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<mods:title id="EC1F0DE8A7B3AC113373D1C88CAC79E0">A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter &amp; Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of ‘ Canthon sensu lato’</mods:title>
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<mods:namePart id="F9FB3D3157095B107BF8B943371EBED0">Cupello, Mario</mods:namePart>
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<emphasis id="1DF1EAFFFFAFFF860EDF0D6009F89101" bold="true" box="[452,726,321,348]" italics="true" pageId="59" pageNumber="60">Sylvicanthon enkerlini</emphasis>
(
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Martínez 
<emphasis id="1DF1EAFFFFAFFF860C420D6308BA9106" box="[857,916,321,347]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
, 1964
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)
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<paragraph id="2F3A36EDFFAFFF860D080D440F0E9122" blockId="59.[452,1136,321,384]" box="[531,1056,357,384]" pageId="59" pageNumber="60">
<figureCitation id="B7BE2A68FFAFFF860D080D44095A9122" box="[531,628,357,383]" captionStart="Fig" captionStartId="20.[189,232,1928,1954]" captionTargetBox="[212,1372,265,1894]" captionTargetId="figure@20.[212,1376,265,1895]" captionTargetPageId="20" captionText="Fig. 6. Head, dorsal view. A. Sylvicanthon enkerlini (Martínez et al., 1964) comb. nov. B. S. candezei (Harold, 1869). C. S. genieri sp. nov. D. S. proseni (Martínez, 1949) stat. et comb. nov. E. S. seag sp. nov. F. S. securus (Schmidt, 1920) comb. nov. G. S. furvus (Schmidt, 1920)." figureDoi="http://doi.org/10.5281/zenodo.3824791" httpUri="https://zenodo.org/record/3824791/files/figure.png" pageId="59" pageNumber="60">Figs 6A</figureCitation>
, 
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B–D, 10B, 11A, 15A, 17A, 20–22
</paragraph>
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<emphasis id="1DF1EAFFFFAFFF860FA60D860AA5919C" box="[189,395,423,449]" italics="true" pageId="59" pageNumber="60">Glaphyrocanthon</emphasis>
(
<emphasis id="1DF1EAFFFFAFFF860E870D860944919C" box="[412,618,423,449]" italics="true" pageId="59" pageNumber="60">Glaphyrocanthon</emphasis>
) 
<emphasis id="1DF1EAFFFFAFFF860D600D8609CD919C" box="[635,739,423,449]" italics="true" pageId="59" pageNumber="60">enkerlini</emphasis>
<bibRefCitation id="4B144B1CFFAFFF860DF10D8608D1919F" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[746,1023,423,450]" pageId="59" pageNumber="60" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFAFFF860C470D8908B8919C" box="[860,918,423,449]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
, 1964: 5
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, 8, 13, 17–21, figs 1–2.
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<emphasis id="1DF1EAFFFFAFFF860FA60DC80A0C925E" box="[189,290,489,515]" italics="true" pageId="59" pageNumber="60">Canthon</emphasis>
(
<emphasis id="1DF1EAFFFFAFFF860E280DCB0AC6925E" box="[307,488,490,515]" italics="true" pageId="59" pageNumber="60">Francmonrosia</emphasis>
) 
<emphasis id="1DF1EAFFFFAFFF860EE10DC8094C925E" box="[506,610,489,515]" italics="true" pageId="59" pageNumber="60">enkerlini</emphasis>
</taxonomicName>
– 
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Halffter &amp; Martínez 
<date id="5B3B102DFFAFFF860C690DCB089F9259" box="[882,945,490,516]" pageId="59" pageNumber="60" value="1977">1977</date>
: 86
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.
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</paragraph>
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<treatmentCitationGroup id="0F9511C3FFAFFF860FA60E2C0953927A" box="[189,637,525,552]" pageId="59" pageNumber="60">
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<emphasis id="1DF1EAFFFFAFFF860FA60E2C0ABF927A" box="[189,401,525,551]" italics="true" pageId="59" pageNumber="60">Canthon enkerlini</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFAFFF860EB40E2C0957927A" author="Krajcik M." box="[431,633,525,551]" page="63" pageId="59" pageNumber="60" year="2012">
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Krajcik 
<date id="5B3B102DFFAFFF860D150E2C0963927A" box="[526,589,525,551]" pageId="59" pageNumber="60" value="2012">2012</date>
: 63
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.
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</paragraph>
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<emphasis id="1DF1EAFFFFAFFF860FA60E100A7B9216" box="[189,341,561,587]" italics="true" pageId="59" pageNumber="60">Sylvicanthon</emphasis>
</taxonomicName>
sp. – 
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<emphasis id="1DF1EAFFFFAFFF860EFB0E1309359216" box="[480,539,561,587]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
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: 91–93
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.
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</paragraph>
</subSubSection>
<subSubSection id="679F6566FFAFFF860FA60E520A2792BC" pageId="59" pageNumber="60" type="etymology">
<paragraph id="2F3A36EDFFAFFF860FA60E520A6A92D0" blockId="59.[189,324,627,653]" box="[189,324,627,653]" pageId="59" pageNumber="60">
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<emphasis id="1DF1EAFFFFAFFF860FA60E520A6A92D0" bold="true" box="[189,324,627,653]" pageId="59" pageNumber="60">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFAFFF860FA60E820A2792BC" blockId="59.[189,1399,674,737]" pageId="59" pageNumber="60">
Eponym after the Mexican entomologist Dieter Enkerlin Schallenmüller (
<date id="5B3B102DFFAFFF860B3D0E830F8092E0" box="[1062,1198,674,701]" pageId="59" pageNumber="60" value="1926" valueMax="1995">
<date id="5B3B102DFFAFFF860B3D0E830F4C92E0" box="[1062,1122,674,701]" pageId="59" pageNumber="60" value="1926" year="1926">1926</date>
– 
<date id="5B3B102DFFAFFF860B680E830F8092E0" box="[1139,1198,674,701]" pageId="59" pageNumber="60" value="1995" year="1995">1995</date>
</date>
) (
<bibRefCitation id="4B144B1CFFAFFF860BDD0E830BD292BC" author="Martinez A. &amp; Halffter G. &amp; Halffter V." pageId="59" pageNumber="60" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFAFFF860A200E820E5992E1" box="[1339,1399,674,700]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
<date id="5B3B102DFFAFFF860FA60EE70BD292BC" box="[189,252,710,737]" pageId="59" pageNumber="60" value="1964">1964</date>
</bibRefCitation>
).
</paragraph>
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<emphasis id="1DF1EAFFFFAFFF860FA60F290A84937F" bold="true" box="[189,426,776,802]" pageId="59" pageNumber="60">Material examined</emphasis>
</heading>
</paragraph>
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</materialsCitation>
</paragraph>
<paragraph id="2F3A36EDFFAFFF860FA60F7D0AB493BC" blockId="59.[189,1398,824,994]" pageId="59" pageNumber="60">
<collectingCountry id="5792767DFFAFFF860FA60F7D0A02932B" box="[189,300,860,886]" name="Brazil" pageId="59" pageNumber="60">BRAZIL</collectingCountry>
: 
<specimenCount id="3983FD64FFAFFF860E220F780A619328" box="[313,335,857,885]" pageId="59" pageNumber="60" type="male">♂</specimenCount>
, 
<collectingRegion id="ED41F80FFFAFFF860E7B0F7A0AF49328" box="[352,474,859,885]" country="Brazil" name="Maranhao" pageId="59" pageNumber="60">Maranhão</collectingRegion>
, 
<collectingMunicipality id="CF5EAC97FFAFFF860EF20F7A097D9328" box="[489,595,859,885]" pageId="59" pageNumber="60">São Luís</collectingMunicipality>
(“ 
<emphasis id="1DF1EAFFFFAFFF860D6F0F7D09FD932B" box="[628,723,860,886]" italics="true" pageId="59" pageNumber="60">
<collectingCountry id="5792767DFFAFFF860D6F0F7D09FD932B" box="[628,723,860,886]" name="Brazil" pageId="59" pageNumber="60">BRASIL</collectingCountry>
</emphasis>
/ 
<emphasis id="1DF1EAFFFFAFFF860DF50F7D08A19328" box="[750,911,859,886]" italics="true" pageId="59" pageNumber="60">
Eº 
<collectingRegion id="ED41F80FFFAFFF860C080F7A08A19328" box="[787,911,859,885]" country="Brazil" name="Maranhao" pageId="59" pageNumber="60">Maranhao</collectingRegion>
</emphasis>
/ 
<location id="2A5A6036FFAFFF860CB70F7D0E02932B" LSID="urn:lsid:plazi:treatment:A72C87FBFFAFFFFD0EDF0D600E599295:2A5A6036FFAFFF860CB70F7D0E02932B" box="[940,1324,859,886]" country="Brazil" municipality="Sao Luis" name="Sao Luiz / Coll. Martinez / Sep." pageId="59" pageNumber="60" stateProvince="Maranhao">
<emphasis id="1DF1EAFFFFAFFF860CB70F7D0F3B9328" box="[940,1045,860,886]" italics="true" pageId="59" pageNumber="60">São Luiz</emphasis>
/ 
<emphasis id="1DF1EAFFFFAFFF860B2B0F7A0FF0932B" box="[1072,1246,859,886]" italics="true" pageId="59" pageNumber="60">Coll. Martínez</emphasis>
/ 
<emphasis id="1DF1EAFFFFAFFF860BE10F7D0E02932B" box="[1274,1324,860,886]" italics="true" pageId="59" pageNumber="60">Sep.</emphasis>
</location>
955”, “ 
<typeStatus id="F03E884FFFAFFF860FDC0F5E0A5093C4" box="[199,382,895,921]" pageId="59" pageNumber="60" type="holotype">HOLOTYPUS</typeStatus>
”, “ 
<taxonomicName id="E8854D6EFFAFFF860EBF0F5E095C93C4" box="[420,626,895,921]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFFAFFF860EBF0F5E095C93C4" box="[420,626,895,921]" italics="true" pageId="59" pageNumber="60">
<location id="2A5A6036FFAFFF860EBF0F5E095C93C4" LSID="urn:lsid:plazi:treatment:A72C87FBFFAFFFFD0EDF0D600E599295:2A5A6036FFAFFF860EBF0F5E095C93C4" box="[420,626,895,921]" country="Brazil" municipality="Sao Luis" name="Glaphyrocanthon" pageId="59" pageNumber="60" stateProvince="Maranhao">Glaphyrocanthon</location>
</emphasis>
</taxonomicName>
/ 
<taxonomicName id="E8854D6EFFAFFF860D940F5E09D993C4" authorityName="Martinez, Halffter &amp; Halffter" authorityYear="1964" box="[655,759,895,921]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFAFFF860D940F5E09D993C4" box="[655,759,895,921]" italics="true" pageId="59" pageNumber="60">enkerlini</emphasis>
</taxonomicName>
/ 
<specimenCount id="3983FD64FFAFFF860C0F0F5C080493C4" box="[788,810,893,921]" pageId="59" pageNumber="60" type="male">♂</specimenCount>
/ 
<emphasis id="1DF1EAFFFFAFFF860C5D0FA1088C93C4" box="[838,930,896,921]" italics="true" pageId="59" pageNumber="60">
<specimenCount id="3983FD64FFAFFF860C5D0FA1084693C4" box="[838,872,896,921]" pageId="59" pageNumber="60" type="generic">sp.</specimenCount>
nov.
</emphasis>
/ 
<emphasis id="1DF1EAFFFFAFFF860CA50FA10F6A93C4" box="[958,1092,896,921]" italics="true" pageId="59" pageNumber="60">
<collectionCode id="4994AE28FFAFFF860CA50FA108F493C4" box="[958,986,896,921]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860CF30FA10F2F93C4" box="[1000,1025,896,921]" country="Finland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15618" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="59" pageNumber="60" type="Herbarium">H</collectionCode>
. y 
<collectionCode id="4994AE28FFAFFF860B3C0FA10F6E93C4" box="[1063,1088,896,921]" country="Finland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15618" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="59" pageNumber="60" type="Herbarium">H</collectionCode>
.
</emphasis>
/ 
<collectionCode id="4994AE28FFAFFF860B450F5E0F5993C4" box="[1118,1143,895,921]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="59" pageNumber="60" type="Herbarium">A</collectionCode>
. 
<location id="2A5A6036FFAFFF860B9E0F5E0FC193C4" LSID="urn:lsid:plazi:treatment:A72C87FBFFAFFFFD0EDF0D600E599295:2A5A6036FFAFFF860B9E0F5E0FC193C4" box="[1157,1263,895,921]" country="Brazil" municipality="Sao Luis" name="Martinez" pageId="59" pageNumber="60" stateProvince="Maranhao">Martínez</location>
det 19 
<emphasis id="1DF1EAFFFFAFFF860A5E0F5E0E4C93C4" box="[1349,1378,895,921]" italics="true" pageId="59" pageNumber="60">63</emphasis>
”, “ 
<emphasis id="1DF1EAFFFFAFFF860FD10F820A6593E0" box="[202,331,931,957]" italics="true" pageId="59" pageNumber="60">FICHADO</emphasis>
”, “MACN-En / 1050”), genital capsule removed and glued in a triangular label (MACN-En / 1050) (
<figureCitation id="B7BE2A68FFAFFF860E3A0FE60AA393BC" box="[289,397,967,993]" captionStart="Fig" captionStartId="60.[189,232,1964,1990]" captionTargetBox="[199,1392,494,1919]" captionTargetId="figure@60.[189,1399,488,1932]" captionTargetPageId="60" captionText="Fig. 21. Sylvicanthon enkerlini (Martínez et al., 1964) comb. nov. A. Dorsal view. B. Ventral view. C. Holotype and its labels." figureDoi="http://doi.org/10.5281/zenodo.3824821" httpUri="https://zenodo.org/record/3824821/files/figure.png" pageId="59" pageNumber="60">
Fig. 21 
<collectionCode id="4994AE28FFAFFF860E6C0FE60AA393BC" box="[375,397,967,993]" country="Denmark" name="University of Copenhagen" pageId="59" pageNumber="60" type="Herbarium">C</collectionCode>
</figureCitation>
).
</paragraph>
<paragraph id="2F3A36EDFFAFFF860FF1082809B0947E" blockId="59.[189,1399,1031,2027]" box="[234,670,1031,1059]" pageId="59" pageNumber="60">
<emphasis id="1DF1EAFFFFAFFF860FF108280AFD947E" bold="true" box="[234,467,1033,1059]" pageId="59" pageNumber="60">Additinal material</emphasis>
(47 ♂♂, 
<specimenCount id="3983FD64FFAFFF860D58082809BC947E" box="[579,658,1033,1059]" pageId="59" pageNumber="60" type="female">
44 
<specimenCount id="3983FD64FFAFFF860D73082809BC947E" box="[616,658,1033,1059]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
)
</paragraph>
<paragraph id="2F3A36EDFFAFFF800FA6080C0F2891CE" blockId="59.[189,1399,1031,2027]" lastBlockId="61.[189,1399,265,404]" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60">
<collectingCountry id="5792767DFFAFFF860FA6080C0A02941A" box="[189,300,1069,1095]" name="Brazil" pageId="59" pageNumber="60">BRAZIL</collectingCountry>
: 
<emphasis id="1DF1EAFFFFAFFF860E2C080C0AAE941A" bold="true" box="[311,384,1069,1095]" pageId="59" pageNumber="60">
<collectingRegion id="ED41F80FFFAFFF860E2C080C0AAE941A" box="[311,384,1069,1095]" country="Brazil" name="Bahia" pageId="59" pageNumber="60">Bahia</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFAFFF860E8B080C0A93941A" box="[400,445,1069,1095]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860EBC080C0A93941A" box="[423,445,1069,1095]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Barreiras, 
<date id="5B3B102DFFAFFF860D5D080C0993941A" box="[582,701,1069,1095]" pageId="59" pageNumber="60" value="1991-10">
Oct. 
<date id="5B3B102DFFAFFF860D9B080C0993941A" box="[640,701,1069,1095]" pageId="59" pageNumber="60" value="1991" year="1991">1991</date>
</date>
, light trap, without collector (
<collectionCode id="4994AE28FFAFFF860B07080C0F5A941A" box="[1052,1140,1069,1095]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860B93080C0F9B941A" box="[1160,1205,1069,1095]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860B84080C0F9B941A" box="[1183,1205,1069,1095]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Barreiras, 
<date id="5B3B102DFFAFFF860A24080C0BD49436" pageId="59" pageNumber="60" value="1991-11">
Nov. 
<date id="5B3B102DFFAFFF860FA608700BD49436" box="[189,250,1105,1131]" pageId="59" pageNumber="60" value="1991" year="1991">1991</date>
</date>
, without collector (
<collectionCode id="4994AE28FFAFFF860EC00870091D9436" box="[475,563,1105,1131]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 2 ♂♂, 
<specimenCount id="3983FD64FFAFFF860D8F087009919436" box="[660,703,1105,1131]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860DB2087009919436" box="[681,703,1105,1131]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Barreiras, 
<date id="5B3B102DFFAFFF860C5D087008969436" box="[838,952,1105,1131]" pageId="59" pageNumber="60" value="2003-01">
Jan. 
<date id="5B3B102DFFAFFF860C60087008969436" box="[891,952,1105,1131]" pageId="59" pageNumber="60" value="2003" year="2003">2003</date>
</date>
, 
<collectionCode id="4994AE28FFAFFF860CD8087008FA9436" box="[963,980,1105,1131]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. Schmidt leg. (
<collectionCode id="4994AE28FFAFFF860B9B08700FF69436" box="[1152,1240,1105,1131]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860BF008700E039436" box="[1259,1325,1105,1131]" pageId="59" pageNumber="60" type="female">
2 
<specimenCount id="3983FD64FFAFFF860A1B08700E039436" box="[1280,1325,1105,1131]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Pilão Arcado, Barra do Brejo, 
<geoCoordinate id="4AB1502AFFAFFF860EF7085209A894D2" box="[492,646,1139,1167]" degrees="10" direction="south" minutes="06.677" orientation="latitude" pageId="59" pageNumber="60" precision="1" value="-10.111283">
10º06.677′ 
<collectionCode id="4994AE28FFAFFF860D6F085409A894D2" box="[628,646,1141,1167]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860D8D0852081794D2" box="[662,825,1139,1167]" degrees="42" direction="west" minutes="53.678" orientation="longitude" pageId="59" pageNumber="60" precision="1" value="-42.894634">
42º53.678′ 
<collectionCode id="4994AE28FFAFFF860C060854081794D2" box="[797,825,1141,1167]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860C53085508B694D2" box="[840,920,1140,1167]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="59" pageNumber="60" unit="m" value="416.0">416 m</quantity>
, 
<date id="5B3B102DFFAFFF860CBC08550F6E94D2" box="[935,1088,1140,1167]" pageId="59" pageNumber="60" value="2005-12-05">
<date id="5B3B102DFFAFFF860CBC085508D994D2" box="[935,1015,1140,1167]" day="05" month="12" monthName="Dec" pageId="59" pageNumber="60">
5 
<date id="5B3B102DFFAFFF860CDA085408D994D2" box="[961,1015,1141,1167]" month="12" monthName="Dec" pageId="59" pageNumber="60" year="2005">Dec.</date>
</date>
<date id="5B3B102DFFAFFF860B1908550F6E94D2" box="[1026,1088,1140,1167]" pageId="59" pageNumber="60" value="2005" year="2005">2005</date>
</date>
, 
<collectionCode id="4994AE28FFAFFF860B5508540F7194D2" box="[1102,1119,1141,1167]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860B7908540F5D94D2" box="[1122,1139,1141,1167]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. Lopes leg. (
<collectionCode id="4994AE28FFAFFF860A0908540E4494D2" box="[1298,1386,1141,1167]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860FA608B90A2E94EE" box="[189,256,1176,1203]" pageId="59" pageNumber="60" type="female">
2 
<specimenCount id="3983FD64FFAFFF860FC808B80A2E94EE" box="[211,256,1177,1203]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Pilão Arcado, Barra do Brejo, 
<geoCoordinate id="4AB1502AFFAFFF860D6908B7082794EF" box="[626,777,1174,1202]" degrees="10" direction="south" minutes="06.677" orientation="latitude" pageId="59" pageNumber="60" precision="1" value="-10.111283">
10º06.677′ 
<collectionCode id="4994AE28FFAFFF860DEC08B9082794EF" box="[759,777,1176,1202]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860C0D08B7089994EE" box="[790,951,1174,1203]" degrees="42" direction="west" minutes="53,678" orientation="longitude" pageId="59" pageNumber="60" precision="1" value="-42.894634">
42º53,678′ 
<collectionCode id="4994AE28FFAFFF860C8008B8089994EE" box="[923,951,1177,1203]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860CD908B90F2194EE" box="[962,1039,1176,1203]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="59" pageNumber="60" unit="m" value="416.0">416 m</quantity>
, 
<date id="5B3B102DFFAFFF860B0008B90F9294EF" box="[1051,1212,1176,1203]" pageId="59" pageNumber="60" value="2006-03-27">
<date id="5B3B102DFFAFFF860B0008B90F5994EE" box="[1051,1143,1176,1203]" day="27" month="03" monthName="Mar" pageId="59" pageNumber="60" year="1927">
27 
<date id="5B3B102DFFAFFF860B5B08B80F5994EE" box="[1088,1143,1177,1203]" month="03" monthName="Mar" pageId="59" pageNumber="60" year="2006">Mar.</date>
</date>
2006
</date>
, 
<collectionCode id="4994AE28FFAFFF860BDC08B80FF294EE" box="[1223,1244,1177,1203]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="59" pageNumber="60" type="Museum">V</collectionCode>
. Mendes leg. (
<collectionCode id="4994AE28FFAFFF860FDE089D0A36948B" box="[197,280,1212,1238]" pageId="59" pageNumber="60">MZFS</collectionCode>
). – 
<emphasis id="1DF1EAFFFFAFFF860E5E089D0AE7948B" bold="true" box="[325,457,1212,1238]" pageId="59" pageNumber="60">
<collectingRegion id="ED41F80FFFAFFF860E5E089D0AE7948B" box="[325,457,1212,1238]" country="Brazil" name="Maranhao" pageId="59" pageNumber="60">Maranhão</collectingRegion>
</emphasis>
: 2 ♂♂, 
<specimenCount id="3983FD64FFAFFF860D36089D095C948A" box="[557,626,1212,1239]" pageId="59" pageNumber="60" type="female">
4 
<specimenCount id="3983FD64FFAFFF860D5E089C095C948A" box="[581,626,1213,1239]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Mirador, Parque Estadual do Mirador, Povoado Pindaíba (Mel), 
<geoCoordinate id="4AB1502AFFAFFF860FA608FF0A6694A7" box="[189,328,1246,1274]" degrees="06" direction="south" minutes="41" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="06" value="-6.685">
06º41′06″ 
<collectionCode id="4994AE28FFAFFF860E2D08C10A6694A7" box="[310,328,1248,1274]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860E4F08FF0AC694A6" box="[340,488,1246,1275]" degrees="45" direction="west" minutes="00" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="26" value="-45.00722">
45º00′26″ 
<collectionCode id="4994AE28FFAFFF860ED708C00AC694A6" box="[460,488,1249,1275]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<date id="5B3B102DFFAFFF860EE908C109B694A7" box="[498,664,1248,1275]" pageId="59" pageNumber="60" value="2010-06-01" valueMax="2010-06-05" valueMin="2010-06-01">
1– 
<date id="5B3B102DFFAFFF860D1408C109B694A7" box="[527,664,1248,1275]" day="05" month="06" pageId="59" pageNumber="60" partOrder="dmy" value="2010-06-05" year="2010">
<date id="5B3B102DFFAFFF860D1408C1097A94A6" box="[527,596,1248,1275]" day="05" month="06" monthName="Jun" pageId="59" pageNumber="60">
5 
<date id="5B3B102DFFAFFF860D3F08C0097A94A6" box="[548,596,1249,1275]" month="06" monthName="Jun" pageId="59" pageNumber="60" year="2010">Jun.</date>
</date>
2010
</date>
</date>
, pitfall with human faeces, 
<collectionCode id="4994AE28FFAFFF860CC908C008CA94A6" box="[978,996,1249,1275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="59" pageNumber="60" type="Herbarium">F</collectionCode>
. Limeira-de-Oliveira, 
<collectionCode id="4994AE28FFAFFF860BFD08C00E2A94A6" box="[1254,1284,1249,1275]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860A1308C00E0894A6" box="[1288,1318,1249,1275]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. Abreu and 
<collectionCode id="4994AE28FFAFFF860FEA09250A2E9543" box="[241,256,1284,1310]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14647" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="59" pageNumber="60" type="Herbarium">J</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860E1F09250A389543" box="[260,278,1284,1310]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
. Pinto leg. (
<collectionCode id="4994AE28FFAFFF860EB20925092F9543" box="[425,513,1284,1310]" pageId="59" pageNumber="60">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFAFFF860D34092509F49543" bold="true" box="[559,730,1284,1310]" pageId="59" pageNumber="60">
<collectingRegion id="ED41F80FFFAFFF860D34092509F49543" box="[559,730,1284,1310]" country="Brazil" name="Minas Gerais" pageId="59" pageNumber="60">Minas Gerais</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFAFFF860DF0092508349543" box="[747,794,1284,1310]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860C1F092508349543" box="[772,794,1284,1310]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Bandeira (“ 
<emphasis id="1DF1EAFFFFAFFF860CAF09240F6A9543" box="[948,1092,1284,1310]" italics="true" pageId="59" pageNumber="60">
<collectionCode id="4994AE28FFAFFF860CAF092408E79543" box="[948,969,1285,1310]" country="Chile" httpUri="http://grbio.org/cool/az7k-p4g9" name="Departamento de Geologia, Universidad de Chile" pageId="59" pageNumber="60">R</collectionCode>
. Bandeira
</emphasis>
”), 
<date id="5B3B102DFFAFFF860B7709250FCF9543" box="[1132,1249,1284,1310]" pageId="59" pageNumber="60" value="1932-01">Jan. 1932</date>
, José Blaser leg. (
<collectionCode id="4994AE28FFAFFF860FEC09090A64951F" box="[247,330,1320,1346]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 1 ♂, Jaíba, Mocambinho, 
<geoCoordinate id="4AB1502AFFAFFF860D9609070801951F" box="[653,815,1318,1346]" degrees="15" direction="south" minutes="09" orientation="latitude" pageId="59" pageNumber="60" precision="1" seconds="03.1" value="-15.150861">
15º09′03.1″ 
<collectionCode id="4994AE28FFAFFF860C0609090801951F" box="[797,815,1320,1346]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860C27090708C6951F" box="[828,1000,1318,1346]" degrees="43" direction="west" minutes="56" orientation="longitude" pageId="59" pageNumber="60" precision="1" seconds="03.4" value="-43.93428">
43º56′03.4″ 
<collectionCode id="4994AE28FFAFFF860CD7090908C6951F" box="[972,1000,1320,1346]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860CE809060F11951F" box="[1011,1087,1319,1346]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="59" pageNumber="60" unit="m" value="450.0">450 m</quantity>
, 
<date id="5B3B102DFFAFFF860B5009090FC5951F" box="[1099,1259,1320,1346]" pageId="59" pageNumber="60" value="2012-03-28">
<date id="5B3B102DFFAFFF860B5009090F88951F" box="[1099,1190,1320,1346]" day="28" month="03" monthName="Mar" pageId="59" pageNumber="60" year="1928">
28 
<date id="5B3B102DFFAFFF860B7409090F88951F" box="[1135,1190,1320,1346]" month="03" monthName="Mar" pageId="59" pageNumber="60" year="2012">Mar.</date>
</date>
2012
</date>
, pitfall with human faeces, 
<collectionCode id="4994AE28FFAFFF860E75096D0AA9953B" box="[366,391,1356,1382]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="59" pageNumber="60" type="Herbarium">A</collectionCode>
. Fialho leg. (
<collectionCode id="4994AE28FFAFFF860D3F096D0952953B" box="[548,636,1356,1382]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860D89096D09EE953B" box="[658,704,1356,1382]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860DB1096D09EE953B" box="[682,704,1356,1382]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Três Marias, 
<date id="5B3B102DFFAFFF860C75096D0F3C953B" box="[878,1042,1356,1382]" pageId="59" pageNumber="60" value="1992-03-17">
<date id="5B3B102DFFAFFF860C75096D08E2953B" box="[878,972,1356,1382]" day="17" month="03" monthName="Mar" pageId="59" pageNumber="60" year="2017">
17 
<date id="5B3B102DFFAFFF860C8E096D08E2953B" box="[917,972,1356,1382]" month="03" monthName="Mar" pageId="59" pageNumber="60" year="1992">Mar.</date>
</date>
1992
</date>
, without collector (
<collectionCode id="4994AE28FFAFFF860BE0096D0E7D953B" box="[1275,1363,1356,1382]" pageId="59" pageNumber="60">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFAFFF860FA6094F0BD095D4" bold="true" box="[189,254,1390,1417]" pageId="59" pageNumber="60">
<collectingRegion id="ED41F80FFFAFFF860FA6094F0BD095D4" box="[189,254,1390,1417]" country="Brazil" name="Piaui" pageId="59" pageNumber="60">Piauí</collectingRegion>
</emphasis>
: 1 ♂ (dissected), 
<specimenCount id="3983FD64FFAFFF860EC9094E093895D7" box="[466,534,1391,1418]" pageId="59" pageNumber="60" type="female">
2 
<specimenCount id="3983FD64FFAFFF860EF20951093895D7" box="[489,534,1392,1418]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Canto do Buriti, 
<date id="5B3B102DFFAFFF860DF6094E089295D4" box="[749,956,1391,1418]" pageId="59" pageNumber="60" value="1991-11-18" valueMax="1991-11-22" valueMin="1991-11-18">
<date id="5B3B102DFFAFFF860DF6094E085995D7" box="[749,887,1391,1418]" day="18" last_day="22" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1918-11-22" year="1918">
18– 
<date id="5B3B102DFFAFFF860C03094E085995D7" box="[792,887,1391,1418]" day="22" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1991-11-22" year="1922">
22 
<date id="5B3B102DFFAFFF860C240951085995D7" box="[831,887,1392,1418]" month="11" monthName="Nov" pageId="59" pageNumber="60" year="1991">Nov.</date>
</date>
</date>
1991
</date>
, 
<collectionCode id="4994AE28FFAFFF860CD2094E08F195D4" box="[969,991,1391,1417]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="59" pageNumber="60" type="Herbarium">C</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860CF8094E08D495D4" box="[995,1018,1391,1417]" country="Chile" httpUri="http://grbio.org/cool/az7k-p4g9" name="Departamento de Geologia, Universidad de Chile" pageId="59" pageNumber="60">R</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860CE6094E0F3E95D4" box="[1021,1040,1391,1417]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="59" pageNumber="60" type="Herbarium">F</collectionCode>
. Brandão leg. (
<collectionCode id="4994AE28FFAFFF860BDF094E0E3995D4" box="[1220,1303,1391,1417]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 2 ♂♂, 
<specimenCount id="3983FD64FFAFFF860FA609B20BC395F3" box="[189,237,1427,1454]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860FCC09B50BC395F3" box="[215,237,1428,1454]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Canto do Buriti, 
<geoCoordinate id="4AB1502AFFAFFF860ECA09B0094295F3" box="[465,620,1425,1454]" degrees="42" direction="west" minutes="48" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="05" value="-42.801388">
42º48′05″ 
<collectionCode id="4994AE28FFAFFF860D4B09B5094295F3" box="[592,620,1428,1454]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860D6009B0082595F0" box="[635,779,1425,1453]" degrees="08" direction="south" minutes="11" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="12" value="-8.1866665">
08º11′12″ 
<collectionCode id="4994AE28FFAFFF860DE209B2082595F0" box="[761,779,1427,1453]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860C0709B2084395F3" box="[796,877,1427,1454]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="59" pageNumber="60" unit="m" value="254.0">254 m</quantity>
, 
<date id="5B3B102DFFAFFF860C6609B20F0E95F0" box="[893,1056,1427,1454]" pageId="59" pageNumber="60" value="2008-null-05">
<date id="5B3B102DFFAFFF860C6609B208F895F3" box="[893,982,1427,1454]" day="05" monthName="Apr" pageId="59" pageNumber="60">
5 
<date id="5B3B102DFFAFFF860C8D09B208F895F3" box="[918,982,1427,1454]" monthName="Apr" pageId="59" pageNumber="60" year="2008">Apr.</date>
</date>
2008
</date>
, trap with dung, Gillett leg. (
<collectionCode id="4994AE28FFAFFF860FDE09960A33958C" box="[197,285,1463,1489]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 1 ♂, Corrente (“ 
<quantity id="E87D9B08FFAFFF860EED09960962958C" box="[502,588,1463,1489]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="km" value="10.0">10 km</quantity>
<collectionCode id="4994AE28FFAFFF860D4D09990945958F" box="[598,619,1464,1490]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="59" pageNumber="60" type="Herbarium">N</collectionCode>
Corrente”), Fazenda Maracujá, 
<date id="5B3B102DFFAFFF860CF709960F92958C" box="[1004,1212,1463,1490]" pageId="59" pageNumber="60" value="1991-11-23" valueMax="1991-11-27" valueMin="1991-11-23">
<date id="5B3B102DFFAFFF860CF709960F59958F" box="[1004,1143,1463,1490]" day="23" last_day="27" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1923-11-27" year="1923">
23– 
<date id="5B3B102DFFAFFF860B0309960F59958F" box="[1048,1143,1463,1490]" day="27" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1991-11-27" year="1927">
27 
<date id="5B3B102DFFAFFF860B2509990F59958F" box="[1086,1143,1464,1490]" month="11" monthName="Nov" pageId="59" pageNumber="60" year="1991">Nov.</date>
</date>
</date>
1991
</date>
, 
<collectionCode id="4994AE28FFAFFF860BD109990FF1958F" box="[1226,1247,1464,1490]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="59" pageNumber="60" type="Herbarium">E</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860BF809990E2F958F" box="[1251,1281,1464,1490]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. Cancello and 
<collectionCode id="4994AE28FFAFFF860FEA09FA0A2095A8" box="[241,270,1499,1525]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860E0909FA0A0895A8" box="[274,294,1499,1525]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="59" pageNumber="60">T</collectionCode>
. Ponte leg. (
<collectionCode id="4994AE28FFAFFF860EA509FA093F95A8" box="[446,529,1499,1525]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 1 ♂, Corrente (“ 
<quantity id="E87D9B08FFAFFF860DF509FA086995A8" box="[750,839,1499,1525]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="km" value="10.0">10 km</quantity>
<collectionCode id="4994AE28FFAFFF860C4A09FA084895A8" box="[849,870,1499,1525]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="59" pageNumber="60" type="Herbarium">N</collectionCode>
Corrente”), Fazenda Maracujá, 
<date id="5B3B102DFFAFFF860BF009FA0BD49644" pageId="59" pageNumber="60" value="1991-11-23" valueMax="1991-11-27" valueMin="1991-11-23">
<date id="5B3B102DFFAFFF860BF009FA0E5995A8" box="[1259,1399,1499,1525]" day="23" last_day="27" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1923-11-27" year="1923">
23– 
<date id="5B3B102DFFAFFF860A0C09FA0E5995A8" box="[1303,1399,1499,1525]" day="27" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1991-11-27" year="1927">
27 
<date id="5B3B102DFFAFFF860A2509FA0E5995A8" box="[1342,1399,1499,1525]" month="11" monthName="Nov" pageId="59" pageNumber="60" year="1991">Nov.</date>
</date>
</date>
1991
</date>
, 
<collectionCode id="4994AE28FFAFFF860E1E09DE0A3B9644" box="[261,277,1535,1561]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860E0209DE0A009644" box="[281,302,1535,1561]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="59" pageNumber="60">T</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860E2A09DE0A6A9644" box="[305,324,1535,1561]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. Amarante and 
<collectionCode id="4994AE28FFAFFF860EED09DE09259644" box="[502,523,1535,1561]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="59" pageNumber="60" type="Herbarium">C</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860D0B09DE090D9644" box="[528,547,1535,1561]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="59" pageNumber="60" type="Herbarium">F</collectionCode>
. Martins leg. (
<collectionCode id="4994AE28FFAFFF860DDC09DE08349644" box="[711,794,1535,1561]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 1 ♂, Floriano, Fazenda Buriti Sol, 
<date id="5B3B102DFFAFFF860BA709DE0E5F9644" box="[1212,1393,1535,1562]" pageId="59" pageNumber="60" value="1991-10-05" valueMax="1991-10-12" valueMin="1991-10-05">
5– 
<date id="5B3B102DFFAFFF860BC109DE0E5F9644" box="[1242,1393,1535,1562]" day="12" month="10" pageId="59" pageNumber="60" partOrder="dmy" value="1991-10-12" year="1991">
<date id="5B3B102DFFAFFF860BC109DE0E019644" box="[1242,1327,1535,1562]" day="12" month="10" monthName="Oct" pageId="59" pageNumber="60" year="2012">
12 
<date id="5B3B102DFFAFFF860BE609DE0E019644" box="[1277,1327,1535,1561]" month="10" monthName="Oct" pageId="59" pageNumber="60" year="1991">Oct.</date>
</date>
1991
</date>
</date>
, 
<collectionCode id="4994AE28FFAFFF860FA60A020BE39660" box="[189,205,1571,1597]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860FCA0A020BC89660" box="[209,230,1571,1597]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="59" pageNumber="60">T</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860FF20A020BD29660" box="[233,252,1571,1597]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. Amarante leg. (
<collectionCode id="4994AE28FFAFFF860EA40A02093C9660" box="[447,530,1571,1597]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 6 ♂♂, 
<specimenCount id="3983FD64FFAFFF860D600A0209EE9660" box="[635,704,1571,1597]" pageId="59" pageNumber="60" type="female">
3 
<specimenCount id="3983FD64FFAFFF860D880A0209EE9660" box="[659,704,1571,1597]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Oeiras (“ 
<quantity id="E87D9B08FFAFFF860C210A0308519661" box="[826,895,1570,1597]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="59" pageNumber="60" unit="km" value="5.0">5 km</quantity>
<collectionCode id="4994AE28FFAFFF860C920A0208B59660" box="[905,923,1571,1597]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="59" pageNumber="60" type="Herbarium">E</collectionCode>
Oeiras”), Fazenda Talhada, 
<date id="5B3B102DFFAFFF860BF60A020BD4963C" pageId="59" pageNumber="60" value="1991-11-13" valueMax="1991-11-17" valueMin="1991-11-13">
<date id="5B3B102DFFAFFF860BF60A020E599660" box="[1261,1399,1571,1597]" day="13" last_day="17" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="2013-11-17" year="2013">
13– 
<date id="5B3B102DFFAFFF860A020A020E599660" box="[1305,1399,1571,1597]" day="17" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1991-11-17" year="2017">
17 
<date id="5B3B102DFFAFFF860A240A020E599660" box="[1343,1399,1571,1597]" month="11" monthName="Nov" pageId="59" pageNumber="60" year="1991">Nov.</date>
</date>
</date>
1991
</date>
, 
<collectionCode id="4994AE28FFAFFF860E130A660A30963C" box="[264,286,1607,1633]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="59" pageNumber="60" type="Herbarium">C</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860E3A0A660A16963C" box="[289,312,1607,1633]" country="Chile" httpUri="http://grbio.org/cool/az7k-p4g9" name="Departamento de Geologia, Universidad de Chile" pageId="59" pageNumber="60">R</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860E270A660A61963C" box="[316,335,1607,1633]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="59" pageNumber="60" type="Herbarium">F</collectionCode>
. Brandão and 
<collectionCode id="4994AE28FFAFFF860EE00A660922963C" box="[507,524,1607,1633]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. Moutinho leg. (
<collectionCode id="4994AE28FFAFFF860DCA0A66080A963C" box="[721,804,1607,1633]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 5 ♂♂, 
<specimenCount id="3983FD64FFAFFF860C950A660895963C" box="[910,955,1607,1633]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860CBE0A660895963C" box="[933,955,1607,1633]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Oeiras (“ 
<quantity id="E87D9B08FFAFFF860B2C0A670F53963D" box="[1079,1149,1606,1633]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="59" pageNumber="60" unit="km" value="5.0">5 km</quantity>
<collectionCode id="4994AE28FFAFFF860B9D0A660FB6963C" box="[1158,1176,1607,1633]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="59" pageNumber="60" type="Herbarium">E</collectionCode>
Oeiras”), Fazenda Talhada, 
<date id="5B3B102DFFAFFF860E3C0A4B0ADD96D9" box="[295,499,1642,1669]" pageId="59" pageNumber="60" value="1991-11-13" valueMax="1991-11-17" valueMin="1991-11-13">
<date id="5B3B102DFFAFFF860E3C0A4B0A9E96D8" box="[295,432,1642,1669]" day="13" last_day="17" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="2013-11-17" year="2013">
13– 
<date id="5B3B102DFFAFFF860E4F0A4B0A9E96D8" box="[340,432,1642,1669]" day="17" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1991-11-17" year="2017">
17 
<date id="5B3B102DFFAFFF860E6C0A4A0A9E96D8" box="[375,432,1643,1669]" month="11" monthName="Nov" pageId="59" pageNumber="60" year="1991">Nov.</date>
</date>
</date>
1991
</date>
, 
<collectionCode id="4994AE28FFAFFF860EE50A4A093D96D8" box="[510,531,1643,1669]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="59" pageNumber="60" type="Herbarium">E</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860D0C0A4A091B96D8" box="[535,565,1643,1669]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. Cancello and 
<collectionCode id="4994AE28FFAFFF860DC40A4A09D296D8" box="[735,764,1643,1669]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="59" pageNumber="60" type="Herbarium">M</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860C1B0A4A083A96D8" box="[768,788,1643,1669]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="59" pageNumber="60">T</collectionCode>
. Ponte leg. (
<collectionCode id="4994AE28FFAFFF860CB90A4B08DB96D9" box="[930,1013,1642,1668]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860B130A4B0F1A96D8" box="[1032,1076,1642,1669]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860B050A4A0F1A96D8" box="[1054,1076,1643,1669]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Oeiras (“ 
<quantity id="E87D9B08FFAFFF860BB00A4B0FC296D9" box="[1195,1260,1642,1669]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="59" pageNumber="60" unit="km" value="5.0">5 km</quantity>
<collectionCode id="4994AE28FFAFFF860BE80A4A0E2B96D8" box="[1267,1285,1643,1669]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="59" pageNumber="60" type="Herbarium">E</collectionCode>
Oeiras”), Fazenda Talhada, 
<date id="5B3B102DFFAFFF860E950AAF097696F5" box="[398,600,1678,1705]" pageId="59" pageNumber="60" value="1991-11-13" valueMax="1991-11-17" valueMin="1991-11-13">
<date id="5B3B102DFFAFFF860E950AAF093B96F4" box="[398,533,1678,1705]" day="13" last_day="17" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="2013-11-17" year="2013">
13– 
<date id="5B3B102DFFAFFF860EA00AAF093B96F4" box="[443,533,1678,1705]" day="17" month="11" monthName="Nov" pageId="59" pageNumber="60" partOrder="dmy" value="1991-11-17" year="2017">
17 
<date id="5B3B102DFFAFFF860EC60AAE093B96F4" box="[477,533,1679,1705]" month="11" monthName="Nov" pageId="59" pageNumber="60" year="1991">Nov.</date>
</date>
</date>
1991
</date>
, 
<collectionCode id="4994AE28FFAFFF860D780AAF095D96F5" box="[611,627,1678,1704]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860D6C0AAF09A296F5" box="[631,652,1678,1704]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="59" pageNumber="60">T</collectionCode>
. 
<collectionCode id="4994AE28FFAFFF860D940AAF098C96F5" box="[655,674,1678,1704]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15763" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="59" pageNumber="60" type="Herbarium">P</collectionCode>
. Amarente leg. (
<collectionCode id="4994AE28FFAFFF860C460AAF089E96F5" box="[861,944,1678,1704]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="59" pageNumber="60" type="Museum">MZSP</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860CD80AAF08C096F4" box="[963,1006,1678,1705]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860CC30AAE08C096F4" box="[984,1006,1679,1705]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Piripiri, Parque Nacional de Sete Cidades, 
<geoCoordinate id="4AB1502AFFAFFF860E370A910A959691" box="[300,443,1712,1740]" degrees="04" direction="south" minutes="05" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="51" value="-4.0975003">
04º05′51″ 
<collectionCode id="4994AE28FFAFFF860EB20A930A959691" box="[425,443,1714,1740]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860ED20A91094C9691" box="[457,610,1712,1740]" degrees="41" direction="west" minutes="42" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="30" value="-41.708336">
41º42′30″ 
<collectionCode id="4994AE28FFAFFF860D5D0A93094C9691" box="[582,610,1714,1740]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860D740A9309939691" box="[623,701,1714,1740]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="59" pageNumber="60" unit="m" value="170.0">170 m</quantity>
, 
<date id="5B3B102DFFAFFF860DD00A9308A79691" box="[715,905,1714,1740]" pageId="59" pageNumber="60" value="2013-02-07" valueMax="2013-02-12" valueMin="2013-02-07">
7– 
<date id="5B3B102DFFAFFF860DF10A9308A79691" box="[746,905,1714,1740]" day="12" month="02" pageId="59" pageNumber="60" partOrder="dmy" value="2013-02-12" year="2013">
<date id="5B3B102DFFAFFF860DF10A93086D9691" box="[746,835,1714,1740]" day="12" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2012">
12 
<date id="5B3B102DFFAFFF860C140A93086D9691" box="[783,835,1714,1740]" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">Feb.</date>
</date>
2013
</date>
</date>
, pitfall with human faeces, Vaz-de-Mello and Grossi leg. (
<collectionCode id="4994AE28FFAFFF860E9B0AF70AF696AD" box="[384,472,1750,1776]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FFAFFF860D310AF7094096AD" box="[554,622,1750,1776]" pageId="59" pageNumber="60" type="female">
4 
<specimenCount id="3983FD64FFAFFF860D5A0AF7094096AD" box="[577,622,1750,1776]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, same collecting data as for preceding but 
<quantity id="E87D9B08FFAFFF860B700AF70F9896AD" box="[1131,1206,1750,1776]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="59" pageNumber="60" unit="m" value="180.0">180 m</quantity>
(
<collectionCode id="4994AE28FFAFFF860BDD0AF70E3096AD" box="[1222,1310,1750,1776]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 9 ♂♂ (1 dissected), 
<specimenCount id="3983FD64FFAFFF860E7F0ADB0A879749" box="[356,425,1786,1812]" pageId="59" pageNumber="60" type="female">
9 
<specimenCount id="3983FD64FFAFFF860E670ADB0A879749" box="[380,425,1786,1812]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Piripiri, Parque Nacional de Sete Cidades, 
<geoCoordinate id="4AB1502AFFAFFF860CA20AD90F669749" box="[953,1096,1784,1812]" degrees="04" direction="south" minutes="05" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="54" value="-4.0983334">
04º05′54″ 
<collectionCode id="4994AE28FFAFFF860B2D0ADB0F669749" box="[1078,1096,1786,1812]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860B4C0AD90FC19749" box="[1111,1263,1784,1812]" degrees="41" direction="west" minutes="42" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="31" value="-41.70861">
41º42′31″ 
<collectionCode id="4994AE28FFAFFF860BC80ADB0FC19749" box="[1235,1263,1786,1812]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860BE60ADB0E659749" box="[1277,1355,1786,1812]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="59" pageNumber="60" unit="m" value="170.0">170 m</quantity>
, 
<date id="5B3B102DFFAFFF860A420ADB0A1A9765" pageId="59" pageNumber="60" value="2013-02-13">
<date id="5B3B102DFFAFFF860A420ADB0BDF976A" day="13" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">
13 
<date id="5B3B102DFFAFFF860FA60B3C0BDF976A" box="[189,241,1821,1847]" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">Feb.</date>
</date>
2013
</date>
, flight interception trap, Vaz-de-Mello and Grossi leg. (
<collectionCode id="4994AE28FFAFFF860CA20B3F0F3F9765" box="[953,1041,1822,1848]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 9 ♂♂, 
<specimenCount id="3983FD64FFAFFF860B6F0B3F0F999765" box="[1140,1207,1822,1848]" pageId="59" pageNumber="60" type="female">
4 
<specimenCount id="3983FD64FFAFFF860B910B3F0F999765" box="[1162,1207,1822,1848]" pageId="59" pageNumber="60">♀♀</specimenCount>
</specimenCount>
, Piripiri, Parque Nacional de Sete Cidades, 
<geoCoordinate id="4AB1502AFFAFFF860EE70B1E09A49706" box="[508,650,1855,1883]" degrees="04" direction="south" minutes="05" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="38" value="-4.093889">
04º05′38″ 
<collectionCode id="4994AE28FFAFFF860D630B6009A49706" box="[632,650,1857,1883]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860D830B1E081E9701" box="[664,816,1855,1884]" degrees="41" direction="west" minutes="42" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="35" value="-41.709724">
41º42′35″ 
<collectionCode id="4994AE28FFAFFF860C0F0B63081E9701" box="[788,816,1858,1884]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860C270B6008A49701" box="[828,906,1857,1884]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="60" unit="m" value="200.0">200 m</quantity>
, 
<date id="5B3B102DFFAFFF860C8C0B600F189706" box="[919,1078,1857,1883]" pageId="59" pageNumber="60" value="2013-02-13">
<date id="5B3B102DFFAFFF860C8C0B6008DF9706" box="[919,1009,1857,1883]" day="13" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">
13 
<date id="5B3B102DFFAFFF860CA60B6008DF9706" box="[957,1009,1857,1883]" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">Feb.</date>
</date>
2013
</date>
, pitfall with human faeces, Vaz-de-Mello and Grossi leg. (
<collectionCode id="4994AE28FFAFFF860D340B4409A99722" box="[559,647,1893,1919]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860D850B4409E297DD" box="[670,716,1893,1920]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860DAD0B4709E297DD" box="[694,716,1894,1920]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Piripiri, Parque Nacional de Sete Cidades, 
<geoCoordinate id="4AB1502AFFAFFF860BFA0B420E5F9722" box="[1249,1393,1891,1919]" degrees="04" direction="south" minutes="06" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="38" value="-4.1105556">
04º06′38″ 
<collectionCode id="4994AE28FFAFFF860A440B440E5F9722" box="[1375,1393,1893,1919]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860FA60BA60A7D97F9" box="[189,339,1927,1956]" degrees="41" direction="west" minutes="44" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="48" value="-41.746666">
41º44′48″ 
<collectionCode id="4994AE28FFAFFF860E2C0BAB0A7D97F9" box="[311,339,1930,1956]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860E450BA80A8497F9" box="[350,426,1929,1956]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="59" pageNumber="60" unit="m" value="180.0">180 m</quantity>
, 
<date id="5B3B102DFFAFFF860EAD0BA8097C97FE" box="[438,594,1929,1955]" pageId="59" pageNumber="60" value="2013-02-12">
<date id="5B3B102DFFAFFF860EAD0BA8092097FE" box="[438,526,1929,1955]" day="12" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2012">
12 
<date id="5B3B102DFFAFFF860EC10BA8092097FE" box="[474,526,1929,1955]" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">Feb.</date>
</date>
2013
</date>
, flight interception trap, Vaz-de-Mello and Grossi leg. (
<collectionCode id="4994AE28FFAFFF860BC30BA80E1E97FE" box="[1240,1328,1929,1955]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFAFFF860A580BA80E4197F9" box="[1347,1391,1929,1956]" pageId="59" pageNumber="60" type="female">
1 
<specimenCount id="3983FD64FFAFFF860A420BAB0E4197F9" box="[1369,1391,1930,1956]" pageId="59" pageNumber="60">♀</specimenCount>
</specimenCount>
, Piripiri, Parque Nacional de Sete Cidades, 
<geoCoordinate id="4AB1502AFFAFFF860DDA0B8A087E979A" box="[705,848,1963,1991]" degrees="04" direction="south" minutes="05" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="03" value="-4.084167">
04º05′03″ 
<collectionCode id="4994AE28FFAFFF860C250B8C087E979A" box="[830,848,1965,1991]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="59" pageNumber="60" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFAFFF860C440B8A08D6979A" box="[863,1016,1963,1991]" degrees="41" direction="west" minutes="42" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="34" value="-41.709446">
41º42′34″ 
<collectionCode id="4994AE28FFAFFF860CC70B8C08D6979A" box="[988,1016,1965,1991]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="59" pageNumber="60" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<quantity id="E87D9B08FFAFFF860B1D0B8C0F7B979A" box="[1030,1109,1965,1991]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="59" pageNumber="60" unit="m" value="190.0">190 m</quantity>
, 
<date id="5B3B102DFFAFFF860B780B8C0E0C979A" box="[1123,1314,1965,1991]" pageId="59" pageNumber="60" value="2013-02-07" valueMax="2013-02-12" valueMin="2013-02-07">
7– 
<date id="5B3B102DFFAFFF860B990B8C0E0C979A" box="[1154,1314,1965,1991]" day="12" month="02" pageId="59" pageNumber="60" partOrder="dmy" value="2013-02-12" year="2013">
<date id="5B3B102DFFAFFF860B990B8C0FF5979A" box="[1154,1243,1965,1991]" day="12" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2012">
12 
<date id="5B3B102DFFAFFF860BB30B8C0FF5979A" box="[1192,1243,1965,1991]" month="02" monthName="Feb" pageId="59" pageNumber="60" year="2013">Feb.</date>
</date>
2013
</date>
</date>
, pitfall with human faeces, Vaz-de-Mello and Grossi leg. (
<collectionCode id="4994AE28FFAFFF860C140BF0084997B6" box="[783,871,2001,2027]" pageId="59" pageNumber="60">CEMT</collectionCode>
); 1 ♂, Ribeiro Gonçalves, Estação Ecológica Uruçuí-Una, 
<geoCoordinate id="4AB1502AFFA9FF800E400D280AED9178" box="[347,451,265,293]" degrees="08" direction="south" minutes="52" orientation="latitude" pageId="61" pageNumber="62" precision="925" value="-8.866667">
08º52′ 
<collectionCode id="4994AE28FFA9FF800EAA0D2A0AED9178" box="[433,451,267,293]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="61" pageNumber="62" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFA9FF800EC80D28096A917B" box="[467,580,265,294]" degrees="44" direction="west" minutes="57" orientation="longitude" pageId="61" pageNumber="62" precision="925" value="-44.95">
44º57′ 
<collectionCode id="4994AE28FFA9FF800D330D2D096A917B" box="[552,580,268,294]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="61" pageNumber="62" type="Herbarium">W</collectionCode>
</geoCoordinate>
, 
<date id="5B3B102DFFA9FF800D480D2A08309178" box="[595,798,267,294]" pageId="61" pageNumber="62" value="2001-01-19" valueMax="2001-01-29" valueMin="2001-01-19">
<date id="5B3B102DFFA9FF800D480D2A09F8917B" box="[595,726,267,294]" day="19" last_day="29" month="01" monthName="Jan" pageId="61" pageNumber="62" partOrder="dmy" value="1919-01-29" year="1919">
19– 
<date id="5B3B102DFFA9FF800D650D2A09F8917B" box="[638,726,267,294]" day="29" month="01" monthName="Jan" pageId="61" pageNumber="62" partOrder="dmy" value="2001-01-29" year="1929">
29 
<date id="5B3B102DFFA9FF800DBC0D2D09F8917B" box="[679,726,268,294]" month="01" monthName="Jan" pageId="61" pageNumber="62" year="2001">Jan.</date>
</date>
</date>
2001
</date>
, 
<collectionCode id="4994AE28FFA9FF800C350D2A08689178" box="[814,838,267,293]" country="Switzerland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15706" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="61" pageNumber="62" type="Herbarium">G</collectionCode>
. 
<collectionCode id="4994AE28FFA9FF800C510D2A084C9178" box="[842,866,267,293]" country="Switzerland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15706" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="61" pageNumber="62" type="Herbarium">G</collectionCode>
. Montingelli leg. (
<collectionCode id="4994AE28FFA9FF800B5D0D2A0FB79178" box="[1094,1177,267,293]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="61" pageNumber="62" type="Museum">MZSP</collectionCode>
); 4 ♂♂, 
<specimenCount id="3983FD64FFA9FF800A120D2A0E16917B" box="[1289,1336,267,294]" pageId="61" pageNumber="62" type="female">
1 
<specimenCount id="3983FD64FFA9FF800A390D2D0E16917B" box="[1314,1336,268,294]" pageId="61" pageNumber="62">♀</specimenCount>
</specimenCount>
, São Raimundo Nonato, Parque Nacional da Serra da Capivara, 
<date id="5B3B102DFFA9FF800C9B0D1108DA9117" box="[896,1012,304,330]" pageId="61" pageNumber="62" value="1999-01">Jan. 1999</date>
, 
<collectionCode id="4994AE28FFA9FF800B190D110F369117" box="[1026,1048,304,330]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="61" pageNumber="62" type="Herbarium">C</collectionCode>
. 
<collectionCode id="4994AE28FFA9FF800B060D110F189117" box="[1053,1078,304,330]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="61" pageNumber="62" type="Herbarium">A</collectionCode>
. Matrangolo leg. (
<collectionCode id="4994AE28FFA9FF800A0A0D110E479117" box="[1297,1385,304,330]" pageId="61" pageNumber="62">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFA9FF800FA60D750A2D9132" box="[189,259,340,367]" pageId="61" pageNumber="62" type="female">
2 
<specimenCount id="3983FD64FFA9FF800FCD0D740A2D9132" box="[214,259,341,367]" pageId="61" pageNumber="62">♀♀</specimenCount>
</specimenCount>
, São Raimundo Nonato, Parque Nacional da Serra da Capivara, Zabelê, 
<date id="5B3B102DFFA9FF800B6E0D750E3B9133" box="[1141,1301,340,367]" pageId="61" pageNumber="62" value="2001-04-12">
<date id="5B3B102DFFA9FF800B6E0D750FE19132" box="[1141,1231,340,367]" day="12" month="04" monthName="Apr" pageId="61" pageNumber="62" year="2012">
12 
<date id="5B3B102DFFA9FF800B800D740FE19132" box="[1179,1231,341,367]" month="04" monthName="Apr" pageId="61" pageNumber="62" year="2001">Apr.</date>
</date>
2001
</date>
, 
<collectionCode id="4994AE28FFA9FF800A380D750E129133" box="[1315,1340,340,366]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="61" pageNumber="62" type="Herbarium">A</collectionCode>
. 
<collectionCode id="4994AE28FFA9FF800A5B0D750E789133" box="[1344,1366,340,366]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="61" pageNumber="62" type="Herbarium">C</collectionCode>
. 
<collectionCode id="4994AE28FFA9FF800A410D750E5D9133" box="[1370,1395,340,366]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="61" pageNumber="62" type="Herbarium">A</collectionCode>
. Moura leg. (
<collectionCode id="4994AE28FFA9FF800E560D580A8B91CE" box="[333,421,377,403]" pageId="61" pageNumber="62">CEMT</collectionCode>
); 2 ♂♂, Teresina, 
<date id="5B3B102DFFA9FF800D600D5809C091C9" box="[635,750,377,404]" pageId="61" pageNumber="62" value="1953-01">Jan. 1953</date>
, Oliveira leg. (
<collectionCode id="4994AE28FFA9FF800C860D5808D491CE" box="[925,1018,377,403]" pageId="61" pageNumber="62">CMNC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="7BFA6665FFA8FF810FA60B8D0AD897B4" ID-DOI="http://doi.org/10.5281/zenodo.3824821" ID-Zenodo-Dep="3824821" httpUri="https://zenodo.org/record/3824821/files/figure.png" pageId="60" pageNumber="61" startId="60.[189,232,1964,1990]" targetBox="[199,1392,494,1919]" targetPageId="60">
<paragraph id="2F3A36EDFFA8FF810FA60B8D0AD897B4" blockId="60.[189,1399,1964,2026]" pageId="60" pageNumber="61">
<emphasis id="1DF1EAFFFFA8FF810FA60B8D0A33979B" bold="true" box="[189,285,1964,1990]" pageId="60" pageNumber="61">Fig. 21.</emphasis>
<taxonomicName id="E8854D6EFFA8FF810E3C0B8D087E979A" authority="(Martinez et al., 1964)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[295,848,1964,1991]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="enkerlini" status="comb. nov.">
<emphasis id="1DF1EAFFFFA8FF810E3C0B8D091C979B" box="[295,562,1964,1990]" italics="true" pageId="60" pageNumber="61">Sylvicanthon enkerlini</emphasis>
(
<bibRefCitation id="4B144B1CFFA8FF810D5E0B8D0866979A" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[581,840,1964,1991]" pageId="60" pageNumber="61" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFA8FF810DA70B8C09D4979B" box="[700,762,1964,1990]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
, 1964
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="06C25784FFA8FF810C400B8D08CF979A" box="[859,993,1964,1991]" pageId="60" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="1DF1EAFFFFA8FF810CF20B8D08D0979B" bold="true" box="[1001,1022,1964,1990]" pageId="60" pageNumber="61">A</emphasis>
. Dorsal view. 
<emphasis id="1DF1EAFFFFA8FF810BA80B8C0FE9979A" bold="true" box="[1203,1223,1965,1991]" pageId="60" pageNumber="61">B</emphasis>
. Ventral view. 
<emphasis id="1DF1EAFFFFA8FF810FA60BEE0BFC97B4" bold="true" box="[189,210,1999,2025]" pageId="60" pageNumber="61">C</emphasis>
. Holotype and its labels.
</paragraph>
</caption>
<subSubSection id="679F6566FFA9FFFD0FA60D9A0E599295" lastPageId="64" lastPageNumber="65" pageId="61" pageNumber="62" type="description">
<paragraph id="2F3A36EDFFA9FF800FA60D9A0A459188" blockId="61.[189,363,443,469]" box="[189,363,443,469]" pageId="61" pageNumber="62">
<heading id="74728181FFA9FF800FA60D9A0A459188" bold="true" box="[189,363,443,469]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="1DF1EAFFFFA9FF800FA60D9A0A459188" bold="true" box="[189,363,443,469]" pageId="61" pageNumber="62">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFA9FF800FA60DCD09169212" blockId="61.[189,1399,491,591]" pageId="61" pageNumber="62">COLOURATION. Dorsum, metaventrite and pygidium entirely black, without metallic reflections; external edge of clypeus occasionally dark brown. Ventral face of legs ranging from black to dark brown; dorsal face of protibiae reddish-brown.</paragraph>
<paragraph id="2F3A36EDFFA9FF800FA60E590FE29330" blockId="61.[189,1399,631,877]" pageId="61" pageNumber="62">
HEAD. Tegument with diffuse shine and strong alveolar microsculpture, which is usually more marked on frons and genae than on clypeus, which can present diffuse microsculpture; micropunctation always evident throughout dorsal surface, more impressed in areas with sparser microsculpture. Clypeus with four large, acute teeth well separated from one another (
<figureCitation id="B7BE2A68FFA9FF800C760EC408E192A2" box="[877,975,741,767]" captionStart="Fig" captionStartId="20.[189,232,1928,1954]" captionTargetBox="[212,1372,265,1894]" captionTargetId="figure@20.[212,1376,265,1895]" captionTargetPageId="20" captionText="Fig. 6. Head, dorsal view. A. Sylvicanthon enkerlini (Martínez et al., 1964) comb. nov. B. S. candezei (Harold, 1869). C. S. genieri sp. nov. D. S. proseni (Martínez, 1949) stat. et comb. nov. E. S. seag sp. nov. F. S. securus (Schmidt, 1920) comb. nov. G. S. furvus (Schmidt, 1920)." figureDoi="http://doi.org/10.5281/zenodo.3824791" httpUri="https://zenodo.org/record/3824791/files/figure.png" pageId="61" pageNumber="62">Fig. 6A</figureCitation>
; in worn specimens, teeth obtuse); external edge (including teeth) clearly folded up; with a single row of setae covering the base of the four teeth. Genae with strong tooth immediately behind clypeal-genal juncture. Posterior edge of head with a fine margin between eyes; occasionally, margin lacking only at the area adjacent to eyes.
</paragraph>
<paragraph id="2F3A36EDFFA9FF800FA60FB7086294A4" blockId="61.[189,1399,917,1273]" pageId="61" pageNumber="62">
THORAX. Pronotum with shiny and lustrous tegument; alveolar microsculpture usually restricted to sides, where it is very dense; centre with strong micropunctation and without microsculpture, or with very diffuse microsculpture. Posterior edge with fine transverse line at centre (usually extending up to the second elytral stria). Hypomeral cavity with tegument densely covered by long yellowish setae and with depressed area close to external margin, the latter with a weak tubercle; posterior part of hypomeron with long, individual setae (around five) aligned longitudinally close to its external edge (
<figureCitation id="B7BE2A68FFA9FF800A1C086D0E47943B" box="[1287,1385,1100,1126]" captionStart="Fig" captionStartId="23.[189,232,1788,1814]" captionTargetBox="[193,1393,858,1750]" captionTargetId="figure@23.[189,1399,858,1763]" captionTargetPageId="23" captionText="Fig. 9. A. Sylvicanthon bridarollii (Martínez, 1949), profemur. Note that, as occurs in the other species of the candezei group, there is no denticulation on the anterior edge, which is, by contrast, completely smooth. B. S. enkerlini (Martínez et al., 1964) comb. nov., profemur.Arrow indicates the denticulation of anterior edge, a condition present in both sexes and unique in the genus. C. S. enkerlini, metepisternum. Arrow indicates the small tubercle at the suture between metepisternum and the lateral region of metaventrite. D. S. enkerlini, hypomeron. Notice the row of long setae present on the lateral of posterior region, a condition seen uniquely in this species in Sylvicanthon." figureDoi="http://doi.org/10.5281/zenodo.3824797" httpUri="https://zenodo.org/record/3824797/files/figure.png" pageId="61" pageNumber="62">Fig. 9D</figureCitation>
). Metepisternum with posterior region, at the suture with metaventrite, with an evident tubercle (
<figureCitation id="B7BE2A68FFA9FF800A1608510E4794D6" box="[1293,1385,1136,1163]" captionStart="Fig" captionStartId="23.[189,232,1788,1814]" captionTargetBox="[193,1393,858,1750]" captionTargetId="figure@23.[189,1399,858,1763]" captionTargetPageId="23" captionText="Fig. 9. A. Sylvicanthon bridarollii (Martínez, 1949), profemur. Note that, as occurs in the other species of the candezei group, there is no denticulation on the anterior edge, which is, by contrast, completely smooth. B. S. enkerlini (Martínez et al., 1964) comb. nov., profemur.Arrow indicates the denticulation of anterior edge, a condition present in both sexes and unique in the genus. C. S. enkerlini, metepisternum. Arrow indicates the small tubercle at the suture between metepisternum and the lateral region of metaventrite. D. S. enkerlini, hypomeron. Notice the row of long setae present on the lateral of posterior region, a condition seen uniquely in this species in Sylvicanthon." figureDoi="http://doi.org/10.5281/zenodo.3824797" httpUri="https://zenodo.org/record/3824797/files/figure.png" pageId="61" pageNumber="62">Fig. 9C</figureCitation>
). Metaventrite entirely glabrous; tegument with strong rivose microsculpture at lateral and anterior regions; at centre, alveolar microsculpture very fine and diffuse, slightly more delimited at posterior region; micropunctation very fine, but distinct at centre.
</paragraph>
<paragraph id="2F3A36EDFFA9FF800FA6090009AB974B" blockId="61.[189,1399,1313,1814]" pageId="61" pageNumber="62">
LEGS. Profemora with tegument with strong alveolar microsculpture at anterior region, and alveolar microsculpture at posterior region; without micropunctation; anterior margin, at apex of profemur, interrupted by row of denticles (
<figureCitation id="B7BE2A68FFA9FF800D2C094B09BA95D9" box="[567,660,1386,1412]" captionStart="Fig" captionStartId="23.[189,232,1788,1814]" captionTargetBox="[193,1393,858,1750]" captionTargetId="figure@23.[189,1399,858,1763]" captionTargetPageId="23" captionText="Fig. 9. A. Sylvicanthon bridarollii (Martínez, 1949), profemur. Note that, as occurs in the other species of the candezei group, there is no denticulation on the anterior edge, which is, by contrast, completely smooth. B. S. enkerlini (Martínez et al., 1964) comb. nov., profemur.Arrow indicates the denticulation of anterior edge, a condition present in both sexes and unique in the genus. C. S. enkerlini, metepisternum. Arrow indicates the small tubercle at the suture between metepisternum and the lateral region of metaventrite. D. S. enkerlini, hypomeron. Notice the row of long setae present on the lateral of posterior region, a condition seen uniquely in this species in Sylvicanthon." figureDoi="http://doi.org/10.5281/zenodo.3824797" httpUri="https://zenodo.org/record/3824797/files/figure.png" pageId="61" pageNumber="62">Fig. 9B</figureCitation>
). Protibiae wide and with internal margin strongly expanded at their apical half (
<figureCitation id="B7BE2A68FFA9FF800E6609AF0AC995F4" box="[381,487,1422,1449]" captionStart="Fig" captionStartId="25.[189,232,1787,1813]" captionTargetBox="[196,1391,645,1731]" captionTargetId="figure@25.[189,1399,645,1755]" captionTargetPageId="25" captionText="Fig. 11. Morphological diversity in the protibiae of Sylvicanthon Halffter &amp; Martínez, 1977. A. S. enkerlini (Martínez et al., 1964) comb. nov. B. S. aequinoctialis (Harold, 1869) comb. nov. C. S. securus (Schmidt, 1920) comb. nov. D. S. obscurus (Schmidt, 1920). E. S. furvus (Schmidt, 1920). F. S. bridarollii (Martínez, 1949) (southern populations). G. S. bridarollii (northern populations). H. S. attenboroughi sp. nov. I. S. edmondsi sp. nov. J. S. candezei (Harold, 1869). Note the variation related to the presence and degree of development of an expansion on the internal edge of protibiae and to the number, size and position of the external teeth." figureDoi="http://doi.org/10.5281/zenodo.3824801" httpUri="https://zenodo.org/record/3824801/files/figure.png" pageId="61" pageNumber="62">Fig. 11A</figureCitation>
); in its apical half, external margin with three large, acute teeth widely separated from one another, the apical two of subequal length and longer than the basal. Mesofemora margined anteriorly only at their basal two-thirds; unmargined portion with a row of very short setae; tegument with strong alveolar microsculpture at anterior region and gradually with more diffuse microsculpture towards posterior and basal regions, where micropunctation is strong. Metafemora margined only anteriorly, posterior margin absent; tegument with strong rivose microsculpture at apical and anterior regions, with microsculpture gradually more diffuse towards posterior region and base; micropunctation present throughout the tegument, more marked in areas of diffuse microsculpture, especially at base, where it can be very dense. Metatarsomeres II and V subequal in length and longer than the others; metatarsomere IV shorter than the others; the entire meso- and metatarsi with a single continuous row of setae throughout its internal margin.
</paragraph>
<paragraph id="2F3A36EDFFA9FF830FA60B1E0E7691C9" blockId="61.[189,1399,1854,2027]" lastBlockId="62.[189,1399,267,404]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">ELYTRA. With nine very narrow visible striae; in general, first six to seven striae well marked, but never carinulate; from them, striae progressively more effaced and interrupted; humeral carina absent. Tegument of interstriae at centre of elytral disc lustrous and without microsculpture (or with very diffuse microsculpture); lateral and apical regions with strong alveolar microsculpture and diffuse shine; micropunctation present throughout tegument, but more distinguishable in areas without microsculpture. ABDOMEN. Ventrite VI with diffuse rivose microsculpture at middle and more clearly marked on the sides; micropunctation absent or very subtle; both sexes without lateral foveae. Pygidium with tegument with diffuse shine and covered by strong alveolar or rivose microsculpture; in some specimens, microsculpture weaker or even absent at apex; micropunctation obliterated by microsculpture and usually indistinct.</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60D9C088791A1" blockId="62.[189,1398,445,508]" pageId="62" pageNumber="63">
AEDEAGUS. Parameres almost as long as phallobase and symmetrical, both faces flat. In lateral view, parameres simple, without any ventral keel or notch (
<figureCitation id="B7BE2A68FFAAFF830C350DC008B291A1" box="[814,924,481,508]" captionStart="Fig" captionStartId="49.[189,232,1895,1921]" captionTargetBox="[189,1386,973,1847]" captionTargetId="figure@49.[189,1399,969,1854]" captionTargetPageId="49" captionText="Fig. 17. Aedeagus morphological diversity in Sylvicanthon. A. S. enkerlini (Martínez et al., 1964) comb. nov. B. S. foveiventris (Schmidt, 1920). C. S. candezei (Harold, 1869) (arrows points to the ventral notch). D. S. genieri sp. nov. (idem). E. S. aequinoctialis (Harold, 1868). F. S. proseni (Martínez, 1949) stat. et comb. nov." figureDoi="http://doi.org/10.5281/zenodo.3824813" httpUri="https://zenodo.org/record/3824813/files/figure.png" pageId="62" pageNumber="63">Fig. 17A</figureCitation>
).
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60E040F3792AA" blockId="62.[189,1399,548,759]" pageId="62" pageNumber="63">
SEXUAL DIMORPHISM. 
<emphasis id="1DF1EAFFFFAAFF830EAF0E040AD19262" bold="true" box="[436,511,549,575]" pageId="62" pageNumber="63">Males</emphasis>
: Protibial spur narrow and bifid at apex, with spiniform projections, the external project longer than the internal one (
<figureCitation id="B7BE2A68FFAAFF830DD90E6808009239" box="[706,814,585,612]" captionStart="Fig" captionStartId="30.[189,232,1790,1816]" captionTargetBox="[196,1383,292,1753]" captionTargetId="figure@30.[196,1383,289,1754]" captionTargetPageId="30" captionText="Fig. 15. Morphological variation and sexual dimorphism in the protibial spur of Sylvicanthon. A. S. enkerlini (Martínez et al., 1964) comb. nov., ♂. B. S. foveiventris (Schmidt, 1920), ♂. C. S. candezei (Harold, 1869), ♂. D. S. genieri sp. nov., ♂. E. S. aequinoctialis (Harold, 1868), ♂. F. S. aequinoctialis comb. nov., ♀. G. S. proseni (Martínez, 1949) stat. et comb. nov., ♂. H. S. proseni stat. et comb. nov., ♀. I. S. bridarollii (Martínez, 1949), ♂. J. S. seag sp. nov., ♂. K. S. edmondsi sp. nov.,♂. L. S. attenboroughi sp. nov., ♂. M. S. obscurus (Schmidt, 1920), ♂. N. S. furvus (Schmidt, 1920), ♂. O. S. securus (Schmidt, 1920) comb. nov., ♂." figureDoi="http://doi.org/10.5281/zenodo.3824809" httpUri="https://zenodo.org/record/3824809/files/figure.png" pageId="62" pageNumber="63">Fig. 15A</figureCitation>
). Ventrite VI strongly narrowed at middle; anterior margin covered only slightly by weak medial expansion of posterior edge of ventrite V. Pygidium very long (length between 1.2 and 
<quantity id="E87D9B08FFAAFF830D390EB209AD92F0" box="[546,643,659,685]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="62" pageNumber="63" unit="mm" value="0.9">0.9 mm</quantity>
). 
<emphasis id="1DF1EAFFFFAAFF830D830EB209D192F0" bold="true" box="[664,767,659,685]" pageId="62" pageNumber="63">Females</emphasis>
: Protibial spur spiniform, simple, and distinctly bent towards external side. Ventrite VI very wide at middle; anterior margin covered by medial flange of posterior edge of ventrite V. Pygidium shorter (between 1.0 and 
<quantity id="E87D9B08FFAAFF830CB50EFC0F2392AA" box="[942,1037,733,759]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="62" pageNumber="63" unit="mm" value="0.7">0.7 mm</quantity>
).
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60F010A5B9367" blockId="62.[189,373,800,826]" box="[189,373,800,826]" pageId="62" pageNumber="63">
<heading id="74728181FFAAFF830FA60F010A5B9367" bold="true" box="[189,373,800,826]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="1DF1EAFFFFAAFF830FA60F010A5B9367" bold="true" box="[189,373,800,826]" pageId="62" pageNumber="63">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60F71083A93EE" blockId="62.[189,1399,847,948]" pageId="62" pageNumber="63">
Males (N = 14). 
<emphasis id="1DF1EAFFFFAAFF830E670F710A839337" bold="true" box="[380,429,848,874]" pageId="62" pageNumber="63">TL:</emphasis>
AV: 6.6 ± 0.64; 
<collectingCountry id="5792767DFFAAFF830D710F7109B09337" box="[618,670,848,874]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 8; MN: 5.8. 
<emphasis id="1DF1EAFFFFAAFF830C2E0F7108409337" bold="true" box="[821,878,848,874]" pageId="62" pageNumber="63">EW:</emphasis>
AV: 4.6 ± 0.38; 
<collectingCountry id="5792767DFFAAFF830B300F710F719337" box="[1067,1119,848,874]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 5,5; MN: 4.1. 
<emphasis id="1DF1EAFFFFAAFF830A160F710E129337" bold="true" box="[1293,1340,848,874]" pageId="62" pageNumber="63">PL:</emphasis>
ME: 2.1 ± 0.2; 
<collectingCountry id="5792767DFFAAFF830E230F540A4293D2" box="[312,364,885,911]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 2.5; MN: 1.8. 
<emphasis id="1DF1EAFFFFAAFF830D3B0F54097893D2" bold="true" box="[544,598,885,911]" pageId="62" pageNumber="63">PW:</emphasis>
AV: 4.0 ± 0.35; 
<collectingCountry id="5792767DFFAAFF830C060F54087F93D2" box="[797,849,885,911]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 4.8; MN: 3.5. 
<emphasis id="1DF1EAFFFFAAFF830B1D0F540F6D93D2" bold="true" box="[1030,1091,885,911]" pageId="62" pageNumber="63">PgL:</emphasis>
AV: 1.0 ± 0.09; 
<collectingCountry id="5792767DFFAAFF830A110F540E1093D2" box="[1290,1342,885,911]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 1.2; MN: 0.9. 
<emphasis id="1DF1EAFFFFAAFF830E340FB80A5B93EE" bold="true" box="[303,373,921,947]" pageId="62" pageNumber="63">PgW:</emphasis>
AV: 1.7 ± 0.23; 
<collectingCountry id="5792767DFFAAFF830D2C0FBB094593E9" box="[567,619,922,948]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 2.2; MN: 1.3.
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60FFD0840941C" blockId="62.[189,1399,988,1089]" pageId="62" pageNumber="63">
Females (N = 15). 
<emphasis id="1DF1EAFFFFAAFF830EB90FFD0AFD93AB" bold="true" box="[418,467,988,1014]" pageId="62" pageNumber="63">TL:</emphasis>
AV: 6.4 ± 0.58; 
<collectingCountry id="5792767DFFAAFF830D840FFC09FD93AA" box="[671,723,989,1015]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 7.3; MN: 5.7. 
<emphasis id="1DF1EAFFFFAAFF830C960FFD08E893AB" bold="true" box="[909,966,988,1014]" pageId="62" pageNumber="63">EW:</emphasis>
AV: 4.5 ± 0.39; 
<collectingCountry id="5792767DFFAAFF830B890FFC0FE893AA" box="[1170,1222,989,1015]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 5.2; MN: 3.9. 
<emphasis id="1DF1EAFFFFAAFF830FA608200BC39446" bold="true" box="[189,237,1025,1051]" pageId="62" pageNumber="63">PL:</emphasis>
AV: 2.0 ± 0.18; 
<collectingCountry id="5792767DFFAAFF830EAA08230ACB9441" box="[433,485,1026,1052]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 2.3; MN: 1.7. 
<emphasis id="1DF1EAFFFFAAFF830D82082009FE9446" bold="true" box="[665,720,1025,1051]" pageId="62" pageNumber="63">PW:</emphasis>
AV: 3.8 ± 0.31; 
<collectingCountry id="5792767DFFAAFF830C8E082308E79441" box="[917,969,1026,1052]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 4.4; MN: 3.3. 
<emphasis id="1DF1EAFFFFAAFF830B6608200F959446" bold="true" box="[1149,1211,1025,1051]" pageId="62" pageNumber="63">PgL:</emphasis>
AV: 0.9 ± 0.08; 
<collectingCountry id="5792767DFFAAFF830FA608070BDF941D" box="[189,241,1062,1088]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 1; MN: 0.7. 
<emphasis id="1DF1EAFFFFAAFF830E9708070AFC941D" bold="true" box="[396,466,1062,1088]" pageId="62" pageNumber="63">PgW:</emphasis>
AV: 1.7 ± 0.11; 
<collectingCountry id="5792767DFFAAFF830D8A080709EB941D" box="[657,709,1062,1088]" name="Mexico" pageId="62" pageNumber="63">MX</collectingCountry>
: 1.9; MN: 1.5.
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60848092C94DE" blockId="62.[189,514,1129,1155]" box="[189,514,1129,1155]" pageId="62" pageNumber="63">
<heading id="74728181FFAAFF830FA60848092C94DE" bold="true" box="[189,514,1129,1155]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="1DF1EAFFFFAAFF830FA60848092C94DE" bold="true" box="[189,514,1129,1155]" pageId="62" pageNumber="63">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA608B80A1C9485" blockId="62.[189,1398,1177,1240]" pageId="62" pageNumber="63">Dry forests between Cerrado, Caatinga, Amazonia and the Atlantic Forest in the Brazilian north- and southeast.</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA609200A699546" blockId="62.[189,327,1281,1307]" box="[189,327,1281,1307]" pageId="62" pageNumber="63">
<heading id="74728181FFAAFF830FA609200A699546" bold="true" box="[189,327,1281,1307]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="1DF1EAFFFFAAFF830FA609200A699546" bold="true" box="[189,327,1281,1307]" pageId="62" pageNumber="63">Ecoregions</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA6091008C49516" blockId="62.[189,1002,1329,1355]" box="[189,1002,1329,1355]" pageId="62" pageNumber="63">
<collectingRegion id="ED41F80FFFAAFF830FA609100A1A9516" box="[189,308,1329,1355]" country="Brazil" name="Maranhao" pageId="62" pageNumber="63">Maranhão</collectingRegion>
Babaçu Forests, Cerrado, Caatinga, 
<collectingRegion id="ED41F80FFFAAFF830DF9091008089516" box="[738,806,1329,1355]" country="Brazil" name="Bahia" pageId="62" pageNumber="63">Bahia</collectingRegion>
Interior Forests.
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA609550AC995D3" blockId="62.[189,487,1396,1422]" box="[189,487,1396,1422]" pageId="62" pageNumber="63">
<emphasis id="1DF1EAFFFFAAFF830FA609550A5795D3" bold="true" box="[189,377,1396,1422]" pageId="62" pageNumber="63">Collecting sites</emphasis>
(
<figureCitation id="B7BE2A68FFAAFF830E9209550AF195D3" box="[393,479,1396,1422]" captionStart="Fig" captionStartId="63.[189,232,1929,1955]" captionTargetBox="[191,1398,709,1889]" captionTargetId="figure@63.[191,1398,709,1889]" captionTargetPageId="63" captionText="Fig. 22. Distribution of Sylvicanthon enkerlini (Martínez et al., 1964) comb. nov. Note that the species occupies transition zones between the Amazonia, Cerrado, Caatinga and Atlantic Forest biomes, regions much dryer and more open than the humid forests occupied by the species of the candezei group." figureDoi="http://doi.org/10.5281/zenodo.3824823" httpUri="https://zenodo.org/record/3824823/files/figure.png" pageId="62" pageNumber="63">Fig. 22</figureCitation>
)
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA609850E599671" blockId="62.[189,1399,1443,1581]" pageId="62" pageNumber="63">
<collectingCountry id="5792767DFFAAFF830FA609850A0995E3" box="[189,295,1444,1470]" name="Brazil" pageId="62" pageNumber="63">BRAZIL</collectingCountry>
. 
<emphasis id="1DF1EAFFFFAAFF830E2609850AEF95E3" bold="true" box="[317,449,1444,1470]" pageId="62" pageNumber="63">
<collectingRegion id="ED41F80FFFAAFF830E2609850AEF95E3" box="[317,449,1444,1470]" country="Brazil" name="Maranhao" pageId="62" pageNumber="63">Maranhão</collectingRegion>
</emphasis>
: São Luís, Mirador (Parque Estadual do Mirador). 
<emphasis id="1DF1EAFFFFAAFF830B4809820FBA95E3" bold="true" box="[1107,1172,1443,1470]" pageId="62" pageNumber="63">
<collectingRegion id="ED41F80FFFAAFF830B4809820FBA95E3" box="[1107,1172,1443,1470]" country="Brazil" name="Piaui" pageId="62" pageNumber="63">Piauí</collectingRegion>
</emphasis>
: Canto do Buriti, Corrente, Floriano, Oeiras, Parnaíba, Piripiri (Parque Nacional de Sete Cidades), Ribeiro Gonçalves (Estação Ecológica de Uruçuí-Una), São Raimundo Nonato (Parque Nacional da Serra da Capivara), Teresina. 
<emphasis id="1DF1EAFFFFAAFF830E350A330A559670" bold="true" box="[302,379,1554,1581]" pageId="62" pageNumber="63">
<collectingRegion id="ED41F80FFFAAFF830E350A330A559670" box="[302,379,1554,1581]" country="Brazil" name="Ceara" pageId="62" pageNumber="63">Ceará</collectingRegion>
</emphasis>
: Caucaia. 
<emphasis id="1DF1EAFFFFAAFF830EEE0A3309109671" bold="true" box="[501,574,1554,1580]" pageId="62" pageNumber="63">
<collectingRegion id="ED41F80FFFAAFF830EEE0A3309109671" box="[501,574,1554,1580]" country="Brazil" name="Bahia" pageId="62" pageNumber="63">Bahia</collectingRegion>
</emphasis>
: Barreiras, Pilão Arcado. 
<emphasis id="1DF1EAFFFFAAFF830C770A330F3A9671" bold="true" box="[876,1044,1554,1580]" pageId="62" pageNumber="63">
<collectingRegion id="ED41F80FFFAAFF830C770A330F3A9671" box="[876,1044,1554,1580]" country="Brazil" name="Minas Gerais" pageId="62" pageNumber="63">Minas Gerais</collectingRegion>
</emphasis>
: Bandeira, Jaíba, Três Marias.
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60A7408379632" blockId="62.[189,793,1621,1648]" box="[189,793,1621,1648]" pageId="62" pageNumber="63">
<heading id="74728181FFAAFF830FA60A7408379632" bold="true" box="[189,793,1621,1648]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="1DF1EAFFFFAAFF830FA60A7408379632" bold="true" box="[189,793,1621,1648]" pageId="62" pageNumber="63">Intraspecific variation and taxonomic discussion</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFAAFF830FA60AA409C197B6" blockId="62.[189,1399,1669,2027]" pageId="62" pageNumber="63">
If, on the one side, 
<taxonomicName id="E8854D6EFFAAFF830E820AA7093396FD" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[409,541,1670,1696]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="enkerlini" status="comb. nov.">
<emphasis id="1DF1EAFFFFAAFF830E820AA7093396FD" box="[409,541,1670,1696]" italics="true" pageId="62" pageNumber="63">S. enkerlini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06C25784FFAAFF830D3F0AA4098A96FD" box="[548,676,1669,1696]" pageId="62" pageNumber="63" rank="species">comb. nov.</taxonomicNameLabel>
seems to be the most isolated species in the genus and with no close relationships to any known 
<taxonomicName id="E8854D6EFFAAFF830D5B0A8B09E19699" authorityName="sensu Tarasov &amp; Dimitrov" authorityYear="2016" box="[576,719,1706,1732]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="tribe" tribe="Deltochilini">Deltochilini</taxonomicName>
, on the other side, the studied populations form a cohesive entity without any noticeable geographical variation. In the same way, little intrapopulational variation was observed. The exceptions are the specimens (two males and one female) collected in 
<date id="5B3B102DFFAAFF830BEC0AD50E1C9753" box="[1271,1330,1780,1806]" pageId="62" pageNumber="63" value="2003">2003</date>
by P. Schmidt in Barreiras (
<collectingRegion id="ED41F80FFFAAFF830EDD0B38093F9769" box="[454,529,1817,1844]" country="Brazil" name="Bahia" pageId="62" pageNumber="63">Bahia</collectingRegion>
): the alveolar microsculpture of the entire tegument is much stronger on them than on the other individuals, being evident even at the centre of the pronotum (where it is diffuse in the other specimens). The two females collected at the same place in 
<date id="5B3B102DFFAAFF830B0E0B430F7C9721" box="[1045,1106,1890,1916]" pageId="62" pageNumber="63" value="1991">1991</date>
, on the other hand, have microsculpture typical of the rest of the distribution of 
<taxonomicName id="E8854D6EFFAAFF830C260BA908EC97FC" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[829,962,1927,1954]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFAAFF830C260BA908EC97FC" box="[829,962,1927,1954]" italics="true" pageId="62" pageNumber="63">S. enkerlini</emphasis>
</taxonomicName>
, as well as the four females collected between 
<date id="5B3B102DFFAAFF830E3D0B8D0AFA979B" box="[294,468,1964,1991]" pageId="62" pageNumber="63" value="2005" valueMax="2006">
2005 and 
<quantity id="E87D9B08FFAAFF830E820B8D0AFA979B" box="[409,468,1964,1990]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="62" pageNumber="63" unit="in" value="2006.0">2006</quantity>
</date>
in the municipality of Pilão Arcado, distant only about 
<quantity id="E87D9B08FFAAFF830B460B8D0F92979B" box="[1117,1212,1964,1990]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="62" pageNumber="63" unit="km" value="300.0">300 km</quantity>
from Barreiras. The reason for that difference is unknown to us.
</paragraph>
<paragraph id="2F3A36EDFFABFF820FA60D2A0A509178" blockId="63.[189,382,267,293]" box="[189,382,267,293]" pageId="63" pageNumber="64">
<heading id="74728181FFABFF820FA60D2A0A509178" bold="true" box="[189,382,267,293]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="1DF1EAFFFFABFF820FA60D2A0A509178" bold="true" box="[189,382,267,293]" pageId="63" pageNumber="64">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFABFFFD0FA60D1809F19240" blockId="63.[189,1399,313,622]" lastBlockId="64.[189,1399,267,541]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<bibRefCitation id="4B144B1CFFABFF820FA60D180AEB9109" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[189,453,313,340]" pageId="63" pageNumber="64" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFABFF820E2A0D1B0A40910E" box="[305,366,313,339]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
(
<date id="5B3B102DFFABFF820E9B0D180A939109" box="[384,445,313,340]" pageId="63" pageNumber="64" value="1964">1964</date>
)
</bibRefCitation>
described 
<taxonomicName id="E8854D6EFFABFF820D510D1B09FC910E" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[586,722,313,340]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFABFF820D510D1B09FC910E" box="[586,722,313,340]" italics="true" pageId="63" pageNumber="64">S. enkerlini</emphasis>
</taxonomicName>
based solely on the male 
<typeStatus id="F03E884FFFABFF820B0B0D180F59910E" box="[1040,1143,313,339]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
collected in a humid forest surrounding a water reservoir in the outskirts of the city of São Luís, Maranhão, 
<collectingCountry id="5792767DFFABFF820BEC0D7D0E11912B" box="[1271,1343,348,374]" name="Brazil" pageId="63" pageNumber="64">Brazil</collectingCountry>
(“ 
<emphasis id="1DF1EAFFFFABFF820A7A0D7C0AC491C7" italics="true" pageId="63" pageNumber="64">el ejemplar holotipo y único</emphasis>
[...] 
<emphasis id="1DF1EAFFFFABFF820D390DA10F1391E0" italics="true" pageId="63" pageNumber="64">fue capturado en las afueras de la ciudad de Sao Luiz, dentro del bosque tropical y húmedo que rodea la reserva de agua potable de esa población</emphasis>
”). As they were aware of only this single geographical record, those authors considered 
<taxonomicName id="E8854D6EFFABFF820C8F0DE60F3791BD" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[916,1049,454,481]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFABFF820C8F0DE60F3791BD" box="[916,1049,454,481]" italics="true" pageId="63" pageNumber="64">S. enkerlini</emphasis>
</taxonomicName>
as being a typical Amazonian species. Nonetheless, over the past 25 years, several collections in transitional regions between the Cerrado, Caatinga and Amazonia, in the Brazilian states of 
<collectingRegion id="ED41F80FFFABFF820C6C0E2D0898927B" box="[887,950,524,550]" country="Brazil" name="Piaui" pageId="63" pageNumber="64">Piauí</collectingRegion>
, 
<collectingRegion id="ED41F80FFFABFF820CD90E2D0F27927B" box="[962,1033,524,550]" country="Brazil" name="Bahia" pageId="63" pageNumber="64">Bahia</collectingRegion>
, 
<collectingRegion id="ED41F80FFFABFF820B0E0E2C0F77927A" box="[1045,1113,525,551]" country="Brazil" name="Ceara" pageId="63" pageNumber="64">Ceará</collectingRegion>
and 
<collectingRegion id="ED41F80FFFABFF820B880E2D0E1D927B" box="[1171,1331,524,550]" country="Brazil" name="Minas Gerais" pageId="63" pageNumber="64">Minas Gerais</collectingRegion>
, have revealed the presence of 
<taxonomicName id="E8854D6EFFABFF820EF80E1109479217" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[483,617,560,586]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFABFF820EF80E1109479217" box="[483,617,560,586]" italics="true" pageId="63" pageNumber="64">S. enkerlini</emphasis>
</taxonomicName>
in that vast area, whereas other collections in regions farther west in the Amazon forest, such as in the state of Pará, have not yielded the species. Therefore, it seems that 
<taxonomicName id="E8854D6EFFD4FFFD0FEF0D2D0A539178" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[244,381,267,294]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFD4FFFD0FEF0D2D0A539178" box="[244,381,267,294]" italics="true" pageId="64" pageNumber="65">S. enkerlini</emphasis>
</taxonomicName>
is actually an inhabitant of open and dry lowland forests typical of the transitional zone between those three biomes, including the Mata dos Cocais, in the transitional zone between the Amazon rainforest and the Caatinga, where the babaçu palm (
<taxonomicName id="E8854D6EFFD4FFFD0CB50D730F919130" authority="Mart." authorityName="Mart." box="[942,1215,338,365]" class="Liliopsida" family="Arecaceae" genus="Attalea" kingdom="Plantae" order="Arecales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="1DF1EAFFFFD4FFFD0CB50D730F5C9131" box="[942,1138,338,364]" italics="true" pageId="64" pageNumber="65">Attalea speciosa</emphasis>
Mart.
</taxonomicName>
) predominates. One specimen was collected in 
<quantity id="E87D9B08FFD4FFFD0D2F0D5409A091D2" box="[564,654,373,399]" metricMagnitude="1" metricUnit="m" metricValue="4.907279999999999" pageId="64" pageNumber="65" unit="in" value="1932.0">
<date id="5B3B102DFFD4FFFD0D2F0D54094191D2" box="[564,623,373,399]" pageId="64" pageNumber="65" value="1932">1932</date>
in
</quantity>
the municipality of Bandeira, 
<collectingRegion id="ED41F80FFFD4FFFD0CE20D540FB491D2" box="[1017,1178,373,399]" country="Brazil" name="Minas Gerais" pageId="64" pageNumber="65">Minas Gerais</collectingRegion>
, an area originally covered by the Atlantic rainforest (ecoregion of 
<collectingRegion id="ED41F80FFFD4FFFD0C1F0DB8086691EE" box="[772,840,409,435]" country="Brazil" name="Bahia" pageId="64" pageNumber="65">Bahia</collectingRegion>
Interior Forests). However, due to the intense anthropogenic activity across this entire ecoregion converting the former humid forest into open areas for agriculture and pasture, it was given the right conditions for the immigration of a fauna typical of Cerrado into that region, 
<taxonomicName id="E8854D6EFFD4FFFD0EFE0E2209449240" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[485,618,515,541]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFD4FFFD0EFE0E2209449240" box="[485,618,515,541]" italics="true" pageId="64" pageNumber="65">S. enkerlini</emphasis>
</taxonomicName>
included.
</paragraph>
<caption id="7BFA6665FFABFF820FA60BA80E0197B6" ID-DOI="http://doi.org/10.5281/zenodo.3824823" ID-Zenodo-Dep="3824823" httpUri="https://zenodo.org/record/3824823/files/figure.png" pageId="63" pageNumber="64" startId="63.[189,232,1929,1955]" targetBox="[191,1398,709,1889]" targetPageId="63">
<paragraph id="2F3A36EDFFABFF820FA60BA80E0197B6" blockId="63.[189,1398,1929,2027]" pageId="63" pageNumber="64">
<emphasis id="1DF1EAFFFFABFF820FA60BA80A3797F9" bold="true" box="[189,281,1929,1956]" pageId="63" pageNumber="64">Fig. 22.</emphasis>
Distribution of 
<taxonomicName id="E8854D6EFFABFF820ECC0BAB08DE97F8" authority="(Martinez et al., 1964)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[471,1008,1930,1957]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="enkerlini" status="comb. nov.">
<emphasis id="1DF1EAFFFFABFF820ECC0BAB09F097F9" box="[471,734,1930,1956]" italics="true" pageId="63" pageNumber="64">Sylvicanthon enkerlini</emphasis>
(
<bibRefCitation id="4B144B1CFFABFF820DF50BAB08C697F8" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[750,1000,1930,1957]" pageId="63" pageNumber="64" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFABFF820C7A0BAA08B397F9" box="[865,925,1930,1956]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
, 1964
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="06C25784FFABFF820CE30BAB0F5497F9" box="[1016,1146,1930,1956]" pageId="63" pageNumber="64" rank="species">comb. nov.</taxonomicNameLabel>
Note that the species occupies transition zones between the Amazonia, Cerrado, Caatinga and Atlantic Forest biomes, regions much dryer and more open than the humid forests occupied by the species of the 
<taxonomicName id="E8854D6EFFABFF820B6E0BF10FF297B7" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1141,1244,2000,2026]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="1DF1EAFFFFABFF820B6E0BF10FF297B7" box="[1141,1244,2000,2026]" italics="true" pageId="63" pageNumber="64">candezei</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="2F3A36EDFFD4FFFD0FA60E650E599295" blockId="64.[189,1399,580,713]" pageId="64" pageNumber="65">
Based on the specimen labels, it is possible to know that 
<taxonomicName id="E8854D6EFFD4FFFD0C560E6408FC9203" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Cupello &amp; Vaz-De" baseAuthorityYear="1964" box="[845,978,580,607]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="enkerlini">
<emphasis id="1DF1EAFFFFD4FFFD0C560E6408FC9203" box="[845,978,580,607]" italics="true" pageId="64" pageNumber="65">S. enkerlini</emphasis>
</taxonomicName>
occurs in altitudes between 170 and 
<quantity id="E87D9B08FFD4FFFD0FA60E460A2792DF" box="[189,265,615,642]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="64" pageNumber="65" unit="m" value="416.0">416 m</quantity>
and is collected between October and June (no records for May) using pitfall traps baited with human faeces as well as flight interception traps and light traps. The only record of flight activity time is that of 
<bibRefCitation id="4B144B1CFFD4FFFD0E140E8F093D9294" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[271,531,686,713]" pageId="64" pageNumber="65" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFD4FFFD0E9B0E8E0A959295" box="[384,443,686,712]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
(
<date id="5B3B102DFFD4FFFD0ED00E8F09249294" box="[459,522,686,713]" pageId="64" pageNumber="65" value="1964">1964</date>
)
</bibRefCitation>
, who said the 
<typeStatus id="F03E884FFFD4FFFD0DA20E8F080E9295" box="[697,800,686,712]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
was collected during the first hours of the evening.
</paragraph>
</subSubSection>
</treatment>
</document>