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<document id="9921C7B5623332D3A68278C5D303736A" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA2477BD2EFF37DFE41882A4DF" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma occidentalis Henderson" docType="treatment" docVersion="7" lastPageNumber="106" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="106" updateTime="1698765326825" updateUser="ExternalLinkService">
<mods:mods id="16FE1A8CCA192F6FB8C92EAC6B6FEB1E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E454EB5B119CC80489AE3216719D0015">
<mods:title id="14B69C4C33BFBB4867EF0162577A8625">A revision of Geonoma (Arecaceae)</mods:title>
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<mods:name id="D31B676F2B87A17672F7DEF9630D6436" type="personal">
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<mods:namePart id="C403A135643DE3C0260FBDA9A6627380">Henderson, Andrew</mods:namePart>
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<mods:typeOfResource id="A86C0ADB38F2BB2E9ABA70CC1105E8B8">text</mods:typeOfResource>
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<mods:title id="F21B5F59E29809D17FB3E9E1D5B0267C">Phytotaxa</mods:title>
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<mods:part id="5CC1A942EC708B1273C59D1960E8B85C">
<mods:date id="7F6CB91E8FE4AE6C323DA4653F2B8FC0">2011</mods:date>
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<mods:number id="ABC05C8C157FA014748B4DB9D0DA7D70">2011-02-18</mods:number>
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<mods:number id="1510601B6D333AA7F7CC01BE5FF213E5">17</mods:number>
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<mods:identifier id="F24C937889D469735FFB2FD18A4439BB" type="DOI">10.11646/phytotaxa.17.1</mods:identifier>
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<treatment id="FA4887FA2477BD2EFF37DFE41882A4DF" ID-DOI="http://doi.org/10.5281/zenodo.5609080" ID-GBIF-Taxon="160342417" ID-Zenodo-Dep="5609080" LSID="urn:lsid:plazi:treatment:FA4887FA2477BD2EFF37DFE41882A4DF" httpUri="http://treatment.plazi.org/id/FA4887FA2477BD2EFF37DFE41882A4DF" lastPageNumber="106" pageId="105" pageNumber="106">
<subSubSection id="3AFB65672477BD2EFF37DFE418D6A2C1" box="[151,1001,421,448]" pageId="105" pageNumber="106" type="nomenclature">
<paragraph id="725E36EC2477BD2EFF37DFE418D6A2C1" blockId="105.[151,1436,421,548]" box="[151,1001,421,448]" pageId="105" pageNumber="106">
<heading id="291681802477BD2EFF37DFE418D6A2C1" bold="true" box="[151,1001,421,448]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="6">
<emphasis id="4095EAFE2477BD2EFF37DFE418D6A2C1" bold="true" box="[151,1001,421,448]" pageId="105" pageNumber="106">
41.
<taxonomicName id="B5E14D6F2477BD2EFF63DFE4183CA2C1" ID-CoL="3FQN9" authority="(Henderson) Henderson" authorityName="Henderson" baseAuthorityName="Henderson" box="[195,771,421,448]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="105" pageNumber="106" phylum="Tracheophyta" rank="species" species="occidentalis">
<emphasis id="4095EAFE2477BD2EFF63DFE41AF1A2C1" bold="true" box="[195,462,421,448]" italics="true" pageId="105" pageNumber="106">Geonoma occidentalis</emphasis>
(Henderson) Henderson
</taxonomicName>
,
<emphasis id="4095EAFE2477BD2EFCB0DFE718D6A2C1" bold="true" box="[784,1001,421,448]" italics="true" pageId="105" pageNumber="106">comb. &amp; stat. nov.</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672477BD2EFF37DF8F1A9BA583" pageId="105" pageNumber="106" type="reference_group">
<paragraph id="725E36EC2477BD2EFF37DF8F1953A125" blockId="105.[151,1436,421,548]" pageId="105" pageNumber="106">
Basionym:
<taxonomicName id="B5E14D6F2477BD2EFEB1DF8F1895A2E5" authority="Henderson (1995: 264)" authorityName="Henderson" authorityPageNumber="264" authorityYear="1995" box="[273,938,461,484]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="105" pageNumber="106" phylum="Tracheophyta" rank="variety" species="brevispatha" variety="occidentale">
<emphasis id="4095EAFE2477BD2EFEB1DF8F1AC3A2E5" box="[273,508,461,484]" italics="true" pageId="105" pageNumber="106">Geonoma brevispatha</emphasis>
var.
<emphasis id="4095EAFE2477BD2EFD8FDF8F1995A2E5" box="[559,682,461,484]" italics="true" pageId="105" pageNumber="106">occidentale</emphasis>
<bibRefCitation id="16704B1D2477BD2EFD10DF8F1895A2E5" author="Henderson, A." box="[688,938,461,484]" journalOrPublisher="Oxford University Press, New York" pageId="105" pageNumber="106" pagination="1 - 362" refId="ref122671" refString="Henderson, A. (1995) Palms of the Amazon, Oxford University Press, New York, pp. 1 - 362." title="Palms of the Amazon" type="book chapter" year="1995">Henderson (1995: 264)</bibRefCitation>
</taxonomicName>
. Type:
<collectingCountry id="0AF6767C2477BD2EFC56DF8F1F01A2E5" box="[1014,1086,461,484]" name="Peru" pageId="105" pageNumber="106">PERU</collectingCountry>
.
<collectingRegion id="B025F80E2477BD2EFBE7DF8F1FD5A2E5" box="[1095,1258,461,484]" country="Peru" name="Madre de Dios" pageId="105" pageNumber="106">Madre de Dios</collectingRegion>
: Río Tambopata, Explorers Inn at junction with Río La Torre,
<geoCoordinate id="17D5502B2477BD2EFD08DFAF183DA105" box="[680,770,493,516]" degrees="12" direction="south" minutes="50" orientation="latitude" pageId="105" pageNumber="106" precision="925" value="-12.833333">12°50S</geoCoordinate>
,
<geoCoordinate id="17D5502B2477BD2EFCADDFAF1850A105" box="[781,879,493,516]" degrees="69" direction="west" minutes="17" orientation="longitude" pageId="105" pageNumber="106" precision="925" value="-69.28333">69°17W</geoCoordinate>
,
<date id="065F102C2477BD2EFCD9DFAF1F04A105" box="[889,1083,493,516]" pageId="105" pageNumber="106" value="1991-11-03">3 November 1991</date>
,
<emphasis id="4095EAFE2477BD2EFBE5DFAF1EA3A105" box="[1093,1436,493,517]" italics="true" pageId="105" pageNumber="106">A. Henderson &amp; F. Chávez 1633</emphasis>
(
<typeStatus id="AD5A884E2477BD2EFF6EDC4F1A12A125" box="[206,301,525,548]" pageId="105" pageNumber="106" type="holotype">holotype</typeStatus>
USM!, isotypes CUZ!, NY!).
</paragraph>
<paragraph id="725E36EC2477BD2EFF37DC161A9BA583" blockId="105.[151,1438,596,2014]" pageId="105" pageNumber="106">
<emphasis id="4095EAFE2477BD2EFF37DC161BDCA16F" box="[151,227,596,622]" italics="true" pageId="105" pageNumber="106">Plants</emphasis>
2.7(1.54.0) m tall; stems 1.6(1.13.0) m tall, 1.3(0.91.7) cm in diameter, solitary or clustered, not cane-like or cane-like; internodes 1.6(0.83.6) cm long, yellowish and smooth.
<emphasis id="4095EAFE2477BD2EFBFCDC3E1F90A194" box="[1116,1199,636,661]" italics="true" pageId="105" pageNumber="106">Leaves</emphasis>
11(518) per stem, irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths 15.0 cm long; petioles 37.4(25.555.0) cm long, drying green or yellowish; rachis 43.8(35.057.5) cm long, 3.5(2.74.4) mm in diameter; veins not raised or slightly raised and triangular in cross-section adaxially; pinnae 4(27) per side of rachis; basal pinna 23.6(17.532.0) m long, 2.8(1.06.0) cm wide, forming an angle of 60(3390)° with the rachis; apical pinna 14.4(11.517.5) cm long, 14.8(8.823.4) cm wide, forming an angle of 36(30 42)° with the rachis.
<emphasis id="4095EAFE2477BD2EFE37DD20197BA07D" box="[407,580,866,892]" italics="true" pageId="105" pageNumber="106">Inflorescences</emphasis>
branched 23 orders; prophylls and peduncular bracts not ribbed with elongate, unbranched fibers, flattened, deciduous or persistent; prophylls 9.7(6.111.0) cm long, short, asymmetrically apiculate, the margins curved around the stem, the surfaces flat with dense, felty, brown tomentum, prophyll equal to and early deciduous with the peduncular bract, the surfaces not ridged, without unequally wide ridges; peduncular bracts
<quantity id="B5199B092477BD2EFD2FDDBF19DDA716" box="[655,738,1021,1047]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="105" pageNumber="106" unit="cm" value="9.5">9.5 cm</quantity>
long, well-developed, inserted 0.3(0.10.5) cm above the prophyll; peduncles 6.4(3.39.5) cm long, 6.8(3.810.2) mm in diameter; rachillae 24(1333), 25.0(15.0 33.5) cm long, 1.8(1.22.6) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown, with faint to pronounced, short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits decussately arranged throughout the rachillae, the groups of pits closely spaced, glabrous internally; proximal lips without a central notch before anthesis, not recurved after anthesis, hood-shaped at anthesis, sometimes splitting post-anthesis; proximal and distal lips drying the same color as the rachillae, joined laterally with no clear gap between them, often forming a raised cupule, the margins not overlapping; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; nonfertilized pistillate flowers deciduous after anthesis; staminodial tubes crenulate or shallowly lobed at the apex, those of non-fertilized pistillate flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE2477BD2EFB64DB8F1E3CA6E6" box="[1220,1283,1485,1511]" italics="true" pageId="105" pageNumber="106">fruits</emphasis>
7.2(6.28.5) mm long, 5.9(5.46.6) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, bumpy from the numerous, subepidermal, tangential, short fibers present, these coming to a point at fruit apices; locular epidermis without operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672477BD2EFF66D8CC1882A4DF" pageId="105" pageNumber="106" type="distribution">
<paragraph id="725E36EC2477BD2EFF66D8CC1FC0A5CE" blockId="105.[151,1438,596,2014]" pageId="105" pageNumber="106">
<emphasis id="4095EAFE2477BD2EFF66D8CC1922A5A9" bold="true" box="[198,541,1678,1704]" pageId="105" pageNumber="106">Distribution and habitat:—</emphasis>
From 4°29
<geoCoordinate id="17D5502B2477BD2EFD16D8CC182BA5A9" box="[694,788,1678,1704]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="105" pageNumber="106" precision="925" value="-17.833334">17°50S</geoCoordinate>
and 62°53
<geoCoordinate id="17D5502B2477BD2EFC0BD8CC1F28A5A9" box="[939,1047,1678,1704]" degrees="73" direction="west" minutes="35" orientation="longitude" pageId="105" pageNumber="106" precision="925" value="-73.583336">73°35W</geoCoordinate>
in the western Amazon region in
<collectingCountry id="0AF6767C2477BD2EFF37D8F71BDDA5CE" box="[151,226,1717,1743]" name="Brazil" pageId="105" pageNumber="106">Brazil</collectingCountry>
,
<collectingCountry id="0AF6767C2477BD2EFF4DD8F71A1AA5CE" box="[237,293,1717,1743]" name="Peru" pageId="105" pageNumber="106">Peru</collectingCountry>
, and
<collectingCountry id="0AF6767C2477BD2EFEC3D8F71A86A5CE" box="[355,441,1717,1743]" name="Bolivia" pageId="105" pageNumber="106">Bolivia</collectingCountry>
at a mean elevation of 312(150950) m in lowland rainforest (
<figureCitation id="EADA2A692477BD2EFB3AD8F71FCEA5CE" box="[1178,1265,1717,1743]" captionStart="FIGURE 28" captionStartId="104.[151,255,1873,1896]" captionTargetBox="[164,1410,848,1856]" captionTargetId="figure@104.[151,1436,828,1873]" captionTargetPageId="104" captionText="FIGURE 28. Distribution maps of Geonoma monospatha, G. mooreana, G. multisecta, and G. occidentalis." figureDoi="http://doi.org/10.5281/zenodo.3538418" httpUri="https://zenodo.org/record/3538418/files/figure.png" pageId="105" pageNumber="106">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="725E36EC2477BD2EFF66D89E1EA7A46B" blockId="105.[151,1438,596,2014]" pageId="105" pageNumber="106">
<emphasis id="4095EAFE2477BD2EFF66D89E1A81A5F7" bold="true" box="[198,446,1756,1782]" pageId="105" pageNumber="106">Taxonomic notes:—</emphasis>
This taxon was treated as a variety of
<taxonomicName id="B5E14D6F2477BD2EFCD5D89E1F48A5F7" authorityName="Barbosa Rodrigues" authorityYear="1879" box="[885,1143,1756,1782]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="105" pageNumber="106" phylum="Tracheophyta" rank="species" species="brevispatha">
<emphasis id="4095EAFE2477BD2EFCD5D89E1F48A5F7" box="[885,1143,1756,1782]" italics="true" pageId="105" pageNumber="106">Geonoma brevispatha</emphasis>
</taxonomicName>
by
<bibRefCitation id="16704B1D2477BD2EFB02D89E1E46A5F7" author="Henderson, A." box="[1186,1401,1756,1782]" journalOrPublisher="Oxford University Press, New York" pageId="105" pageNumber="106" pagination="1 - 362" refId="ref122671" refString="Henderson, A. (1995) Palms of the Amazon, Oxford University Press, New York, pp. 1 - 362." title="Palms of the Amazon" type="book chapter" year="1995">Henderson (1995)</bibRefCitation>
. In the present study, this name is treated as an excluded name, and the taxon to which Henderson was referring is called
<taxonomicName id="B5E14D6F2477BD2EFF44D96B1992A442" authorityName="Henderson" authorityYear="2011" baseAuthorityName="Wendland ex Drude" box="[228,685,1833,1859]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="105" pageNumber="106" phylum="Tracheophyta" rank="subSpecies" species="pohliana" subSpecies="weddelliana">
<emphasis id="4095EAFE2477BD2EFF44D96B1AFAA442" box="[228,453,1833,1859]" italics="true" pageId="105" pageNumber="106">Geonoma pohliana</emphasis>
subsp.
<emphasis id="4095EAFE2477BD2EFDBED96B1992A442" box="[542,685,1833,1859]" italics="true" pageId="105" pageNumber="106">weddelliana</emphasis>
</taxonomicName>
. The similarity between the two taxa is superficial and there are many differences between them, notably in the prophylls (not available to
<bibRefCitation id="16704B1D2477BD2EFC5ED9121FF7A46B" author="Henderson, A." box="[1022,1224,1872,1898]" journalOrPublisher="Oxford University Press, New York" pageId="105" pageNumber="106" pagination="1 - 362" refId="ref122671" refString="Henderson, A. (1995) Palms of the Amazon, Oxford University Press, New York, pp. 1 - 362." title="Palms of the Amazon" type="book chapter" year="1995">Henderson, 1995</bibRefCitation>
) and in the fruits.
</paragraph>
<paragraph id="725E36EC2477BD2EFF66D9341882A4DF" blockId="105.[151,1438,596,2014]" pageId="105" pageNumber="106">
<emphasis id="4095EAFE2477BD2EFF66D9341AC7A491" bold="true" box="[198,504,1910,1936]" pageId="105" pageNumber="106">Subspecific variation:—</emphasis>
Two traits vary within this species (stem branching, stem
<typeStatus id="AD5A884E2477BD2EFB61D9341FC7A491" box="[1217,1272,1910,1936]" pageId="105" pageNumber="106">type</typeStatus>
). There is no geographic disjunction—the gap in
<collectingCountry id="0AF6767C2477BD2EFD9BD9DF19AEA4B6" box="[571,657,1949,1975]" name="Bolivia" pageId="105" pageNumber="106">Bolivia</collectingCountry>
is likely to be an artifact of insufficient collecting. There are three outlying specimens from
<collectingCountry id="0AF6767C2477BD2EFE61D9861937A4DF" box="[449,520,1988,2014]" name="Brazil" pageId="105" pageNumber="106">Brazil</collectingCountry>
(
<collectingRegion id="B025F80E2477BD2EFDB8D986196AA4DF" box="[536,597,1988,2014]" country="Brazil" name="Acre" pageId="105" pageNumber="106">Acre</collectingRegion>
,
<collectingRegion id="B025F80E2477BD2EFDC1D98619EAA4DF" box="[609,725,1988,2014]" country="Brazil" name="Rondonia" pageId="105" pageNumber="106">Rondônia</collectingRegion>
) and
<collectingCountry id="0AF6767C2477BD2EFCB6D9861873A4DF" box="[790,844,1988,2014]" name="Peru" pageId="105" pageNumber="106">Peru</collectingCountry>
(
<collectingRegion id="B025F80E2477BD2EFCFCD986188EA4DF" box="[860,945,1988,2014]" country="Peru" name="Loreto" pageId="105" pageNumber="106">Loreto</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>