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1585B48EDD105D544958FD9EFD5809CC" lastPageId="27" lastPageNumber="92" pageId="23" pageNumber="88" type="discussion">
<paragraph id="5D20E705DD105D584958FD9EFC590A72" blockId="23.[811,1457,535,2028]" box="[813,917,535,561]" pageId="23" pageNumber="88">FINDINGS</paragraph>
<paragraph id="5D20E705DD105D584958FDBFFB660BED" blockId="23.[811,1457,535,2028]" pageId="23" pageNumber="88">
Bailey obtained both a photo and a drawing of the
<typeStatus id="822459A7DD105D584F34FDBEFAA30A0C" box="[1345,1391,567,593]" pageId="23" pageNumber="88">type</typeStatus>
(both printed in
<bibRefCitation id="390E9AF4DD105D5849DEFDDFFBA00A2D" author="BAILEY L. H." box="[939,1132,598,624]" pageId="23" pageNumber="88" pagination="137 - 200" refId="ref16205" refString="BAILEY L. H. 1923. - Certain cultivated Rubi. Gentes Herbarum 1: 137 - 200. https: // hdl. handle. net / 2027 / inu. 30000046319301" type="journal article" year="1923">Bailey 1923: 194</bibRefCitation>
f, the drawing also in
<bibRefCitation id="390E9AF4DD105D584F18FDDFFC6A0ACD" author="BAILEY L. H." pageId="23" pageNumber="88" refId="ref16166" refString="BAILEY L. H. 1898. - Sketch of the Evolution of our Native Fruits. MacMillan, New York, 465 p. https: // doi. org / 10.5962 / bhl. title. 19869" type="book" year="1898">Bailey 1898: 382</bibRefCitation>
). Initially (1898), he considered it as a variety of
<taxonomicName id="9A9F9C86DD105D584939FD1FFC700AF2" authorityName="Link (Link" authorityYear="1822" box="[844,956,662,687]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="argutus">
<emphasis id="6FEB3B17DD105D584939FD1FFC700AF2" box="[844,956,662,687]" italics="true" pageId="23" pageNumber="88">R. argutus</emphasis>
</taxonomicName>
. Later he dealt with it as a separate species. Though he noted that the specimen has some glands, he inserted it in a group of mainly glandless brambles under the name
<emphasis id="6FEB3B17DD105D5849E8FD7CFC2B0B52" box="[925,999,757,783]" italics="true" pageId="23" pageNumber="88">Floridi</emphasis>
(
<bibRefCitation id="390E9AF4DD105D58498DFD7CFB770B52" author="BAILEY L. H." box="[1016,1211,757,784]" pageId="23" pageNumber="88" pagination="137 - 200" refId="ref16205" refString="BAILEY L. H. 1923. - Certain cultivated Rubi. Gentes Herbarum 1: 137 - 200. https: // hdl. handle. net / 2027 / inu. 30000046319301" type="journal article" year="1923">Bailey 1923: 193</bibRefCitation>
), later identified with the
<emphasis id="6FEB3B17DD105D584923FC9CFC7E0B72" box="[854,946,789,815]" italics="true" pageId="23" pageNumber="88">Frondosi</emphasis>
(
<bibRefCitation id="390E9AF4DD105D5849B5FC9CFBB20B6D" author="BAILEY L. H." box="[960,1150,789,816]" pageId="23" pageNumber="88" pagination="137 - 200" refId="ref16205" refString="BAILEY L. H. 1923. - Certain cultivated Rubi. Gentes Herbarum 1: 137 - 200. https: // hdl. handle. net / 2027 / inu. 30000046319301" type="journal article" year="1923">Bailey 1923: 265</bibRefCitation>
, 268). Finally (
<bibRefCitation id="390E9AF4DD105D584F50FC9CFC900B0D" author="BAILEY L. H." pageId="23" pageNumber="88" pagination="229 - 477" refId="ref16321" refString="BAILEY L. H. 1943. - Rubus in North America. Flagellares. Gentes Herbarum 5: 229 - 477. https: // hdl. handle. net / 2027 / uc 1.31210001328085" type="journal article" year="1943">Bailey 1943: 603</bibRefCitation>
), he postponed his efforts to identify it. So did
<bibRefCitation id="390E9AF4DD105D584F05FCBCFC070B2D" author="DAVIS H. A. &amp; FULLER A. M. &amp; DAVIS T." pageId="23" pageNumber="88" pagination="235 - 266" refId="ref17164" refString="DAVIS H. A., FULLER A. M. &amp; DAVIS T. 1969 b. - Contributions toward the Revision of the Eubati of Eastern North America. V. Arguti. Castanea 34 (3): 235 - 266. https: // www. jstor. org / stable / 4032358" type="journal article" year="1969">
Davis
<emphasis id="6FEB3B17DD105D584958FCDFFCA90B32" box="[813,869,853,879]" italics="true" pageId="23" pageNumber="88">et al.</emphasis>
(1969b)
</bibRefCitation>
, who wrote they considered it as a
<emphasis id="6FEB3B17DD105D584F1FFCDFFCB50BD2" italics="true" pageId="23" pageNumber="88">nomen nudum</emphasis>
(obviously meaning
<emphasis id="6FEB3B17DD105D584E16FCFFFB0E0BD2" box="[1123,1218,886,911]" italics="true" pageId="23" pageNumber="88">incertum</emphasis>
or
<emphasis id="6FEB3B17DD105D584E87FCFCFAA80BD2" box="[1266,1380,885,911]" italics="true" pageId="23" pageNumber="88">ambiguum</emphasis>
, since the species was validly published).
</paragraph>
<paragraph id="5D20E705DD105D584936FC3CFA610CD2" blockId="23.[811,1457,535,2028]" pageId="23" pageNumber="88">
Though
<bibRefCitation id="390E9AF4DD105D5849D7FC3CFB4C0B92" author="DAVIS H. A. &amp; FULLER A. M. &amp; DAVIS T." box="[930,1152,949,975]" pageId="23" pageNumber="88" pagination="235 - 266" refId="ref17164" refString="DAVIS H. A., FULLER A. M. &amp; DAVIS T. 1969 b. - Contributions toward the Revision of the Eubati of Eastern North America. V. Arguti. Castanea 34 (3): 235 - 266. https: // www. jstor. org / stable / 4032358" type="journal article" year="1969">
Davis
<emphasis id="6FEB3B17DD105D58499CFC3CFBD20B92" box="[1001,1054,949,975]" italics="true" pageId="23" pageNumber="88">et al.</emphasis>
(1969b)
</bibRefCitation>
discussed
<taxonomicName id="9A9F9C86DD105D584E82FC3CFAAA0B92" box="[1271,1382,949,975]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="floridus">
<emphasis id="6FEB3B17DD105D584E82FC3CFAAA0B92" box="[1271,1382,949,975]" italics="true" pageId="23" pageNumber="88">R. floridus</emphasis>
</taxonomicName>
under the
<emphasis id="6FEB3B17DD105D584920FC5CFC550BB2" box="[853,921,981,1007]" italics="true" pageId="23" pageNumber="88">Arguti</emphasis>
, they suggested that it might fit better within the
<emphasis id="6FEB3B17DD105D58495EFC7DFC0C0C53" box="[811,960,1012,1038]" italics="true" pageId="23" pageNumber="88">Alleghenienses</emphasis>
because
<bibRefCitation id="390E9AF4DD105D584E5FFC7DFB300C52" author="TRATTINNICK L." box="[1066,1276,1012,1039]" pageId="23" pageNumber="88" refId="ref19304" refString="TRATTINNICK L. 1823. - Rosacearum monographia 3. Heubner, Vindobonae, xvi + 171 p." type="book" year="1823">Trattinnick (1823)</bibRefCitation>
explicitly mentioned its glands. We examined a high-resolution image of the
<typeStatus id="822459A7DD105D584920FBBCFC490C12" box="[853,901,1077,1103]" pageId="23" pageNumber="88">type</typeStatus>
(
<figureCitation id="C5A4FB80DD105D5849E3FBBDFC2F0C13" box="[918,995,1076,1102]" captionStart="FIG" captionStartId="26.[559,570,2008,2025]" captionTargetBox="[245,1342,215,1915]" captionTargetId="figure-23@26.[245,1342,215,1915]" captionTargetPageId="26" captionText="FIG. 11. — Rubus floridus Tratt. holotype, W(W0079959)." figureDoi="http://doi.org/10.5281/zenodo.4680808" httpUri="https://zenodo.org/record/4680808/files/figure.png" pageId="23" pageNumber="88">Fig. 11</figureCitation>
), and it does bear long stipitate glands. If
<taxonomicName id="9A9F9C86DD105D584959FBDCFC5B0C33" box="[812,919,1108,1134]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="floridus">
<emphasis id="6FEB3B17DD105D584959FBDCFC5B0C33" box="[812,919,1108,1134]" italics="true" pageId="23" pageNumber="88">R. floridus</emphasis>
</taxonomicName>
is truly a highbush blackberry, this would support Davis
<emphasis id="6FEB3B17DD105D584901FBFCFC650CD3" box="[884,937,1140,1166]" italics="true" pageId="23" pageNumber="88">et al.</emphasis>
s (1969b) suggested transfer to
<emphasis id="6FEB3B17DD105D584F65FBFDFA690CD3" box="[1296,1445,1140,1166]" italics="true" pageId="23" pageNumber="88">Alleghenienses</emphasis>
.
</paragraph>
<paragraph id="5D20E705DD105D584936FB1DFBF20E70" blockId="23.[811,1457,535,2028]" pageId="23" pageNumber="88">
Before accepting this option, the possibility that it belongs to another section, especially the
<emphasis id="6FEB3B17DD105D584F78FB3DFA6B0C93" box="[1293,1447,1204,1230]" italics="true" pageId="23" pageNumber="88">Procumbentes</emphasis>
, must be excluded. Though
<bibRefCitation id="390E9AF4DD105D584E14FB5DFAFF0CB3" author="TRATTINNICK L." box="[1121,1331,1236,1262]" pageId="23" pageNumber="88" refId="ref19304" refString="TRATTINNICK L. 1823. - Rosacearum monographia 3. Heubner, Vindobonae, xvi + 171 p." type="book" year="1823">Trattinnick (1823)</bibRefCitation>
wrote that the stems are recti;
<bibRefCitation id="390E9AF4DD105D584E6CFB7AFB270D53" author="BAILEY L. H." box="[1049,1259,1267,1294]" pageId="23" pageNumber="88" pagination="591 - 835" refId="ref16395" refString="BAILEY L. H. 1945. - Rubus in North America. Eubatus. Arguti. Gentes Herbarum 5: 591 - 835. https: // hdl. handle. net / 2027 / uc 1.31210001328085" type="journal article" year="1945">Bailey (1945: 604)</bibRefCitation>
might have been correct when he stated that this only means straight, not erect. However, the way the inflorescences are positioned on the floricane makes clear that the stem was more-orless erect. On procumbent canes, the angles relative to the subtending cane would not have been so acute. The inflorescences were growing in the same direction as the stem. Consequently, the
<typeStatus id="822459A7DD105D584E70FA5AFBF90DB0" box="[1029,1077,1491,1517]" pageId="23" pageNumber="88">type</typeStatus>
was most likely collected from a highbush blackberry, and, because of its stipitate glands, in section
<emphasis id="6FEB3B17DD105D5849D7F99BFBFB0E71" box="[930,1079,1554,1580]" italics="true" pageId="23" pageNumber="88">Alleghenienses</emphasis>
.
</paragraph>
<paragraph id="5D20E705DD105D584936F9BBFBCA0FF1" blockId="23.[811,1457,535,2028]" pageId="23" pageNumber="88">
<bibRefCitation id="390E9AF4DD105D584936F9BBFBEB0E10" author="DAVIS H. A. &amp; FULLER A. M. &amp; DAVIS T." box="[835,1063,1586,1613]" pageId="23" pageNumber="88" pagination="235 - 266" refId="ref17164" refString="DAVIS H. A., FULLER A. M. &amp; DAVIS T. 1969 b. - Contributions toward the Revision of the Eubati of Eastern North America. V. Arguti. Castanea 34 (3): 235 - 266. https: // www. jstor. org / stable / 4032358" type="journal article" year="1969">
Davis
<emphasis id="6FEB3B17DD105D5849F9F9BAFC080E11" box="[908,964,1586,1612]" italics="true" pageId="23" pageNumber="88">et al.</emphasis>
(1969b)
</bibRefCitation>
suggested that it would belong to the
<taxonomicName id="9A9F9C86DD105D584922F9DAFB970E31" authority="L. H. Bailey" authorityName="L. H. Bailey" box="[855,1115,1618,1645]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="alumnus">
<emphasis id="6FEB3B17DD105D584922F9DAFC1E0E31" box="[855,978,1618,1644]" italics="true" pageId="23" pageNumber="88">R. alumnus</emphasis>
L.H. Bailey
</taxonomicName>
complex if it is situated in the
<emphasis id="6FEB3B17DD105D584958F9FBFC0D0ED1" box="[813,961,1650,1676]" italics="true" pageId="23" pageNumber="88">Alleghenienses</emphasis>
. Among the currently recognized taxa in that section, it is true that the flaring inflorescences displayed by the
<typeStatus id="822459A7DD105D58490DF93AFC640E90" box="[888,936,1715,1741]" pageId="23" pageNumber="88">type</typeStatus>
are typically found only in
<taxonomicName id="9A9F9C86DD105D584EA0F93AFA830E91" box="[1237,1359,1714,1740]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="alumnus">
<emphasis id="6FEB3B17DD105D584EA0F93AFA830E91" box="[1237,1359,1714,1740]" italics="true" pageId="23" pageNumber="88">R. alumnus</emphasis>
</taxonomicName>
. But the size of its calyx lobes is smaller than those of
<taxonomicName id="9A9F9C86DD105D584F58F95AFA640EB1" box="[1325,1448,1746,1772]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="alumnus">
<emphasis id="6FEB3B17DD105D584F58F95AFA640EB1" box="[1325,1448,1746,1772]" italics="true" pageId="23" pageNumber="88">R. alumnus</emphasis>
</taxonomicName>
, and its inflorescence length and hooked prickles also would be atypical for that species (
<bibRefCitation id="390E9AF4DD105D584E1EF89BFAF50F71" author="WIDRLECHNER M. P." box="[1131,1337,1810,1836]" pageId="23" pageNumber="88" pagination="979 - 1004" refId="ref19856" refString="WIDRLECHNER M. P. 2013. - Rubus L. (raspberry, blackberry, bramble), in YATSKIEVYCH G. (ed.), Steyermark's Flora of Missouri. Vol. 3. Missouri Botanical Garden Press and Missouri Department of Conservation, St. Louis and Jefferson City, MO: 976 - 977, 979 - 1004. https: // lib. dr. iastate. edu / bot _ pubs / 84" type="journal article" year="2013">Widrlechner 2013</bibRefCitation>
). After an extensive search, we could find no other known member of the
<emphasis id="6FEB3B17DD105D584909F8D8FBDA0F36" box="[892,1046,1873,1899]" italics="true" pageId="23" pageNumber="88">Alleghenienses</emphasis>
that is a good match only a few specimens resembling most of the characters presented in the
<taxonomicName id="9A9F9C86DD105D58492DF81BFC0B0FF6" box="[856,967,1937,1963]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="23" pageNumber="88" phylum="Tracheophyta" rank="species" species="floridus">
<emphasis id="6FEB3B17DD105D58492DF81BFC0B0FF6" box="[856,967,1937,1963]" italics="true" pageId="23" pageNumber="88">R. floridus</emphasis>
</taxonomicName>
<typeStatus id="822459A7DD105D5849BAF81BFBCE0FF1" box="[975,1026,1938,1964]" pageId="23" pageNumber="88">type</typeStatus>
.
</paragraph>
<paragraph id="5D20E705DD105D544931F838FD5809CC" blockId="23.[811,1457,535,2028]" lastBlockId="27.[131,774,215,401]" lastPageId="27" lastPageNumber="92" pageId="23" pageNumber="88">
Returning to the other option, some large, mounding species of
<emphasis id="6FEB3B17DD105D5849D3F858FBF10FB6" box="[934,1085,2001,2027]" italics="true" pageId="23" pageNumber="88">Procumbentes</emphasis>
produce side branches that get caught in vegetation and display determinate development and hooked prickles resembling those of the
<taxonomicName id="9A9F9C86DD1C5D5448E6FF71FCCA094C" box="[659,774,247,273]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="27" pageNumber="92" phylum="Tracheophyta" rank="species" species="floridus">
<emphasis id="6FEB3B17DD1C5D5448E6FF71FCCA094C" box="[659,774,247,273]" italics="true" pageId="27" pageNumber="92">R. floridus</emphasis>
</taxonomicName>
<typeStatus id="822459A7DD1C5D544AF1FE91FF74096F" box="[132,184,280,306]" pageId="27" pageNumber="92">type</typeStatus>
. Thus, we also conducted a search of the mounding, glandular
<emphasis id="6FEB3B17DD1C5D544A82FEBEFE45090C" box="[247,393,311,337]" italics="true" pageId="27" pageNumber="92">Procumbentes</emphasis>
. Of this group, the closest match is
<taxonomicName id="9A9F9C86DD1C5D544AEAFED1FE6E092C" authority="L. H. Bailey" authorityName="L. H. Bailey" box="[159,418,343,369]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="27" pageNumber="92" phylum="Tracheophyta" rank="species" species="grimesii">
<emphasis id="6FEB3B17DD1C5D544AEAFED1FED8092C" box="[159,276,344,369]" italics="true" pageId="27" pageNumber="92">R. grimesii</emphasis>
L.H. Bailey
</taxonomicName>
, a connection noted in passing by
<bibRefCitation id="390E9AF4DD1C5D544AD2FEFEFEF309CC" author="BAILEY L. H." box="[167,319,374,401]" pageId="27" pageNumber="92" pagination="591 - 835" refId="ref16395" refString="BAILEY L. H. 1945. - Rubus in North America. Eubatus. Arguti. Gentes Herbarum 5: 591 - 835. https: // hdl. handle. net / 2027 / uc 1.31210001328085" type="journal article" year="1945">Bailey (1945)</bibRefCitation>
but never critically examined.
</paragraph>
</subSubSection>
</treatment>
</document>