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<mods:title id="7DD0B9A167842CAD6DDF821C33AB425B">A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter &amp; Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of ‘ Canthon sensu lato’</mods:title>
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<mods:namePart id="EFF76B67627F0D68EE1470A8B7055A22">Cupello, Mario</mods:namePart>
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<mods:affiliation id="6EC61C8CEF83387B648E42C6A2F5452C">Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, UFRJ, Quinta da Boa Vista, São Cristóvão, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:namePart id="4D2AC0FADFA95CC3C42A848FF5B61E1A">Vaz-De, Fernando Z.</mods:namePart>
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<emphasis id="1DF1EAFFFF3CFF150E6A0DAB090191F8" bold="true" box="[369,559,394,421]" italics="true" pageId="168" pageNumber="169">Canthon cobosi</emphasis>
(
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)
</taxonomicName>
revalidation and comb. nov.
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<figureCitation id="B7BE2A68FF3CFF150DFD0D8E08609194" box="[742,846,431,457]" captionStart="Fig" captionStartId="169.[189,232,1961,1987]" captionTargetBox="[350,1246,265,1923]" captionTargetId="figure@169.[313,1275,265,1935]" captionTargetPageId="169" captionText="Fig. 46. A. Canthon cobosi (Pereira &amp; Martínez, 1960) stat. et comb. nov., holotype. B–C. Canthon machadoi (Martínez &amp; Pereira, 1967) comb. nov. B. Paratype 2. C. Paratype 4." figureDoi="http://doi.org/10.5281/zenodo.3824878" httpUri="https://zenodo.org/record/3824878/files/figure.png" pageId="168" pageNumber="169">Fig. 46A</figureCitation>
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<emphasis id="1DF1EAFFFF3CFF150FA60DD20AA59250" box="[189,395,499,525]" italics="true" pageId="168" pageNumber="169">Glaphyrocanthon</emphasis>
(
<emphasis id="1DF1EAFFFF3CFF150E870DD209449250" box="[412,618,499,525]" italics="true" pageId="168" pageNumber="169">Glaphyrocanthon</emphasis>
) 
<emphasis id="1DF1EAFFFF3CFF150D600DD209E69250" box="[635,712,499,525]" italics="true" pageId="168" pageNumber="169">cobosi</emphasis>
<bibRefCitation id="4B144B1CFF3CFF150DD40DD20F7D9253" author="Pereira F. S. &amp; Martinez A." box="[719,1107,499,526]" pageId="168" pageNumber="169" pagination="37 - 55" refId="ref144264" refString="Pereira F. S. &amp; Martinez A. 1960. Notas escarabeidologicas - II. Revista Brasileira de Entomologia 9: 37 - 55." type="journal article" year="1960">Pereira &amp; Martínez, 1960: 45–47</bibRefCitation>
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<taxonomicName id="E8854D6EFF3CFF150FA60E1709E6920D" ID-CoL="3G5RX" authorityName="Pereira &amp; Martinez" authorityYear="1960" box="[189,712,566,593]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="cobosi" subGenus="Glaphyrocanthon">
<emphasis id="1DF1EAFFFF3CFF150FA60E170AA5920D" box="[189,395,566,592]" italics="true" pageId="168" pageNumber="169">Glaphyrocanthon</emphasis>
(
<emphasis id="1DF1EAFFFF3CFF150E870E170944920D" box="[412,618,566,592]" italics="true" pageId="168" pageNumber="169">Glaphyrocanthon</emphasis>
) 
<emphasis id="1DF1EAFFFF3CFF150D600E1709E6920D" box="[635,712,566,592]" italics="true" pageId="168" pageNumber="169">cobosi</emphasis>
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– 
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<bibRefCitation id="4B144B1CFF3CFF150DFD0E1708DA920C" author="Martinez A. &amp; Halffter G. &amp; Halffter V." box="[742,1012,566,593]" pageId="168" pageNumber="169" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFF3CFF150C4C0E1608BD920D" box="[855,915,566,592]" italics="true" pageId="168" pageNumber="169">et al.</emphasis>
1964: 5
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, 8, 11, 14. — 
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<bibRefCitation id="4B144B1CFF3CFF150B870E170A459228" author="Vulcano M. A. &amp; Pereira F. S." pageId="168" pageNumber="169" pagination="570 - 685" refId="ref147746" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey 15 (2): 570 - 685. Available from https: // biodiversitylibrary. org / page / 45986646 [accessed 30 May 2018]." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 662</bibRefCitation>
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; 
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<bibRefCitation id="4B144B1CFF3CFF150E660E7A0AD7922B" author="Vulcano M. A. &amp; Pereira F. S." box="[381,505,603,630]" pageId="168" pageNumber="169" pagination="533 - 603" refId="ref147845" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5 (Zoologia): 533 - 603." type="journal article" year="1967">1967: 561</bibRefCitation>
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. — 
<treatmentCitation id="AE2410FCFF3CFF150D210E7A08839228" author="Halffter G. &amp; Martinez A." box="[570,941,603,629]" page="63" pageId="168" pageNumber="169" year="1977">
<bibRefCitation id="4B144B1CFF3CFF150D210E7A08839228" author="Halffter G. &amp; Martinez A." box="[570,941,603,629]" pageId="168" pageNumber="169" pagination="29 - 107" refId="ref136250" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 63</bibRefCitation>
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(as junior synonym of 
<taxonomicName id="E8854D6EFF3CFF150BC40E7A0A4D92C7" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="xanthopus">
<emphasis id="1DF1EAFFFF3CFF150BC40E7A0A4D92C7" italics="true" pageId="168" pageNumber="169">Sylvicanthon xanthopus</emphasis>
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).
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<emphasis id="1DF1EAFFFF3CFF150FA60EE50A6A9283" bold="true" box="[189,324,708,734]" pageId="168" pageNumber="169">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFF3CFF150FA60ED50A549369" blockId="168.[189,1399,756,820]" pageId="168" pageNumber="169">
The specific name refers to the Spanish entomologist Antonio Cobos Sánchez (
<date id="5B3B102DFF3CFF150B780ED50FC49353" box="[1123,1258,756,782]" pageId="168" pageNumber="169" value="1922" valueMax="1998">1922–1998</date>
) (
<bibRefCitation id="4B144B1CFF3CFF150A190ED50A439369" author="Pereira F. S. &amp; Martinez A." pageId="168" pageNumber="169" pagination="37 - 55" refId="ref144264" refString="Pereira F. S. &amp; Martinez A. 1960. Notas escarabeidologicas - II. Revista Brasileira de Entomologia 9: 37 - 55." type="journal article" year="1960">Pereira &amp; Martínez 1960</bibRefCitation>
).
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<emphasis id="1DF1EAFFFF3CFF150FA60F7D0A84932B" bold="true" box="[189,426,860,886]" pageId="168" pageNumber="169">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="9FED3CB0FF3CFF150FF10FAC0E7B944A" ID-GBIF-Occurrence="2614776311" collectionCode="MACN" collectorName="Sta. Cruz &amp; Pcia. Cordillera &amp; Las Juntas &amp; Maldonado B &amp; Coll. Martinez &amp; Oct. &amp; Glaph." country="Bolivia" determinerName="A. Martinez" location="Santa Cruz" pageId="168" pageNumber="169" specimenCount="2" specimenCount-female="2" stateProvince="Santa Cruz" typeStatus="holotype">
<paragraph id="2F3A36EDFF3CFF150FF10FAC0A7293FA" blockId="168.[189,1398,909,1047]" box="[234,348,909,935]" pageId="168" pageNumber="169">
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<typeStatus id="F03E884FFF3CFF150FF10FAC0A7293FA" box="[234,348,909,935]" pageId="168" pageNumber="169" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="2F3A36EDFF3CFF150FA60F930E7B944A" blockId="168.[189,1398,909,1047]" pageId="168" pageNumber="169">
<collectingCountry id="5792767DFF3CFF150FA60F930A149391" box="[189,314,946,972]" name="Bolivia" pageId="168" pageNumber="169">BOLIVIA</collectingCountry>
: 
<specimenCount id="3983FD64FF3CFF150E5C0F930A739391" box="[327,349,946,972]" pageId="168" pageNumber="169" type="female">♀</specimenCount>
, 
<collectingRegion id="ED41F80FFF3CFF150E740F930ADB9391" box="[367,501,946,972]" country="Bolivia" name="Santa Cruz" pageId="168" pageNumber="169">Santa Cruz</collectingRegion>
, Cordillera, Las Juntas, (“ 
<emphasis id="1DF1EAFFFF3CFF150C2A0F93088E9391" box="[817,928,946,972]" italics="true" pageId="168" pageNumber="169">
<collectingCountry id="5792767DFF3CFF150C2A0F93088E9391" box="[817,928,946,972]" name="Bolivia" pageId="168" pageNumber="169">BOLIVIA</collectingCountry>
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150CA00F930F7D9391" box="[955,1107,946,972]" italics="true" pageId="168" pageNumber="169">
Dº 
<collectorName id="8270533BFF3CFF150CF80F930F7D9391" box="[995,1107,946,972]" pageId="168" pageNumber="169">Sta. Cruz</collectorName>
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150B740F920E1F9391" box="[1135,1329,946,972]" italics="true" pageId="168" pageNumber="169">
<collectorName id="8270533BFF3CFF150B740F920E1F9391" box="[1135,1329,946,972]" pageId="168" pageNumber="169">Pcia. Cordillera</collectorName>
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/ 
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<collectorName id="8270533BFF3CFF150A570F920A2493AC" pageId="168" pageNumber="169">Las Juntas</collectorName>
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150E3B0FF60AC393AC" box="[288,493,983,1009]" italics="true" pageId="168" pageNumber="169">
<collectorName id="8270533BFF3CFF150E3B0FF60AEA93AC" box="[288,452,983,1009]" pageId="168" pageNumber="169">Maldonado B</collectorName>
:!!
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150D1F0FF6098193AF" box="[516,687,983,1010]" italics="true" pageId="168" pageNumber="169">
<collectorName id="8270533BFF3CFF150D1F0FF6098193AF" box="[516,687,983,1010]" pageId="168" pageNumber="169">Coll. Martínez</collectorName>
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150DDD0FF6080093AC" box="[710,814,983,1009]" italics="true" pageId="168" pageNumber="169">
<collectorName id="8270533BFF3CFF150DDD0FF609DB93AC" box="[710,757,983,1009]" pageId="168" pageNumber="169">Oct.</collectorName>
-948
</emphasis>
”, “ 
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”, “ 
<taxonomicName id="E8854D6EFF3CFF150B360FF60FD593AC" box="[1069,1275,983,1009]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFF3CFF150B360FF60FD593AC" box="[1069,1275,983,1009]" italics="true" pageId="168" pageNumber="169">Glaphyrocanthon</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFF3CFF150A090FF60E5893AC" box="[1298,1398,983,1009]" italics="true" pageId="168" pageNumber="169">
(
<collectorName id="8270533BFF3CFF150A000FF60E4093AC" box="[1307,1390,983,1009]" pageId="168" pageNumber="169">Glaph.</collectorName>
)
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150FD70FDD0A76944B" box="[204,344,1020,1046]" italics="true" pageId="168" pageNumber="169">
<taxonomicName id="E8854D6EFF3CFF150FD70FDD0A37944B" authority="Cupello &amp; Vaz-De, 2018" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" box="[204,281,1020,1046]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="cobosi" status="sp. nov.">cobosi</taxonomicName>
<taxonomicNameLabel id="06C25784FF3CFF150E3B0FDC0A76944B" box="[288,344,1021,1046]" pageId="168" pageNumber="169" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
/ 
<emphasis id="1DF1EAFFFF3CFF150E740FDB0AAB944B" box="[367,389,1018,1046]" italics="true" pageId="168" pageNumber="169">
<specimenCount id="3983FD64FF3CFF150E740FDB0AAB944B" box="[367,389,1018,1046]" pageId="168" pageNumber="169" type="female">♀</specimenCount>
</emphasis>
/ 
<determinerName id="DAE6DA29FF3CFF150E810FDD0906944B" box="[410,552,1020,1046]" pageId="168" pageNumber="169">A. Martínez</determinerName>
det. 19 
<emphasis id="1DF1EAFFFF3CFF150D640FDD09B2944B" box="[639,668,1020,1046]" italics="true" pageId="168" pageNumber="169">56</emphasis>
”, “ 
<emphasis id="1DF1EAFFFF3CFF150DDE0FDC0868944A" box="[709,838,1021,1047]" italics="true" pageId="168" pageNumber="169">FICHADO</emphasis>
”, “MACN-En / 1009”) (
<collectionCode id="4994AE28FF3CFF150B730FDD0FEA944B" box="[1128,1220,1020,1046]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="168" pageNumber="169" type="Museum">MACN</collectionCode>
) (
<figureCitation id="B7BE2A68FF3CFF150BC70FDD0E69944A" box="[1244,1351,1020,1047]" captionStart="Fig" captionStartId="166.[189,232,1998,2024]" captionTargetBox="[192,1395,1280,1946]" captionTargetId="figure@166.[189,1399,1277,1949]" captionTargetPageId="166" captionText="Fig. 45. Canthon xanthopus Blanchard, 1846. A. Lectotype and its labels. B. An ordinary specimen." figureDoi="http://doi.org/10.5281/zenodo.3824875" httpUri="https://zenodo.org/record/3824875/files/figure.png" pageId="168" pageNumber="169">Fig. 45B</figureCitation>
).
</paragraph>
</materialsCitation>
<materialsCitation id="9FED3CB0FF3CFF150FF1081E0FE094F9" ID-GBIF-Occurrence="2614776310" pageId="168" pageNumber="169">
<paragraph id="2F3A36EDFF3CFF150FF1081E09829407" blockId="168.[189,1398,1086,1188]" box="[234,684,1086,1114]" pageId="168" pageNumber="169">
<emphasis id="1DF1EAFFFF3CFF150FF1081E0ACF9404" bold="true" box="[234,481,1087,1113]" pageId="168" pageNumber="169">Additional material</emphasis>
(22 ♂♂, 
<specimenCount id="3983FD64FF3CFF150D49081E098E9407" box="[594,672,1087,1114]" pageId="168" pageNumber="169" type="female">26 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="2F3A36EDFF3CFF150FA608440FE094F9" blockId="168.[189,1398,1086,1188]" pageId="168" pageNumber="169">
<collectingCountry id="5792767DFF3CFF150FA608440A439422" box="[189,365,1125,1151]" name="Argentina" pageId="168" pageNumber="169">ARGENTINA</collectingCountry>
: 
<emphasis id="1DF1EAFFFF3CFF150E6308450AEE9423" bold="true" box="[376,448,1124,1150]" pageId="168" pageNumber="169">
<collectingRegion id="ED41F80FFF3CFF150E6308450AEE9423" box="[376,448,1124,1150]" country="Argentina" name="Jujuy" pageId="168" pageNumber="169">Jujuy</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FF3CFF150ECB08440AD29422" box="[464,508,1125,1151]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Parque Nacional Calilegua, Águas Negras, 
<quantity id="E87D9B08FF3CFF150B1008450F769422" box="[1035,1112,1124,1151]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="168" pageNumber="169" unit="m" value="500.0">500 m</quantity>
, 
<date id="5B3B102DFF3CFF150B7E08440E2B9422" box="[1125,1285,1125,1151]" pageId="168" pageNumber="169" value="1987-12-18">18 Dec. 1987</date>
, S. and J. Peck leg. (
<collectionCode id="4994AE28FF3CFF150E2308AB0ABB94F9" box="[312,405,1162,1188]" pageId="168" pageNumber="169">CMNC</collectionCode>
); 3 ♂♂ (1 dissected), 
<specimenCount id="3983FD64FF3CFF150D8D08A809F794F9" box="[662,729,1161,1188]" pageId="168" pageNumber="169" type="female">5 ♀♀</specimenCount>
, Santa Barbara, Yuto, 
<date id="5B3B102DFF3CFF150CFB08A80F7894F9" box="[992,1110,1161,1188]" pageId="168" pageNumber="169" value="1982-02">Feb. 1982</date>
(
<collectionCode id="4994AE28FF3CFF150B7E08AB0FEC94F9" box="[1125,1218,1162,1188]" pageId="168" pageNumber="169">CMNC</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="2F3A36EDFF3CFF150FA608EC094E9526" blockId="168.[189,1399,1227,1404]" pageId="168" pageNumber="169">
<collectingCountry id="5792767DFF3CFF150FA608EC0A1494BA" box="[189,314,1229,1255]" name="Bolivia" pageId="168" pageNumber="169">BOLIVIA</collectingCountry>
: 
<emphasis id="1DF1EAFFFF3CFF150E5208EC0AF394BA" bold="true" box="[329,477,1229,1255]" pageId="168" pageNumber="169">
<collectingRegion id="ED41F80FFF3CFF150E5208EC0AF394BA" box="[329,477,1229,1255]" country="Bolivia" name="Santa Cruz" pageId="168" pageNumber="169">Santa Cruz</collectingRegion>
</emphasis>
: 1 ♂, Cordillera, Parabanó (
<collectionCode id="4994AE28FF3CFF150C2508EC08B894BA" box="[830,918,1229,1255]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150CB508EC08F094BA" box="[942,990,1229,1255]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Cordillera, Río Seco, 
<date id="5B3B102DFF3CFF150BE608EC0E5994BA" box="[1277,1399,1229,1255]" pageId="168" pageNumber="169" value="1961-02">Feb. 1961</date>
(
<collectionCode id="4994AE28FF3CFF150FDE08D30A0C9551" box="[197,290,1266,1292]" pageId="168" pageNumber="169">CMNC</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FF3CFF150E6B08D30A9C9551" box="[368,434,1266,1292]" pageId="168" pageNumber="169" type="female">4 ♀♀</specimenCount>
, Cordillera, Río Seco, 
<date id="5B3B102DFF3CFF150DA608D3081C9551" box="[701,818,1266,1292]" pageId="168" pageNumber="169" value="1962-02">Feb. 1962</date>
(
<collectionCode id="4994AE28FF3CFF150C5A08D308B09551" box="[833,926,1266,1292]" pageId="168" pageNumber="169">CMNC</collectionCode>
); 1 ♂, 
<collectingRegion id="ED41F80FFF3CFF150CF708D30FD59551" box="[1004,1275,1266,1292]" country="Bolivia" name="Santa Cruz" pageId="168" pageNumber="169">Santa Cruz de la Sierra</collectingRegion>
, Río Piraí, 
<date id="5B3B102DFF3CFF150FA609360A1B956C" box="[189,309,1303,1329]" pageId="168" pageNumber="169" value="1964-09">Sep. 1964</date>
(?) (
<collectionCode id="4994AE28FF3CFF150E6A09360AE0956C" box="[369,462,1303,1329]" pageId="168" pageNumber="169">CMNC</collectionCode>
); 2 ♂♂, 
<specimenCount id="3983FD64FF3CFF150D20093609AE956F" box="[571,640,1303,1330]" pageId="168" pageNumber="169" type="female">2 ♀♀</specimenCount>
, Tentarembei, 
<geoCoordinate id="4AB1502AFF3CFF150C2F093408B4956C" box="[820,922,1301,1329]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="168" pageNumber="169" precision="925" value="-18.466667">18º28′ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150CB209340F37956F" box="[937,1049,1301,1330]" degrees="62" direction="west" minutes="05" orientation="longitude" pageId="168" pageNumber="169" precision="925" value="-62.083332">62º05′ W</geoCoordinate>
, 
<quantity id="E87D9B08FF3CFF150B3C09360F58956F" box="[1063,1142,1303,1330]" metricMagnitude="2" metricUnit="m" metricValue="3.18" pageId="168" pageNumber="169" unit="m" value="318.0">318 m</quantity>
, 
<date id="5B3B102DFF3CFF150B9E09360E37956C" box="[1157,1305,1303,1329]" pageId="168" pageNumber="169" value="2010-02-06">6 Feb. 2010</date>
, human faeces, Vidaurre 
<emphasis id="1DF1EAFFFF3CFF150E98091C0A90950B" box="[387,446,1340,1366]" italics="true" pageId="168" pageNumber="169">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FF3CFF150D1A091D0977950B" box="[513,601,1340,1366]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 3 ♂♂, Sara, Santa Rosa del Sara, 
<date id="5B3B102DFF3CFF150CE6091C0F56950B" box="[1021,1144,1340,1367]" pageId="168" pageNumber="169" value="1969-11">Nov. 1969</date>
(
<collectionCode id="4994AE28FF3CFF150B93091D0FCB950B" box="[1160,1253,1340,1366]" pageId="168" pageNumber="169">CMNC</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150BE1091D0E08950A" box="[1274,1318,1340,1367]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Santa Rosa del Sara, 
<date id="5B3B102DFF3CFF150E7609430AC99526" box="[365,487,1377,1404]" pageId="168" pageNumber="169" value="1972-11">Nov. 1972</date>
(
<collectionCode id="4994AE28FF3CFF150EED0940097D9526" box="[502,595,1377,1403]" pageId="168" pageNumber="169">CMNC</collectionCode>
).
</paragraph>
<paragraph id="2F3A36EDFF3CFF150FA609840F4797B6" blockId="168.[189,1399,1443,2027]" pageId="168" pageNumber="169">
<collectingCountry id="5792767DFF3CFF150FA609840A0295E2" box="[189,300,1445,1471]" name="Brazil" pageId="168" pageNumber="169">BRAZIL</collectingCountry>
: 
<emphasis id="1DF1EAFFFF3CFF150E2D09840AF895E3" bold="true" box="[310,470,1444,1471]" pageId="168" pageNumber="169">
<collectingRegion id="ED41F80FFF3CFF150E2D09840AF895E3" box="[310,470,1444,1471]" country="Brazil" name="Mato Grosso" pageId="168" pageNumber="169">Mato Grosso</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FF3CFF150EFD0985090695E2" box="[486,552,1444,1471]" pageId="168" pageNumber="169" type="female">5 ♀♀</specimenCount>
, Barra do Bugres, Rio Currupiras, 
<geoCoordinate id="4AB1502AFF3CFF150CA409820F4C95E2" box="[959,1122,1443,1471]" degrees="15" direction="south" minutes="10" orientation="latitude" pageId="168" pageNumber="169" precision="1" seconds="08.2" value="-15.168944">15º10′08.2″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150B7509820E3495E2" box="[1134,1306,1443,1471]" degrees="56" direction="west" minutes="50" orientation="longitude" pageId="168" pageNumber="169" precision="1" seconds="22.7" value="-56.839638">56º50′22.7″ W</geoCoordinate>
, 
<quantity id="E87D9B08FF3CFF150A3E09850E5F95E2" box="[1317,1393,1444,1471]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="168" pageNumber="169" unit="m" value="296.0">296 m</quantity>
, 
<date id="5B3B102DFF3CFF150FA609EB0A7495B9" box="[189,346,1481,1508]" pageId="168" pageNumber="169" value="2010-02-28">28 Feb. 2010</date>
, pitfall with human faeces, J.G. Mota-Souza leg. (
<collectionCode id="4994AE28FF3CFF150CB909EB08D495B9" box="[930,1018,1482,1508]" pageId="168" pageNumber="169">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFF3CFF150B3809EB0E3195BE" bold="true" box="[1059,1311,1481,1508]" pageId="168" pageNumber="169">
<collectingRegion id="ED41F80FFF3CFF150B3809EB0E3195BE" box="[1059,1311,1481,1508]" country="Brazil" name="Mato Grosso do Sul" pageId="168" pageNumber="169">Mato Grosso Do Sul</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FF3CFF150A3609EB0E5E95B9" box="[1325,1392,1482,1508]" pageId="168" pageNumber="169" type="female">2 ♀♀</specimenCount>
, Aquidauana, Piraputanga, 
<date id="5B3B102DFF3CFF150EEC09CE09BD9654" box="[503,659,1519,1545]" pageId="168" pageNumber="169" value="2000-06-17">17 Jun. 2000</date>
, V. Lopes leg. (
<collectionCode id="4994AE28FF3CFF150C5609CE088B9654" box="[845,933,1519,1545]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, Bodoquena, Fazenda Califórnia, 
<geoCoordinate id="4AB1502AFF3CFF150FA60A330A619673" box="[189,335,1554,1582]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="05" value="-20.684721">20º41′05″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150E7B0A330AD59673" box="[352,507,1554,1582]" degrees="56" direction="west" minutes="51" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="33" value="-56.859165">56º51′33″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150D100A3509A79673" box="[523,649,1556,1582]" pageId="168" pageNumber="169" value="2011-03">Mar. 2011</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150B2E0A350FA39673" box="[1077,1165,1556,1582]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, Bodoquena, Fazenda Califórnia, 
<geoCoordinate id="4AB1502AFF3CFF150EAB0A16096E960E" box="[432,576,1591,1619]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="08" value="-20.685555">20º41′08″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150D540A1609C6960E" box="[591,744,1591,1619]" degrees="56" direction="west" minutes="51" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="31" value="-56.85861">56º51′31″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150DED0A18085C960E" box="[758,882,1593,1619]" pageId="168" pageNumber="169" value="2011-03">Mar. 2011</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150A090A180E44960E" box="[1298,1386,1593,1619]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150FA60A7F0BC59625" box="[189,235,1630,1656]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Bonito, Fazenda Palmeirinhas, 
<geoCoordinate id="4AB1502AFF3CFF150D690A7D08249625" box="[626,778,1628,1656]" degrees="21" direction="south" minutes="11.269" orientation="latitude" pageId="168" pageNumber="169" precision="1" value="-21.187817">21º11.269′ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150C020A7D08959625" box="[793,955,1628,1656]" degrees="56" direction="west" minutes="33.701" orientation="longitude" pageId="168" pageNumber="169" precision="1" value="-56.561684">56º33.701′ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150CD20A7F0F669625" box="[969,1096,1630,1656]" pageId="168" pageNumber="169" value="2009-11">Nov. 2009</date>
, banana, F.O. Roque leg. (
<collectionCode id="4994AE28FF3CFF150FDE0AA20A3396C0" box="[197,285,1667,1693]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FF3CFF150E7C0AA20A8696C3" box="[359,424,1667,1694]" pageId="168" pageNumber="169" type="female">2 ♀♀</specimenCount>
, Bonito, [Hotel] Cabanas, 
<geoCoordinate id="4AB1502AFF3CFF150DC70AA0084996C0" box="[732,871,1665,1693]" degrees="21" direction="south" minutes="10" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="15" value="-21.170834">21º10′15″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150C690AA00F2996C3" box="[882,1031,1665,1694]" degrees="56" direction="west" minutes="26" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="22" value="-56.439445">56º26′22″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150B0A0AA50FA596C0" box="[1041,1163,1667,1694]" pageId="168" pageNumber="169" value="2010-12">Dec. 2010</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150E4F0A890A82969F" box="[340,428,1704,1730]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150EDB0A890AC5969E" box="[448,491,1704,1731]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Bonito, Hotel Cabanas, 
<geoCoordinate id="4AB1502AFF3CFF150C0B0A8708B2969F" box="[784,924,1702,1730]" degrees="21" direction="south" minutes="10" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="17" value="-21.171389">21º10′17″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150CB20A870F10969E" box="[937,1086,1702,1731]" degrees="56" direction="west" minutes="26" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="26" value="-56.440556">56º26′26″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150B520A880FEB969F" box="[1097,1221,1704,1731]" pageId="168" pageNumber="169" value="2010-12">Dec. 2010</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150E8D0AEC0AC096BA" box="[406,494,1741,1767]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150D180AEC091E96B5" box="[515,560,1741,1768]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Bonito, Hotel Cabanas, 
<geoCoordinate id="4AB1502AFF3CFF150C470AEA08C596BA" box="[860,1003,1739,1767]" degrees="21" direction="south" minutes="10" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="20" value="-21.172222">21º10′20″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150CE20AEA0FBE96B5" box="[1017,1168,1739,1768]" degrees="56" direction="west" minutes="26" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="37" value="-56.44361">56º26′37″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150B860AEF0E3496BA" box="[1181,1306,1741,1768]" pageId="168" pageNumber="169" value="2010-12">Dec. 2010</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150EF40AD309699751" box="[495,583,1778,1804]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150D470AD309A79750" box="[604,649,1778,1805]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Bonito, Hotel Cabanas, 
<geoCoordinate id="4AB1502AFF3CFF150CAE0AD10F6D9751" box="[949,1091,1776,1804]" degrees="21" direction="south" minutes="10" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="21" value="-21.172499">21º10′21″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150B4A0AD10FC79750" box="[1105,1257,1776,1805]" degrees="56" direction="west" minutes="26" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="34" value="-56.44278">56º26′34″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150BEE0AD20E5C9751" box="[1269,1394,1778,1805]" pageId="168" pageNumber="169" value="2010-12">Dec. 2010</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150D500B36098D976C" box="[587,675,1815,1841]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, Corguinho, Quinta do Sol, 
<geoCoordinate id="4AB1502AFF3CFF150B270B340FE5976C" box="[1084,1227,1813,1841]" degrees="19" direction="south" minutes="49" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="57" value="-19.8325">19º49′57″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150BC10B340E5C976F" box="[1242,1394,1813,1842]" degrees="54" direction="west" minutes="49" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="45" value="-54.829166">54º49′45″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150FA60B1D0A1A970B" box="[189,308,1852,1878]" pageId="168" pageNumber="169" value="2011-02">Feb. 2011</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150DDE0B1D0833970B" box="[709,797,1852,1878]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, Corumbá, 
<date id="5B3B102DFF3CFF150CF30B1C0F4A970B" box="[1000,1124,1852,1879]" pageId="168" pageNumber="169" value="2007-12">Dec. 2007</date>
, C. Aoki leg. (
<collectionCode id="4994AE28FF3CFF150A160B1D0E44970B" box="[1293,1386,1852,1878]" pageId="168" pageNumber="169">AMBC</collectionCode>
); 1 ♂, same collecting data as for preceding (
<collectionCode id="4994AE28FF3CFF150DD80B4308359721" box="[707,795,1890,1916]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FF3CFF150C760B4308B49721" box="[877,922,1890,1916]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Corumbá, Passo do Lontra, 
<date id="5B3B102DFF3CFF150BEE0B430E5C9721" box="[1269,1394,1889,1916]" pageId="168" pageNumber="169" value="2005-12">Dec. 2005</date>
, J.N.C. Louzada leg. (
<collectionCode id="4994AE28FF3CFF150EA80BA6092597FC" box="[435,523,1927,1953]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, Ivinhema, 
<geoCoordinate id="4AB1502AFF3CFF150DCE0BA4084F97FC" box="[725,865,1925,1953]" degrees="22" direction="south" minutes="31" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="41" value="-22.528055">22º31′41″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150C750BA40F2D97FC" box="[878,1027,1925,1953]" degrees="53" direction="west" minutes="53" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="38" value="-53.89389">53º53′38″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150B150BA60FA697FC" box="[1038,1160,1927,1953]" pageId="168" pageNumber="169" value="2011-03">Mar. 2011</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150E430B8D0A9E979B" box="[344,432,1964,1990]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FF3CFF150D180B8D091E979B" box="[515,560,1964,1990]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Jardim, 
<date id="5B3B102DFF3CFF150DBB0B8D0839979B" box="[672,791,1964,1990]" pageId="168" pageNumber="169" value="2000-06">Jun. 2000</date>
, V. Lopes leg. (
<collectionCode id="4994AE28FF3CFF150CCA0B8D0F07979B" box="[977,1065,1964,1990]" pageId="168" pageNumber="169">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FF3CFF150B250B8D0F42979B" box="[1086,1132,1964,1990]" pageId="168" pageNumber="169" type="female">1 ♀</specimenCount>
, Jardim, Rio da Prata, 
<geoCoordinate id="4AB1502AFF3CFF150FA60BEE0A6497B6" box="[189,330,1999,2027]" degrees="21" direction="south" minutes="25" orientation="latitude" pageId="168" pageNumber="169" precision="15" seconds="40" value="-21.427776">21º25′40″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFF3CFF150E4C0BEE0AC097B6" box="[343,494,1999,2027]" degrees="56" direction="west" minutes="27" orientation="longitude" pageId="168" pageNumber="169" precision="15" seconds="00" value="-56.45">56º27′00″ W</geoCoordinate>
, 
<date id="5B3B102DFF3CFF150EE20BF0095D97B6" box="[505,627,2001,2027]" pageId="168" pageNumber="169" value="2011-03">Mar. 2011</date>
, human faeces, L.O. Bavutti leg. (
<collectionCode id="4994AE28FF3CFF150B1F0BF00F7297B6" box="[1028,1116,2001,2027]" pageId="168" pageNumber="169">CEMT</collectionCode>
).
</paragraph>
<caption id="7BFA6665FF3DFF140FA60B880F7297B6" ID-DOI="http://doi.org/10.5281/zenodo.3824878" ID-Zenodo-Dep="3824878" httpUri="https://zenodo.org/record/3824878/files/figure.png" pageId="169" pageNumber="170" startId="169.[189,232,1961,1987]" targetBox="[350,1246,265,1923]" targetPageId="169">
<paragraph id="2F3A36EDFF3DFF140FA60B880F7297B6" blockId="169.[189,1398,1961,2028]" pageId="169" pageNumber="170">
<emphasis id="1DF1EAFFFF3DFF140FA60B880A15979E" bold="true" box="[189,315,1961,1987]" pageId="169" pageNumber="170">Fig. 46. A</emphasis>
. 
<taxonomicName id="E8854D6EFF3DFF140E570B8808769799" authority="(Pereira &amp; Martinez, 1960)" baseAuthorityName="Pereira &amp; Martinez" baseAuthorityYear="1960" box="[332,856,1961,1988]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="1DF1EAFFFF3DFF140E570B880926979E" box="[332,520,1961,1987]" italics="true" pageId="169" pageNumber="170">Canthon cobosi</emphasis>
(
<bibRefCitation id="4B144B1CFF3DFF140D020B88087E9799" author="Pereira F. S. &amp; Martinez A." box="[537,848,1961,1988]" pageId="169" pageNumber="170" pagination="37 - 55" refId="ref144264" refString="Pereira F. S. &amp; Martinez A. 1960. Notas escarabeidologicas - II. Revista Brasileira de Entomologia 9: 37 - 55." type="journal article" year="1960">Pereira &amp; Martínez, 1960</bibRefCitation>
)
</taxonomicName>
stat. et comb. nov., holotype. 
<emphasis id="1DF1EAFFFF3DFF140BD20B880E2E979E" bold="true" box="[1225,1280,1961,1987]" pageId="169" pageNumber="170">B–C</emphasis>
. 
<taxonomicName id="E8854D6EFF3DFF140A0A0B88095997B1" authority="(Martinez &amp; Pereira, 1967)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Martinez &amp; Pereira" baseAuthorityYear="1967" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="machadoi" status="comb. nov.">
<emphasis id="1DF1EAFFFF3DFF140A0A0B880A1F97B6" italics="true" pageId="169" pageNumber="170">Canthon machadoi</emphasis>
(
<bibRefCitation id="4B144B1CFF3DFF140E5B0BF0094197B1" author="Martinez A. &amp; Pereira F. S." box="[320,623,2001,2028]" pageId="169" pageNumber="170" pagination="53 - 69" refId="ref140364" refString="Martinez A. &amp; Pereira F. S. 1967. Notas escarabeidologicas III (Col. Scarabaeidae-Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez &amp; Pereira, 1967</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="06C25784FF3DFF140D640BF0082E97B6" box="[639,768,2001,2027]" pageId="169" pageNumber="170" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="1DF1EAFFFF3DFF140C1C0BF0083597B6" bold="true" box="[775,795,2001,2027]" pageId="169" pageNumber="170">B</emphasis>
. Paratype 2. 
<emphasis id="1DF1EAFFFF3DFF140CAE0BF108E497B7" bold="true" box="[949,970,2000,2026]" pageId="169" pageNumber="170">C</emphasis>
. Paratype 4.
</paragraph>
</caption>
<paragraph id="2F3A36EDFF3EFF170FA60D2A09239114" blockId="170.[189,1399,266,330]" pageId="170" pageNumber="171">
<collectingCountry id="5792767DFF3EFF170FA60D2A0A739178" box="[189,349,267,293]" name="Paraguay" pageId="170" pageNumber="171">PARAGUAY</collectingCountry>
: 
<emphasis id="1DF1EAFFFF3EFF170E730D2A0AC49178" bold="true" box="[360,490,267,293]" pageId="170" pageNumber="171">
<collectingRegion id="ED41F80FFF3EFF170E730D2A0AC49178" box="[360,490,267,293]" country="Paraguay" name="Cordillera" pageId="170" pageNumber="171">Cordillera</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FF3EFF170EE00D2A0909917B" box="[507,551,267,294]" pageId="170" pageNumber="171" type="female">1 ♀</specimenCount>
, 
<date id="5B3B102DFF3EFF170D2C0D2A0993917B" box="[567,701,267,294]" pageId="170" pageNumber="171" value="2005-07-05">5 Jul. 2005</date>
, Naraujo leg. (
<collectionCode id="4994AE28FF3EFF170C770D2A08EA9178" box="[876,964,267,293]" pageId="170" pageNumber="171">CEMT</collectionCode>
); 2 ♂♂, Caacupé (“ 
<emphasis id="1DF1EAFFFF3EFF170BB40D2D0E039178" box="[1199,1325,267,294]" italics="true" pageId="170" pageNumber="171">Dº Capital</emphasis>
/ 
<emphasis id="1DF1EAFFFF3EFF170A5D0D2D0BD59117" italics="true" pageId="170" pageNumber="171">Caa Cupé</emphasis>
”), 
<date id="5B3B102DFF3EFF170E3B0D0E0ABA9114" box="[288,404,303,329]" pageId="170" pageNumber="171" value="1979-10">Oct. 1979</date>
(
<collectionCode id="4994AE28FF3EFF170EBF0D0E092F9114" box="[420,513,303,329]" pageId="170" pageNumber="171">CMNC</collectionCode>
).
</paragraph>
<paragraph id="2F3A36EDFF3EFF170FA60D500A5791D6" blockId="170.[189,377,369,395]" box="[189,377,369,395]" pageId="170" pageNumber="171">
<heading id="74728181FF3EFF170FA60D500A5791D6" bold="true" box="[189,377,369,395]" fontSize="11" level="3" pageId="170" pageNumber="171" reason="3">
<emphasis id="1DF1EAFFFF3EFF170FA60D500A5791D6" bold="true" box="[189,377,369,395]" pageId="170" pageNumber="171">Collecting sites</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFF3EFF170FA60D8009D69183" blockId="170.[189,1398,416,478]" pageId="170" pageNumber="171">
<collectingCountry id="5792767DFF3EFF170FA60D800A0591E6" box="[189,299,417,443]" name="Brazil" pageId="170" pageNumber="171">BRAZIL</collectingCountry>
. 
<emphasis id="1DF1EAFFFF3EFF170E2F0D800AFD91E7" bold="true" box="[308,467,416,443]" pageId="170" pageNumber="171">
<collectingRegion id="ED41F80FFF3EFF170E2F0D800AFD91E7" box="[308,467,416,443]" country="Brazil" name="Mato Grosso" pageId="170" pageNumber="171">Mato Grosso</collectingRegion>
</emphasis>
: Barra do Bugres. 
<emphasis id="1DF1EAFFFF3EFF170DB00D8008B091E7" bold="true" box="[683,926,416,443]" pageId="170" pageNumber="171">
<collectingRegion id="ED41F80FFF3EFF170DB00D8008B091E7" box="[683,926,416,443]" country="Brazil" name="Mato Grosso do Sul" pageId="170" pageNumber="171">Mato Grosso do Sul</collectingRegion>
</emphasis>
: Aquidauana (Piraputanga), Bodoquena, Bonito, Corguinho, Corumbá, Ivinhema, Jardim.
</paragraph>
<paragraph id="2F3A36EDFF3EFF170FA60E260A009219" blockId="170.[189,1399,518,580]" pageId="170" pageNumber="171">
<collectingCountry id="5792767DFF3EFF170FA60E260A17927C" box="[189,313,519,545]" name="Bolivia" pageId="170" pageNumber="171">BOLIVIA</collectingCountry>
. 
<emphasis id="1DF1EAFFFF3EFF170E5D0E270AF9927D" bold="true" box="[326,471,518,544]" pageId="170" pageNumber="171">
<collectingRegion id="ED41F80FFF3EFF170E5D0E270AF9927D" box="[326,471,518,544]" country="Bolivia" name="Santa Cruz" pageId="170" pageNumber="171">Santa Cruz</collectingRegion>
</emphasis>
: Cordillera (Las Juntas, Río Seco), 
<collectingRegion id="ED41F80FFF3EFF170C9E0E260FB3927D" box="[901,1181,518,545]" country="Bolivia" name="Santa Cruz" pageId="170" pageNumber="171">Santa Cruz de la Sierra</collectingRegion>
, Sara (Santa Rosa del Sara).
</paragraph>
<paragraph id="2F3A36EDFF3EFF170FA60E4C094A92DA" blockId="170.[189,612,620,647]" box="[189,612,620,647]" pageId="170" pageNumber="171">
<collectingCountry id="5792767DFF3EFF170FA60E4C0A7592DA" box="[189,347,621,647]" name="Paraguay" pageId="170" pageNumber="171">PARAGUAY</collectingCountry>
. 
<emphasis id="1DF1EAFFFF3EFF170E7E0E4D0AC992DB" bold="true" box="[357,487,620,646]" pageId="170" pageNumber="171">
<collectingRegion id="ED41F80FFF3EFF170E7E0E4D0AC992DB" box="[357,487,620,646]" country="Paraguay" name="Cordillera" pageId="170" pageNumber="171">Cordillera</collectingRegion>
</emphasis>
: Caacupé.
</paragraph>
<paragraph id="2F3A36EDFF3EFF170FA60E8E08EA9295" blockId="170.[189,964,686,713]" box="[189,964,686,713]" pageId="170" pageNumber="171">
<collectingCountry id="5792767DFF3EFF170FA60E8E0A429294" box="[189,364,687,713]" name="Argentina" pageId="170" pageNumber="171">ARGENTINA</collectingCountry>
. 
<emphasis id="1DF1EAFFFF3EFF170E6C0E8F0A919295" bold="true" box="[375,447,686,712]" pageId="170" pageNumber="171">
<collectingRegion id="ED41F80FFF3EFF170E6C0E8F0A919295" box="[375,447,686,712]" country="Argentina" name="Jujuy" pageId="170" pageNumber="171">Jujuy</collectingRegion>
</emphasis>
: Parque Nacional Calilegua, Santa Barbara.
</paragraph>
</subSubSection>
<subSubSection id="679F6566FF3EFF170FA60ED10ADA9551" pageId="170" pageNumber="171" type="discussion">
<paragraph id="2F3A36EDFF3EFF170FA60ED10A6A9357" blockId="170.[189,324,752,778]" box="[189,324,752,778]" pageId="170" pageNumber="171">
<heading id="74728181FF3EFF170FA60ED10A6A9357" bold="true" box="[189,324,752,778]" fontSize="11" level="3" pageId="170" pageNumber="171" reason="3">
<emphasis id="1DF1EAFFFF3EFF170FA60ED10A6A9357" bold="true" box="[189,324,752,778]" pageId="170" pageNumber="171">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFF3EFF170FA60F010ADA9551" blockId="170.[189,1399,800,1292]" pageId="170" pageNumber="171">
From the only geographical record ever published for 
<taxonomicName id="E8854D6EFF3EFF170C4E0F0108E99367" baseAuthorityName="Pereira &amp; Martinez" baseAuthorityYear="1960" box="[853,967,800,826]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="1DF1EAFFFF3EFF170C4E0F0108E99367" box="[853,967,800,826]" italics="true" pageId="170" pageNumber="171">C. cobosi</emphasis>
</taxonomicName>
– the locality of Las Juntas, in the province of Cordillera, 
<collectingRegion id="ED41F80FFF3EFF170EC60F6509489303" box="[477,614,836,862]" country="Bolivia" name="Santa Cruz" pageId="170" pageNumber="171">Santa Cruz</collectingRegion>
, 
<collectingCountry id="5792767DFF3EFF170D6D0F6509E29303" box="[630,716,836,862]" name="Bolivia" pageId="170" pageNumber="171">Bolivia</collectingCountry>
–, the known distribution of this species is widened to encompass areas also in 
<collectingCountry id="5792767DFF3EFF170ECE0F49093393DF" box="[469,541,872,898]" name="Brazil" pageId="170" pageNumber="171">Brazil</collectingCountry>
(
<collectingRegion id="ED41F80FFF3EFF170D320F4909EF93DF" box="[553,705,872,898]" country="Brazil" name="Mato Grosso" pageId="170" pageNumber="171">Mato Grosso</collectingRegion>
and 
<collectingRegion id="ED41F80FFF3EFF170DED0F4908F193DE" box="[758,991,872,899]" country="Brazil" name="Mato Grosso do Sul" pageId="170" pageNumber="171">Mato Grosso do Sul</collectingRegion>
), 
<collectingCountry id="5792767DFF3EFF170CEA0F490F7093DF" box="[1009,1118,872,898]" name="Paraguay" pageId="170" pageNumber="171">Paraguay</collectingCountry>
and northern 
<collectingCountry id="5792767DFF3EFF170BE10F490E5D93DF" box="[1274,1395,872,898]" name="Argentina" pageId="170" pageNumber="171">Argentina</collectingCountry>
. There is, however, a noteworthy geographical variation that, when studied in more detail, may reveal that what we are considering to be a single species represents, in fact, two: specimens from 
<collectingCountry id="5792767DFF3EFF170A3C0F8E0E5C9394" box="[1319,1394,943,969]" name="Brazil" pageId="170" pageNumber="171">Brazil</collectingCountry>
, in general, are darker, with few coppery reflections, while individuals from 
<collectingCountry id="5792767DFF3EFF170B7F0FF20F9393B0" box="[1124,1213,979,1005]" name="Bolivia" pageId="170" pageNumber="171">Bolivia</collectingCountry>
, 
<collectingCountry id="5792767DFF3EFF170BD70FF20E6F93B0" box="[1228,1345,979,1005]" name="Argentina" pageId="170" pageNumber="171">Argentina</collectingCountry>
and 
<collectingCountry id="5792767DFF3EFF170FA60FD90A04944F" box="[189,298,1016,1042]" name="Paraguay" pageId="170" pageNumber="171">Paraguay</collectingCountry>
show a coppery colouration with greenish reflections (
<figureCitation id="B7BE2A68FF3EFF170CAD0FD60F0B944F" box="[950,1061,1015,1042]" captionStart="Fig" captionStartId="169.[189,232,1961,1987]" captionTargetBox="[350,1246,265,1923]" captionTargetId="figure@169.[313,1275,265,1935]" captionTargetPageId="169" captionText="Fig. 46. A. Canthon cobosi (Pereira &amp; Martínez, 1960) stat. et comb. nov., holotype. B–C. Canthon machadoi (Martínez &amp; Pereira, 1967) comb. nov. B. Paratype 2. C. Paratype 4." figureDoi="http://doi.org/10.5281/zenodo.3824878" httpUri="https://zenodo.org/record/3824878/files/figure.png" pageId="170" pageNumber="171">Fig. 46A</figureCitation>
). As no other morphological variations were found and as there is chromatic variation even among individuals from 
<collectingCountry id="5792767DFF3EFF170BC1083A0E0C9468" box="[1242,1314,1051,1077]" name="Brazil" pageId="170" pageNumber="171">Brazil</collectingCountry>
(being some of them more coppery than others), we do not consider we have at this moment enough evidence for the existence of more than a single species. Nonetheless, we recognize that more research is needed in order to have a deeper understanding of the inter- and intrapopulational variation of 
<taxonomicName id="E8854D6EFF3EFF170BD608A60E1094FC" baseAuthorityName="Pereira &amp; Martinez" baseAuthorityYear="1960" box="[1229,1342,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="1DF1EAFFFF3EFF170BD608A60E1094FC" box="[1229,1342,1159,1185]" italics="true" pageId="170" pageNumber="171">C. cobosi</emphasis>
</taxonomicName>
. For details on the revalidation of this species from its synonymy with 
<taxonomicName id="E8854D6EFF3EFF170CAD088A0F7E9499" box="[950,1104,1194,1221]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="xanthopus">
<emphasis id="1DF1EAFFFF3EFF170CAD088A0F7E9499" box="[950,1104,1194,1221]" italics="true" pageId="170" pageNumber="171">C. xanthopus</emphasis>
</taxonomicName>
, see the discussion under this latter species, while for details on its transference to 
<taxonomicName id="E8854D6EFF3EFF170C7408EF08FA94B5" authorityName=", Harold" authorityYear="1868" box="[879,980,1230,1256]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFF3EFF170C7408EF08FA94B5" box="[879,980,1230,1256]" italics="true" pageId="170" pageNumber="171">Canthon</emphasis>
</taxonomicName>
, see section ‘Species composition’ under 
<taxonomicName id="E8854D6EFF3EFF170E1C08D30AB19551" authorityName="Halffter &amp; Martinez" authorityYear="1977" box="[263,415,1266,1292]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFF3EFF170E1C08D30AB19551" box="[263,415,1266,1292]" italics="true" pageId="170" pageNumber="171">Sylvicanthon</emphasis>
</taxonomicName>
above.
</paragraph>
</subSubSection>
</treatment>
</document>