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<document id="21D71BF62B855D3FA2F4DE12BB6D5A25" ID-DOI="10.11646/zootaxa.5020.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5223173" ID-ZooBank="4FF882DF-334F-49C8-A576-4192B5F2654C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629388374973" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2021" docId="FA08E75DAA378056FDC034A9902A6889" docLanguage="en" docName="zootaxa.5020.2.1.pdf" docOrigin="Zootaxa 5020 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habetia spada Brunner 1898" docType="treatment" docVersion="3" lastPageNumber="209" masterDocId="06319F25AA3F805EFD57366B91686D58" masterDocTitle="Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="209" updateTime="1699071125630" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CEBA0907534E3056890AE28348B9FE16">Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)</mods:title>
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<mods:namePart id="F0A87316ABEE163C630FC9E092FC97BB">Ingrisch, Sigfrid</mods:namePart>
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<mods:typeOfResource id="868C0DAED4AEB814A8C56EF255626784">text</mods:typeOfResource>
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<mods:title id="40F09089E35C55D84E25287A6D237A8F">Zootaxa</mods:title>
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<mods:date id="6EA17CF75AD06506A505FFEC050E80FE">2021</mods:date>
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<mods:number id="12588498968D3BA9CEBD9B41843F1C6B">2021-08-12</mods:number>
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<mods:identifier id="BE82EB56B10E3F08E1593D1A21AFDF06" type="ISSN">1175-5326</mods:identifier>
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<treatment id="FA08E75DAA378056FDC034A9902A6889" ID-DOI="http://doi.org/10.5281/zenodo.5223203" ID-Zenodo-Dep="5223203" LSID="urn:lsid:plazi:treatment:FA08E75DAA378056FDC034A9902A6889" httpUri="http://treatment.plazi.org/id/FA08E75DAA378056FDC034A9902A6889" lastPageNumber="209" pageId="8" pageNumber="209">
<subSubSection id="3ABB05C0AA378056FDC034A993D66F84" box="[151,702,706,733]" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="721E564BAA378056FDC034A993D66F84" blockId="8.[151,702,706,769]" box="[151,702,706,733]" pageId="8" pageNumber="209">
<heading id="2956E127AA378056FDC034A993D66F84" bold="true" box="[151,702,706,733]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
<emphasis id="40D58A59AA378056FDC034A993D66F84" bold="true" box="[151,702,706,733]" pageId="8" pageNumber="209">
<taxonomicName id="B5A12DC8AA378056FDC034A9902D6F85" ID-CoL="3J4HQ" authorityName="Brunner" authorityYear="1898" box="[151,325,706,733]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056FDC034A9902D6F85" bold="true" box="[151,325,706,733]" italics="true" pageId="8" pageNumber="209">Habetia spada</emphasis>
</taxonomicName>
(Brunner v.
<bibRefCitation id="16302BBAAA378056FCB434A893DD6F84" author="Brunner von Wattenwyl, C." box="[483,693,706,733]" pageId="8" pageNumber="209" pagination="193 - 288" refId="ref42428" refString="Brunner von Wattenwyl, C. (1898) Orthopteren des Malayischen Archipels, gesammelt von Prof. Dr. W. Kukenthal in den Jahren 1893 und 1894. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, M 24, 193 - 288, pls. 16 - 20." type="journal article" year="1898">Wattenwyl, 1898</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378056FDC0348D90A76E59" box="[151,463,742,769]" pageId="8" pageNumber="209" type="description">
<paragraph id="721E564BAA378056FDC0348D90A76E59" blockId="8.[151,702,706,769]" box="[151,463,742,769]" pageId="8" pageNumber="209">
<figureCitation id="EA9A4ACEAA378056FDC0348D919B6E59" box="[151,243,742,769]" captionStart="FIGURE 1" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[163,1425,181,1920]" captionTargetId="figure-19@10.[162,1425,181,1920]" captionTargetPageId="10" captionText="FIGURE 1. Habetia sp. females: habitus lateral view; species with very long ovipositor: A. H. spada (Brunner, 1898) holotype; B. H. tuta sp. nov. (Keram River); C. H. pedala sp. nov. (Toricelli Mts); D. H. pallida sp. nov. (Sentani); E–F. H. multispinulosa Griffini, 1908 (E holotype, F Rigo Dorom). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223175" httpUri="https://zenodo.org/record/5223175/files/figure.png" pageId="8" pageNumber="209">Figs 1A</figureCitation>
,
<figureCitation id="EA9A4ACEAA378056FDA8348C904B6E59" box="[255,291,743,769]" captionStart="FIGURE 3" captionStartId="17.[151,250,1860,1884]" captionTargetBox="[177,1409,182,1837]" captionTargetId="figure-15@17.[177,1409,181,1837]" captionTargetPageId="17" captionText="FIGURE 3. A, Habetia pilleata sp. nov. male (Wau 1200m); B, H. pallida sp. nov. female (Hollandia); C, H. tuta sp. nov. male (Quelllager); D–F, H. curvata sp. nov. (D, male Albatros Bivak, E–F, female Prauwenbivak); G, H. spada (Brunner, 1898) female holotype (Konstantinhafen); H, H. multispinulosa Griffini, 1908 female holotype (Dilo); I, H. pedala sp. nov. female (Toricelli Mountains).—Frons (A–E, G–I), female dorsal view (F). Scales 5 mm (A–E, G–I), 10 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5223179" httpUri="https://zenodo.org/record/5223179/files/figure.png" pageId="8" pageNumber="209">3G</figureCitation>
,
<figureCitation id="EA9A4ACEAA378056FC78348D903B6E59" box="[303,339,742,769]" captionStart="FIGURE 5" captionStartId="19.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1620]" captionTargetId="figure-15@19.[151,1436,181,1621]" captionTargetPageId="19" captionText="FIGURE 5. Female subgenital plate of Habetia species in oblique ventral (A, J), ventral (C, E, G, L), apico-ventral (B, D, G, K), apical (H), and lateral (F, I) view.—A, H. spada holotype; B–C, H. multispinulosa (B holotype, C from Dorom), D, H. pedala sp. nov. (Toricelli Mts); E–F, H. tuta sp. nov. (Keram river); G, H. curvata sp. nov. (Prauwenbivak); H–L, H. pallida sp. nov. (H–I Jutefa Bay, J Bewani, K Ifar, L Hollandia).—Scales 1 mm; A, D, G not to scale." figureDoi="http://doi.org/10.5281/zenodo.5223183" httpUri="https://zenodo.org/record/5223183/files/figure.png" pageId="8" pageNumber="209">5A</figureCitation>
, Map 1(1).
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378056FDC0354491916EC9" pageId="8" pageNumber="209" type="materials_examined">
<paragraph id="721E564BAA378056FDC0354491916EC9" blockId="8.[151,1437,814,1489]" pageId="8" pageNumber="209">
<materialsCitation id="C2C95C16AA378056FDC0354491916EC9" collectionCode="NMW" country="Papua New Guinea" latitude="-5.5" location="Naturhistorisches Museum Wien" longLatPrecision="1305" longitude="145.93333" municipality="Konstantinhafen" pageId="8" pageNumber="209" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="40D58A59AA378056FDC03544901E6E10" bold="true" box="[151,374,814,841]" pageId="8" pageNumber="209">
<typeStatus id="AD1AE8E9AA378056FDC03544906C6E11" box="[151,260,815,841]" pageId="8" pageNumber="209" type="holotype">Holotype</typeStatus>
(female):
</emphasis>
<collectingCountry id="0AB616DBAA378056FC2B3544933E6E10" box="[380,598,814,841]" name="Papua New Guinea" pageId="8" pageNumber="209">Papua New Guinea</collectingCountry>
,
<collectingRegion id="B06598A9AA378056FF36354593A86E10" box="[609,704,814,840]" country="Papua New Guinea" name="Madang" pageId="8" pageNumber="209">Madang</collectingRegion>
,
<collectingMunicipality id="927ACC31AA378056FF9D354592EB6E10" box="[714,899,814,840]" pageId="8" pageNumber="209">Konstantinhafen</collectingMunicipality>
[= Enke Port, the label reads “Konstantinshafen Deutsch-Neuguinea”], (
<geoCoordinate id="1795308CAA378056FCFC353990936E35" box="[427,507,850,877]" degrees="5" direction="south" minutes="30" orientation="latitude" pageId="8" pageNumber="209" precision="925" value="-5.5">5°30’S</geoCoordinate>
,
<geoCoordinate id="1795308CAA378056FF47353993176E35" box="[528,639,850,877]" degrees="145" direction="east" minutes="56" orientation="longitude" pageId="8" pageNumber="209" precision="925" value="145.93333">145°56’E</geoCoordinate>
), coll. BrW. 20.597—depository:
<location id="777E0090AA378056F94B353994F56E34" LSID="urn:lsid:plazi:treatment:FA08E75DAA378056FDC034A9902A6889:777E0090AA378056F94B353994F56E34" box="[1052,1437,850,877]" country="Papua New Guinea" latitude="-5.5" longLatPrecision="1305" longitude="145.93333" municipality="Konstantinhafen" name="Naturhistorisches Museum Wien" pageId="8" pageNumber="209" stateProvince="Madang">Naturhistorisches Museum Wien</location>
(
<collectionCode id="14B0CE8EAA378056FDC8351C91856EC9" box="[159,237,887,913]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="209">NMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378056FD9035F193556930" pageId="8" pageNumber="209" type="diagnosis">
<paragraph id="721E564BAA378056FD9035F193556930" blockId="8.[151,1437,814,1489]" pageId="8" pageNumber="209">
<emphasis id="40D58A59AA378056FD9035F1902A6EEC" bold="true" box="[199,322,922,948]" pageId="8" pageNumber="209">Diagnosis.</emphasis>
<taxonomicName id="B5A12DC8AA378056FC1C35F090DB6EEC" authorityName="Brunner" authorityYear="1898" box="[331,435,922,948]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056FC1C35F090DB6EEC" box="[331,435,922,948]" italics="true" pageId="8" pageNumber="209">H. spada</emphasis>
</taxonomicName>
differs from other species by the shape of the female subgenital plate that is flattened and has the lateral margins not markedly upcurved, the apical margin is wide-triangularly incised and the spines at apico-lateral angles are nearly rectangularly upcurved. The females from Toricelli Mountains that Karny identified as
<taxonomicName id="B5A12DC8AA378056FDE1326C90736978" authorityName="Brunner" authorityYear="1898" box="[182,283,1030,1056]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056FDE1326C90736978" box="[182,283,1030,1056]" italics="true" pageId="8" pageNumber="209">H. spada</emphasis>
</taxonomicName>
, however, differ significantly from the
<typeStatus id="AD1AE8E9AA378056FF80326C926F6979" box="[727,775,1031,1057]" pageId="8" pageNumber="209">type</typeStatus>
of
<taxonomicName id="B5A12DC8AA378056FE7A326C92FB6978" authorityName="Brunner" authorityYear="1898" box="[813,915,1030,1056]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056FE7A326C92FB6978" box="[813,915,1030,1056]" italics="true" pageId="8" pageNumber="209">H. spada</emphasis>
</taxonomicName>
by the shape of the subgenital plate despite of the general similarity between both taxa as described below under
<taxonomicName id="B5A12DC8AA378056FE203240928C691C" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[887,996,1066,1092]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="pedala" status="sp. nov.">
<emphasis id="40D58A59AA378056FE203240928C691C" box="[887,996,1066,1092]" italics="true" pageId="8" pageNumber="209">H. pedala</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA378056FEBD3240952A691D" bold="true" box="[1002,1090,1067,1093]" pageId="8" pageNumber="209">
<taxonomicNameLabel id="5BE63722AA378056FEBD3240952A691D" box="[1002,1090,1067,1093]" pageId="8" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which has a domed subgenital plate while in
<taxonomicName id="B5A12DC8AA378056FC60322490F46930" authorityName="Brunner" authorityYear="1898" box="[311,412,1102,1128]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA378056FC60322490F46930" box="[311,412,1102,1128]" italics="true" pageId="8" pageNumber="209">H. spada</emphasis>
</taxonomicName>
it is flattened.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA378056FD903219902A6889" pageId="8" pageNumber="209" type="description">
<paragraph id="721E564BAA378056FD90321995BA698D" blockId="8.[151,1437,814,1489]" pageId="8" pageNumber="209">
<emphasis id="40D58A59AA378056FD903219903169D4" bold="true" box="[199,345,1138,1164]" pageId="8" pageNumber="209">Description.</emphasis>
Index fastigium verticis: eye diameter 1.70:
<quantity id="B559FBAEAA378056FE36321992A069D5" box="[865,968,1138,1165]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" pageId="8" pageNumber="209" unit="mm" value="1.96">1.96 mm</quantity>
. Coloration of face of light general color; mandibles fully black; antennal scrobae and tip of fastigium frontis light brown (
<figureCitation id="EA9A4ACEAA378056F95C32FD950C69E9" box="[1035,1124,1174,1201]" captionStart="FIGURE 3" captionStartId="17.[151,250,1860,1884]" captionTargetBox="[177,1409,182,1837]" captionTargetId="figure-15@17.[177,1409,181,1837]" captionTargetPageId="17" captionText="FIGURE 3. A, Habetia pilleata sp. nov. male (Wau 1200m); B, H. pallida sp. nov. female (Hollandia); C, H. tuta sp. nov. male (Quelllager); D–F, H. curvata sp. nov. (D, male Albatros Bivak, E–F, female Prauwenbivak); G, H. spada (Brunner, 1898) female holotype (Konstantinhafen); H, H. multispinulosa Griffini, 1908 female holotype (Dilo); I, H. pedala sp. nov. female (Toricelli Mountains).—Frons (A–E, G–I), female dorsal view (F). Scales 5 mm (A–E, G–I), 10 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5223179" httpUri="https://zenodo.org/record/5223179/files/figure.png" pageId="8" pageNumber="209">Fig. 3G</figureCitation>
). Femora with the following number of spines on ventral margins: Femur spines: (1) a 6 p 8; (2) a 7 p 3; (3) a 12–14 p 13–14.
</paragraph>
<paragraph id="721E564BAA378056FD9032B5901069A0" blockId="8.[151,1437,814,1489]" box="[199,376,1246,1272]" pageId="8" pageNumber="209">Male unknown.</paragraph>
<paragraph id="721E564BAA378056FD90336990C668D1" blockId="8.[151,1437,814,1489]" pageId="8" pageNumber="209">
Female. Subgenital plate rather flat with converging lateral margins and with a medial furrow; at posterior-lateral angles with up-bent, acute cones; apical margin between cones wide, triangularly incised (
<figureCitation id="EA9A4ACEAA378056F9E7334D94666819" box="[1200,1294,1318,1345]" captionStart="FIGURE 5" captionStartId="19.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1620]" captionTargetId="figure-15@19.[151,1436,181,1621]" captionTargetPageId="19" captionText="FIGURE 5. Female subgenital plate of Habetia species in oblique ventral (A, J), ventral (C, E, G, L), apico-ventral (B, D, G, K), apical (H), and lateral (F, I) view.—A, H. spada holotype; B–C, H. multispinulosa (B holotype, C from Dorom), D, H. pedala sp. nov. (Toricelli Mts); E–F, H. tuta sp. nov. (Keram river); G, H. curvata sp. nov. (Prauwenbivak); H–L, H. pallida sp. nov. (H–I Jutefa Bay, J Bewani, K Ifar, L Hollandia).—Scales 1 mm; A, D, G not to scale." figureDoi="http://doi.org/10.5281/zenodo.5223183" httpUri="https://zenodo.org/record/5223183/files/figure.png" pageId="8" pageNumber="209">Fig. 5A</figureCitation>
). Ovipositor about one and half time as long as body; behind basal constriction straight with dorsal valves in basal half only faintly convex (
<figureCitation id="EA9A4ACEAA378056FC10330590C868D1" box="[327,416,1390,1417]" captionStart="FIGURE 1" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[163,1425,181,1920]" captionTargetId="figure-19@10.[162,1425,181,1920]" captionTargetPageId="10" captionText="FIGURE 1. Habetia sp. females: habitus lateral view; species with very long ovipositor: A. H. spada (Brunner, 1898) holotype; B. H. tuta sp. nov. (Keram River); C. H. pedala sp. nov. (Toricelli Mts); D. H. pallida sp. nov. (Sentani); E–F. H. multispinulosa Griffini, 1908 (E holotype, F Rigo Dorom). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223175" httpUri="https://zenodo.org/record/5223175/files/figure.png" pageId="8" pageNumber="209">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="721E564BAA378056FD9033F8902A6889" blockId="8.[151,1437,814,1489]" pageId="8" pageNumber="209">
Measurements (
<specimenCount id="64A79DC2AA378056FCD733F8908F68F5" box="[384,487,1426,1453]" pageId="8" pageNumber="209" type="female">1 female</specimenCount>
).—Body w/o wings: 42; pronotum: 9.3; tegmen: 38; hind femur: 28.5; ovipositor length:
<quantity id="B559FBAEAA378056FDBB33DD90566889" box="[236,318,1462,1489]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="8" pageNumber="209" unit="mm" value="63.0">63 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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