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<document id="94118888FEA1470AA32A68095B245244" ID-DOI="10.11646/zootaxa.5020.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5223173" ID-ZooBank="4FF882DF-334F-49C8-A576-4192B5F2654C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629388374973" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2021" docId="FA08E75DAA378057FDC030499440699D" docLanguage="en" docName="zootaxa.5020.2.1.pdf" docOrigin="Zootaxa 5020 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habetia imitatrix Karny 1912" docType="treatment" docVersion="3" lastPageNumber="210" masterDocId="06319F25AA3F805EFD57366B91686D58" masterDocTitle="Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="209" updateTime="1699071125630" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B1DCF6B913B2860BC287C71A75AE6611">Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)</mods:title>
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<treatment id="FA08E75DAA378057FDC030499440699D" ID-DOI="http://doi.org/10.5281/zenodo.5223209" ID-Zenodo-Dep="5223209" LSID="urn:lsid:plazi:treatment:FA08E75DAA378057FDC030499440699D" httpUri="http://treatment.plazi.org/id/FA08E75DAA378057FDC030499440699D" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<subSubSection id="3ABB05C0AA378056FDC0304993606B64" box="[151,520,1570,1597]" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="721E564BAA378056FDC0304993606B64" blockId="8.[151,623,1570,1633]" box="[151,520,1570,1597]" pageId="8" pageNumber="209">
<heading id="2956E127AA378056FDC0304993606B64" bold="true" box="[151,520,1570,1597]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
<taxonomicName id="B5A12DC8AA378056FDC0304993606B64" ID-CoL="3J4HN" authority="Karny, 1912" authorityName="Karny" authorityYear="1912" box="[151,520,1570,1597]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA378056FDC0304993606B64" bold="true" box="[151,520,1570,1597]" pageId="8" pageNumber="209">
<emphasis id="40D58A59AA378056FDC03049900F6B64" bold="true" box="[151,359,1570,1597]" italics="true" pageId="8" pageNumber="209">Habetia imitatrix</emphasis>
Karny, 1912
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378056FDC0302D93076B39" box="[151,623,1606,1633]" pageId="8" pageNumber="209" type="description">
<paragraph id="721E564BAA378056FDC0302D93076B39" blockId="8.[151,623,1570,1633]" box="[151,623,1606,1633]" pageId="8" pageNumber="209">
<figureCitation id="EA9A4ACEAA378056FDC0302D919B6B39" box="[151,243,1606,1633]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (AB, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); CE, Habetia pedala sp. nov. (Toricelli Mts); FH, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (AC, FG), 1 mm (DE, H). Phallus removed in D, FG." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="8" pageNumber="209">Figs 8A</figureCitation>
,
<figureCitation id="EA9A4ACEAA378056FDA8302C90456B39" box="[255,301,1607,1633]" captionStart="FIGURE 11" captionStartId="27.[151,250,1579,1603]" captionTargetBox="[151,1436,181,1555]" captionTargetId="figure-15@27.[151,1436,181,1555]" captionTargetPageId="27" captionText="FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (AE), 1 mm (FK)." figureDoi="http://doi.org/10.5281/zenodo.5223195" httpUri="https://zenodo.org/record/5223195/files/figure.png" pageId="8" pageNumber="209">11E</figureCitation>
,
<figureCitation id="EA9A4ACEAA378056FC6E302C90166B39" box="[313,382,1607,1633]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: AC, H. tuta sp. nov. (A Quelllager, BC Töpferfluss); DE, H. pedala sp. nov. (Toricelli Mts); FI, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="8" pageNumber="209">12FI</figureCitation>
,
<figureCitation id="EA9A4ACEAA378056FCDD302C90DE6B39" box="[394,438,1607,1633]" captionStart="FIGURE 14" captionStartId="31.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1594]" captionTargetId="figure-15@31.[151,1436,181,1595]" captionTargetPageId="31" captionText="FIGURE 14. AE: Titillators of Habetia bivittata sp. nov. (AB in situ, male from Goroka, CE dissected, male from Moife).— A, abdominal apex in lateral view; B, view through the gap of the subgenital plate on the apical widenings of the titillators; CE, titillators of male phallus in ventro-apical (C), lateral (D), and dorso-proximal (E) views.—F: Habetia imitatrix Karny, 1912 male subgenital plate with apical projection in lateral view.—Abbreviations: aw apical widening, b base of titillator, dc dorsal cap, dp dorso-apical projection, ls lateral sclerite, pp proximal projection, ti titillator stem, ts testes, vp ventral projection; st stylus of subgenital plate. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223205" httpUri="https://zenodo.org/record/5223205/files/figure.png" pageId="8" pageNumber="209">14F</figureCitation>
,
<figureCitation id="EA9A4ACEAA378056FC96302C909C6B39" box="[449,500,1607,1633]" captionStart="FIGURE 18" captionStartId="35.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-15@35.[151,1436,181,1786]" captionTargetPageId="35" captionText="FIGURE 18. Stridulatory files on underside of left tegmen of males of Habetia species: A, H. tuta sp. nov. (Lake Chambri); B, H. pallida sp. nov. (Waris); C, H. pedala sp. nov. (Toricelli Mts); D, H. imitatrix Karny, 1912 (Bongu); E, H. dentata sp. nov. (Saidor); F, H. pilleata sp. nov. (Wau 1200 m); GH, H. wau sp. nov. (G Nami Creek; H Wau).—Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223217" httpUri="https://zenodo.org/record/5223217/files/figure.png" pageId="8" pageNumber="209">18D</figureCitation>
, Map 1(2).
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378056FDC030E4944D6B95" pageId="8" pageNumber="209" type="materials_examined">
<paragraph id="721E564BAA378056FDC030E4944D6B95" blockId="8.[151,1437,1678,2029]" pageId="8" pageNumber="209">
<materialsCitation id="C2C95C16AA378056FDC030E4944D6B95" collectionCode="SMTD" collectorName="K. Wilhelm-Land" country="Papua New Guinea" latitude="-5.483333" location="Madang" longLatPrecision="1305" longitude="145.81667" pageId="8" pageNumber="209" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="40D58A59AA378056FDC030E4900B6BF0" bold="true" box="[151,355,1678,1705]" pageId="8" pageNumber="209">
<typeStatus id="AD1AE8E9AA378056FDC030E4906C6BF1" box="[151,260,1679,1705]" pageId="8" pageNumber="209" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="0AB616DBAA378056FC3A30E493276BF0" box="[365,591,1678,1705]" name="Papua New Guinea" pageId="8" pageNumber="209">Papua New Guinea</collectingCountry>
:
<collectingRegion id="B06598A9AA378056FF0B30E593D36BF0" box="[604,699,1678,1704]" country="Papua New Guinea" name="Madang" pageId="8" pageNumber="209">Madang</collectingRegion>
, labeled “
<collectorName id="DF54339DAA378056FE7930E492936BF0" box="[814,1019,1678,1705]" pageId="8" pageNumber="209">K. Wilhelm-Land</collectorName>
, Bongu”, (
<geoCoordinate id="1795308CAA378056F92E30E595A16BF1" box="[1145,1225,1678,1705]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="8" pageNumber="209" precision="925" value="-5.483333">5°29S</geoCoordinate>
,
<geoCoordinate id="1795308CAA378056F98F30E5942F6BF1" box="[1240,1351,1678,1705]" degrees="145" direction="east" minutes="49" orientation="longitude" pageId="8" pageNumber="209" precision="925" value="145.81667">145°49E</geoCoordinate>
), about 1910, collector and date not stated—depository: Senckenberg Museum für Tierkunde Dresden (
<collectionCode id="14B0CE8EAA378056F99230D894716B95" box="[1221,1305,1715,1741]" pageId="8" pageNumber="209">SMTD</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378057FD9030BD92706D8D" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209" type="discussion">
<paragraph id="721E564BAA378057FD9030BD92706D8D" blockId="8.[151,1437,1678,2029]" lastBlockId="9.[151,1437,151,1222]" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<emphasis id="40D58A59AA378056FD9030BD90246BA8" bold="true" box="[199,332,1750,1776]" pageId="8" pageNumber="209">Discussion.</emphasis>
The only known specimen of
<taxonomicName id="B5A12DC8AA378056FFF530BC924D6BA8" authorityName="Karny" authorityYear="1912" box="[674,805,1751,1776]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA378056FFF530BC924D6BA8" box="[674,805,1751,1776]" italics="true" pageId="8" pageNumber="209">H. imitatrix</emphasis>
</taxonomicName>
, the male
<typeStatus id="AD1AE8E9AA378056FECC30BD95686BA8" box="[923,1024,1750,1776]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
, is in general characters very similar to the type of
<taxonomicName id="B5A12DC8AA378056FC6D309090C96A4C" authorityName="Brunner" authorityYear="1898" box="[314,417,1786,1812]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056FC6D309090C96A4C" box="[314,417,1786,1812]" italics="true" pageId="8" pageNumber="209">H. spada</emphasis>
</taxonomicName>
, a female. Both taxa are known from a single specimen each. The type localities of both taxa are located very close to each other or are maybe identic: Konstantinhafen has been one of the old German settlements in New
<collectingCountry id="0AB616DBAA378056FC2C312990A76A04" box="[379,463,1858,1884]" name="Guinea-Bissau" pageId="8" pageNumber="209">Guinea</collectingCountry>
, while Bongu is not a locality but the name of a tribe living in the area about
<quantity id="B559FBAEAA378056F804312894F46A04" box="[1363,1436,1858,1885]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="209" unit="km" value="30.0">30 km</quantity>
South of Madang (
<bibRefCitation id="16302BBAAA378056FC3F310D93526AD9" author="Hempenstall, P. J." box="[360,570,1894,1921]" pageId="8" pageNumber="209" pagination="163 - 198" refId="ref42787" refString="Hempenstall, P. J. (2016) Chapter 7. The Mainland: New Guineans under Company and Empire. In: Pacific Islanders Under German Rule: A Study in the Meaning of Colonial Resistance. ANU Press, Canberra, pp. 163 - 198. Online Version. Available from: http: // www. jstor. org / stable / j. ctt 1 dgn 5 vg. 17 (accessed 20 July 20210" type="book chapter" year="2016">Hempenstall 2016</bibRefCitation>
), that is close to Enke Port. Karny (1912) might have been not aware of these facts when he described
<taxonomicName id="B5A12DC8AA378056FCF831E0935A6AFC" authorityName="Karny" authorityYear="1912" box="[431,562,1931,1956]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA378056FCF831E0935A6AFC" box="[431,562,1931,1956]" italics="true" pageId="8" pageNumber="209">H. imitatrix</emphasis>
</taxonomicName>
. This raises the question if the female type of
<taxonomicName id="B5A12DC8AA378056F91231E095C36AFC" authorityName="Brunner" authorityYear="1898" box="[1093,1195,1930,1956]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056F91231E095C36AFC" box="[1093,1195,1930,1956]" italics="true" pageId="8" pageNumber="209">H. spada</emphasis>
</taxonomicName>
and the male type of
<taxonomicName id="B5A12DC8AA378056FDC031C490726A90" authorityName="Karny" authorityYear="1912" box="[151,282,1967,1992]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA378056FDC031C490726A90" box="[151,282,1967,1992]" italics="true" pageId="8" pageNumber="209">H. imitatrix</emphasis>
</taxonomicName>
might belong to the same species. However, there are localities that house two different species of the genus
<taxonomicName id="B5A12DC8AA378056FD8931B990506AB4" authorityName="Kirby" authorityYear="1906" box="[222,312,2002,2028]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="40D58A59AA378056FD8931B990506AB4" box="[222,312,2002,2028]" italics="true" pageId="8" pageNumber="209">Habetia</emphasis>
</taxonomicName>
. Moreover, both taxa are known from a single specimen, and these specimens are of the opposite sex. It is thus better to wait with any nomenclatural changes until more specimens of both taxa become available to solve this question, preferably male and female found together.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA368057FD9036B590236C3D" pageId="9" pageNumber="210" type="diagnosis">
<paragraph id="721E564BAA368057FD9036B590236C3D" blockId="9.[151,1437,151,1222]" pageId="9" pageNumber="210">
<emphasis id="40D58A59AA368057FD9036B5902A6DA0" bold="true" box="[199,322,222,248]" pageId="9" pageNumber="210">Diagnosis.</emphasis>
The male of
<taxonomicName id="B5A12DC8AA368057FCB636B4930D6DA0" authorityName="Karny" authorityYear="1912" box="[481,613,223,248]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA368057FCB636B4930D6DA0" box="[481,613,223,248]" italics="true" pageId="9" pageNumber="210">H. imitatrix</emphasis>
</taxonomicName>
is unique for the shape of the subgenital plate that has the central area prolonged behind and that area shaped like a hood that carries on apico-dorsal margin a row of teeth of decreasing size toward end, and the styli project laterally from the subgenital plate not from apical margin as in all other species (
<figureCitation id="EA9A4ACEAA368057FDC83720906C6C3D" box="[159,260,331,357]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: AC, H. tuta sp. nov. (A Quelllager, BC Töpferfluss); DE, H. pedala sp. nov. (Toricelli Mts); FI, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="9" pageNumber="210">Figs 12F</figureCitation>
,
<figureCitation id="EA9A4ACEAA368057FC58372090566C3D" box="[271,318,331,357]" captionStart="FIGURE 14" captionStartId="31.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1594]" captionTargetId="figure-15@31.[151,1436,181,1595]" captionTargetPageId="31" captionText="FIGURE 14. AE: Titillators of Habetia bivittata sp. nov. (AB in situ, male from Goroka, CE dissected, male from Moife).— A, abdominal apex in lateral view; B, view through the gap of the subgenital plate on the apical widenings of the titillators; CE, titillators of male phallus in ventro-apical (C), lateral (D), and dorso-proximal (E) views.—F: Habetia imitatrix Karny, 1912 male subgenital plate with apical projection in lateral view.—Abbreviations: aw apical widening, b base of titillator, dc dorsal cap, dp dorso-apical projection, ls lateral sclerite, pp proximal projection, ti titillator stem, ts testes, vp ventral projection; st stylus of subgenital plate. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223205" httpUri="https://zenodo.org/record/5223205/files/figure.png" pageId="9" pageNumber="210">14F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA368057FD9037059440699D" pageId="9" pageNumber="210" type="description">
<paragraph id="721E564BAA368057FD90370592146C89" blockId="9.[151,1437,151,1222]" pageId="9" pageNumber="210">
<emphasis id="40D58A59AA368057FD90370590316CD0" bold="true" box="[199,345,366,392]" pageId="9" pageNumber="210">Description.</emphasis>
General habitus of the genus. Coloration of face of general color; mandibles fully black; antennal scrobae with indistinctly darkened spot at ventro-internal angle. Femora with the following number of spines on ventral margins: (1) a 6 p 6; (2) a 7 p 3; (3) a 912 p ca 11 (n = 1).
</paragraph>
<paragraph id="721E564BAA368057FD9037B095586925" blockId="9.[151,1437,151,1222]" pageId="9" pageNumber="210">
Male. Stridulatory file on underside of left tegmen with a distinct step in distal area; file length
<quantity id="B559FBAEAA368057F9A837B1940B6CAD" box="[1279,1379,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="210" unit="mm" value="2.5">2.50 mm</quantity>
with 78 teeth, of which narrow curved basal area
<quantity id="B559FBAEAA368057FFD63795938C6F41" box="[641,740,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="210" unit="mm" value="0.25">0.25 mm</quantity>
with 8 teeth, central area
<quantity id="B559FBAEAA368057F9573795950B6F41" box="[1024,1123,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="9" pageNumber="210" unit="mm" value="1.58">1.58 mm</quantity>
with 37 teeth, step
<quantity id="B559FBAEAA368057F86D379594F56F41" box="[1338,1437,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="210" unit="mm" value="0.15">0.15 mm</quantity>
with 5 teeth, narrow apical area
<quantity id="B559FBAEAA368057FCA33449933E6F65" box="[500,598,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="210" unit="mm" value="0.54">0.54 mm</quantity>
with 28 teeth (
<figureCitation id="EA9A4ACEAA368057FFA0344892356F65" box="[759,861,547,573]" captionStart="FIGURE 18" captionStartId="35.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-15@35.[151,1436,181,1786]" captionTargetPageId="35" captionText="FIGURE 18. Stridulatory files on underside of left tegmen of males of Habetia species: A, H. tuta sp. nov. (Lake Chambri); B, H. pallida sp. nov. (Waris); C, H. pedala sp. nov. (Toricelli Mts); D, H. imitatrix Karny, 1912 (Bongu); E, H. dentata sp. nov. (Saidor); F, H. pilleata sp. nov. (Wau 1200 m); GH, H. wau sp. nov. (G Nami Creek; H Wau).—Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223217" httpUri="https://zenodo.org/record/5223217/files/figure.png" pageId="9" pageNumber="210">Fig. 18D</figureCitation>
). Tenth abdominal tergite with convex surface; from hind margin with a long and narrow oval incision closed from behind by projecting acute angles from tip of resulting gap. Cerci moderately curved and little narrowing towards end; from little before mid-length with a dorso-ventrally compressed inner ridge that widens to a wide and compressed internal process, which divides towards end into two compressed, wide, and little curved lobes, the proximal branch little larger than the distal branch, both with a minute denticle at tip, the proximal branch also with a dorsal carina from base of process; inner surface of cercus between these projections and upper margin little concave and terminated dorsally by a narrow rim (
<figureCitation id="EA9A4ACEAA368057F9E4349094676E4D" box="[1203,1295,763,789]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (AB, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); CE, Habetia pedala sp. nov. (Toricelli Mts); FH, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (AC, FG), 1 mm (DE, H). Phallus removed in D, FG." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="9" pageNumber="210">Fig. 8A</figureCitation>
). Subgenital plate roughly semi-circular with a long and narrow medial projection; in ventral view deeply incised from base and membranous between resulting oblique anterior margins; sclerotized area with convex and later concave lateral margins forming a long and narrow apical projection with a fine medial carina and acute and laterally compressed tip; this rather high and towards ventral margin laterally compressed projection obviously results from a widened fusion of the dorsal lateral margins with the ventral, medial carina, it is in lateral view obliquely truncate at end and carries 3 larger transverse teeth in upper and a few minute denticles in lower area (
<figureCitation id="EA9A4ACEAA368057F96835B895CF6EB5" box="[1087,1191,979,1005]" captionStart="FIGURE 11" captionStartId="27.[151,250,1579,1603]" captionTargetBox="[151,1436,181,1555]" captionTargetId="figure-15@27.[151,1436,181,1555]" captionTargetPageId="27" captionText="FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (AE), 1 mm (FK)." figureDoi="http://doi.org/10.5281/zenodo.5223195" httpUri="https://zenodo.org/record/5223195/files/figure.png" pageId="9" pageNumber="210">Figs 11E</figureCitation>
,
<figureCitation id="EA9A4ACEAA368057F9E435B895886EB5" box="[1203,1248,979,1005]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: AC, H. tuta sp. nov. (A Quelllager, BC Töpferfluss); DE, H. pedala sp. nov. (Toricelli Mts); FI, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="9" pageNumber="210">12F</figureCitation>
,
<figureCitation id="EA9A4ACEAA368057F9BB35B894736EB5" box="[1260,1307,979,1005]" captionStart="FIGURE 14" captionStartId="31.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1594]" captionTargetId="figure-15@31.[151,1436,181,1595]" captionTargetPageId="31" captionText="FIGURE 14. AE: Titillators of Habetia bivittata sp. nov. (AB in situ, male from Goroka, CE dissected, male from Moife).— A, abdominal apex in lateral view; B, view through the gap of the subgenital plate on the apical widenings of the titillators; CE, titillators of male phallus in ventro-apical (C), lateral (D), and dorso-proximal (E) views.—F: Habetia imitatrix Karny, 1912 male subgenital plate with apical projection in lateral view.—Abbreviations: aw apical widening, b base of titillator, dc dorsal cap, dp dorso-apical projection, ls lateral sclerite, pp proximal projection, ti titillator stem, ts testes, vp ventral projection; st stylus of subgenital plate. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223205" httpUri="https://zenodo.org/record/5223205/files/figure.png" pageId="9" pageNumber="210">14F</figureCitation>
). Titillators mostly black; at base little recurved and with narrowing margins, then straight and approaching each other; before performing a curvature slightly widened and diverging, then performing a more than 180° curve, in apical area straight with parallel margins and truncate but with shortly recurved tip (
<figureCitation id="EA9A4ACEAA368057FE9A3254953A6901" box="[973,1106,1087,1113]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: AC, H. tuta sp. nov. (A Quelllager, BC Töpferfluss); DE, H. pedala sp. nov. (Toricelli Mts); FI, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="9" pageNumber="210">Figs 12GI</figureCitation>
); lateral sclerites elongate, of same color as titillators, with rounded ends and a small notch in about mid-length.
</paragraph>
<paragraph id="721E564BAA368057FD9032EC90F969F9" blockId="9.[151,1437,151,1222]" box="[199,401,1159,1185]" pageId="9" pageNumber="210">Female unknown.</paragraph>
<paragraph id="721E564BAA368057FD9032C09440699D" blockId="9.[151,1437,151,1222]" box="[199,1320,1194,1222]" pageId="9" pageNumber="210">
Measurements (
<specimenCount id="64A79DC2AA368057FC2A32C090A0699E" box="[381,456,1195,1222]" pageId="9" pageNumber="210" type="male">1 male</specimenCount>
).—Body w/o wings: 35; pronotum: 9.3; tegmen: 33.5; hind femur:
<quantity id="B559FBAEAA368057F9EA32C1944C699D" box="[1213,1316,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="9" pageNumber="210" unit="mm" value="24.5">24.5 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>