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<document id="8BDDCCCE04012A598F04B0FEB48E1716" ID-DOI="10.5281/zenodo.208769" ID-GBIF-Dataset="c44a46a0-62ed-4553-99a9-97603714d0f4" ID-ISSN="1175-5326" ID-Zenodo-Dep="208769" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460503201734" checkinUser="plazi" docAuthor="Çiftçi, Mustafa Cemal, Hasbenli, Abdullah &amp; Canpolat, Derya" docDate="2012" docId="E6209671FF8E050FFF07B2FEFAF566CF" docLanguage="en" docName="zt03408p053.pdf" docOrigin="Zootaxa 3408" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empis (Polyblepharis) nabucco Ciftci, sp. nov." docType="treatment" docVersion="9" lastPageNumber="49" masterDocId="1A19EE09FF8F050DFF90B004FFA8610B" masterDocTitle="Two new species of Empis (Diptera: Empididae) from Turkey" masterLastPageNumber="53" masterPageNumber="47" pageNumber="48" updateTime="1698312289020" updateUser="plazi">
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<mods:title id="335627538EC08E8EB132D29F30E4C895">Two new species of Empis (Diptera: Empididae) from Turkey</mods:title>
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<mods:namePart id="BC07A0DE595C0FFA851D01AE6BF2745A">Çiftçi, Mustafa Cemal</mods:namePart>
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<mods:namePart id="038A8A5D4C14F9D1B9DEDF214F5D88C3">Hasbenli, Abdullah</mods:namePart>
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<mods:namePart id="4CD2D0C64ED62364258E8A2308A16C65">Canpolat, Derya</mods:namePart>
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<mods:title id="3DBAE30870C12BC6EEA3164EE5FF7680">Zootaxa</mods:title>
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<mods:part id="AE0F569DDD53B63B42638309A5E6ADB4">
<mods:date id="1107F98EA777710538EF0A0C48CB641C">2012</mods:date>
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<mods:number id="82E151E8A0883AEB536B1696EB86603B">3408</mods:number>
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<treatment id="E6209671FF8E050FFF07B2FEFAF566CF" ID-DOI="http://doi.org/10.5281/zenodo.6175502" ID-GBIF-Taxon="119521773" ID-Zenodo-Dep="6175502" LSID="urn:lsid:plazi:treatment:E6209671FF8E050FFF07B2FEFAF566CF" httpUri="http://treatment.plazi.org/id/E6209671FF8E050FFF07B2FEFAF566CF" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48">
<subSubSection id="269374ECFF8E050CFF07B2FEFEBA623E" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="6E362767FF8E050CFF07B2FEFD6C621F" blockId="1.[151,708,762,821]" box="[151,708,762,788]" pageId="1" pageNumber="48">
<heading id="357E900BFF8E050CFF07B2FEFD6C621F" bold="true" box="[151,708,762,788]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="5CFDFB75FF8E050CFF07B2FEFD6C621F" bold="true" box="[151,708,762,788]" pageId="1" pageNumber="48">
<taxonomicName id="A9895CE4FF8E050CFF07B2FEFDF7621F" authority="Ciftci" authorityName="Ciftci" box="[151,607,762,788]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="nabucco" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="5CFDFB75FF8E050CFF07B2FEFF4B621F" bold="true" box="[151,227,762,788]" italics="true" pageId="1" pageNumber="48">Empis</emphasis>
(
<emphasis id="5CFDFB75FF8E050CFF64B2FEFE32621F" bold="true" box="[244,410,762,788]" italics="true" pageId="1" pageNumber="48">Polyblepharis</emphasis>
)
<emphasis id="5CFDFB75FF8E050CFE3CB2FEFDBB621F" bold="true" box="[428,531,762,788]" italics="true" pageId="1" pageNumber="48">nabucco</emphasis>
Çiftçi
</taxonomicName>
<taxonomicNameLabel id="47CE460EFF8E050CFDF7B2FEFD6C621F" box="[615,708,762,788]" pageId="1" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6E362767FF8E050CFF07B319FEBA623E" blockId="1.[151,708,762,821]" box="[151,274,797,821]" pageId="1" pageNumber="48">
(
<figureCitation id="F6B23BE2FF8E050CFF0FB319FEA1623E" box="[159,265,797,821]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,720,743]" captionTargetBox="[161,1423,238,674]" captionTargetId="figure@2.[151,1436,193,699]" captionTargetPageId="2" captionText="FIGURES 1 3. Empis (Polyblepharis) nabucco 1. Genitalia; 2. Dorsal lamella (cercus) in dorsal view; 3. Aedeagus (phallus). Scale = 0.3 mm." httpUri="https://zenodo.org/record/208770/files/figure.png" pageId="1" pageNumber="48">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="269374ECFF8E050CFF07B361FB0662CE" pageId="1" pageNumber="48" type="diagnosis">
<paragraph id="6E362767FF8E050CFF07B361FB0662CE" blockId="1.[151,1437,869,1973]" pageId="1" pageNumber="48">
<emphasis id="5CFDFB75FF8E050CFF07B361FEB96275" bold="true" box="[151,273,869,894]" pageId="1" pageNumber="48">Diagnosis.</emphasis>
Medium to large, body
<quantity id="A9718A82FF8E050CFD8DB361FD0B6276" box="[541,675,869,893]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.6" metricValueMin="5.7" pageId="1" pageNumber="48" unit="mm" value="6.65" valueMax="7.6" valueMin="5.7">5.77.6 mm</quantity>
long, legs and halteres black. Hairs on thorax and abdomen whitish, scutum with long dense hairs leaving only inner stripes bare, acrostichals and dorsocentrals mostly white, humeral bristle distinct. Female scutum with few short hairs, legs without pennation or flattened hairs.
</paragraph>
</subSubSection>
<subSubSection id="269374ECFF8E050FFF59B3D5FA8D654B" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48" type="description">
<paragraph id="6E362767FF8E050CFF59B3D5FB066572" blockId="1.[151,1437,869,1973]" pageId="1" pageNumber="48">
<emphasis id="5CFDFB75FF8E050CFF59B3D5FEF362E1" bold="true" box="[201,347,977,1002]" pageId="1" pageNumber="48">Description.</emphasis>
<emphasis id="5CFDFB75FF8E050CFEF2B3D5FE3462E2" box="[354,412,977,1001]" italics="true" pageId="1" pageNumber="48">Male</emphasis>
. Eyes meeting on frons for long distance, upper ommatidia enlarged. Face dull grey, lower edge shinning black. Pair of ocellar bristles as long as other occipital bristles. Occiput grey, upper occipital bristles black, lower part of occiput with thin whitish hairs. Antenna black, basal segments (scape and pedicel) with black bristly hairs as long as corresponding segments, stylus as long as half-length of 3rd antennal segment (postpedicel). Palpus yellow, basally black with sparse bristly hairs. Labrum 1.5X as long as height of head.
</paragraph>
<paragraph id="6E362767FF8E050CFF59B481FDE26492" blockId="1.[151,1437,869,1973]" pageId="1" pageNumber="48">
Thorax grey, scutum with 4 dull black stripes, 2 narrow inner stripes separating acrostichals and dorsocentrals, outer stripes
<date id="1A3701A7FF8E050CFEBAB4ADFEE565CA" box="[298,333,1193,1217]" pageId="1" pageNumber="48">2X</date>
broader than inner stripes. Scutum with long dense whitish hairs as long as length of 1st (scape) and 2nd (pedicel) antennal segments leaving only inner stripes bare. Acrostichals and dorsocentrals whitish and as long as other hairs on scutum, acrostichals irregularly 35 serial, dorsocentrals multiserial also with row of black, slightly thicker bristles and ending with 2 pairs of long prescutellar bristles. Humeri (postpronotum) covered with whitish hairs, humeral bristle black anteriorly with 34 smaller bristles. Large thoracic bristles black, 1 intrahumeral, 3 notopleural, 1 supra-alar, 1 postalar, 24 scutellar bristles. Prothorax with whitish hairs. Metapleural fan (laterotergal bristles) long and whitish.
</paragraph>
<paragraph id="6E362767FF8E050CFF59B5A1FCC864EA" blockId="1.[151,1437,869,1973]" pageId="1" pageNumber="48">Wing almost clear, all veins dark and distinct, only wing base yellowish. Costal bristle absent. Squama yellow with white fringes. Haltere with black knob and brownish stem.</paragraph>
<paragraph id="6E362767FF8E050CFF59B5EAFC13679E" blockId="1.[151,1437,869,1973]" pageId="1" pageNumber="48">Legs long and slender, shinning black to blackish brown except for greyish dusted coxae. Coxae with long whitish hairs and 45 outer black bristles. Femora with short black hairs, mid and hind femora ventrally with row of spine-like bristles. Tibiae ventrally with short adpressed hairs, dorsal hairs bristle-like, slightly longer than ventral hairs. Mid tibia anteroventrally with row of short bristles. Tarsal segments with dense short hairs and apical circlet of bristles. Basitarsi (tarsomere 1) ventrally with spine-like bristles.</paragraph>
<paragraph id="6E362767FF8E050CFF59B6A5FE8766BE" blockId="1.[151,1437,869,1973]" pageId="1" pageNumber="48">
Abdomen grey, terga much darker and slightly brownish pollinose in dorsal view. Terga 5 and 6 black and brownish pollinose, sternum 8 and genitalia shinning black with slightly brownish dusting. Abdominal hairs whitish, lateral hairs on anterior segments long. Terga 24 and 6 laterally with 1, tergum 5 laterally with 2 marginal bristles. Sternum 8 with sparse hairs and long and thick hind marginal bristles. Genitalia (terminalia) (
<figureCitation id="F6B23BE2FF8E050CFAB8B709FA3B662E" box="[1320,1427,1805,1829]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,720,743]" captionTargetBox="[161,1423,238,674]" captionTargetId="figure@2.[151,1436,193,699]" captionTargetPageId="2" captionText="FIGURES 1 3. Empis (Polyblepharis) nabucco 1. Genitalia; 2. Dorsal lamella (cercus) in dorsal view; 3. Aedeagus (phallus). Scale = 0.3 mm." httpUri="https://zenodo.org/record/208770/files/figure.png" pageId="1" pageNumber="48">Figs. 13</figureCitation>
) large and black with black marginal hairs, lower hind edge of lateral lamella (epandrium) brownish. Lateral lamella (epandrium) large, almost square, posterior edge oval (
<figureCitation id="F6B23BE2FF8E050CFD6DB751FC976666" box="[765,831,1877,1901]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,720,743]" captionTargetBox="[161,1423,238,674]" captionTargetId="figure@2.[151,1436,193,699]" captionTargetPageId="2" captionText="FIGURES 1 3. Empis (Polyblepharis) nabucco 1. Genitalia; 2. Dorsal lamella (cercus) in dorsal view; 3. Aedeagus (phallus). Scale = 0.3 mm." httpUri="https://zenodo.org/record/208770/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
). Dorsal lamella (cercus) with short black hairs, anteriorly bilobed; inner lobe very narrowed and finger-like in dorsal view; inner edge on apical half serrate in dorsal view (
<figureCitation id="F6B23BE2FF8E050CFF4DB799FE8A66BE" box="[221,290,1949,1973]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,720,743]" captionTargetBox="[161,1423,238,674]" captionTargetId="figure@2.[151,1436,193,699]" captionTargetPageId="2" captionText="FIGURES 1 3. Empis (Polyblepharis) nabucco 1. Genitalia; 2. Dorsal lamella (cercus) in dorsal view; 3. Aedeagus (phallus). Scale = 0.3 mm." httpUri="https://zenodo.org/record/208770/files/figure.png" pageId="1" pageNumber="48">Fig. 2</figureCitation>
).
</paragraph>
<caption id="3AF677EFFF8D050FFF07B2D4FE95620E" httpUri="https://zenodo.org/record/208770/files/figure.png" pageId="2" pageNumber="49" targetBox="[161,1423,238,674]" targetPageId="2">
<paragraph id="6E362767FF8D050FFF07B2D4FE95620E" blockId="2.[151,1436,720,773]" pageId="2" pageNumber="49">
<emphasis id="5CFDFB75FF8D050FFF07B2D4FE9263ED" bold="true" box="[151,314,720,743]" pageId="2" pageNumber="49">FIGURES 13.</emphasis>
<taxonomicName id="A9895CE4FF8D050FFED1B2D6FD2A63ED" box="[321,642,720,743]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nabucco" subGenus="Polyblepharis">
<emphasis id="5CFDFB75FF8D050FFED1B2D6FE2863EC" box="[321,384,722,743]" italics="true" pageId="2" pageNumber="49">Empis</emphasis>
(
<emphasis id="5CFDFB75FF8D050FFE1FB2D4FDB563ED" box="[399,541,720,742]" italics="true" pageId="2" pageNumber="49">Polyblepharis</emphasis>
)
<emphasis id="5CFDFB75FF8D050FFDBCB2D4FD2A63ED" box="[556,642,720,742]" italics="true" pageId="2" pageNumber="49">nabucco</emphasis>
</taxonomicName>
<emphasis id="5CFDFB75FF8D050FFD19B2D5FD3E63ED" bold="true" box="[649,662,721,742]" pageId="2" pageNumber="49">1</emphasis>
. Genitalia;
<emphasis id="5CFDFB75FF8D050FFC9DB2D5FCB163ED" bold="true" box="[781,793,721,742]" pageId="2" pageNumber="49">2</emphasis>
. Dorsal lamella (cercus) in dorsal view;
<emphasis id="5CFDFB75FF8D050FFB27B2D5FB6C63ED" bold="true" box="[1207,1220,721,742]" pageId="2" pageNumber="49">3</emphasis>
. Aedeagus (phallus). Scale = 0.3 mm.
</paragraph>
</caption>
<paragraph id="6E362767FF8D050FFF59B328FCE16263" blockId="2.[151,1436,812,1989]" pageId="2" pageNumber="49">
<typeStatus id="B13299C5FF8D050FFF59B328FE99624F" box="[201,305,812,836]" pageId="2" pageNumber="49" type="holotype">Holotype</typeStatus>
male body length:
<quantity id="A9718A82FF8D050FFD82B328FDC6624F" box="[530,622,812,836]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="49" unit="mm" value="6.8">6.8 mm</quantity>
, wing length:
<quantity id="A9718A82FF8D050FFC83B328FCC7624F" box="[787,879,812,836]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="49" unit="mm" value="7.1">7.1 mm</quantity>
.
<typeStatus id="B13299C5FF8D050FFCECB329FC41624E" box="[892,1001,813,837]" pageId="2" pageNumber="49" type="paratype">Paratypes</typeStatus>
body length: male
<quantity id="A9718A82FF8D050FFB5BB328FAEA624F" box="[1227,1346,812,836]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.0" metricValueMin="5.7" pageId="2" pageNumber="49" unit="mm" value="6.35" valueMax="7.0" valueMin="5.7">5.77 mm</quantity>
; female
<quantity id="A9718A82FF8D050FFF07B354FEA36263" box="[151,267,848,872]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="7.4" metricValueMin="7.0" pageId="2" pageNumber="49" unit="mm" value="7.2" valueMax="7.4" valueMin="7.0">77.4 mm</quantity>
, wing length: male
<quantity id="A9718A82FF8D050FFE78B354FDF46263" box="[488,604,848,872]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="7.2" metricValueMin="6.0" pageId="2" pageNumber="49" unit="mm" value="6.6" valueMax="7.2" valueMin="6.0">67.2 mm</quantity>
; female
<quantity id="A9718A82FF8D050FFD2BB354FCED6263" box="[699,837,848,872]" metricMagnitude="-3" metricUnit="m" metricValue="7.449999999999999" metricValueMax="7.6" metricValueMin="7.3" pageId="2" pageNumber="49" unit="mm" value="7.449999999999999" valueMax="7.6" valueMin="7.3">7.37.6 mm</quantity>
.
</paragraph>
<paragraph id="6E362767FF8D050FFF59B371FA8D654B" blockId="2.[151,1436,812,1989]" pageId="2" pageNumber="49">
<emphasis id="5CFDFB75FF8D050FFF59B371FEB56286" box="[201,285,885,909]" italics="true" pageId="2" pageNumber="49">Female</emphasis>
. All main differential characters as in male but hairs and bristles shorter. Frons grey and wide. Whitish thin hairs on thorax and head considerably reduced. Dense whitish hairs on mesonotum in male reduced to few short hairs in female. Acrostichals and dorsocentral bristles black, very short and spine-like. Large thoracic bristles as in male but shorter. Wings clearly brownish. Legs long, slender and shinning black, without pennation and flattened hairs. Hairs and bristles on coxae short, ventral spine-like bristles on mid femur very shortened. Abdomen dull light grey with very short and sparse hairs, no hind marginal bristles. Last segments shinning black.
</paragraph>
</subSubSection>
<subSubSection id="269374ECFF8D050FFF59B449FE2B648E" pageId="2" pageNumber="49" type="discussion">
<paragraph id="6E362767FF8D050FFF59B449FE2B648E" blockId="2.[151,1436,812,1989]" pageId="2" pageNumber="49">
<emphasis id="5CFDFB75FF8D050FFF59B449FE95656E" bold="true" box="[201,317,1101,1125]" pageId="2" pageNumber="49">Remarks.</emphasis>
<taxonomicName id="A9895CE4FF8D050FFED5B44AFE5B656E" box="[325,499,1101,1125]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nabucco">
<emphasis id="5CFDFB75FF8D050FFED5B44AFE5B656E" box="[325,499,1101,1125]" italics="true" pageId="2" pageNumber="49">Empis nabucco</emphasis>
</taxonomicName>
is closely related to a small group of species (
<taxonomicName id="A9895CE4FF8D050FFB98B44AFB64656F" authority="Collin" authorityName="Collin" box="[1032,1228,1100,1125]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="syriaca">
<emphasis id="5CFDFB75FF8D050FFB98B44AFBD2656E" box="[1032,1146,1102,1125]" italics="true" pageId="2" pageNumber="49">E. syriaca</emphasis>
Collin
</taxonomicName>
,
<taxonomicName id="A9895CE4FF8D050FFB49B44AFA34656F" authority="Loew" authorityName="Loew" box="[1241,1436,1101,1125]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dedecor">
<emphasis id="5CFDFB75FF8D050FFB49B44AFAFB656E" box="[1241,1363,1101,1125]" italics="true" pageId="2" pageNumber="49">E. dedecor</emphasis>
Loew
</taxonomicName>
and
<taxonomicName id="A9895CE4FF8D050FFF56B475FE286583" authority="Chvala" authorityName="Chvala" box="[198,384,1136,1160]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="5CFDFB75FF8D050FFF56B475FE806583" box="[198,296,1136,1160]" italics="true" pageId="2" pageNumber="49">E. engeli</emphasis>
Chvála
</taxonomicName>
) within
<taxonomicName id="A9895CE4FF8D050FFE4DB475FD316583" box="[477,665,1136,1160]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="haemorrhoica">
<emphasis id="5CFDFB75FF8D050FFE4DB475FD316583" box="[477,665,1136,1160]" italics="true" pageId="2" pageNumber="49">E. haemorrhoica</emphasis>
</taxonomicName>
-group due to the densely haired mesonotum, black legs and halteres, whitish abdominal hairs and metapleural fan (laterotergal bristles).
<taxonomicName id="A9895CE4FF8D050FFC0AB492FBE065A6" box="[922,1096,1173,1197]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nabucco">
<emphasis id="5CFDFB75FF8D050FFC0AB492FBE065A6" box="[922,1096,1173,1197]" italics="true" pageId="2" pageNumber="49">Empis nabucco</emphasis>
</taxonomicName>
mostly resembles
<taxonomicName id="A9895CE4FF8D050FFAB0B492FA3465A6" box="[1312,1436,1173,1197]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dedecor">
<emphasis id="5CFDFB75FF8D050FFAB0B492FA3465A6" box="[1312,1436,1173,1197]" italics="true" pageId="2" pageNumber="49">E. dedecor</emphasis>
</taxonomicName>
and
<taxonomicName id="A9895CE4FF8D050FFF57B4BDFE9265DB" box="[199,314,1209,1232]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="syriaca">
<emphasis id="5CFDFB75FF8D050FFF57B4BDFE9265DB" box="[199,314,1209,1232]" italics="true" pageId="2" pageNumber="49">E. syriaca</emphasis>
</taxonomicName>
having long and dense pale hair-like acrostichals and dorsocentrals, but the latter two species have the humeral bristle not differentiated.
<taxonomicName id="A9895CE4FF8D050FFDAFB4DAFD4265FE" box="[575,746,1245,1269]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nabucco">
<emphasis id="5CFDFB75FF8D050FFDAFB4DAFD4265FE" box="[575,746,1245,1269]" italics="true" pageId="2" pageNumber="49">Empis nabucco</emphasis>
</taxonomicName>
shares distinct humeral bristles as in
<taxonomicName id="A9895CE4FF8D050FFB1DB4DAFB5865FE" box="[1165,1264,1245,1269]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="5CFDFB75FF8D050FFB1DB4DAFB5865FE" box="[1165,1264,1245,1269]" italics="true" pageId="2" pageNumber="49">E. engeli</emphasis>
</taxonomicName>
, but with additional 34 smaller anterior bristles. Female legs of
<taxonomicName id="A9895CE4FF8D050FFD4BB505FCF36413" box="[731,859,1280,1304]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nabucco">
<emphasis id="5CFDFB75FF8D050FFD4BB505FCF36413" box="[731,859,1280,1304]" italics="true" pageId="2" pageNumber="49">E. nabucco</emphasis>
</taxonomicName>
are similar to
<taxonomicName id="A9895CE4FF8D050FFB92B505FBD46413" box="[1026,1148,1280,1304]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dedecor">
<emphasis id="5CFDFB75FF8D050FFB92B505FBD46413" box="[1026,1148,1280,1304]" italics="true" pageId="2" pageNumber="49">E. dedecor</emphasis>
</taxonomicName>
in lacking pennate setae, but the female of
<taxonomicName id="A9895CE4FF8D050FFEFBB522FE446436" box="[363,492,1317,1341]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nabucco">
<emphasis id="5CFDFB75FF8D050FFEFBB522FE446436" box="[363,492,1317,1341]" italics="true" pageId="2" pageNumber="49">E. nabucco</emphasis>
</taxonomicName>
is clearly differentiated from the female of
<taxonomicName id="A9895CE4FF8D050FFC63B522FBC76436" box="[1011,1135,1317,1341]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dedecor">
<emphasis id="5CFDFB75FF8D050FFC63B522FBC76436" box="[1011,1135,1317,1341]" italics="true" pageId="2" pageNumber="49">E. dedecor</emphasis>
</taxonomicName>
by the short and reduced mesonotal hairs, spine-like acrostichals and dorsocentrals and completely grey abdomen except for the last shinning black segments.
</paragraph>
</subSubSection>
<subSubSection id="269374ECFF8D050FFF59B595FBBE6677" pageId="2" pageNumber="49" type="materials_examined">
<paragraph id="6E362767FF8D050FFF59B595FBBE6677" blockId="2.[151,1436,812,1989]" pageId="2" pageNumber="49">
<emphasis id="5CFDFB75FF8D050FFF59B595FEC664A2" bold="true" box="[201,366,1424,1449]" pageId="2" pageNumber="49">
<typeStatus id="B13299C5FF8D050FFF59B595FEAA64A2" box="[201,258,1425,1449]" pageId="2" pageNumber="49">Type</typeStatus>
material
</emphasis>
.
<emphasis id="5CFDFB75FF8D050FFEEFB594FDB664A2" bold="true" box="[383,542,1424,1449]" pageId="2" pageNumber="49">
<typeStatus id="B13299C5FF8D050FFEEFB594FDB664A2" box="[383,542,1424,1449]" pageId="2" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
3 (spn13685):
<emphasis id="5CFDFB75FF8D050FFD41B595FCE664A2" bold="true" box="[721,846,1425,1449]" pageId="2" pageNumber="49">
<collectingCountry id="169E67F7FF8D050FFD41B595FCE064A2" box="[721,840,1425,1449]" name="Turkey" pageId="2" pageNumber="49">TURKEY</collectingCountry>
.
</emphasis>
Ankara, Bala, northwest of Bahçekaradalak village,
<quantity id="A9718A82FF8D050FFF07B5B0FF5864C7" box="[151,240,1460,1484]" metricMagnitude="3" metricUnit="m" metricValue="1.212" pageId="2" pageNumber="49" unit="m" value="1212.0">1212 m</quantity>
,
<date id="1A3701A7FF8D050FFF6DB5B0FED064C7" box="[253,376,1460,1484]" pageId="2" pageNumber="49" value="2011-06-04">04.vi.2011</date>
.
<emphasis id="5CFDFB75FF8D050FFE14B5B0FD8064C6" bold="true" box="[388,552,1460,1485]" pageId="2" pageNumber="49">
<typeStatus id="B13299C5FF8D050FFE14B5B0FD8064C6" box="[388,552,1460,1485]" pageId="2" pageNumber="49" type="paratype">PARATYPES</typeStatus>
</emphasis>
(spn12648, 13649, 13686-13759):
<emphasis id="5CFDFB75FF8D050FFC2CB5B1FB9064C6" bold="true" box="[956,1080,1461,1485]" pageId="2" pageNumber="49">
<collectingCountry id="169E67F7FF8D050FFC2CB5B1FB9B64C6" box="[956,1075,1461,1485]" name="Turkey" pageId="2" pageNumber="49">TURKEY</collectingCountry>
.
</emphasis>
3Ƥ, Ankara, Bala, S of Abazlı village,
<quantity id="A9718A82FF8D050FFF65B5DCFEE664FB" box="[245,334,1496,1520]" metricMagnitude="3" metricUnit="m" metricValue="1.133" pageId="2" pageNumber="49" unit="m" value="1133.0">1133 m</quantity>
,
<date id="1A3701A7FF8D050FFECEB5DCFE7064FB" box="[350,472,1496,1520]" pageId="2" pageNumber="49" value="2011-06-04">04.vi.2011</date>
; 33, 20Ƥ, Ankara, Bala, NW of Bahçekaradalak village,
<quantity id="A9718A82FF8D050FFBEEB5DCFB7164FB" box="[1150,1241,1496,1520]" metricMagnitude="3" metricUnit="m" metricValue="1.212" pageId="2" pageNumber="49" unit="m" value="1212.0">1212 m</quantity>
,
<date id="1A3701A7FF8D050FFB78B5DCFACB64FB" box="[1256,1379,1496,1520]" pageId="2" pageNumber="49" value="2011-06-04">04.vi.2011</date>
; 4Ƥ, Ankara, Bala, N of Bahçekaradalak village,
<quantity id="A9718A82FF8D050FFD1FB5F8FD4F671F" box="[655,743,1532,1556]" metricMagnitude="3" metricUnit="m" metricValue="1.216" pageId="2" pageNumber="49" unit="m" value="1216.0">1216 m</quantity>
,
<date id="1A3701A7FF8D050FFD64B5F8FCC5671F" box="[756,877,1532,1556]" pageId="2" pageNumber="49" value="2011-06-04">04.vi.2011</date>
; 1Ƥ, Yozgat, Akdaġmadeni, W of Bozhüyük village,
<quantity id="A9718A82FF8D050FFF44B624FE846733" box="[212,300,1568,1592]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="2" pageNumber="49" unit="m" value="1785.0">1785 m</quantity>
,
<date id="1A3701A7FF8D050FFEA8B624FE1A6733" box="[312,434,1568,1592]" pageId="2" pageNumber="49" value="2011-06-07">07.vi.2011</date>
; 3Ƥ, Sivas, Şarkışla, S of Hocabey village,
<quantity id="A9718A82FF8D050FFC31B624FC516733" box="[929,1017,1568,1592]" metricMagnitude="3" metricUnit="m" metricValue="1.453" pageId="2" pageNumber="49" unit="m" value="1453.0">1453 m</quantity>
,
<date id="1A3701A7FF8D050FFB95B624FB286733" box="[1029,1152,1568,1592]" pageId="2" pageNumber="49" value="2011-06-07">07.vi.2011</date>
; 13, 3Ƥ, Sivas, Şarkışla, NW of Gaziköy village,
<quantity id="A9718A82FF8D050FFE55B640FD8A6757" box="[453,546,1604,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.389" pageId="2" pageNumber="49" unit="m" value="1389.0">1389 m</quantity>
,
<date id="1A3701A7FF8D050FFDA5B640FD076757" box="[565,687,1604,1628]" pageId="2" pageNumber="49" value="2011-06-07">07.vi.2011</date>
; 23, 8Ƥ, Sivas, Merkez, N of Büyükyüreġil village,
<quantity id="A9718A82FF8D050FFAAAB640FA3F6757" box="[1338,1431,1604,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.441" pageId="2" pageNumber="49" unit="m" value="1441.0">1441 m</quantity>
,
<date id="1A3701A7FF8D050FFF07B66CFEB9678B" box="[151,273,1640,1664]" pageId="2" pageNumber="49" value="2011-06-08">08.vi.2011</date>
; 2Ƥ, Sivas, Merkez, N of Kartalca village,
<quantity id="A9718A82FF8D050FFCB6B66CFC2B678B" box="[806,899,1640,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.366" pageId="2" pageNumber="49" unit="m" value="1366.0">1366 m</quantity>
,
<date id="1A3701A7FF8D050FFC05B66CFBA7678B" box="[917,1039,1640,1664]" pageId="2" pageNumber="49" value="2011-06-08">08.vi.2011</date>
; 4Ƥ, Sivas, Zara, W of Mahmutaġaçiftliġi village,
<quantity id="A9718A82FF8D050FFEE0B688FE6E67AF" box="[368,454,1676,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="2" pageNumber="49" unit="m" value="1412.0">1412 m</quantity>
,
<date id="1A3701A7FF8D050FFE41B688FDE367AF" box="[465,587,1676,1700]" pageId="2" pageNumber="49" value="2011-06-09">09.vi.2011</date>
; 33, Sivas, Hafik, W of Topçuyeniköy village,
<quantity id="A9718A82FF8D050FFBCCB688FB1A67AF" box="[1116,1202,1676,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="2" pageNumber="49" unit="m" value="1438.0">1438 m</quantity>
,
<date id="1A3701A7FF8D050FFB2DB688FA9F67AF" box="[1213,1335,1676,1700]" pageId="2" pageNumber="49" value="2011-06-09">09.vi.2011</date>
; 13, 2Ƥ, Sivas, Hafik, N of Bakımlı village,
<quantity id="A9718A82FF8D050FFDD0B6B4FD3367C3" box="[576,667,1712,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.374" pageId="2" pageNumber="49" unit="m" value="1374.0">1374 m</quantity>
,
<date id="1A3701A7FF8D050FFD3BB6B4FC8E67C3" box="[683,806,1712,1736]" pageId="2" pageNumber="49" value="2011-06-09">09.vi.2011</date>
; 23, Sivas, Merkez, SE of Karayün village,
<quantity id="A9718A82FF8D050FFAABB6B4FA3F67C3" box="[1339,1431,1712,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.4769999999999999" pageId="2" pageNumber="49" unit="m" value="1477.0">1477 m</quantity>
,
<date id="1A3701A7FF8D050FFF07B6D0FEB067E7" box="[151,280,1748,1772]" pageId="2" pageNumber="49" value="2011-09-06">09.06.2011</date>
; 1Ƥ, Sivas, Merkez, E of Yeniboġazkesen village,
<quantity id="A9718A82FF8D050FFCC1B6D0FC0067E7" box="[849,936,1748,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.399" pageId="2" pageNumber="49" unit="m" value="1399.0">1399 m</quantity>
,
<date id="1A3701A7FF8D050FFC24B6D0FB9C67E7" box="[948,1076,1748,1772]" pageId="2" pageNumber="49" value="2011-09-06">09.06.2011</date>
; 1Ƥ, Sivas, Hafik, E of Akpınar village,
<quantity id="A9718A82FF8D050FFF61B6FCFEEF661B" box="[241,327,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="2" pageNumber="49" unit="m" value="1481.0">1481 m</quantity>
,
<date id="1A3701A7FF8D050FFEC3B6FCFE65661B" box="[339,461,1784,1808]" pageId="2" pageNumber="49" value="2011-06-10">10.vi.2011</date>
; 2Ƥ, Sivas, İmranlı, W of Kılıçköy,
<quantity id="A9718A82FF8D050FFCCFB6FCFC1E661B" box="[863,950,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.577" pageId="2" pageNumber="49" unit="m" value="1577.0">1577 m</quantity>
,
<date id="1A3701A7FF8D050FFC51B6FCFB93661B" box="[961,1083,1784,1808]" pageId="2" pageNumber="49" value="2011-06-10">10.vi.2011</date>
; 1Ƥ, Sivas, İmranlı, SE of Yaylacık village,
<quantity id="A9718A82FF8D050FFEA1B718FE21663F" box="[305,393,1820,1844]" metricMagnitude="3" metricUnit="m" metricValue="1.623" pageId="2" pageNumber="49" unit="m" value="1623.0">1623 m</quantity>
,
<date id="1A3701A7FF8D050FFE06B718FDB8663F" box="[406,528,1820,1844]" pageId="2" pageNumber="49" value="2011-06-10">10.vi.2011</date>
; 4Ƥ, Sivas, İmranlı, W of Piredede village,
<quantity id="A9718A82FF8D050FFB95B718FBF5663F" box="[1029,1117,1820,1844]" metricMagnitude="3" metricUnit="m" metricValue="1.7269999999999999" pageId="2" pageNumber="49" unit="m" value="1727.0">1727 m</quantity>
,
<date id="1A3701A7FF8D050FFBFBB718FB4D663F" box="[1131,1253,1820,1844]" pageId="2" pageNumber="49" value="2011-06-10">10.vi.2011</date>
; 13, 2Ƥ, Erzincan, Refahiye, NW of Kandilköy village,
<quantity id="A9718A82FF8D050FFDE1B744FD626653" box="[625,714,1856,1880]" metricMagnitude="3" metricUnit="m" metricValue="2.138" pageId="2" pageNumber="49" unit="m" value="2138.0">2138 m</quantity>
,
<date id="1A3701A7FF8D050FFD47B744FCF86653" box="[727,848,1856,1880]" pageId="2" pageNumber="49" value="2011-06-11">11.vi.2011</date>
; 13, Sivas, İmranlı, S of Toklucak village,
<quantity id="A9718A82FF8D050FFAAFB744FA3F6653" box="[1343,1431,1856,1880]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="49" unit="m" value="1700.0">1700 m</quantity>
,
<date id="1A3701A7FF8D050FFF07B760FEB86677" box="[151,272,1892,1916]" pageId="2" pageNumber="49" value="2011-06-11">11.vi.2011</date>
; 1Ƥ, Erzincan, Refahiye, SE of Yurtbaşı village,
<quantity id="A9718A82FF8D050FFCA5B760FC246677" box="[821,908,1892,1916]" metricMagnitude="3" metricUnit="m" metricValue="1.728" pageId="2" pageNumber="49" unit="m" value="1728.0">1728 m</quantity>
,
<date id="1A3701A7FF8D050FFC08B760FBBA6677" box="[920,1042,1892,1916]" pageId="2" pageNumber="49" value="2011-06-21">21.vi.2011</date>
.
</paragraph>
</subSubSection>
<subSubSection id="269374ECFF8D050FFF59B78CFAF566CF" pageId="2" pageNumber="49" type="etymology">
<paragraph id="6E362767FF8D050FFF59B78CFAF566CF" blockId="2.[151,1436,812,1989]" pageId="2" pageNumber="49">
<emphasis id="5CFDFB75FF8D050FFF59B78CFE3766AA" bold="true" box="[201,415,1928,1953]" pageId="2" pageNumber="49">Derivatio nominis</emphasis>
. This species is named after the
<taxonomicName id="A9895CE4FF8D050FFC9FB78CFCDA66AB" box="[783,882,1928,1952]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="nabucco" status="sp. nov.">Nabucco</taxonomicName>
Gas Pipeline Project, the environmental baseline field studies for which provided the opportunity and financial support for the collection of the
<typeStatus id="B13299C5FF8D050FFB27B7A9FB4F66CE" box="[1207,1255,1965,1989]" pageId="2" pageNumber="49">type</typeStatus>
materials.
</paragraph>
</subSubSection>
</treatment>
</document>