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<document id="63D7A234FEB4FE19F726F77A48CF31EB" ID-DOI="10.11646/zootaxa.3895.3.2" ID-GBIF-Dataset="2074457c-3dbe-4179-9ed8-5c6f57f161dd" ID-ISSN="1175-5326" ID-Zenodo-Dep="287623" ID-ZooBank="44081058-1ACC-4DFE-AF20-096062256345" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461000773178" checkinUser="plazi" docAuthor="Back, Jinwook &amp; Lee, Wonchoel" docDate="2014" docId="DB0B5C67FF98FFE6CBA1FD0419188B7A" docLanguage="en" docName="zt03895p366.pdf" docOrigin="Zootaxa 3895 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wellsopsyllus (Wellsopsyllus) egregius Back &amp; Lee, 2014, sp. nov." docType="treatment" docVersion="6" lastPageNumber="354" masterDocId="2732241FFF9AFFEECB36FFEC183A8C66" masterDocTitle="Two new species of the genus Wellsopsyllus (Copepoda; Harpacticoida; Paramesochridae) from the Yellow Sea" masterLastPageNumber="366" masterPageNumber="346" pageNumber="348" updateTime="1698564508390" updateUser="plazi">
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<mods:title id="C5F4DDA172F4B46CCE6C237DD0B272C0">Two new species of the genus Wellsopsyllus (Copepoda; Harpacticoida; Paramesochridae) from the Yellow Sea</mods:title>
</mods:titleInfo>
<mods:name id="3096FCBFC150F1299C62F42F763B6C97" type="personal">
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<mods:namePart id="BD15B2F5FFE0DCDA7BFB4D403025ED4D">Back, Jinwook</mods:namePart>
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<mods:namePart id="3F1E408CD54E94D9E3D9A4865AB0F599">Lee, Wonchoel</mods:namePart>
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<mods:title id="5E74601AF742067577CBDFF52C4001BE">Zootaxa</mods:title>
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<mods:date id="D1ADF336629879AC485DCBFD3F83AA1D">2014</mods:date>
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<mods:number id="45719B816B62FBA61CD44738BB1BBE27">3895</mods:number>
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<mods:identifier id="DB9DD1FDBAD7E65C90843643D2EEA217" type="DOI">10.11646/zootaxa.3895.3.2</mods:identifier>
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<treatment id="DB0B5C67FF98FFE6CBA1FD0419188B7A" ID-DOI="http://doi.org/10.5281/zenodo.6144387" ID-GBIF-Taxon="119598025" ID-Zenodo-Dep="6144387" LSID="urn:lsid:plazi:treatment:DB0B5C67FF98FFE6CBA1FD0419188B7A" httpUri="http://treatment.plazi.org/id/DB0B5C67FF98FFE6CBA1FD0419188B7A" lastPageId="8" lastPageNumber="354" pageId="2" pageNumber="348">
<subSubSection id="1BB8BEFAFF98FFECCBA1FD0419148F43" pageId="2" pageNumber="348" type="nomenclature">
<paragraph id="531DED71FF98FFECCBA1FD041AF38F65" blockId="2.[151,713,744,805]" box="[151,713,744,771]" pageId="2" pageNumber="348">
<heading id="08555A1DFF98FFECCBA1FD041AF38F65" bold="true" box="[151,713,744,771]" fontSize="11" level="1" pageId="2" pageNumber="348" reason="1">
<emphasis id="61D63163FF98FFECCBA1FD041AF38F65" bold="true" box="[151,713,744,771]" pageId="2" pageNumber="348">
<taxonomicName id="94A296F2FF98FFECCBA1FD041A598F64" box="[151,611,744,771]" class="Maxillopoda" family="Paramesochridae" genus="Wellsopsyllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="348" phylum="Arthropoda" rank="species" species="egregius" status="sp. nov." subGenus="Wellsopsyllus">
<emphasis id="61D63163FF98FFECCBA1FD0419038F64" bold="true" box="[151,313,744,770]" italics="true" pageId="2" pageNumber="348">Wellsopsyllus</emphasis>
(
<emphasis id="61D63163FF98FFECCA7CFD0419D68F64" bold="true" box="[330,492,744,770]" italics="true" pageId="2" pageNumber="348">Wellsopsyllus</emphasis>
)
<emphasis id="61D63163FF98FFECCACBFD041A598F64" bold="true" box="[509,611,744,770]" italics="true" pageId="2" pageNumber="348">egregius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AE58C18FF98FFECC95DFD051AF38F65" box="[619,713,745,771]" pageId="2" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="531DED71FF98FFECCBA1FCE019148F43" blockId="2.[151,713,744,805]" box="[151,302,780,805]" pageId="2" pageNumber="348">(Figures 26)</paragraph>
</subSubSection>
<subSubSection id="1BB8BEFAFF98FFECCBA1FCB8198E8F9A" pageId="2" pageNumber="348" type="materials_examined">
<paragraph id="531DED71FF98FFECCBA1FCB81BC58FE9" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">
<emphasis id="61D63163FF98FFECCBA1FCB8190D8F0B" bold="true" box="[151,311,852,877]" pageId="2" pageNumber="348">
<typeStatus id="8C1953D3FF98FFECCBA1FCB818E88F0B" box="[151,210,852,877]" pageId="2" pageNumber="348">Type</typeStatus>
locality.
</emphasis>
Gosapo Beach on the Byeonsan peninsula, on the west (Yellow Sea) coast of
<collectingCountry id="2BB5ADE1FF98FFECCF8FFCB91D3B8F0A" box="[1209,1281,853,876]" name="South Korea" pageId="2" pageNumber="348">Korea</collectingCountry>
. Coordinates:
<geoCoordinate id="36968BB6FF98FFECCBA1FC9B190B8FF6" box="[151,305,887,912]" direction="north" orientation="latitude" pageId="2" pageNumber="348" precision="15" value="35.66417">35° 39' 51 ''N</geoCoordinate>
, 126° 30' 35'E (
<figureCitation id="CB99F1F4FF98FFECCAD1FC9B1A7C8FF6" box="[487,582,887,912]" captionStart="FIGURE 1" captionStartId="1.[151,250,1704,1726]" captionTargetBox="[183,1399,923,1671]" captionTargetId="figure@1.[176,1411,916,1682]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the family Paramesochridae in Korea." httpUri="https://zenodo.org/record/287624/files/figure.png" pageId="2" pageNumber="348">Figure 1</figureCitation>
). Sediment
<typeStatus id="8C1953D3FF98FFECC9F9FC941B3E8FF6" box="[719,772,888,912]" pageId="2" pageNumber="348">type</typeStatus>
: sand. Salinity 33 psu.
</paragraph>
<paragraph id="531DED71FF98FFECCBF1FC70198E8F9A" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">
<emphasis id="61D63163FF98FFECCBF1FC70198E8FD3" bold="true" box="[199,436,924,949]" pageId="2" pageNumber="348">Material examined.</emphasis>
<typeStatus id="8C1953D3FF98FFECCA89FC701A108FD3" box="[447,554,924,949]" pageId="2" pageNumber="348" type="holotype">Holotype</typeStatus>
, 1♀ (
<collectionCode id="35B375B4FF98FFECC959FC701AF48FD2" box="[623,718,924,948]" pageId="2" pageNumber="348">MABIK</collectionCode>
CR00218007) dissected on 7 slides.
<typeStatus id="8C1953D3FF98FFECCFBEFC711CC38FD3" box="[1160,1273,925,949]" pageId="2" pageNumber="348" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="45A426F8FF98FFECCE31FC701D108FD2" box="[1287,1322,924,948]" pageId="2" pageNumber="348" type="generic">1♂</specimenCount>
(
<collectionCode id="35B375B4FF98FFECCE0BFC701DA68FD2" box="[1341,1436,924,948]" pageId="2" pageNumber="348">MABIK</collectionCode>
CR00218008) dissected on 3 slides, 5 ♀♀ (
<collectionCode id="35B375B4FF98FFECC9AFFC2C1AC28FBE" box="[665,760,960,984]" pageId="2" pageNumber="348">MABIK</collectionCode>
CR00218009) and 2 ♂♂ (
<collectionCode id="35B375B4FF98FFECCF03FC2C1CAE8FBE" box="[1077,1172,960,984]" pageId="2" pageNumber="348">MABIK</collectionCode>
CR00218010) in 70% alcohol.
<date id="271CCBB1FF98FFECCBC1FC08198B8F9A" box="[247,433,996,1021]" pageId="2" pageNumber="348" value="2010-10-03">October 3, 2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="1BB8BEFAFF98FFE6CBF1FBEB1C5F8AB5" lastPageId="8" lastPageNumber="354" pageId="2" pageNumber="348" type="description">
<paragraph id="531DED71FF98FFECCBF1FBEB1C8C88EB" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">
<emphasis id="61D63163FF98FFECCBF1FBEB19EB8846" bold="true" box="[199,465,1031,1056]" pageId="2" pageNumber="348">Description of female.</emphasis>
Total body (Figure 2AB) length, 462 µm (n = 6, mean = 460 µm), measured from anterior tip of rostrum of cephalic shield to posterior margin of caudal rami (Figure 2AB)); largest width measured at posterior margin of cephalic shield: 51 µm. Body cylindrical, depressed dorsoventrally; urosome gradually tapering posteriorly (smallest width of last somite: 21 µm) with a weak hyaline frill.
</paragraph>
<paragraph id="531DED71FF98FFECCBF1FB741941889E" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">Rostrum (Figure 2A), small, fused to cephalothorax, without sensilla. Cephalothorax (Figure 2AB), bellshaped; pleural areas weakly developed and posterolateral angles rounded; posterior margin, smooth, with a distinct hyaline frill.</paragraph>
<paragraph id="531DED71FF98FFECCBF1FAE81A8B89CA" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">
Genital somite and first abdominal somite (Figure 2AB), completely fused forming a genital double-somite; genital field located mid-ventrally approximately halfway along the length of the genital double-somite. Copulatory pore (
<figureCitation id="CB99F1F4FF98FFECCA6DFAA019818903" box="[347,443,1356,1381]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[189,1415,224,1921]" captionTargetId="figure@6.[168,1419,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Wellsopsyllus (Wellsopsyllus) egregius sp. nov.: (A) female P 5; (B) female P 6 and genital field; (C) male P 5; (D) male P 6." httpUri="https://zenodo.org/record/287626/files/figure.png" pageId="2" pageNumber="348">Figure 5</figureCitation>
B) located near posterior border of genital field, covered by a small process and enclosed within a depression. P6 (
<figureCitation id="CB99F1F4FF98FFECCAFDFA831A0E89EE" box="[459,564,1391,1416]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[189,1415,224,1921]" captionTargetId="figure@6.[168,1419,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Wellsopsyllus (Wellsopsyllus) egregius sp. nov.: (A) female P 5; (B) female P 6 and genital field; (C) male P 5; (D) male P 6." httpUri="https://zenodo.org/record/287626/files/figure.png" pageId="2" pageNumber="348">Figure 5</figureCitation>
B) represented by a small process with 1 bare and 1 unipinnate setae. Pseudoperulum, weakly developed (Figure 2A).
</paragraph>
<paragraph id="531DED71FF98FFECCBF1FA5419418A05" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">CR (Figure 2D1D2), juxtaposed, approximately 2.8 times longer than greatest width, posterior inner margin acute, distal margin rounded inwardly, and with a small process near seta V. Each ramus armed with 6 setae: seta I bare; setae II bare; seta III blunt, ornamented with stout spinules; seta IV pinnate; seta V blunt, pinnate; seta VI not observed and probably represented by a small process; seta VII bi-articulate at base and arising sub-distally from inner dorsal surface.</paragraph>
<paragraph id="531DED71FF98FFECCBF1F9831B008B71" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">A1 (Figure 2C), 8-segmented, short, robust. Segment-1 longest with a blunt process; segment-4 with a subcylindrical process with 1 long slender seta fused basally to aesthetasc; segment-6 with 1 slender bare seta arising from a ventral sub-cylindrical process. Armature formula: 1-[1 pinnate], 2-[7 bare + 2 pinnate], 3-[2 bare + 4 pinnate], 4-[1 bare + (1 + ae)], 5-[2 bare], 6-[2 bare], 7-[3 bare], 8-[4 bare + acrothek]; apical acrothek consisting of a well-developed aesthetasc fused basally to 2 slender setae.</paragraph>
<paragraph id="531DED71FF98FFECCBF1F8CF195E8B89" blockId="2.[151,1437,852,2031]" pageId="2" pageNumber="348">
A2 (
<figureCitation id="CB99F1F4FF98FFECCBC1F8CE196C8B5D" box="[247,342,1826,1851]" captionStart="FIGURE 3" captionStartId="4.[151,250,1949,1971]" captionTargetBox="[176,1412,204,1925]" captionTargetId="figure@4.[151,1436,193,1928]" captionTargetPageId="4" captionText="FIGURE 3. Wellsopsyllus (Wellsopsyllus) egregius sp. nov., female: (A) Antenna; (B) B 1: gnathobase of the mandible; B 2: mandibular palp; (C) maxilla; (D) maxilliped." httpUri="https://zenodo.org/record/287625/files/figure.png" pageId="2" pageNumber="348">Figure 3</figureCitation>
A), 4-segmented, comprising coxa, basis, 1-segmented exp, and 2-segmented free enp; coxa small and bare; basis approximately 1.7 times longer than wide, without any surface ornamentation; proximal endopodal segment with 1 pinnate abexopodal seta; distal endopodal segment ornamented with 2 rows of spinules on surface and armed with 1 lateral bare seta, 2 bare spine-like setae sub-distally, 5 geniculate distal setae, one of which was fused basally to long slender seta; exp 1-segmented, with 2 bare setae laterally, 1 bare seta at distal corner, and 1 bifid seta apically.
</paragraph>
<paragraph id="531DED71FF99FFEDCBA1F84D1A668BB0" blockId="3.[151,1436,1953,2009]" pageId="3" pageNumber="349">
<emphasis id="61D63163FF99FFEDCBA1F84D192E8BD0" bold="true" box="[151,276,1953,1975]" pageId="3" pageNumber="349">FIGURE 2.</emphasis>
<taxonomicName id="94A296F2FF99FFEDCA2AF84D1A9F8BD1" box="[284,677,1953,1975]" class="Maxillopoda" family="Paramesochridae" genus="Wellsopsyllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="349" phylum="Arthropoda" rank="species" species="egregius" status="sp. nov." subGenus="Wellsopsyllus">
<emphasis id="61D63163FF99FFEDCA2AF84D199F8BD1" box="[284,421,1953,1975]" italics="true" pageId="3" pageNumber="349">Wellsopsyllus</emphasis>
(
<emphasis id="61D63163FF99FFEDCA80F84D1A058BD1" box="[438,575,1953,1975]" italics="true" pageId="3" pageNumber="349">Wellsopsyllus</emphasis>
)
<emphasis id="61D63163FF99FFEDC979F84E1A9F8BD1" box="[591,677,1954,1975]" italics="true" pageId="3" pageNumber="349">egregius</emphasis>
</taxonomicName>
<emphasis id="61D63163FF99FFEDC998F84D1B3A8BD0" bold="true" box="[686,768,1953,1974]" pageId="3" pageNumber="349">
<taxonomicNameLabel id="7AE58C18FF99FFEDC998F84D1B3A8BD0" box="[686,768,1953,1974]" pageId="3" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) Habitus, dorsal; (B) habitus, lateral; (C) antennule; (D) D1: left caudal rami, dorsal; D2: seta VII.
</paragraph>
<caption id="07DDBDF9FF9EFFEACBA1F8711A528BB4" httpUri="https://zenodo.org/record/287625/files/figure.png" pageId="4" pageNumber="350" targetBox="[176,1412,204,1925]" targetPageId="4">
<paragraph id="531DED71FF9EFFEACBA1F8711A528BB4" blockId="4.[151,1436,1949,2002]" pageId="4" pageNumber="350">
<emphasis id="61D63163FF9EFFEACBA1F871192E8BD4" bold="true" box="[151,276,1949,1971]" pageId="4" pageNumber="350">FIGURE 3.</emphasis>
<taxonomicName id="94A296F2FF9EFFEACA2BF8711A9C8BD5" box="[285,678,1949,1971]" class="Maxillopoda" family="Paramesochridae" genus="Wellsopsyllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="350" phylum="Arthropoda" rank="species" species="egregius" status="sp. nov." subGenus="Wellsopsyllus">
<emphasis id="61D63163FF9EFFEACA2BF871199C8BD5" box="[285,422,1949,1971]" italics="true" pageId="4" pageNumber="350">Wellsopsyllus</emphasis>
(
<emphasis id="61D63163FF9EFFEACA80F8711A058BD5" box="[438,575,1949,1971]" italics="true" pageId="4" pageNumber="350">Wellsopsyllus</emphasis>
)
<emphasis id="61D63163FF9EFFEAC966F8721A9C8BD5" box="[592,678,1950,1971]" italics="true" pageId="4" pageNumber="350">egregius</emphasis>
</taxonomicName>
<emphasis id="61D63163FF9EFFEAC998F8721B3B8BD5" bold="true" box="[686,769,1950,1971]" pageId="4" pageNumber="350">
<taxonomicNameLabel id="7AE58C18FF9EFFEAC998F8721B3B8BD5" box="[686,769,1950,1971]" pageId="4" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) Antenna; (B) B1: gnathobase of the mandible; B2: mandibular palp; (C) maxilla; (D) maxilliped.
</paragraph>
</caption>
<paragraph id="531DED71FF9FFFEBCBA1F84D1CAB8BD1" blockId="5.[151,1169,1953,1975]" box="[151,1169,1953,1975]" pageId="5" pageNumber="351">
<emphasis id="61D63163FF9FFFEBCBA1F84D19288BD0" bold="true" box="[151,274,1953,1975]" pageId="5" pageNumber="351">FIGURE 4.</emphasis>
<taxonomicName id="94A296F2FF9FFFEBCA2FF84D1AA48BD1" box="[281,670,1953,1975]" class="Maxillopoda" family="Paramesochridae" genus="Wellsopsyllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="351" phylum="Arthropoda" rank="species" species="egregius" status="sp. nov." subGenus="Wellsopsyllus">
<emphasis id="61D63163FF9FFFEBCA2FF84D19988BD1" box="[281,418,1953,1975]" italics="true" pageId="5" pageNumber="351">Wellsopsyllus</emphasis>
(
<emphasis id="61D63163FF9FFFEBCA86F84D1A038BD1" box="[432,569,1953,1975]" italics="true" pageId="5" pageNumber="351">Wellsopsyllus</emphasis>
)
<emphasis id="61D63163FF9FFFEBC97EF84E1AA48BD1" box="[584,670,1954,1975]" italics="true" pageId="5" pageNumber="351">egregius</emphasis>
</taxonomicName>
<emphasis id="61D63163FF9FFFEBC992F84D1ACF8BD0" bold="true" box="[676,757,1953,1974]" pageId="5" pageNumber="351">
<taxonomicNameLabel id="7AE58C18FF9FFFEBC992F84D1ACF8BD0" box="[676,757,1953,1974]" pageId="5" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) P1; (B) P2; (C) P3; (D) P4.
</paragraph>
<caption id="07DDBDF9FF9CFFE8CBA1F84D18CA8BB0" httpUri="https://zenodo.org/record/287626/files/figure.png" pageId="6" pageNumber="352" targetBox="[189,1415,224,1921]" targetPageId="6">
<paragraph id="531DED71FF9CFFE8CBA1F84D18CA8BB0" blockId="6.[151,1436,1953,2006]" pageId="6" pageNumber="352">
<emphasis id="61D63163FF9CFFE8CBA1F84D19298BD0" bold="true" box="[151,275,1953,1975]" pageId="6" pageNumber="352">FIGURE 5.</emphasis>
<taxonomicName id="94A296F2FF9CFFE8CA2CF84D1A9B8BD1" box="[282,673,1953,1975]" class="Maxillopoda" family="Paramesochridae" genus="Wellsopsyllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="egregius" status="sp. nov." subGenus="Wellsopsyllus">
<emphasis id="61D63163FF9CFFE8CA2CF84D19998BD1" box="[282,419,1953,1975]" italics="true" pageId="6" pageNumber="352">Wellsopsyllus</emphasis>
(
<emphasis id="61D63163FF9CFFE8CA84F84D1A018BD1" box="[434,571,1953,1975]" italics="true" pageId="6" pageNumber="352">Wellsopsyllus</emphasis>
)
<emphasis id="61D63163FF9CFFE8C97DF84E1A9B8BD1" box="[587,673,1954,1975]" italics="true" pageId="6" pageNumber="352">egregius</emphasis>
</taxonomicName>
<emphasis id="61D63163FF9CFFE8C99EF84D1AC38BD0" bold="true" box="[680,761,1953,1974]" pageId="6" pageNumber="352">
<taxonomicNameLabel id="7AE58C18FF9CFFE8C99EF84D1AC38BD0" box="[680,761,1953,1974]" pageId="6" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) female P5; (B) female P6 and genital field; (C) male P5; (D) male P6.
</paragraph>
</caption>
<paragraph id="531DED71FF9DFFE9CBA1F84D1A788BBE" blockId="7.[151,1436,1953,2012]" pageId="7" pageNumber="353">
<emphasis id="61D63163FF9DFFE9CBA1F84D19288BD0" bold="true" box="[151,274,1953,1975]" pageId="7" pageNumber="353">FIGURE 6.</emphasis>
<taxonomicName id="94A296F2FF9DFFE9CA2FF84D1AA78BD1" box="[281,669,1953,1975]" class="Maxillopoda" family="Paramesochridae" genus="Wellsopsyllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="353" phylum="Arthropoda" rank="species" species="egregius" status="sp. nov." subGenus="Wellsopsyllus">
<emphasis id="61D63163FF9DFFE9CA2FF84D19988BD1" box="[281,418,1953,1975]" italics="true" pageId="7" pageNumber="353">Wellsopsyllus</emphasis>
(
<emphasis id="61D63163FF9DFFE9CA86F84D1A038BD1" box="[432,569,1953,1975]" italics="true" pageId="7" pageNumber="353">Wellsopsyllus</emphasis>
)
<emphasis id="61D63163FF9DFFE9C971F84E1AA78BD1" box="[583,669,1954,1975]" italics="true" pageId="7" pageNumber="353">egregius</emphasis>
</taxonomicName>
<emphasis id="61D63163FF9DFFE9C992F84D1ACE8BD0" bold="true" box="[676,756,1953,1974]" pageId="7" pageNumber="353">
<taxonomicNameLabel id="7AE58C18FF9DFFE9C992F84D1ACE8BD0" box="[676,756,1953,1974]" pageId="7" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) male habitus, dorsal; (B) male antennule; B1: segments 1 to 4; B2: segments 5 to 7; (C) female maxillule.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FF7B1C928D79" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">
Mandible (
<figureCitation id="CB99F1F4FF92FFE6CA77FF7B199A8CD6" box="[321,416,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1949,1971]" captionTargetBox="[176,1412,204,1925]" captionTargetId="figure@4.[151,1436,193,1928]" captionTargetPageId="4" captionText="FIGURE 3. Wellsopsyllus (Wellsopsyllus) egregius sp. nov., female: (A) Antenna; (B) B 1: gnathobase of the mandible; B 2: mandibular palp; (C) maxilla; (D) maxilliped." httpUri="https://zenodo.org/record/287625/files/figure.png" pageId="8" pageNumber="354">Figure 3</figureCitation>
B1B2), coxa with well-developed gnathobase bearing 1 small bare seta at the dorsal corner and 7 major blunt overlapping teeth; palp biramous, comprising basis, 1-segmented exp and 2-segmented enp; basis widening distally, without armature; exp with 2 distal bare setae; enp elongated, approximately 3 times as long as exp; enp-1 with 2 bare sub-distal setae; enp-2 with 6 basally fused bare setae at apex.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FEC61B438DC9" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">Maxillule (Figure 6C), praecoxal arthrite well developed, with 6 spines, 1 bare seta at distal corner, 1 slender seta on lateral margin, and 2 juxtaposed slender setae on anterior surface; coxa with cylindrical endite bearing 4 bare setae; basis cylindrical; endites fused, with 1 pinnate and 5 bare setae; exp 1-segmented, small, with 2 bare setae; enp 1-segmented, elongate, with 1 pinnate and 3 bare setae.</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FE5619A78E59" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">
Maxilla (
<figureCitation id="CB99F1F4FF92FFE6CA1BFE5619B48DB5" box="[301,398,442,467]" captionStart="FIGURE 3" captionStartId="4.[151,250,1949,1971]" captionTargetBox="[176,1412,204,1925]" captionTargetId="figure@4.[151,1436,193,1928]" captionTargetPageId="4" captionText="FIGURE 3. Wellsopsyllus (Wellsopsyllus) egregius sp. nov., female: (A) Antenna; (B) B 1: gnathobase of the mandible; B 2: mandibular palp; (C) maxilla; (D) maxilliped." httpUri="https://zenodo.org/record/287625/files/figure.png" pageId="8" pageNumber="354">Figure 3</figureCitation>
C), syncoxa with 3 cylindrical endites; praecoxal and proximal endites with 1 pinnate and 1 bare setae; distal coxal endite with 1 bare and 2 pinnate setae; allobasis with 1 strong claw-like spine, 1 pinnate and 1 bare setae distally, and 1 pinnate seta near base of enp; enp 1-segmented, with 1 claw-like spine, 3 pinnate setae, and 1 bare seta distally.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FDA61C688EE1" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">
Maxilliped (
<figureCitation id="CB99F1F4FF92FFE6CA62FDA6198D8E05" box="[340,439,586,611]" captionStart="FIGURE 3" captionStartId="4.[151,250,1949,1971]" captionTargetBox="[176,1412,204,1925]" captionTargetId="figure@4.[151,1436,193,1928]" captionTargetPageId="4" captionText="FIGURE 3. Wellsopsyllus (Wellsopsyllus) egregius sp. nov., female: (A) Antenna; (B) B 1: gnathobase of the mandible; B 2: mandibular palp; (C) maxilla; (D) maxilliped." httpUri="https://zenodo.org/record/287625/files/figure.png" pageId="8" pageNumber="354">Figure 3</figureCitation>
D) 4-segmented, comprising syncoxa, basis, and 2-segmented enp; syncoxa and basis bare; enp-1 with 1 stout seta distally; enp-2 with 2 lateral bare and 2 geniculate setae.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FD7F1B688F5D" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">P1 (Figure 4A), basis with 1 outer and 1 inner bare setae; exp-1 approximately 1.8 times longer than exp-2, the former with 1 long pinnate seta near outer distal corner and ornamented with a row of long spinules horizontally in middle of segment; exp-3 with 4 long pinnate setae distally; enp prehensile; enp-1 elongate, about 1.2 times longer than exp, bare, inner distal corner forming a small projection; enp-2 small, with 1 geniculate seta apically and ornamented with a row of spinules around outer distal margin.</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FCAB1BE88FC1" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">P2P3 (Figure 4BC), basis with 1 outer bare seta and ornamented with 1 pore and 1 row of spinules in middle. exp 3-segmented; exp-1 and 2 with 1 outer spine; exp-3 with 1 spine and 1 geniculate seta; enp 1- segmented, small, without ornamentation, represented by a small process.</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FC5F1AD38874" blockId="8.[151,1436,151,1043]" pageId="8" pageNumber="354">P4 (Figure 4D), basis with 1 outer bare seta and ornamented with 1 pore and 1 row of spinules in middle. exp 3-segmented; exp-1 and 2 with 1 outer spine at distal corner; exp-3 with 1 naked outer spine and 1 geniculate seta; enp 1-segmented, with 1 stout unipinnate apical seta.</paragraph>
<paragraph id="531DED71FF92FFE6CBA9FBC21C2688A7" pageId="8" pageNumber="354">
<table id="21A21FD1FF920011CBA9FBC21C6888A7" box="[159,1106,1070,1217]" gridcols="3" gridrows="4" pageId="8" pageNumber="354">
<tr id="ED92EF33FF920011CBA9FBC21C688822" box="[159,1106,1070,1092]" gridrow="0" pageId="8" pageNumber="354">
<th id="AE43864FFF920011CBA9FBC218838822" box="[159,185,1070,1092]" gridcol="0" gridrow="0" pageId="8" pageNumber="354">?</th>
<th id="AE43864FFF920011C97CFBC21AA38822" box="[586,665,1070,1092]" gridcol="1" gridrow="0" pageId="8" pageNumber="354">Exopod</th>
<th id="AE43864FFF920011C8C0FBC21C688822" box="[1014,1106,1070,1092]" gridcol="2" gridrow="0" pageId="8" pageNumber="354">Endopod</th>
</tr>
<tr id="ED92EF33FF920011CBA9FBB41C688808" box="[159,1106,1112,1134]" gridrow="1" pageId="8" pageNumber="354">
<th id="AE43864FFF920011CBA9FBB418838808" box="[159,185,1112,1134]" gridcol="0" gridrow="1" pageId="8" pageNumber="354">P2</th>
<td id="AE43864FFF920011C97CFBB41AA38808" box="[586,665,1112,1134]" gridcol="1" gridrow="1" pageId="8" pageNumber="354">0.0.0 11</td>
<td id="AE43864FFF920011C8C0FBB41C688808" box="[1014,1106,1112,1134]" gridcol="2" gridrow="1" pageId="8" pageNumber="354">0</td>
</tr>
<tr id="ED92EF33FF920011CBA9FB6D1C6888F1" box="[159,1106,1153,1175]" gridrow="2" pageId="8" pageNumber="354">
<th id="AE43864FFF920011CBA9FB6D188388F1" box="[159,185,1153,1175]" gridcol="0" gridrow="2" pageId="8" pageNumber="354">P3</th>
<td id="AE43864FFF920011C97CFB6D1AA388F1" box="[586,665,1153,1175]" gridcol="1" gridrow="2" pageId="8" pageNumber="354">0.0.0 11</td>
<td id="AE43864FFF920011C8C0FB6D1C6888F1" box="[1014,1106,1153,1175]" gridcol="2" gridrow="2" pageId="8" pageNumber="354">0</td>
</tr>
<tr id="ED92EF33FF920011CBA9FB471C6888A7" box="[159,1106,1195,1217]" gridrow="3" pageId="8" pageNumber="354">
<th id="AE43864FFF920011CBA9FB47188388A7" box="[159,185,1195,1217]" gridcol="0" gridrow="3" pageId="8" pageNumber="354">P4</th>
<td id="AE43864FFF920011C97CFB471AA388A7" box="[586,665,1195,1217]" gridcol="1" gridrow="3" pageId="8" pageNumber="354">0.0.0 11</td>
<td id="AE43864FFF920011C8C0FB471C6888A7" box="[1014,1106,1195,1217]" gridcol="2" gridrow="3" pageId="8" pageNumber="354">0 10</td>
</tr>
</table>
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FAE51BE8890E" blockId="8.[151,1436,1288,1820]" pageId="8" pageNumber="354">
Benp of P5 (
<figureCitation id="CB99F1F4FF92FFE6CA54FAE419FC8947" box="[354,454,1288,1313]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[189,1415,224,1921]" captionTargetId="figure@6.[168,1419,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Wellsopsyllus (Wellsopsyllus) egregius sp. nov.: (A) female P 5; (B) female P 6 and genital field; (C) male P 5; (D) male P 6." httpUri="https://zenodo.org/record/287626/files/figure.png" pageId="8" pageNumber="354">Figure 5</figureCitation>
A) confluent, with 1 naked basal seta, an acute outer process, and one pore; fused endopodal lobes with a small bilobed process in middle; each endopodal lobe with 2 bare setae; exp clearly separated from benp, small, quadrilateral, with 2 bare and 1 pinnate setae.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1FA981CFD899F" blockId="8.[151,1436,1288,1820]" pageId="8" pageNumber="354">
<emphasis id="61D63163FF92FFE6CBF1FA98198089EB" bold="true" box="[199,442,1396,1421]" pageId="8" pageNumber="354">Description of male.</emphasis>
Body (Figure 6A), slightly smaller and more slender than female, body length 385 µm (measured from anterior margin of cephalic shield to posterior margin of caudal rami: n = 3, mean = 381 µm), largest width measured at posterior margin of cephalic shield: 44 µm; general body shape and ornamentation as in female except for separation of genital somite; additional sexual dimorphism in A1, P5, and P6.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1F9E81C6C8A01" blockId="8.[151,1436,1288,1820]" pageId="8" pageNumber="354">A1 (Figure 6B1B2), 7-segmented, subchirocer; armature formula: 1[1 pinnate], 2[8 bare + 2 pinnate], 3[5 bare + 1 pinnate], 4[1 bare + 1 pinnate], 5[6 bare + 1 modified stout + (1 + ae)], 6[4 bare], 7[6 bare + acrothek], apical acrothek consisting of small aesthetasc fused basally to 2 bare setae.</paragraph>
<paragraph id="531DED71FF92FFE6CBF1F99F1BB38AC9" blockId="8.[151,1436,1288,1820]" pageId="8" pageNumber="354">
P5 (
<figureCitation id="CB99F1F4FF92FFE6CBC2F99F196F8AEA" box="[244,341,1651,1676]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[189,1415,224,1921]" captionTargetId="figure@6.[168,1419,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Wellsopsyllus (Wellsopsyllus) egregius sp. nov.: (A) female P 5; (B) female P 6 and genital field; (C) male P 5; (D) male P 6." httpUri="https://zenodo.org/record/287626/files/figure.png" pageId="8" pageNumber="354">Figure 5</figureCitation>
C), benp of P5 confluent, with 1 bare basal seta, 1 pore, and acute outer process; endopodal lobe flat without armature; exp discrete, with 2 bare and 1 pinnate setae.
</paragraph>
<paragraph id="531DED71FF92FFE6CBF1F9571C5F8AB5" blockId="8.[151,1436,1288,1820]" box="[199,1125,1723,1748]" pageId="8" pageNumber="354">
P6 (
<figureCitation id="CB99F1F4FF92FFE6CBC5F95719698AB2" box="[243,339,1723,1748]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[189,1415,224,1921]" captionTargetId="figure@6.[168,1419,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Wellsopsyllus (Wellsopsyllus) egregius sp. nov.: (A) female P 5; (B) female P 6 and genital field; (C) male P 5; (D) male P 6." httpUri="https://zenodo.org/record/287626/files/figure.png" pageId="8" pageNumber="354">Figure 5</figureCitation>
D), pair of P6 asymmetrical, not confluent; each P6 with 1 naked seta.
</paragraph>
</subSubSection>
<subSubSection id="1BB8BEFAFF92FFE6CBF1F93219188B7A" pageId="8" pageNumber="354" type="etymology">
<paragraph id="531DED71FF92FFE6CBF1F93219188B7A" blockId="8.[151,1436,1288,1820]" pageId="8" pageNumber="354">
<emphasis id="61D63163FF92FFE6CBF1F93219748A91" bold="true" box="[199,334,1758,1783]" pageId="8" pageNumber="354">Etymology.</emphasis>
The specific name
<emphasis id="61D63163FF92FFE6C90BF9331A9C8A90" box="[573,678,1759,1782]" italics="true" pageId="8" pageNumber="354">egregius</emphasis>
means shocking in Latin and refers to the unusual shape of the caudal rami.
</paragraph>
</subSubSection>
</treatment>
</document>