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<document id="B5B54DC0C6FC0692BB94000A028981D7" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA2475BD2AFF37D86C1E2EA29B" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma operculata Henderson 2011, sp. nov." docType="treatment" docVersion="7" lastPageNumber="110" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="108" updateTime="1698765326825" updateUser="ExternalLinkService">
<mods:mods id="CE7C469434AEE223A3BF757042609F68" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6EEF8776A2DBAC6D869B480B9B75795">
<mods:title id="0FAB9B86E529E8D4253BFCDAE98A334A">A revision of Geonoma (Arecaceae)</mods:title>
</mods:titleInfo>
<mods:name id="1D2176F81949519BF33D1688B286896D" type="personal">
<mods:role id="10B8ED6F141E01A3F1BAD9F2B07E981A">
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<mods:namePart id="51F977C9D620C3A1A786092786361A0D">Henderson, Andrew</mods:namePart>
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<mods:title id="419FB4C51A2AEF27076A68A60B2D9AB8">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="6CE2AE973915DF4334528E95520A9F32">
<mods:date id="B74B95CBA2797A4F049375BAA378B531">2011</mods:date>
<mods:detail id="D120B4B282D5B09A6F77D721BD0DDCA7" type="pubDate">
<mods:number id="1266CB6709813BFA7D54CED1AEC6C5A4">2011-02-18</mods:number>
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<mods:number id="97F6B8A648471EF544FA673FC79721A5">17</mods:number>
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<mods:classification id="66A9B2E590E6C2C0549F74326F424037">journal article</mods:classification>
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<mods:identifier id="9549E887D4F1BD6F839FB656A0DF1DEB" type="DOI">10.11646/phytotaxa.17.1</mods:identifier>
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<mods:identifier id="E24E5F1AF3BEA42EAA776E8577B6122E" type="ISSN">1179-3163</mods:identifier>
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<treatment id="FA4887FA2475BD2AFF37D86C1E2EA29B" ID-DOI="http://doi.org/10.5281/zenodo.5609086" ID-GBIF-Taxon="160342418" ID-Zenodo-Dep="5609086" LSID="urn:lsid:plazi:treatment:FA4887FA2475BD2AFF37D86C1E2EA29B" httpUri="http://treatment.plazi.org/id/FA4887FA2475BD2AFF37D86C1E2EA29B" lastPageId="109" lastPageNumber="110" pageId="107" pageNumber="108">
<subSubSection id="3AFB65672475BD2CFF37D86C18E8A549" box="[151,983,1581,1608]" pageId="107" pageNumber="108" type="nomenclature">
<paragraph id="725E36EC2475BD2CFF37D86C18E8A549" blockId="107.[151,983,1581,1608]" box="[151,983,1581,1608]" pageId="107" pageNumber="108">
<emphasis id="4095EAFE2475BD2CFF37D86C1986A549" bold="true" box="[151,697,1581,1608]" pageId="107" pageNumber="108">
44.
<taxonomicName id="B5E14D6F2475BD2CFF63D86C196EA549" ID-CoL="6KBQC" authority="Henderson" authorityName="Henderson" authorityYear="2011" box="[195,593,1581,1608]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="operculata" status="sp. nov.">
<emphasis id="4095EAFE2475BD2CFF63D86C1A80A549" bold="true" box="[195,447,1581,1608]" italics="true" pageId="107" pageNumber="108">Geonoma operculata</emphasis>
Henderson
</taxonomicName>
,
<emphasis id="4095EAFE2475BD2CFDFED86C1986A549" bold="true" box="[606,697,1582,1608]" italics="true" pageId="107" pageNumber="108">
<taxonomicNameLabel id="5BA657852475BD2CFDFED86C1986A549" box="[606,697,1582,1608]" pageId="107" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
(Appendix IV, Plate 52)
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672475BD2BFF37D83E18A7A01D" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108" type="description">
<paragraph id="725E36EC2475BD2CFF37D83E1912A592" blockId="107.[151,557,1660,1683]" box="[151,557,1660,1683]" pageId="107" pageNumber="108">
<emphasis id="4095EAFE2475BD2CFF37D83E1912A592" box="[151,557,1660,1683]" italics="true" pageId="107" pageNumber="108">
<taxonomicName id="B5E14D6F2475BD2CFF37D83E1A46A592" box="[151,377,1660,1683]" class="Liliopsida" family="Arecaceae" genus="Asterogyne" kingdom="Plantae" order="Arecales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="subSpecies" species="speciebus" subSpecies="affinibus">A speciebus affinibus</taxonomicName>
operculo differt.
</emphasis>
</paragraph>
<paragraph id="725E36EC2475BD2CFF37D8FE1909A412" blockId="107.[151,1436,1724,1812]" pageId="107" pageNumber="108">
Type:
<collectingCountry id="0AF6767C2475BD2CFF7BD8FE1A42A5D2" box="[219,381,1724,1747]" name="Venezuela" pageId="107" pageNumber="108">VENEZUELA</collectingCountry>
.
<collectingRegion id="B025F80E2475BD2CFE28D8FE1AD8A5D2" box="[392,487,1724,1747]" country="Venezuela" name="Miranda" pageId="107" pageNumber="108">Miranda</collectingRegion>
: Cordillera de la Costa, al noreste de Guatire, Fila Juan Torres-Fila Las Perdices, por el río Guayabal hacia el pueblo Guayabal,
<geoCoordinate id="17D5502B2475BD2CFD2CD89E19D5A5F2" box="[652,746,1756,1779]" degrees="10" direction="north" minutes="31" orientation="latitude" pageId="107" pageNumber="108" precision="925" value="10.516666">10°31N</geoCoordinate>
,
<geoCoordinate id="17D5502B2475BD2CFD5AD89E1863A5F2" box="[762,860,1756,1779]" degrees="66" direction="west" minutes="20" orientation="longitude" pageId="107" pageNumber="108" precision="925" value="-66.333336">66°20W</geoCoordinate>
,
<quantity id="B5199B092475BD2CFCCAD89E18D6A5F2" box="[874,1001,1756,1779]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="107" pageNumber="108" unit="m" value="800.0" valueMax="900.0" valueMin="700.0">700900 m</quantity>
,
<date id="065F102C2475BD2CFC58D89E1FD6A5F2" box="[1016,1257,1756,1779]" pageId="107" pageNumber="108" value="1993-02-19" valueMax="1993-02-22" valueMin="1993-02-19">1922 February 1993</date>
,
<emphasis id="4095EAFE2475BD2CFB57D89F1EA3A5F2" box="[1271,1436,1756,1780]" italics="true" pageId="107" pageNumber="108">W. Meier 3401</emphasis>
(
<typeStatus id="AD5A884E2475BD2CFF6ED8BE1A12A412" box="[206,301,1788,1811]" pageId="107" pageNumber="108" type="holotype">holotype</typeStatus>
NY!, isotype VEN
<emphasis id="4095EAFE2475BD2CFDA1D8BF1919A415" box="[513,550,1789,1812]" italics="true" pageId="107" pageNumber="108">n.v.</emphasis>
).
</paragraph>
<paragraph id="725E36EC2475BD2BFF37D97B18A7A01D" blockId="107.[151,1436,1849,2030]" lastBlockId="108.[151,1437,152,874]" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108">
<emphasis id="4095EAFE2475BD2CFF37D97B1BDDA452" box="[151,226,1849,1875]" italics="true" pageId="107" pageNumber="108">Plants</emphasis>
2.0 m tall; stems height and in diameter no data; internodes yellowish and smooth.
<emphasis id="4095EAFE2475BD2CFB66D9781E27A452" box="[1222,1304,1850,1875]" italics="true" pageId="107" pageNumber="108">Leaves</emphasis>
irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths no data; petioles drying green or yellowish; rachis 23.0 cm long; veins not raised or slightly raised and triangular in cross-section adaxially; pinnae 3 per side of rachis; basal pinna length, width and angle no data; apical pinna
<quantity id="B5199B092475BD2CFB3CD9EF1FC3A4C6" box="[1180,1276,1965,1991]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="107" pageNumber="108" unit="cm" value="12.5">12.5 cm</quantity>
long, 9.0 cm wide, forming an angle of 34° with the rachis.
<emphasis id="4095EAFE2475BD2CFD6DD9961846A4EF" box="[717,889,2004,2030]" italics="true" pageId="107" pageNumber="108">Inflorescences</emphasis>
branched 2 orders; prophylls and peduncular bracts not ribbed with elongate, unbranched fibers, flattened, deciduous or persistent; prophylls length
<quantity id="B5199B092472BD2BFAC1DEDA1EA3A3B3" box="[1377,1436,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="108" pageNumber="109" unit="cm" value="4.0">4 cm</quantity>
long, short, asymmetrically apiculate, the margins curved around the stem, the surfaces flat with dense, felty, brown tomentum, prophyll equal to and early deciduous with the peduncular bract, the surfaces not ridged, without unequally wide ridges; peduncular bracts no data; peduncles 12.0 cm long,
<quantity id="B5199B092472BD2BFB02DF4E1E3FA227" box="[1186,1280,268,294]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="108" pageNumber="109" unit="mm" value="3.2">3.2 mm</quantity>
in diameter; rachillae 37, 15.0 cm long,
<quantity id="B5199B092472BD2BFE49DF701978A24D" box="[489,583,306,332]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="108" pageNumber="109" unit="mm" value="0.8">0.8 mm</quantity>
in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown, with faint to pronounced, short, transverse ridges, filiform with extended narrowed sections between the flower pits; flower pits alternately arranged (sometimes distorted by twisting and contracting of rachillae), glabrous internally; proximal lips without a central notch before anthesis, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, joined to form a raised cupule, the margins not overlapping; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; non-fertilized pistillate flowers no data; staminodial tubes crenulate or shallowly lobed at the apex, persistence no data;
<emphasis id="4095EAFE2472BD2BFD54DCCC180DA1A9" box="[756,818,654,680]" italics="true" pageId="108" pageNumber="109">fruits</emphasis>
<quantity id="B5199B092472BD2BFC9ADCCC18ABA1A9" box="[826,916,654,680]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="108" pageNumber="109" unit="mm" value="7.5">7.5 mm</quantity>
long,
<quantity id="B5199B092472BD2BFC7EDCCC1F08A1A9" box="[990,1079,654,680]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="108" pageNumber="109" unit="mm" value="5.8">5.8 mm</quantity>
in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, bumpy from the numerous, subepidermal, tangential, short fibers present, these coming to a point at fruit apices; locular epidermis with operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672472BD2AFF66DD6B1E2EA29B" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="distribution">
<paragraph id="725E36EC2472BD2BFF66DD6B19DEA06B" blockId="108.[151,1437,152,874]" pageId="108" pageNumber="109">
<emphasis id="4095EAFE2472BD2BFF66DD6B191FA042" bold="true" box="[198,544,809,835]" pageId="108" pageNumber="109">Distribution and habitat:—</emphasis>
At
<geoCoordinate id="17D5502B2472BD2BFDE6DD6B1994A042" box="[582,683,809,835]" degrees="10" direction="north" minutes="31" orientation="latitude" pageId="108" pageNumber="109" precision="925" value="10.516666">10°31N</geoCoordinate>
and
<geoCoordinate id="17D5502B2472BD2BFD46DD6B186DA042" box="[742,850,809,835]" degrees="66" direction="west" minutes="20" orientation="longitude" pageId="108" pageNumber="109" precision="925" value="-66.333336">66°20W</geoCoordinate>
in the Coastal Cordillera in
<collectingCountry id="0AF6767C2472BD2BFB04DD6B1E22A042" box="[1188,1309,809,835]" name="Venezuela" pageId="108" pageNumber="109">Venezuela</collectingCountry>
(
<collectingRegion id="B025F80E2472BD2BFA8DDD6B1EABA042" box="[1325,1428,809,835]" country="Venezuela" name="Miranda" pageId="108" pageNumber="109">Miranda</collectingRegion>
) at
<quantity id="B5199B092472BD2BFF13DD121BC2A06B" box="[179,253,848,874]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="108" pageNumber="109" unit="m" value="800.0">800 m</quantity>
elevation in lowland rainforest (
<figureCitation id="EADA2A692472BD2BFDDBDD1219ECA06B" box="[635,723,848,874]" captionStart="FIGURE 29" captionStartId="108.[151,259,1964,1987]" captionTargetBox="[167,1424,925,1942]" captionTargetId="figure@108.[151,1436,907,1962]" captionTargetPageId="108" captionText="FIGURE 29. Distribution maps of Geonoma oldemanii, G. oligoclona, G. operculata, and G. orbignyana subsp. orbignyana." figureDoi="http://doi.org/10.5281/zenodo.3538420" httpUri="https://zenodo.org/record/3538420/files/figure.png" pageId="108" pageNumber="109">Fig. 29</figureCitation>
).
</paragraph>
<caption id="269E66642472BD2BFF37D9EE1A26A4E7" ID-DOI="http://doi.org/10.5281/zenodo.3538420" ID-Zenodo-Dep="3538420" httpUri="https://zenodo.org/record/3538420/files/figure.png" pageId="108" pageNumber="109" startId="108.[151,259,1964,1987]" targetBox="[167,1424,925,1942]" targetPageId="108">
<paragraph id="725E36EC2472BD2BFF37D9EE1A26A4E7" blockId="108.[151,1437,1964,2022]" pageId="108" pageNumber="109">
<emphasis id="4095EAFE2472BD2BFF37D9EE1A18A4C2" bold="true" box="[151,295,1964,1987]" pageId="108" pageNumber="109">FIGURE 29</emphasis>
. Distribution maps of
<taxonomicName id="B5E14D6F2472BD2BFD8DD9EE182BA4C2" authorityName="de Granville" authorityYear="1975" box="[557,788,1964,1987]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="oldemanii">
<emphasis id="4095EAFE2472BD2BFD8DD9EE182BA4C2" box="[557,788,1964,1987]" italics="true" pageId="108" pageNumber="109">Geonoma oldemanii</emphasis>
</taxonomicName>
,
<taxonomicName id="B5E14D6F2472BD2BFC84D9EE18FEA4C2" authorityName="Trail" authorityYear="1876" box="[804,961,1964,1987]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="oligoclona">
<emphasis id="4095EAFE2472BD2BFC84D9EE180AA4C2" box="[804,821,1964,1987]" italics="true" pageId="108" pageNumber="109">G</emphasis>
.
<emphasis id="4095EAFE2472BD2BFCE4D9EE18FEA4C2" box="[836,961,1964,1987]" italics="true" pageId="108" pageNumber="109">oligoclona</emphasis>
</taxonomicName>
<emphasis id="4095EAFE2472BD2BFC61D9EE18FAA4C2" box="[961,965,1964,1987]" italics="true" pageId="108" pageNumber="109">,</emphasis>
<taxonomicName id="B5E14D6F2472BD2BFC6ED9EE1F53A4C2" authorityName="Henderson" authorityYear="2011" box="[974,1132,1964,1987]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="operculata">
<emphasis id="4095EAFE2472BD2BFC6ED9EE18E0A4C2" box="[974,991,1964,1987]" italics="true" pageId="108" pageNumber="109">G</emphasis>
.
<emphasis id="4095EAFE2472BD2BFC4ED9EE1F53A4C2" box="[1006,1132,1964,1987]" italics="true" pageId="108" pageNumber="109">operculata</emphasis>
</taxonomicName>
<emphasis id="4095EAFE2472BD2BFBCCD9EE1F4FA4C2" box="[1132,1136,1964,1987]" italics="true" pageId="108" pageNumber="109">,</emphasis>
and
<taxonomicName id="B5E14D6F2472BD2BFB09D9EE1A2DA4E7" authority="subsp. orbignyana" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="subSpecies" species="orbignyana" subSpecies="orbignyana">
<emphasis id="4095EAFE2472BD2BFB09D9EE1F85A4C2" box="[1193,1210,1964,1987]" italics="true" pageId="108" pageNumber="109">G</emphasis>
.
<emphasis id="4095EAFE2472BD2BFB6AD9EE1E72A4C2" box="[1226,1357,1964,1987]" italics="true" pageId="108" pageNumber="109">orbignyana</emphasis>
subsp.
<emphasis id="4095EAFE2472BD2BFF37D98D1A2DA4E7" box="[151,274,1999,2022]" italics="true" pageId="108" pageNumber="109">orbignyana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="725E36EC2473BD2AFF66DEDA1FD4A272" blockId="109.[151,1437,152,410]" pageId="109" pageNumber="110">
<emphasis id="4095EAFE2473BD2AFF66DEDA1AFEA3B3" bold="true" box="[198,449,152,178]" pageId="109" pageNumber="110">Taxonomic notes:—</emphasis>
This species is based on a single specimen. This is very similar to specimens of
<taxonomicName id="B5E14D6F2473BD2AFA23DEDA1BD1A3D9" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="braunii">
<emphasis id="4095EAFE2473BD2AFA23DEDA1BD1A3D9" italics="true" pageId="109" pageNumber="110">G. braunii</emphasis>
</taxonomicName>
, and in fact it was identified as such by
<bibRefCitation id="16704B1D2473BD2AFD73DEFC18BDA3D9" author="Stauffer, F." box="[723,898,190,216]" journalOrPublisher="Acta Botanica Venezuelica" pageId="109" pageNumber="110" pagination="1 - 10" part="20" refId="ref124409" refString="Stauffer, F. (1997) Estudio morfologico y taxonomico de Geonoma spinescens H. Wendl. ex Burret (Arecaceae) y descripcion de una nueva variedad. Acta Botanica Venezuelica 20: 1 - 10." title="Estudio morfologico y taxonomico de Geonoma spinescens H. Wendl. ex Burret (Arecaceae) y descripcion de una nueva variedad" type="journal article" year="1997">Stauffer (1997</bibRefCitation>
, as
<taxonomicName id="B5E14D6F2473BD2AFC10DEFC1FDCA3D9" authorityName="Stauffer" authorityYear="1997" box="[944,1251,190,217]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="variety" species="spinescens" variety="braunii">
<emphasis id="4095EAFE2473BD2AFC10DEFC1F73A3D8" box="[944,1100,190,217]" italics="true" pageId="109" pageNumber="110">G. spinescens</emphasis>
var.
<emphasis id="4095EAFE2473BD2AFB2CDEFC1FDCA3D9" box="[1164,1251,190,216]" italics="true" pageId="109" pageNumber="110">braunii</emphasis>
</taxonomicName>
). However, the fruits of the specimen (in a separate packet, see Appendix IV, Plate 52) have an operculum, and because of this the specimen is here recognized as a separate species. It is the only species outside of the
<taxonomicName id="B5E14D6F2473BD2AFA86DF4E1A36A24C" authorityName="Wendland ex Spruce" authorityYear="1871" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="congesta">
<emphasis id="4095EAFE2473BD2AFA86DF4E1A36A24C" italics="true" pageId="109" pageNumber="110">Geonoma congesta</emphasis>
</taxonomicName>
clade to have fruits with an operculum. Apart from the operculum, the specimen bears no resemblance to species in the
<taxonomicName id="B5E14D6F2473BD2AFE54DF1B1943A272" authorityName="Wendland ex Spruce" authorityYear="1871" box="[500,636,345,371]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="congesta">
<emphasis id="4095EAFE2473BD2AFE54DF1B1943A272" box="[500,636,345,371]" italics="true" pageId="109" pageNumber="110">G. congesta</emphasis>
</taxonomicName>
clade but appears similar to specimens of
<taxonomicName id="B5E14D6F2473BD2AFBCEDF1B1FDBA272" box="[1134,1252,345,371]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="braunii">
<emphasis id="4095EAFE2473BD2AFBCEDF1B1FDBA272" box="[1134,1252,345,371]" italics="true" pageId="109" pageNumber="110">G. braunii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="725E36EC2473BD2AFF66DFC21E2EA29B" blockId="109.[151,1437,152,410]" box="[198,1297,384,410]" pageId="109" pageNumber="110">
<emphasis id="4095EAFE2473BD2AFF66DFC21ACBA29B" bold="true" box="[198,500,384,410]" pageId="109" pageNumber="110">Subspecific variation:—</emphasis>
No trait varies within this species and only one specimen is known.
</paragraph>
</subSubSection>
</treatment>
</document>