treatments-xml/data/7B/78/0A/7B780A3C0911FF9C98D7A48AC2B16392.xml

<document id="4C4F6124823BBF23A1277ABAEAE61558" ID-DOI="10.11646/zootaxa.5507.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13757735" ID-ZooBank="3B1CC6CC-4300-4B1E-B257-3A1BBF034E4F" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1726221476130" checkinUser="plazi" docAuthor="Denzer, Wolfgang &amp; Tillack, Frank" docDate="2024" docId="7B780A3C0911FF9C98D7A48AC2B16392" docLanguage="en" docName="zootaxa.5507.2.6.pdf" docOrigin="Zootaxa 5507 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agama gutturosa Merrem 1819" docType="treatment" docVersion="1" lastPageNumber="348" masterDocId="874172440918FF909840A759C302613B" masterDocTitle="The original description of Agama gutturosa Merrem, 1819 and its nomenclatural implication on the genus Bronchocela Kaup, 1827 (Squamata: Agamidae)" masterLastPageNumber="355" masterPageNumber="337" pageNumber="346" updateTime="1726666419292" updateUser="julia" zenodo-license-document="CLOSED">
<mods:mods id="86ACC8853EC10638747C70C9DB4CA030" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="873CF75CC6A99EB9800683184812BD79">
<mods:title id="CA2C80250344D6FB3337EC76A5F0C7A4">The original description of Agama gutturosa Merrem, 1819 and its nomenclatural implication on the genus Bronchocela Kaup, 1827 (Squamata: Agamidae)</mods:title>
</mods:titleInfo>
<mods:name id="AB4C1FB6D5F97EEE23A322CC6958E806" type="personal">
<mods:role id="14AEFB92D61EF819B127A139DF841822">
<mods:roleTerm id="95D773EF857DA9C15AC2082C3420B7A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8813AA7C0AE21669FB9D6F060442E42">Denzer, Wolfgang</mods:namePart>
<mods:nameIdentifier id="3F4686A0F99357D554084C40DCBB0516" type="ORCID">0000-0003-2619-4555</mods:nameIdentifier>
</mods:name>
<mods:name id="79B1E9677B958B74822E4DCB76D01EE1" type="personal">
<mods:role id="009383009280E07D4FDA90BF941B3B70">
<mods:roleTerm id="FC880E31EE148064DE0F9FD5870BBE26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="901385CA05A0418AC8AFC5625320C42E">Tillack, Frank</mods:namePart>
<mods:nameIdentifier id="1DCAF19F904CF8F8BC9F52626514F30A" type="ORCID">0000-0003-1418-6531</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="EA60404E5FE777DD96DA35A150730C65">text</mods:typeOfResource>
<mods:relatedItem id="099CF0FD0D48DEE3A23A5856994C4068" type="host">
<mods:titleInfo id="9C81B69DEEB861BED4C2BFDBEE036BF7">
<mods:title id="262C2C71C1DBCD6660AC0662FA78EC90">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="244729DF4C1223F7889356D02B0177C0">
<mods:date id="26FE41C0F55969286976BA124AEE7056">2024</mods:date>
<mods:detail id="5BD97FC90B0C4AB558632E95E78887D1" type="pubDate">
<mods:number id="DDF684B4E91EA85A8512B4CEC5E016B3">2024-09-12</mods:number>
</mods:detail>
<mods:detail id="2576B83EC1A91F22CDC535DB86C37AAE" type="volume">
<mods:number id="9F1A66311BEAA41D141EEBB8C9B2F5DF">5507</mods:number>
</mods:detail>
<mods:detail id="CC6E42076D01EAD7EDD8BC9EBCB6FE87" type="issue">
<mods:number id="4BFAFADE5E16CC243F86CD6D0C69AAF2">2</mods:number>
</mods:detail>
<mods:extent id="D0493FC06CB721FF2AC769439494E982" unit="page">
<mods:start id="5BD9D286939195A8BC9B3EAD072F810B">337</mods:start>
<mods:end id="0F280DD892CCAD45F3B8DD06FCAD69FC">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="7585910840C0FE950A1E4EE985E3ECF9">
<mods:url id="84549AC504C0CC8E6838A8E064EA17DE">http://dx.doi.org/10.11646/zootaxa.5507.2.6</mods:url>
</mods:location>
<mods:classification id="6FE3F9FA3F297DFC4B4965AC87CA8668">journal article</mods:classification>
<mods:identifier id="1009624BEB084B66F41073C33CE42FE2" type="DOI">10.11646/zootaxa.5507.2.6</mods:identifier>
<mods:identifier id="CE2E5FED36F13AD325BBE9F3DE48E795" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FD6C1CF82BDC0043D89E73415E8A81ED" type="Zenodo-Dep">13757735</mods:identifier>
<mods:identifier id="0CEEA84EBA33A1E94766A46536B31C7F" type="ZooBank">3B1CC6CC-4300-4B1E-B257-3A1BBF034E4F</mods:identifier>
</mods:mods>
<treatment id="7B780A3C0911FF9C98D7A48AC2B16392" LSID="urn:lsid:plazi:treatment:7B780A3C0911FF9C98D7A48AC2B16392" httpUri="http://treatment.plazi.org/id/7B780A3C0911FF9C98D7A48AC2B16392" lastPageId="12" lastPageNumber="348" pageId="9" pageNumber="346">
<subSubSection id="BBCBE8A10911FF9998D7A48AC07462D7" box="[151,886,978,1005]" pageId="9" pageNumber="346" type="nomenclature">
<paragraph id="F36EBB2A0911FF9998D7A48AC07462D7" blockId="9.[151,886,978,1005]" box="[151,886,978,1005]" pageId="9" pageNumber="346">
<heading id="A8260C460911FF9998D7A48AC07462D7" bold="true" box="[151,886,978,1005]" fontSize="11" level="1" pageId="9" pageNumber="346" reason="1">
<emphasis id="C1A567380911FF9998D7A48AC07462D7" bold="true" box="[151,886,978,1005]" pageId="9" pageNumber="346">
The original description of 
<taxonomicName id="34D1C0A90911FF9999ACA48AC07462D7" ID-CoL="65K35" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[492,886,978,1005]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380911FF9999ACA48AC1BC62D6" bold="true" box="[492,702,979,1005]" italics="true" pageId="9" pageNumber="346">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0911FF999A85A48AC07462D7" author="Merrem, B." box="[709,886,978,1005]" pageId="9" pageNumber="346" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBCBE8A10911FF9998D7A342C7C466AE" pageId="9" pageNumber="346" type="description">
<paragraph id="F36EBB2A0911FF9998D7A342C2436546" blockId="9.[151,1436,1051,1149]" pageId="9" pageNumber="346">
In contrast to the comparatively short diagnosis in 
<bibRefCitation id="9740C6DB0911FF999A9FA342C08F650E" author="Merrem, B." box="[735,909,1051,1077]" pageId="9" pageNumber="346" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem (1820)</bibRefCitation>
and the short statement mentioned above, the species is described on pp. 165 (right column) and 166 (left column) in 
<bibRefCitation id="9740C6DB0911FF999BE5A366C74D6562" author="Merrem, B." box="[933,1103,1087,1113]" pageId="9" pageNumber="346" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
in more detail [our translation from German]:
</paragraph>
<paragraph id="F36EBB2A0911FF9998B2A3F2C09B663E" blockId="9.[242,1346,1195,1797]" pageId="9" pageNumber="346">
“ 
<taxonomicName id="34D1C0A90911FF9998BEA3F2C2C265FF" box="[254,448,1195,1221]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380911FF9998BEA3F2C2C265FF" box="[254,448,1195,1221]" italics="true" pageId="9" pageNumber="346">Agama gutturosa</emphasis>
</taxonomicName>
. Goitrus 
<taxonomicName id="34D1C0A90911FF999A6BA3F2C18265FE" box="[555,640,1195,1221]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="genus">Agama</taxonomicName>
. Seba Thes. I. tab. 89. fig. 1. 2. Linné and later Lacepède and Daudin list this illustration as 
<taxonomicName id="34D1C0A90911FF999A0DA396C1F965D2" box="[589,763,1231,1257]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380911FF999A0DA396C1F965D2" box="[589,763,1231,1257]" italics="true" pageId="9" pageNumber="346">Agama Calotes</emphasis>
</taxonomicName>
; however, with much confidence I can state that it belongs to another, by no systematicist listed species, as I had both specimens illustrated by Seba [at my disposal] from the collection of the Graf von Borcke and have described [them]. It differs from the 
<taxonomicName id="34D1C0A90911FF999918A262C2AC646E" box="[344,430,1339,1365]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="genus">
<emphasis id="C1A567380911FF999918A262C2AC646E" box="[344,430,1339,1365]" italics="true" pageId="9" pageNumber="346">Calotes</emphasis>
</taxonomicName>
agama by a stronger goitre [gular pouch], by the absence of spines on the occiput, by keeled scales [that are] directed backwards on the plate [top] of the head and by dorsal scales directed downwards and backwards, as well as their considerably sized scales and their keeled ventral scales from the true agama (
<taxonomicName id="34D1C0A90911FF999AC0A2FEC01A64FA" authorityName="Daudin" authorityYear="1802" box="[640,792,1447,1473]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="species" species="colonorum">
<emphasis id="C1A567380911FF999AC0A2FEC01A64FA" box="[640,792,1447,1473]" italics="true" pageId="9" pageNumber="346">A. colonorum</emphasis>
</taxonomicName>
[now 
<taxonomicName id="34D1C0A90911FF999B1FA2FEC70364FB" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[863,1025,1447,1473]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="species" species="agama">
<emphasis id="C1A567380911FF999B1FA2FEC70364FB" box="[863,1025,1447,1473]" italics="true" pageId="9" pageNumber="346">Agama agama</emphasis>
</taxonomicName>
]). A gular fold is not present and the pouch, formed by the throat, extends onto the chest, and is compressed like in the iguana. That it appears thick and convex in Seba’s first figure results from cloth that it was stuffed with. The body is triangular and the back sharp; across it runs a row of soft, tooth-like blades, that are longest on the neck and continuously decrease in size towards the base of the tail and disappear on the tail itself. The colour of 
<specimenCount id="E5D770A30911FF9999A2A102C17C674E" box="[482,638,1627,1653]" count="1" pageId="9" pageNumber="346" type="generic">one specimen</specimenCount>
is light blue, that of the other grey, both underneath whitish. Whitish spots form on both sides and the legs more or less distinctive parts of irregular bands. The length is 19 Zoll, of which 15 Zoll is contributed by the tail. Teeth are moderately large, pyramidal, slightly compressed and differ from other agamids studied by me through a small tooth on each side not far from the root. According to Seba it inhabits 
<collectingCountry id="8BC6FBBA0911FF999B71A1B2C08E663E" box="[817,908,1771,1797]" name="Mexico" pageId="9" pageNumber="346">Mexico</collectingCountry>
.”
</paragraph>
<paragraph id="F36EBB2A0911FF999887A06AC7C466AE" blockId="9.[151,1436,1843,1941]" pageId="9" pageNumber="346">
Not only is the description in 
<bibRefCitation id="9740C6DB0911FF999A6CA06AC1DE6676" author="Merrem, B." box="[556,732,1843,1869]" pageId="9" pageNumber="346" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem (1820)</bibRefCitation>
much shorter, he also changed his taxonomic approach and considered 
<taxonomicName id="34D1C0A90911FF999958A00EC265664A" box="[280,359,1879,1905]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="genus">
<emphasis id="C1A567380911FF999958A00EC265664A" box="[280,359,1879,1905]" italics="true" pageId="9" pageNumber="346">Agama</emphasis>
</taxonomicName>
as a subgenus of 
<taxonomicName id="34D1C0A90911FF999A6DA00EC181664A" box="[557,643,1879,1905]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="genus">
<emphasis id="C1A567380911FF999A6DA00EC181664A" box="[557,643,1879,1905]" italics="true" pageId="9" pageNumber="346">Calotes</emphasis>
</taxonomicName>
. Consequently, he renamed the species as “ 
<emphasis id="C1A567380911FF999C2CA00EC68A664B" box="[1132,1416,1879,1905]" italics="true" pageId="9" pageNumber="346">
<taxonomicName id="34D1C0A90911FF999C2CA00EC7C0664A" box="[1132,1218,1879,1905]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="genus">Calotes</taxonomicName>
<taxonomicName id="34D1C0A90911FF999C88A00EC68A664B" box="[1224,1416,1879,1905]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="9" pageNumber="346" phylum="Chordata" rank="species" species="gutturosa">Agama gutturosa</taxonomicName>
</emphasis>
”. As 
<bibRefCitation id="9740C6DB0911FF9998FDA022C26B66AE" author="Merrem, B." box="[189,361,1915,1941]" pageId="9" pageNumber="346" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
precedes 
<bibRefCitation id="9740C6DB0911FF999999A022C18766AE" author="Merrem, B." box="[473,645,1915,1941]" pageId="9" pageNumber="346" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem (1820)</bibRefCitation>
this action does not pose a nomenclatural problem.
</paragraph>
</subSubSection>
<subSubSection id="BBCBE8A10912FF9B98D7A7CEC77B66AE" lastPageId="11" lastPageNumber="348" pageId="10" pageNumber="347" type="discussion">
<paragraph id="F36EBB2A0912FF9A98D7A7CEC100618A" blockId="10.[151,514,151,177]" box="[151,514,151,177]" pageId="10" pageNumber="347">
<heading id="A8260C460912FF9A98D7A7CEC100618A" bold="true" box="[151,514,151,177]" fontSize="11" level="1" pageId="10" pageNumber="347" reason="1">
<emphasis id="C1A567380912FF9A98D7A7CEC100618A" bold="true" box="[151,514,151,177]" pageId="10" pageNumber="347">Nomenclatural consequences</emphasis>
</heading>
</paragraph>
<paragraph id="F36EBB2A0912FF9A98D7A7B9C6536026" blockId="10.[151,1437,223,754]" pageId="10" pageNumber="347">
In an attempt to stabilise the nomenclature surrounding 
<taxonomicName id="34D1C0A90912FF9A9B50A786C09861C2" box="[784,922,223,249]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9B50A786C09861C2" box="[784,922,223,249]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90912FF9A9B92A786C62B61C2" authority=", Figueroa (2021)" authorityName="Figueroa" authorityYear="2021" box="[978,1321,223,250]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9B92A786C76661C2" box="[978,1124,223,249]" italics="true" pageId="10" pageNumber="347">A. cristatella</emphasis>
, 
<bibRefCitation id="9740C6DB0912FF9A9C33A786C62B61C2" author="Figueroa, A." box="[1139,1321,223,249]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
</taxonomicName>
discussed the matter on the incorrect assumption that 
<bibRefCitation id="9740C6DB0912FF9A9A3DA65DC02B6026" author="Merrem, B." box="[637,809,259,286]" pageId="10" pageNumber="347" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem (1820)</bibRefCitation>
and 
<bibRefCitation id="9740C6DB0912FF9A9B20A65AC0EB6026" author="Kuhl, H." box="[864,1001,259,285]" pageId="10" pageNumber="347" refId="ref13590" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Hermannschen Buchhandlung, Frankfurt, (ii) + 151 + (1) pp., 7 pls." type="journal volume" year="1820">Kuhl (1820)</bibRefCitation>
were published simultaneously.
</paragraph>
<paragraph id="F36EBB2A0912FF9A9887A67EC17A6322" blockId="10.[151,1437,223,754]" pageId="10" pageNumber="347">
First, it must be acknowledged that 
<bibRefCitation id="9740C6DB0912FF9A9A16A671C000607A" author="Merrem, B." box="[598,770,295,322]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
based his description on only 
<specimenCount id="E5D770A30912FF9A9C15A671C7FC607A" box="[1109,1278,295,322]" count="2" pageId="10" pageNumber="347" type="generic">two specimens</specimenCount>
, namely those that came from Seba’s collection. This contrasts with his 1820 description where he listed further illustrations and descriptions that would typically constitute additional 
<typeStatus id="2C6A05880912FF9A9AB7A629C05B60B1" box="[759,857,368,394]" pageId="10" pageNumber="347" type="syntype">syntypes</typeStatus>
and were treated as such by 
<bibRefCitation id="9740C6DB0912FF9A9CDAA636C65060B2" author="Figueroa, A." box="[1178,1362,367,393]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
. In his analysis, 
<bibRefCitation id="9740C6DB0912FF9A98BEA6CAC2B06096" author="Figueroa, A." box="[254,434,403,429]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
designated a 
<typeStatus id="2C6A05880912FF9A9A08A6CAC1AD6096" box="[584,687,403,429]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
from what he considered the 
<typeStatus id="2C6A05880912FF9A9BB5A6CDC74E6095" box="[1013,1100,404,430]" pageId="10" pageNumber="347" type="syntype">syntype</typeStatus>
series of 
<taxonomicName id="34D1C0A90912FF9A9CF5A6CAC63F6096" box="[1205,1341,403,429]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9CF5A6CAC63F6096" box="[1205,1341,403,429]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
, namely Seba pl. 89 fig. 1, 
<taxonomicName id="34D1C0A90912FF9A9924A6E1C10860EA" baseAuthorityName="Latreille" baseAuthorityYear="1802" box="[356,522,439,465]" class="Squamata" family="Iguanidae" genus="Iguana" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A9924A6E1C10860EA" box="[356,522,439,465]" italics="true" pageId="10" pageNumber="347">Iguana calotes</emphasis>
</taxonomicName>
(
<bibRefCitation id="9740C6DB0912FF9A9A58A6EEC1F660EA" author="Latreille, P. A." box="[536,756,439,465]" pageId="10" pageNumber="347" pagination="1 - 280" refId="ref13627" refString="Latreille, P. A. (1802) N. S. Sonnini de Manoncourt and P. A. Latreille. Histoire Naturelle des Reptiles, avec Figures Dessinnees d'apres Nature. Vol. 1. Deterville, Paris, i - xx + 1 - 280." type="book chapter" year="1802">Latreille 1802: 260</bibRefCitation>
, fig. 1 on plate facing p. 260 [now 
<taxonomicName id="34D1C0A90912FF9A9C3BA6EEC68060EA" authorityName=", Kaup" authorityYear="1827" box="[1147,1410,439,465]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9C3BA6EEC68060EA" box="[1147,1410,439,465]" italics="true" pageId="10" pageNumber="347">Bronchocela cristatella</emphasis>
</taxonomicName>
]); Seba pl. 89 fig. 2 [considered by 
<bibRefCitation id="9740C6DB0912FF9A9A51A682C1C560CE" author="Figueroa, A." box="[529,711,475,501]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
to represent 
<taxonomicName id="34D1C0A90912FF9A9B1CA682C70A60CE" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[860,1032,475,501]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A9B1CA682C70A60CE" box="[860,1032,475,501]" italics="true" pageId="10" pageNumber="347">Calotes calotes</emphasis>
</taxonomicName>
] and 
<taxonomicName id="34D1C0A90912FF9A9C0AA682C7F260CE" box="[1098,1264,475,501]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A9C0AA682C7F260CE" box="[1098,1264,475,501]" italics="true" pageId="10" pageNumber="347">Agama calotes</emphasis>
</taxonomicName>
(
<bibRefCitation id="9740C6DB0912FF9A9D40A682C69A60CE" author="Daudin, F. M." box="[1280,1432,475,501]" pageId="10" pageNumber="347" pagination="29 - 45" refId="ref12247" refString="Daudin, F. M. (1802) Histoire naturelle, generale et particuliere des reptiles; Ouvrage faisant suite aux Oeuvres de Leclerc de Buffon, et partie du cours complet d'histoire naturelle redige par C. S. Sonnini, membre de plusieurs societes savantes. Tome Troisieme. F. Dufart, Paris, 452 pp., pls. 29 - 45. https: // doi. org / 10.5962 / bhl. title. 70894" type="book chapter" year="1802">Daudin 1802</bibRefCitation>
: vol. III, 361, pl. 43 [now 
<taxonomicName id="34D1C0A90912FF9A99F2A6A6C15C6322" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[434,606,511,537]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A99F2A6A6C15C6322" box="[434,606,511,537]" italics="true" pageId="10" pageNumber="347">Calotes calotes</emphasis>
</taxonomicName>
]).
</paragraph>
<paragraph id="F36EBB2A0912FF9A9887A57AC64063C9" blockId="10.[151,1437,223,754]" pageId="10" pageNumber="347">
<bibRefCitation id="9740C6DB0912FF9A9887A57AC27F6306" author="Figueroa, A." box="[199,381,547,573]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
designated the same figure (
<bibRefCitation id="9740C6DB0912FF9A9AFFA57AC04B6306" author="Seba, A." box="[703,841,547,573]" pageId="10" pageNumber="347" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734)</bibRefCitation>
pl. 89 fig. 1) as 
<typeStatus id="2C6A05880912FF9A9C42A57AC7766306" box="[1026,1140,547,573]" pageId="10" pageNumber="347" type="lectotype">lectotypes</typeStatus>
for both 
<taxonomicName id="34D1C0A90912FF9A9C9CA57AC66F6306" authorityName=", Kaup" authorityYear="1827" box="[1244,1389,547,573]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9C9CA57AC66F6306" box="[1244,1389,547,573]" italics="true" pageId="10" pageNumber="347">A. cristatella</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90912FF9A98D7A51EC223635A" box="[151,289,583,609]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A98D7A51EC223635A" box="[151,289,583,609]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
. This 
<typeStatus id="2C6A05880912FF9A9929A51EC2D2635A" box="[361,464,583,609]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
designation complies with Art. 74 of the 
<emphasis id="C1A567380912FF9A9BE5A51EC0E2635A" box="[933,992,583,609]" italics="true" pageId="10" pageNumber="347">Code</emphasis>
and cannot normally be undone as Art. 74.1.1 states that “no later designation of a 
<typeStatus id="2C6A05880912FF9A9AC1A532C1EA63BE" box="[641,744,619,645]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
has any validity”. But our discovery of an earlier publication puts his actions into question. Art. 72.1.1. of the 
<emphasis id="C1A567380912FF9A9AF5A5D6C1F26392" box="[693,752,655,681]" italics="true" pageId="10" pageNumber="347">Code</emphasis>
defines the type series as “all specimens on which the author established a nominal species-group taxon”.As 
<bibRefCitation id="9740C6DB0912FF9A9AE5A5EAC05B63F6" author="Figueroa, A." box="[677,857,691,717]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
was referring to “ 
<taxonomicName id="34D1C0A90912FF9A9C5DA5EAC77163F6" box="[1053,1139,691,717]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9C5DA5EAC77163F6" box="[1053,1139,691,717]" italics="true" pageId="10" pageNumber="347">Calotes</emphasis>
</taxonomicName>
<taxonomicName id="34D1C0A90912FF9A9C37A5EAC3D363CA" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9C37A5EAC63463F6" box="[1143,1334,691,717]" italics="true" pageId="10" pageNumber="347">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9D7BA5EDC3D363CA" author="Merrem, B." pageId="10" pageNumber="347" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem, 1820</bibRefCitation>
</taxonomicName>
” and not to the original publication, i.e., 
<taxonomicName id="34D1C0A90912FF9A9ADDA58EC70A63CA" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[669,1032,727,754]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9ADDA58EC05F63CA" box="[669,861,727,753]" italics="true" pageId="10" pageNumber="347">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9B25A581C70A63CA" author="Merrem, B." box="[869,1032,727,754]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
, this has two consequences:
</paragraph>
<paragraph id="F36EBB2A0912FF9A98B2A446C29962F2" blockId="10.[242,1345,799,969]" pageId="10" pageNumber="347">
1. 
<bibRefCitation id="9740C6DB0912FF9A994DA446C2CF6202" author="Figueroa, A." box="[269,461,799,825]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa’s (2021)</bibRefCitation>
<typeStatus id="2C6A05880912FF9A9993A446C1346202" box="[467,566,799,825]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
designation for 
<taxonomicName id="34D1C0A90912FF9A9AA5A446C06B6202" box="[741,873,799,825]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9AA5A446C06B6202" box="[741,873,799,825]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="9740C6DB0912FF9A9B37A446C0EC6202" author="Seba, A." box="[887,1006,799,825]" pageId="10" pageNumber="347" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba 1734</bibRefCitation>
: pl. 89 fig. 1) becomes invalid. 2. The illustrations of 
<taxonomicName id="34D1C0A90912FF9A99A8A41AC18F6266" box="[488,653,835,861]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A99A8A41AC18F6266" box="[488,653,835,861]" italics="true" pageId="10" pageNumber="347">Agama calotes</emphasis>
</taxonomicName>
(
<bibRefCitation id="9740C6DB0912FF9A9ADBA41AC0306266" author="Daudin, F. M." box="[667,818,835,861]" pageId="10" pageNumber="347" pagination="29 - 45" refId="ref12247" refString="Daudin, F. M. (1802) Histoire naturelle, generale et particuliere des reptiles; Ouvrage faisant suite aux Oeuvres de Leclerc de Buffon, et partie du cours complet d'histoire naturelle redige par C. S. Sonnini, membre de plusieurs societes savantes. Tome Troisieme. F. Dufart, Paris, 452 pp., pls. 29 - 45. https: // doi. org / 10.5962 / bhl. title. 70894" type="book chapter" year="1802">Daudin 1802</bibRefCitation>
: vol. III, 361, pl. 43) [now 
<taxonomicName id="34D1C0A90912FF9A9C1EA41AC60B6266" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1118,1289,835,861]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A9C1EA41AC60B6266" box="[1118,1289,835,861]" italics="true" pageId="10" pageNumber="347">Calotes calotes</emphasis>
</taxonomicName>
] and 
<taxonomicName id="34D1C0A90912FF9A98B2A431C18E62BA" authority="(Latreille 1802: 260)" baseAuthorityName="Latreille" baseAuthorityPageNumber="260" baseAuthorityYear="1802" box="[242,652,871,897]" class="Squamata" family="Iguanidae" genus="Iguana" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380912FF9A98B2A431C29A62BA" box="[242,408,871,897]" italics="true" pageId="10" pageNumber="347">Iguana calotes</emphasis>
(
<bibRefCitation id="9740C6DB0912FF9A99E8A43EC18762BA" author="Latreille, P. A." box="[424,645,871,897]" pageId="10" pageNumber="347" pagination="1 - 280" refId="ref13627" refString="Latreille, P. A. (1802) N. S. Sonnini de Manoncourt and P. A. Latreille. Histoire Naturelle des Reptiles, avec Figures Dessinnees d'apres Nature. Vol. 1. Deterville, Paris, i - xx + 1 - 280." type="book chapter" year="1802">Latreille 1802: 260</bibRefCitation>
)
</taxonomicName>
[now 
<taxonomicName id="34D1C0A90912FF9A9A96A43EC0DC62BA" authorityName=", Kaup" authorityYear="1827" box="[726,990,871,897]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9A96A43EC0DC62BA" box="[726,990,871,897]" italics="true" pageId="10" pageNumber="347">Bronchocela cristatella</emphasis>
</taxonomicName>
] were not explicitly mentioned by 
<bibRefCitation id="9740C6DB0912FF9A9955A4D5C2C3629E" author="Merrem, B." box="[277,449,907,934]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
and therefore do not qualify as 
<typeStatus id="2C6A05880912FF9A9B67A4D5C08F629D" box="[807,909,908,934]" pageId="10" pageNumber="347" type="syntype">syntypes</typeStatus>
. Consequently, they lose their status as 
<typeStatus id="2C6A05880912FF9A98B2A4F6C29562F2" box="[242,407,943,969]" pageId="10" pageNumber="347" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
<paragraph id="F36EBB2A0912FF9A9887A4A1C7FC65FD" blockId="10.[151,1437,1015,1978]" pageId="10" pageNumber="347">
In the earlier work, 
<bibRefCitation id="9740C6DB0912FF9A99DBA4A1C140652A" author="Merrem, B." box="[411,578,1015,1042]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
also listed “Seba Thes.[aurus] I tab. 89 fig. 1” under 
<taxonomicName id="34D1C0A90912FF9A9C3BA4AEC634652A" box="[1147,1334,1015,1041]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9C3BA4AEC634652A" box="[1147,1334,1015,1041]" italics="true" pageId="10" pageNumber="347">Agama gutturosa</emphasis>
</taxonomicName>
such that it qualifies as a 
<typeStatus id="2C6A05880912FF9A9904A345C29E650D" box="[324,412,1052,1078]" pageId="10" pageNumber="347" type="syntype">syntype</typeStatus>
, but he clearly described the specimen in figure 
<quantity id="342916CF0912FF9A9BEEA342C0DA650E" box="[942,984,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="347" unit="in" value="2.0">2 in</quantity>
more detail. Already, Seba’s illustrations are unambiguously showing different species. The specimen in fig. 1 does not have a dorsal crest (see also 
<bibRefCitation id="9740C6DB0912FF9A9D5BA319C6966562" author="Kaup, J." box="[1307,1428,1087,1114]" pageId="10" pageNumber="347" pagination="610 - 625" refId="ref13494" refString="Kaup, J. (1827) Zoologische Monographien. Isis von Oken, 20 (6 - 7), 610 - 625." type="journal article" year="1827">Kaup 1827</bibRefCitation>
) and represents 
<taxonomicName id="34D1C0A90912FF9A997CA33DC2C86546" authorityName=", Kaup" authorityYear="1827" box="[316,458,1123,1149]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A997CA33DC2C86546" box="[316,458,1123,1149]" italics="true" pageId="10" pageNumber="347">B. cristatella</emphasis>
</taxonomicName>
; whereas the specimen in fig. 2 has a well-developed dorsal crest and does not represent 
<taxonomicName id="34D1C0A90912FF9A98D7A3D1C227659A" authorityName=", Kaup" authorityYear="1827" box="[151,293,1159,1185]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A98D7A3D1C227659A" box="[151,293,1159,1185]" italics="true" pageId="10" pageNumber="347">B. cristatella</emphasis>
</taxonomicName>
. 
<bibRefCitation id="9740C6DB0912FF9A9973A3D1C2D9659A" author="Merrem, B." box="[307,475,1159,1186]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
stated that he had both of Seba’s specimens on hand and gives the colouration for both. Therefore, although they may represent two different taxa, both specimens have the status of 
<typeStatus id="2C6A05880912FF9A9CD7A3F5C7F965FD" box="[1175,1275,1196,1222]" pageId="10" pageNumber="347" type="syntype">syntypes</typeStatus>
.
</paragraph>
<paragraph id="F36EBB2A0912FF9A9887A389C21864FA" blockId="10.[151,1437,1015,1978]" pageId="10" pageNumber="347">
To clarify the situation and to stabilise nomenclature, we hereby designate the illustration from 
<bibRefCitation id="9740C6DB0912FF9A9D54A396C69A65D2" author="Seba, A." box="[1300,1432,1231,1257]" pageId="10" pageNumber="347" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734</bibRefCitation>
, plate 89 figure 2) as the 
<typeStatus id="2C6A05880912FF9A99E5A3AAC10E6436" box="[421,524,1267,1293]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="34D1C0A90912FF9A9A70A3AAC0946436" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[560,918,1267,1294]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9A70A3AAC1F26436" box="[560,752,1267,1293]" italics="true" pageId="10" pageNumber="347">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9AB7A3ADC0946436" author="Merrem, B." box="[759,918,1267,1294]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
according to Art. 74.1 of the 
<emphasis id="C1A567380912FF9A9C9EA3AAC61B6436" box="[1246,1305,1267,1293]" italics="true" pageId="10" pageNumber="347">Code</emphasis>
(see 
<figureCitation id="6BEAA7AF0912FF9A9D13A3AAC3A6640A" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-19@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The original illustrations from Seba (1734). Upper panel: Lectotype of Agama cristatella Kuhl, 1820. Lower panel: Lectotype of Agama gutturosa Merrem, 1819." figureDoi="http://doi.org/10.5281/zenodo.13757737" httpUri="https://zenodo.org/record/13757737/files/figure.png" pageId="10" pageNumber="347">Figure 1</figureCitation>
). Through our 
<typeStatus id="2C6A05880912FF9A9911A24EC2BA640A" box="[337,440,1303,1329]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
designation for 
<taxonomicName id="34D1C0A90912FF9A9A30A24EC0DE640A" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[624,988,1303,1330]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9A30A24EC033640A" box="[624,817,1303,1329]" italics="true" pageId="10" pageNumber="347">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9B78A241C0DE640A" author="Merrem, B." box="[824,988,1303,1330]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
, the specimen in 
<bibRefCitation id="9740C6DB0912FF9A9CE0A24EC639640A" author="Seba, A." box="[1184,1339,1303,1329]" pageId="10" pageNumber="347" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba’s (1734)</bibRefCitation>
plate 89 figure 1 becomes a 
<typeStatus id="2C6A05880912FF9A9938A262C10D646E" box="[376,527,1339,1365]" pageId="10" pageNumber="347" type="paralectotype">paralectotype</typeStatus>
and has no longer any name-bearing function. Consequently, 
<taxonomicName id="34D1C0A90912FF9A9C93A262C2196442" authority="Kuhl, 1820" authorityName="Kuhl" authorityYear="1820" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9C93A262C69F646E" box="[1235,1437,1339,1365]" italics="true" pageId="10" pageNumber="347">Agama cristatella</emphasis>
<bibRefCitation id="9740C6DB0912FF9A98D7A206C2196442" author="Kuhl, H." box="[151,283,1375,1401]" pageId="10" pageNumber="347" refId="ref13590" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Hermannschen Buchhandlung, Frankfurt, (ii) + 151 + (1) pp., 7 pls." type="journal volume" year="1820">Kuhl, 1820</bibRefCitation>
</taxonomicName>
, whose 
<typeStatus id="2C6A05880912FF9A9939A206C2E26442" box="[377,480,1375,1401]" pageId="10" pageNumber="347" type="lectotype">lectotype</typeStatus>
is the specimen in 
<bibRefCitation id="9740C6DB0912FF9A9A83A206C0626442" author="Seba, A." box="[707,864,1375,1401]" pageId="10" pageNumber="347" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba’s (1734)</bibRefCitation>
plate 89 figure 1 (
<bibRefCitation id="9740C6DB0912FF9A9C7AA206C7E76442" author="Figueroa, A." box="[1082,1253,1375,1401]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa 2021</bibRefCitation>
), is not a junior synonym of 
<taxonomicName id="34D1C0A90912FF9A9962A2DAC18B64A6" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[290,649,1411,1438]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9962A2DAC2E064A6" box="[290,482,1411,1437]" italics="true" pageId="10" pageNumber="347">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0912FF9A99A9A2DDC18B64A6" author="Merrem, B." box="[489,649,1411,1438]" pageId="10" pageNumber="347" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
and the currently accepted name 
<taxonomicName id="34D1C0A90912FF9A9BBFA2DAC69E64A6" authority="(Kuhl, 1820)" baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[1023,1436,1411,1437]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9BBFA2DAC60464A6" box="[1023,1286,1411,1437]" italics="true" pageId="10" pageNumber="347">Bronchocela cristatella</emphasis>
(
<bibRefCitation id="9740C6DB0912FF9A9D54A2DAC69664A6" author="Kuhl, H." box="[1300,1428,1411,1437]" pageId="10" pageNumber="347" refId="ref13590" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Hermannschen Buchhandlung, Frankfurt, (ii) + 151 + (1) pp., 7 pls." type="journal volume" year="1820">Kuhl, 1820</bibRefCitation>
)
</taxonomicName>
can prevail.
</paragraph>
<paragraph id="F36EBB2A0912FF9A9887A292C1666682" blockId="10.[151,1437,1015,1978]" pageId="10" pageNumber="347">
<bibRefCitation id="9740C6DB0912FF9A9887A292C28264DE" author="Figueroa, A." box="[199,384,1483,1509]" pageId="10" pageNumber="347" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
also relied on 
<bibRefCitation id="9740C6DB0912FF9A9A74A292C00064DE" author="Fitzinger, L." box="[564,770,1483,1509]" pageId="10" pageNumber="347" refId="ref12976" refString="Fitzinger, L. (1843) Systema Reptilium. Fasciculus Primus. Amblyglossae. Braumuller et Seidel, Vindobonae [Vienna], 106 + (vi) + 3 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger’s (1843)</bibRefCitation>
<typeStatus id="2C6A05880912FF9A9B4DA295C03F64DD" box="[781,829,1484,1510]" pageId="10" pageNumber="347">type</typeStatus>
species designation, who he considered as the First Reviser, and designated 
<taxonomicName id="34D1C0A90912FF9A99E4A2B6C1D36732" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" box="[420,721,1519,1546]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A99E4A2B6C12E6732" box="[420,556,1519,1545]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9A72A2A9C1D36732" author="Merrem, B." box="[562,721,1519,1546]" pageId="10" pageNumber="347" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem, 1820</bibRefCitation>
</taxonomicName>
as the 
<typeStatus id="2C6A05880912FF9A9B5BA2A9C0496731" box="[795,843,1520,1546]" pageId="10" pageNumber="347">type</typeStatus>
species of 
<taxonomicName id="34D1C0A90912FF9A9B85A2B6C7DF6732" authority="Kaup, 1827" authorityName="Kaup" authorityYear="1827" box="[965,1245,1519,1546]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9B85A2B6C7516732" box="[965,1107,1519,1545]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9C19A2A9C7DF6732" author="Kaup, J." box="[1113,1245,1519,1546]" pageId="10" pageNumber="347" pagination="610 - 625" refId="ref13494" refString="Kaup, J. (1827) Zoologische Monographien. Isis von Oken, 20 (6 - 7), 610 - 625." type="journal article" year="1827">Kaup, 1827</bibRefCitation>
</taxonomicName>
. 
<bibRefCitation id="9740C6DB0912FF9A9CA7A2B6C69E6732" author="Fitzinger, L." box="[1255,1436,1519,1545]" pageId="10" pageNumber="347" refId="ref12976" refString="Fitzinger, L. (1843) Systema Reptilium. Fasciculus Primus. Amblyglossae. Braumuller et Seidel, Vindobonae [Vienna], 106 + (vi) + 3 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger (1843)</bibRefCitation>
divided the genus into three subgenera (
<taxonomicName id="34D1C0A90912FF9A9A25A14AC1F16716" authorityName="Kaup" authorityYear="1827" box="[613,755,1555,1581]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9A25A14AC1F16716" box="[613,755,1555,1581]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
</taxonomicName>
, 
<taxonomicName id="34D1C0A90912FF9A9B43A14AC0846716" box="[771,902,1555,1581]" pageId="10" pageNumber="347" rank="subGenus" subGenus="Lophodeira">
<emphasis id="C1A567380912FF9A9B43A14AC0846716" box="[771,902,1555,1581]" italics="true" pageId="10" pageNumber="347">Lophodeira</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90912FF9A9B81A14AC7616716" box="[961,1123,1555,1581]" class="Squamata" family="Agamidae" genus="Pseudocalotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9B81A14AC7616716" box="[961,1123,1555,1581]" italics="true" pageId="10" pageNumber="347">Pseudocalotes</emphasis>
</taxonomicName>
). He included 
<taxonomicName id="34D1C0A90912FF9A9D50A14AC69E6716" box="[1296,1436,1555,1581]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9D50A14AC69E6716" box="[1296,1436,1555,1581]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90912FF9A9886A16EC25C676A" box="[198,350,1591,1617]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="intermedia">
<emphasis id="C1A567380912FF9A9886A16EC25C676A" box="[198,350,1591,1617]" italics="true" pageId="10" pageNumber="347">A. intermedia</emphasis>
</taxonomicName>
in the subgenus 
<taxonomicName id="34D1C0A90912FF9A9A58A16EC1A4676A" box="[536,678,1591,1617]" pageId="10" pageNumber="347" rank="subGenus" subGenus="Bronchocela">
<emphasis id="C1A567380912FF9A9A58A16EC1A4676A" box="[536,678,1591,1617]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
</taxonomicName>
by redefining the original genus diagnosis of 
<taxonomicName id="34D1C0A90912FF9A9CE4A16EC630676A" authorityName="Kaup" authorityYear="1827" box="[1188,1330,1591,1617]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9CE4A16EC630676A" box="[1188,1330,1591,1617]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
</taxonomicName>
such that 
<taxonomicName id="34D1C0A90912FF9A98D7A102C22B674E" baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[151,297,1627,1653]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A98D7A102C22B674E" box="[151,297,1627,1653]" italics="true" pageId="10" pageNumber="347">A. cristatella</emphasis>
</taxonomicName>
was excluded. The latter was placed in the newly defined subgenus 
<taxonomicName id="34D1C0A90912FF9A9C75A102C7BA674E" box="[1077,1208,1627,1653]" pageId="10" pageNumber="347" rank="subGenus" subGenus="Lophodeira">
<emphasis id="C1A567380912FF9A9C75A102C7BA674E" box="[1077,1208,1627,1653]" italics="true" pageId="10" pageNumber="347">Lophodeira</emphasis>
</taxonomicName>
. Although 
<bibRefCitation id="9740C6DB0912FF9A9D77A102C20C67A1" author="Fitzinger, L." pageId="10" pageNumber="347" refId="ref12976" refString="Fitzinger, L. (1843) Systema Reptilium. Fasciculus Primus. Amblyglossae. Braumuller et Seidel, Vindobonae [Vienna], 106 + (vi) + 3 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger (1843: 15)</bibRefCitation>
stated that 
<taxonomicName id="34D1C0A90912FF9A99D1A126C11967A2" box="[401,539,1663,1689]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A99D1A126C11967A2" box="[401,539,1663,1689]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
is the “ 
<typeStatus id="2C6A05880912FF9A9A33A126C1BE67A2" box="[627,700,1663,1689]" pageId="10" pageNumber="347">Typus</typeStatus>
” of 
<taxonomicName id="34D1C0A90912FF9A9AADA126C07967A2" authorityName="Kaup" authorityYear="1827" box="[749,891,1663,1689]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9AADA126C07967A2" box="[749,891,1663,1689]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
</taxonomicName>
, this nomenclatural action is questionable if not invalid. According to Art. 69.4 of the 
<emphasis id="C1A567380912FF9A9A01A1FAC17E6786" box="[577,636,1699,1725]" italics="true" pageId="10" pageNumber="347">Code</emphasis>
, “Fixation by elimination” is excluded, i.e., “elimination of all but one of the originally included nominal species from a nominal genus or subgenus does not in itself constitute 
<typeStatus id="2C6A05880912FF9A9D2CA191C69E67D9" box="[1388,1436,1736,1762]" pageId="10" pageNumber="347">type</typeStatus>
fixation.” Note that 
<taxonomicName id="34D1C0A90912FF9A993EA1B2C11A663E" box="[382,536,1771,1797]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="intermedia">
<emphasis id="C1A567380912FF9A993EA1B2C11A663E" box="[382,536,1771,1797]" italics="true" pageId="10" pageNumber="347">A. intermedia</emphasis>
</taxonomicName>
was not included when 
<bibRefCitation id="9740C6DB0912FF9A9B73A1B5C0C1663E" author="Kaup, J." box="[819,963,1771,1798]" pageId="10" pageNumber="347" pagination="610 - 625" refId="ref13494" refString="Kaup, J. (1827) Zoologische Monographien. Isis von Oken, 20 (6 - 7), 610 - 625." type="journal article" year="1827">Kaup (1827)</bibRefCitation>
erected the genus and therefore does not qualify as a 
<typeStatus id="2C6A05880912FF9A9961A049C2536611" box="[289,337,1808,1834]" pageId="10" pageNumber="347">type</typeStatus>
species. 
<bibRefCitation id="9740C6DB0912FF9A99F8A056C1726612" author="Fitzinger, L." box="[440,624,1807,1833]" pageId="10" pageNumber="347" refId="ref12976" refString="Fitzinger, L. (1843) Systema Reptilium. Fasciculus Primus. Amblyglossae. Braumuller et Seidel, Vindobonae [Vienna], 106 + (vi) + 3 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger (1843)</bibRefCitation>
eliminated Kaup’s 
<taxonomicName id="34D1C0A90912FF9A9B0CA056C7566612" baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[844,1108,1807,1833]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9B0CA056C7566612" box="[844,1108,1807,1833]" italics="true" pageId="10" pageNumber="347">Bronchocela cristatella</emphasis>
</taxonomicName>
from that genus and was left with 
<taxonomicName id="34D1C0A90912FF9A9893A06DC25C6676" box="[211,350,1844,1869]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9893A06DC25C6676" box="[211,350,1844,1869]" italics="true" pageId="10" pageNumber="347">B. gutturosa</emphasis>
</taxonomicName>
as the 
<typeStatus id="2C6A05880912FF9A99F5A06DC2E76675" box="[437,485,1844,1870]" pageId="10" pageNumber="347">type</typeStatus>
species. Technically, Fitzinger’s designation of 
<taxonomicName id="34D1C0A90912FF9A9C4EA06AC7986676" box="[1038,1178,1843,1869]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380912FF9A9C4EA06AC7986676" box="[1038,1178,1843,1869]" italics="true" pageId="10" pageNumber="347">A. gutturosa</emphasis>
</taxonomicName>
as the 
<typeStatus id="2C6A05880912FF9A9CB1A06DC6236675" box="[1265,1313,1844,1870]" pageId="10" pageNumber="347">type</typeStatus>
species of 
<taxonomicName id="34D1C0A90912FF9A98D7A00EC227664A" authorityName="Kaup" authorityYear="1827" box="[151,293,1879,1905]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A98D7A00EC227664A" box="[151,293,1879,1905]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
</taxonomicName>
, by elimination, must be seen as invalid. Consequently, the next available 
<typeStatus id="2C6A05880912FF9A9C21A001C7936649" box="[1121,1169,1880,1906]" pageId="10" pageNumber="347">type</typeStatus>
species designation has to be taken into account. Therefore, we follow 
<bibRefCitation id="9740C6DB0912FF9A9AEDA022C04066AD" author="Smith, M. A." box="[685,834,1915,1942]" pageId="10" pageNumber="347" refId="ref14553" refString="Smith, M. A. (1935) The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. Vol. II. Sauria. Taylor and Francis, London, xiii + 440 + (1) pp., 1 pl., 2 maps." type="book" year="1935">Smith (1935)</bibRefCitation>
who designated 
<taxonomicName id="34D1C0A90912FF9A9C43A022C65166AE" authority="Kuhl, 1820" authorityName="Kuhl" authorityYear="1820" box="[1027,1363,1915,1941]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380912FF9A9C43A022C7CE66AE" box="[1027,1228,1915,1941]" italics="true" pageId="10" pageNumber="347">Agama cristatella</emphasis>
<bibRefCitation id="9740C6DB0912FF9A9C94A022C65166AE" author="Kuhl, H." box="[1236,1363,1915,1941]" pageId="10" pageNumber="347" refId="ref13590" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Hermannschen Buchhandlung, Frankfurt, (ii) + 151 + (1) pp., 7 pls." type="journal volume" year="1820">Kuhl, 1820</bibRefCitation>
</taxonomicName>
as the 
<typeStatus id="2C6A05880912FF9A98D7A0F9C3C56681" box="[151,199,1952,1978]" pageId="10" pageNumber="347">type</typeStatus>
species of 
<taxonomicName id="34D1C0A90912FF9A9905A0C6C15D6682" authority="Kaup, 1827" authorityName="Kaup" authorityYear="1827" box="[325,607,1951,1978]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="10" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="C1A567380912FF9A9905A0C6C2D16682" box="[325,467,1951,1977]" italics="true" pageId="10" pageNumber="347">Bronchocela</emphasis>
<bibRefCitation id="9740C6DB0912FF9A999AA0F9C15D6682" author="Kaup, J." box="[474,607,1951,1978]" pageId="10" pageNumber="347" pagination="610 - 625" refId="ref13494" refString="Kaup, J. (1827) Zoologische Monographien. Isis von Oken, 20 (6 - 7), 610 - 625." type="journal article" year="1827">Kaup, 1827</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="F36EBB2A0913FF9B98D7A7CEC15B618A" blockId="11.[151,601,151,177]" box="[151,601,151,177]" pageId="11" pageNumber="348">
<heading id="A8260C460913FF9B98D7A7CEC15B618A" bold="true" box="[151,601,151,177]" fontSize="11" level="1" pageId="11" pageNumber="348" reason="1">
<emphasis id="C1A567380913FF9B98D7A7CEC15B618A" bold="true" box="[151,601,151,177]" pageId="11" pageNumber="348">The identity of Merrem’s specimens</emphasis>
</heading>
</paragraph>
<paragraph id="F36EBB2A0913FF9B98D7A786C09460CE" blockId="11.[151,1437,223,2014]" pageId="11" pageNumber="348">
A very important statement in 
<bibRefCitation id="9740C6DB0913FF9B99B0A7B9C19F61C2" author="Merrem, B." box="[496,669,223,250]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
is that he examined the specimens depicted in 
<bibRefCitation id="9740C6DB0913FF9B9CF2A786C63861C2" author="Seba, A." box="[1202,1338,223,249]" pageId="11" pageNumber="348" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734)</bibRefCitation>
and that they were present in the von Borcke collection. A large part of von Borcke’s collection came to ZMB in 1817 and contained 
<specimenCount id="E5D770A30913FF9B994FA671C2BA607A" box="[271,440,295,322]" count="2" pageId="11" pageNumber="348" type="generic">two specimens</specimenCount>
identified later as 
<taxonomicName id="34D1C0A90913FF9B9AD6A67EC0A2607A" baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[662,928,295,321]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380913FF9B9AD6A67EC0A2607A" box="[662,928,295,321]" italics="true" pageId="11" pageNumber="348">Bronchocela cristatella</emphasis>
</taxonomicName>
(ZMB 680) and 
<taxonomicName id="34D1C0A90913FF9B9C2CA67EC64E607A" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[1132,1356,295,321]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B9C2CA67EC64E607A" box="[1132,1356,295,321]" italics="true" pageId="11" pageNumber="348">Bronchocela jubata</emphasis>
</taxonomicName>
(ZMB 686), respectively. In their type catalogue, 
<bibRefCitation id="9740C6DB0913FF9B9ACEA615C07E605E" author="Denzer, W. &amp; Gunther, R. &amp; Manthey, U." box="[654,892,331,358]" pageId="11" pageNumber="348" pagination="309 - 332" refId="ref12440" refString="Denzer, W., Gunther, R. &amp; Manthey, U. (1997) Kommentierter Typenkatalog der Agamen (Reptilia: Squamata: Agamidae) des Museums fur Naturkunde der Humboldt-Universitat zu Berlin (ehemals Zoologisches Museum Berlin). Mitteilungen aus dem Zoologischen Museum in Berlin, 73, 309 - 332. https: // doi. org / 10.1002 / mmnz. 19970730209" type="journal article" year="1997">
Denzer 
<emphasis id="C1A567380913FF9B9AAAA615C024605E" box="[746,806,331,357]" italics="true" pageId="11" pageNumber="348">et al.</emphasis>
(1997)
</bibRefCitation>
did not list ZMB 680 as a type and 
<bibRefCitation id="9740C6DB0913FF9B9D78A615C24460B2" author="Bauer, A. M. &amp; Gunther, R." pageId="11" pageNumber="348" pagination="167 - 185" refId="ref11369" refString="Bauer, A. M. &amp; Gunther, R. (2013) Origin and identity of the von Borcke collection of amphibians and reptiles in the Museum fur Naturkunde in Berlin: A cache of Seba specimens? Zoosystematics and Evolution, 89 (1), 167 - 185. https: // doi. org / 10.1002 / zoos. 201300005" type="journal article" year="2013">Bauer &amp; Günther (2013)</bibRefCitation>
reported the 
<taxonomicName id="34D1C0A90913FF9B999DA629C17260B2" baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[477,624,367,393]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380913FF9B999DA629C17260B2" box="[477,624,367,393]" italics="true" pageId="11" pageNumber="348">B. cristatella</emphasis>
</taxonomicName>
specimen as “not located”. A new search in the collection revealed that the specimen is still present. However, ZMB 680 has its tail broken twice and does not match the illustration in 
<bibRefCitation id="9740C6DB0913FF9B9D26A6CAC3E760EA" author="Seba, A." pageId="11" pageNumber="348" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734)</bibRefCitation>
. Furthermore, our measurements do not agree with the measurements given by 
<bibRefCitation id="9740C6DB0913FF9B9C31A6E1C62360EA" author="Merrem, B." box="[1137,1313,439,466]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
. ZMB 680 can therefore be excluded as the 
<typeStatus id="2C6A05880913FF9B9A46A685C15F60CD" box="[518,605,476,502]" pageId="11" pageNumber="348" type="syntype">syntype</typeStatus>
depicted in Seba’s figure 1.
</paragraph>
<paragraph id="F36EBB2A0913FF9B9887A6A6C18864FA" blockId="11.[151,1437,223,2014]" pageId="11" pageNumber="348">
The 
<taxonomicName id="34D1C0A90913FF9B98BBA559C2616322" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[251,355,511,537]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B98BBA559C2616322" box="[251,355,511,537]" italics="true" pageId="11" pageNumber="348">B. jubata</emphasis>
</taxonomicName>
specimen (ZMB 686) was reported by 
<bibRefCitation id="9740C6DB0913FF9B9B63A559C7386322" author="Bauer, A. M. &amp; Gunther, R." box="[803,1082,511,538]" pageId="11" pageNumber="348" pagination="167 - 185" refId="ref11369" refString="Bauer, A. M. &amp; Gunther, R. (2013) Origin and identity of the von Borcke collection of amphibians and reptiles in the Museum fur Naturkunde in Berlin: A cache of Seba specimens? Zoosystematics and Evolution, 89 (1), 167 - 185. https: // doi. org / 10.1002 / zoos. 201300005" type="journal article" year="2013">Bauer &amp; Günther (2013)</bibRefCitation>
as “no match to Seba figures”. We re-examined the specimen (ZMB 686) and compared it to Seba’s plate 89 figure 2 as well as to 
<bibRefCitation id="9740C6DB0913FF9B9C9FA57AC69E6306" author="Merrem, B." box="[1247,1436,547,573]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem’s (1819)</bibRefCitation>
description and we were surprised that it was rather a good match to Seba’s plate 89 figure 
<quantity id="342916CF0913FF9B9CCDA51EC7BA635A" box="[1165,1208,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="348" unit="in" value="2.0">2 in</quantity>
general appearance. In particular, the specimen agrees with Merrem’s description regarding the presence of a dorsal crest consisting of “tooth like leafs” (lanceolate scales) that decrease in height towards the base of the tail. There exist, however, some differences between Seba’s figures and Merrem’s description. The direction of the scales in Seba’s figure is backwards pointing, a character typical for 
<taxonomicName id="34D1C0A90913FF9B9AC6A58EC01663C9" box="[646,788,727,754]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C1A567380913FF9B9AC6A58EC1DE63CA" box="[646,732,727,753]" italics="true" pageId="11" pageNumber="348">Calotes</emphasis>
spp.
</taxonomicName>
, instead of backwards and downwards as it is typical for 
<taxonomicName id="34D1C0A90913FF9B98D7A5A2C258622D" box="[151,346,763,790]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C1A567380913FF9B98D7A5A2C227622E" box="[151,293,763,789]" italics="true" pageId="11" pageNumber="348">Bronchocela</emphasis>
spp.
</taxonomicName>
Also the tail lengths (Seba pl. 89 fig. 1 and fig.2) do not reflect their actual size (TL/SVL&gt; 
<quantity id="342916CF0913FF9B9D14A5A2C67C622E" box="[1364,1406,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="11" pageNumber="348" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="34D1C0A90913FF9B9DC4A5A5C20B6202" baseAuthorityName="Kuhl" baseAuthorityYear="1820" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380913FF9B9DC4A5A5C20B6202" italics="true" pageId="11" pageNumber="348">B. cristatella</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90913FF9B997FA479C2CB6202" box="[319,457,800,825]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B997FA479C2CB6202" box="[319,457,800,825]" italics="true" pageId="11" pageNumber="348">B. gutturosa</emphasis>
</taxonomicName>
) as they only account for slightly more than double the snout-vent-length. We consider these differences as a lack of attention to detail by the illustrator. Both of these features were correctly described in 
<bibRefCitation id="9740C6DB0913FF9B98F4A431C26162BA" author="Merrem, B." box="[180,355,871,898]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
. The total length of 
<specimenCount id="E5D770A30913FF9B9A04A431C1DF62BA" box="[580,733,871,898]" count="1" pageId="11" pageNumber="348" type="generic">one specimen</specimenCount>
reported by 
<bibRefCitation id="9740C6DB0913FF9B9B2BA431C71562BA" author="Merrem, B." box="[875,1047,871,898]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
was 19 Zoll of which the tail (TL) measured 15 Zoll (1 Zoll = 1 French Pouce = 
<quantity id="342916CF0913FF9B9AD8A4D2C00C629D" box="[664,782,907,934]" metricMagnitude="-2" metricUnit="m" metricValue="2.707" pageId="11" pageNumber="348" unit="mm" value="27.07">27.07 mm</quantity>
: 19 Zoll = 
<quantity id="342916CF0913FF9B9BC7A4D2C0E6629D" box="[903,996,907,934]" metricMagnitude="-1" metricUnit="m" metricValue="5.14" pageId="11" pageNumber="348" unit="mm" value="514.0">514 mm</quantity>
(TotL); 15 Zoll = 
<quantity id="342916CF0913FF9B9CF1A4D2C60C629D" box="[1201,1294,907,934]" metricMagnitude="-1" metricUnit="m" metricValue="4.06" pageId="11" pageNumber="348" unit="mm" value="406.0">406 mm</quantity>
(TL); 4 Zoll = 
<quantity id="342916CF0913FF9B98EDA4F6C20862F1" box="[173,266,943,970]" metricMagnitude="-1" metricUnit="m" metricValue="1.08" pageId="11" pageNumber="348" unit="mm" value="108.0">108 mm</quantity>
(SVL); alternatively, 1 Zoll 
<collectingRegion id="311575C80913FF9B9A07A4E9C18862F1" box="[583,650,944,970]" country="Germany" name="Hessen" pageId="11" pageNumber="348">Hesse</collectingRegion>
= 
<quantity id="342916CF0913FF9B9AE7A4F6C1FB62F1" box="[679,761,943,970]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="348" unit="mm" value="25.0">25 mm</quantity>
; 19 Zoll = 
<quantity id="342916CF0913FF9B9B31A4F6C0CC62F1" box="[881,974,943,970]" metricMagnitude="-1" metricUnit="m" metricValue="4.82" pageId="11" pageNumber="348" unit="mm" value="482.0">482 mm</quantity>
(TotL); 15 Zoll = 
<quantity id="342916CF0913FF9B9CD9A4F6C7F462F1" box="[1177,1270,943,970]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="11" pageNumber="348" unit="mm" value="381.0">381 mm</quantity>
(TL); TL/SVL = 406/108 = 3.76). The value of 
<quantity id="342916CF0913FF9B9A51A48AC16062D5" box="[529,610,979,1006]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="348" unit="mm" value="25.0">25 mm</quantity>
per Zoll was only introduced in 
<collectingRegion id="311575C80913FF9B9B9DA48DC72262D5" box="[989,1056,980,1006]" country="Germany" name="Hessen" pageId="11" pageNumber="348">Hesse</collectingRegion>
(where Merrem resided) in 1821 and we assume that 
<bibRefCitation id="9740C6DB0913FF9B993AA4A1C125652A" author="Merrem, B." box="[378,551,1015,1042]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
referred to the French measurement system. In any case, our measurements of ZMB 686 (TotL = 
<quantity id="342916CF0913FF9B9929A342C2CB650D" box="[361,457,1051,1078]" metricMagnitude="-1" metricUnit="m" metricValue="5.4799999999999995" pageId="11" pageNumber="348" unit="mm" value="548.0">548 mm</quantity>
, SVL = 
<quantity id="342916CF0913FF9B9A60A342C182650D" box="[544,640,1051,1078]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="348" unit="mm" value="120.0">120 mm</quantity>
; TL = 
<quantity id="342916CF0913FF9B9A8BA342C029650D" box="[715,811,1051,1078]" metricMagnitude="-1" metricUnit="m" metricValue="4.279999999999999" pageId="11" pageNumber="348" unit="mm" value="428.0">428 mm</quantity>
; TL/SVL = 3.57) differ from those reported by 
<bibRefCitation id="9740C6DB0913FF9B9D02A345C3E76562" author="Merrem, B." pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
. Another feature that is discernible in Seba’s plate 89 figure 2, but not mentioned by 
<bibRefCitation id="9740C6DB0913FF9B9CEBA319C6586562" author="Merrem, B." box="[1195,1370,1087,1114]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
, is an unusually formed gular sac that appears to be rather spherical/globular, something that is clearly developed in ZMB 686. However, 
<bibRefCitation id="9740C6DB0913FF9B9900A3D1C2E9659A" author="Merrem, B." box="[320,491,1159,1186]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
stated that the throat of 
<specimenCount id="E5D770A30913FF9B9AB4A3D1C08E659A" box="[756,908,1159,1186]" count="1" pageId="11" pageNumber="348" type="generic">one specimen</specimenCount>
(referring to plate 89 fig. 
<quantity id="342916CF0913FF9B9CE9A3DEC7D1659A" box="[1193,1235,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="348" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="9740C6DB0913FF9B9C98A3DEC663659A" author="Seba, A." box="[1240,1377,1159,1185]" pageId="11" pageNumber="348" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734)</bibRefCitation>
, “der ersten” = the first) was filled with cloth such that the gular sac appeared convex. While this feature is visible in 
<bibRefCitation id="9740C6DB0913FF9B9D26A3F2C3DF65D2" author="Seba, A." pageId="11" pageNumber="348" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734</bibRefCitation>
: plate 89 figure 1) (= 
<taxonomicName id="34D1C0A90913FF9B9997A396C1F365D2" authority="Kuhl, 1820" authorityName="Kuhl" authorityYear="1820" box="[471,753,1231,1257]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380913FF9B9997A396C16B65D2" box="[471,617,1231,1257]" italics="true" pageId="11" pageNumber="348">A. cristatella</emphasis>
<bibRefCitation id="9740C6DB0913FF9B9A31A396C1F365D2" author="Kuhl, H." box="[625,753,1231,1257]" pageId="11" pageNumber="348" refId="ref13590" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Hermannschen Buchhandlung, Frankfurt, (ii) + 151 + (1) pp., 7 pls." type="journal volume" year="1820">Kuhl, 1820</bibRefCitation>
</taxonomicName>
) it is much more pronounced in 
<bibRefCitation id="9740C6DB0913FF9B9C27A396C7E965D2" author="Seba, A." box="[1127,1259,1231,1257]" pageId="11" pageNumber="348" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734</bibRefCitation>
: plate 89 figure 2) (= 
<taxonomicName id="34D1C0A90913FF9B9896A3AAC10E6436" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[214,524,1267,1294]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B9896A3AAC2626436" box="[214,352,1267,1293]" italics="true" pageId="11" pageNumber="348">A. gutturosa</emphasis>
<bibRefCitation id="9740C6DB0913FF9B9928A3ADC10E6436" author="Merrem, B." box="[360,524,1267,1294]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
). Apparently, it was custom in early collections to emphasise anatomical features to produce a more “natural” live appearance. We did not find any cloth in ZMB 686 but the unnaturally formed gular pouch lead us to the assumption that it had been stuffed at an earlier stage (
<figureCitation id="6BEAA7AF0913FF9B9C67A262C789646E" box="[1063,1163,1339,1365]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[151,1436,181,1880]" captionTargetId="figure-27@4.[151,1436,181,1880]" captionTargetPageId="4" captionText="FIGURE 2. Bronchocela jubata (ZMB 686, von Borcke’s specimen). Upper panel: lateral view. Lower panel: ventral view. Please note the unnaturally formed gular region." figureDoi="http://doi.org/10.5281/zenodo.13757741" httpUri="https://zenodo.org/record/13757741/files/figure.png" pageId="11" pageNumber="348">Figure 2</figureCitation>
). As 
<bibRefCitation id="9740C6DB0913FF9B9C85A265C673646E" author="Merrem, B." box="[1221,1393,1339,1366]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
did not mention this particular feature for the specimen he considered to represent plate 89 figure 
<quantity id="342916CF0913FF9B9CF3A206C7DC6442" box="[1203,1246,1375,1401]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="348" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="9740C6DB0913FF9B9CA5A206C66F6442" author="Seba, A." box="[1253,1389,1375,1401]" pageId="11" pageNumber="348" refId="ref14390" refString="Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Tomus I. J. Wetstenium, &amp; Gul. Smith, &amp; Janssonio-Waesbergios, Amstelaedami [Amsterdam], xxxii + 178 pp., 111 pls. https: // doi. org / 10.5962 / bhl. title. 14110" type="book" year="1734">Seba (1734)</bibRefCitation>
and because our measurements differ from Merrem’s we do not treat ZMB 686 as a 
<typeStatus id="2C6A05880913FF9B9C58A2DDC76D64A5" box="[1048,1135,1412,1438]" pageId="11" pageNumber="348" type="syntype">syntype</typeStatus>
of 
<emphasis id="C1A567380913FF9B9CD5A2DDC64764A6" box="[1173,1349,1411,1438]" italics="true" pageId="11" pageNumber="348">
C. A.. 
<taxonomicName id="34D1C0A90913FF9B9C9BA2DDC64764A6" box="[1243,1349,1412,1437]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">gutturosa</taxonomicName>
</emphasis>
despite its otherwise good match with Seba’s figure.
</paragraph>
<paragraph id="F36EBB2A0913FF9B9887A292C77B66AE" blockId="11.[151,1437,223,2014]" pageId="11" pageNumber="348">
<bibRefCitation id="9740C6DB0913FF9B9887A292C28764DE" author="Merrem, B." box="[199,389,1483,1509]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem’s (1819)</bibRefCitation>
description of 
<taxonomicName id="34D1C0A90913FF9B9A6DA292C1EF64DE" box="[557,749,1483,1509]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B9A6DA292C1EF64DE" box="[557,749,1483,1509]" italics="true" pageId="11" pageNumber="348">Agama gutturosa</emphasis>
</taxonomicName>
contained several characters that facilitate a comparison with currently accepted taxa. The absence of a spine on the occiput and dorsal scales directed backwards and downwards clearly differentiates the species from 
<taxonomicName id="34D1C0A90913FF9B9A01A14AC1956716" box="[577,663,1555,1581]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="genus">
<emphasis id="C1A567380913FF9B9A01A14AC1956716" box="[577,663,1555,1581]" italics="true" pageId="11" pageNumber="348">Calotes</emphasis>
</taxonomicName>
and places the species into the genus 
<taxonomicName id="34D1C0A90913FF9B9C7CA14AC7C86716" authorityName="Kaup" authorityYear="1827" box="[1084,1226,1555,1581]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="genus">
<emphasis id="C1A567380913FF9B9C7CA14AC7C86716" box="[1084,1226,1555,1581]" italics="true" pageId="11" pageNumber="348">Bronchocela</emphasis>
</taxonomicName>
. If 
<bibRefCitation id="9740C6DB0913FF9B9CB1A14DC69E6716" author="Merrem, B." box="[1265,1436,1555,1582]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
had Seba’s specimen at hand, then any assumption that Seba’s plate 89 fig. 2 shows 
<taxonomicName id="34D1C0A90913FF9B9C24A16EC610676A" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1124,1298,1591,1617]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="calotes">
<emphasis id="C1A567380913FF9B9C24A16EC610676A" box="[1124,1298,1591,1617]" italics="true" pageId="11" pageNumber="348">Calotes calotes</emphasis>
</taxonomicName>
is therefore unfounded. The presence of a well-developed dorsal crest helps to exclude 
<taxonomicName id="34D1C0A90913FF9B9BACA102C7F6674E" authorityName="Kuhl" authorityYear="1820" box="[1004,1268,1627,1653]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380913FF9B9BACA102C7F6674E" box="[1004,1268,1627,1653]" italics="true" pageId="11" pageNumber="348">Bronchocela cristatella</emphasis>
</taxonomicName>
as this species has at best a few enlarged scales on the dorsal ridge which is otherwise serrated. Within the genus 
<taxonomicName id="34D1C0A90913FF9B9D47A126C69767A2" authorityName="Kaup" authorityYear="1827" box="[1287,1429,1663,1689]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="genus">
<emphasis id="C1A567380913FF9B9D47A126C69767A2" box="[1287,1429,1663,1689]" italics="true" pageId="11" pageNumber="348">Bronchocela</emphasis>
</taxonomicName>
, other than the Javanese 
<taxonomicName id="34D1C0A90913FF9B99E1A1FDC1056786" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[417,519,1699,1725]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B99E1A1FDC1056786" box="[417,519,1699,1725]" italics="true" pageId="11" pageNumber="348">B. jubata</emphasis>
</taxonomicName>
, the Philippine 
<taxonomicName id="34D1C0A90913FF9B9AF7A1FDC0516786" authorityName="Gray" authorityYear="1845" box="[695,851,1700,1725]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="marmorata">
<emphasis id="C1A567380913FF9B9AF7A1FDC0516786" box="[695,851,1700,1725]" italics="true" pageId="11" pageNumber="348">B. marmorata</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90913FF9B9BC9A1FDC7196786" authorityName="Gray" authorityYear="1845" box="[905,1051,1699,1725]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="celebensis">
<emphasis id="C1A567380913FF9B9BC9A1FDC7196786" box="[905,1051,1699,1725]" italics="true" pageId="11" pageNumber="348">B. celebensis</emphasis>
</taxonomicName>
from 
<collectingRegion id="311575C80913FF9B9C1EA1FAC7C16786" box="[1118,1219,1699,1725]" country="Indonesia" name="Sulawesi Utara" pageId="11" pageNumber="348">Sulawesi</collectingRegion>
possess an anterior dorsal crest of elongated scales. However, 
<taxonomicName id="34D1C0A90913FF9B9AC5A191C1ED67DA" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[645,751,1735,1761]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B9AC5A191C1ED67DA" box="[645,751,1735,1761]" italics="true" pageId="11" pageNumber="348">B. jubata</emphasis>
</taxonomicName>
is the only species where the crest runs across most of the back and decreases continuously towards the base of the tail as described by 
<bibRefCitation id="9740C6DB0913FF9B9C4BA1B5C7B9663E" author="Merrem, B." box="[1035,1211,1771,1798]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
. Further to this, the gular pouch of 
<taxonomicName id="34D1C0A90913FF9B9904A049C2D56612" authorityName="Gray" authorityYear="1845" box="[324,471,1807,1833]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="celebensis">
<emphasis id="C1A567380913FF9B9904A049C2D56612" box="[324,471,1807,1833]" italics="true" pageId="11" pageNumber="348">B. celebensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="34D1C0A90913FF9B9A4FA049C1AF6612" authorityName="Gray" authorityYear="1845" box="[527,685,1808,1833]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="marmorata">
<emphasis id="C1A567380913FF9B9A4FA049C1AF6612" box="[527,685,1808,1833]" italics="true" pageId="11" pageNumber="348">B. marmorata</emphasis>
</taxonomicName>
is smaller than in 
<taxonomicName id="34D1C0A90913FF9B9BC0A056C7086612" box="[896,1034,1807,1833]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B9BC0A056C7086612" box="[896,1034,1807,1833]" italics="true" pageId="11" pageNumber="348">A. gutturosa</emphasis>
</taxonomicName>
. 
<bibRefCitation id="9740C6DB0913FF9B9C59A056C7DB6612" author="Merrem, B." box="[1049,1241,1807,1833]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem’s (1819)</bibRefCitation>
description of 
<taxonomicName id="34D1C0A90913FF9B9DC4A056C2036676" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B9DC4A056C2036676" italics="true" pageId="11" pageNumber="348">A. gutturosa</emphasis>
</taxonomicName>
and the von Borcke specimen (ZMB 686) agree well with the remaining 
<specimenCount id="E5D770A30913FF9B9C10A06DC7E76675" box="[1104,1253,1844,1870]" count="2" pageId="11" pageNumber="348" type="generic" typeStatus="syntype">two syntypes</specimenCount>
of 
<taxonomicName id="34D1C0A90913FF9B9D4EA06AC3DD664A" authorityName="Dumeril &amp; Bibron" authorityYear="1837" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B9D4EA06AC3DD664A" italics="true" pageId="11" pageNumber="348">Bronchocela jubata</emphasis>
</taxonomicName>
(MNHN-RA 2542 and MNHN-RA 2543, see 
<bibRefCitation id="9740C6DB0913FF9B9AAFA001C085664A" author="Brygoo, E. R." box="[751,903,1879,1906]" pageId="11" pageNumber="348" pagination="1 - 56" refId="ref11986" refString="Brygoo, E. R. (1988) Les types d'Agamides (Reptiles, Sauriens) du Museum national d'Histoire naturelle catalogue critique. Bulletin du Museum national d'Histoire naturelle, Paris, Series 4 A, 10 (Supplement 3), 1 - 56. https: // doi. org / 10.5962 / p. 287636" type="journal article" year="1988">Brygoo 1988</bibRefCitation>
). We therefore conclude, that 
<bibRefCitation id="9740C6DB0913FF9B9C9CA00EC69E664A" author="Merrem, B." box="[1244,1436,1879,1905]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem’s (1819)</bibRefCitation>
description of 
<taxonomicName id="34D1C0A90913FF9B9979A022C2FB66AE" box="[313,505,1915,1941]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B9979A022C2FB66AE" box="[313,505,1915,1941]" italics="true" pageId="11" pageNumber="348">Agama gutturosa</emphasis>
</taxonomicName>
represents 
<taxonomicName id="34D1C0A90913FF9B9A39A022C77766AE" authority="Dumeril &amp; Bibron, 1837" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[633,1141,1915,1941]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B9A39A022C05466AE" box="[633,854,1915,1941]" italics="true" pageId="11" pageNumber="348">Bronchocela jubata</emphasis>
<bibRefCitation id="9740C6DB0913FF9B9B1DA022C77766AE" author="Dumeril, A. M. C. &amp; Bibron, G." box="[861,1141,1915,1941]" pageId="11" pageNumber="348" refId="ref12679" refString="Dumeril, A. M. C. &amp; Bibron, G. (1837) Erpetologie generale ou histoire naturelle complete des Reptiles. Tome quatrieme. Contenant l'histoire de quarante-six genres et de cent quarante-six especes de la famille des iguaniens, de l'ordre des Sauriens. Vol. IV. Librairie Encyclopedique de Roret, Paris, ii + 572 pp." type="book" year="1837">Duméril &amp; Bibron, 1837</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BBCBE8A10913FF9C9887A0C6C2B16392" lastPageId="12" lastPageNumber="349" pageId="11" pageNumber="348" type="description">
<paragraph id="F36EBB2A0913FF9C9887A0C6C2B16392" blockId="11.[151,1437,223,2014]" lastBlockId="12.[151,1437,151,681]" lastPageId="12" lastPageNumber="349" pageId="11" pageNumber="348">
Consequently, 
<taxonomicName id="34D1C0A90913FF9B992EA0C6C1E76682" authority="Merrem (1819)" authorityName="Merrem" authorityYear="1819" box="[366,741,1951,1978]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380913FF9B992EA0C6C1326682" box="[366,560,1951,1977]" italics="true" pageId="11" pageNumber="348">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0913FF9B9A78A0F9C1E76682" author="Merrem, B." box="[568,741,1951,1978]" pageId="11" pageNumber="348" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem (1819)</bibRefCitation>
</taxonomicName>
becomes a senior subjective synonym of 
<taxonomicName id="34D1C0A90913FF9B9CFFA0C6C2A666E6" authority="Dumeril &amp; Bibron, 1837" authorityName="Dumeril &amp; Bibron" authorityYear="1837" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9B9CFFA0C6C69F6682" box="[1215,1437,1951,1977]" italics="true" pageId="11" pageNumber="348">Bronchocela jubata</emphasis>
<bibRefCitation id="9740C6DB0913FF9B98D7A09AC2A666E6" author="Dumeril, A. M. C. &amp; Bibron, G." box="[151,420,1987,2013]" pageId="11" pageNumber="348" refId="ref12679" refString="Dumeril, A. M. C. &amp; Bibron, G. (1837) Erpetologie generale ou histoire naturelle complete des Reptiles. Tome quatrieme. Contenant l'histoire de quarante-six genres et de cent quarante-six especes de la famille des iguaniens, de l'ordre des Sauriens. Vol. IV. Librairie Encyclopedique de Roret, Paris, ii + 572 pp." type="book" year="1837">Duméril &amp; Bibron, 1837</bibRefCitation>
</taxonomicName>
and would have nomenclatural priority such that the species currently known as 
<taxonomicName id="34D1C0A90913FF9C9D4EA09AC2F8618A" authority="Dumeril &amp; Bibron, 1837" authorityName="Dumeril &amp; Bibron" authorityYear="1837" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" lastPageId="12" lastPageNumber="349" pageId="11" pageNumber="348" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380913FF9C9D4EA09AC3DD618A" italics="true" lastPageId="12" lastPageNumber="349" pageId="11" pageNumber="348">Bronchocela jubata</emphasis>
<bibRefCitation id="9740C6DB0914FF9C98A6A7CEC2F8618A" author="Dumeril, A. M. C. &amp; Bibron, G." box="[230,506,151,177]" pageId="12" pageNumber="349" refId="ref12679" refString="Dumeril, A. M. C. &amp; Bibron, G. (1837) Erpetologie generale ou histoire naturelle complete des Reptiles. Tome quatrieme. Contenant l'histoire de quarante-six genres et de cent quarante-six especes de la famille des iguaniens, de l'ordre des Sauriens. Vol. IV. Librairie Encyclopedique de Roret, Paris, ii + 572 pp." type="book" year="1837">Duméril &amp; Bibron, 1837</bibRefCitation>
</taxonomicName>
should be called “ 
<taxonomicName id="34D1C0A90914FF9C9A8AA7CEC777618A" authority="Merrem, 1819" authorityName="Merrem" authorityYear="1819" box="[714,1141,151,177]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380914FF9C9A8AA7CEC0CB618B" box="[714,969,151,177]" italics="true" pageId="12" pageNumber="349">Bronchocela gutturosa</emphasis>
<bibRefCitation id="9740C6DB0914FF9C9B90A7CEC777618A" author="Merrem, B." box="[976,1141,151,177]" pageId="12" pageNumber="349" pagination="162 - 168" refId="ref13992" refString="Merrem, B. (1819) Agama. in Ersch J. S. &amp; Gruber J. G. (1819) Allgemeine Encyclopadie der Wissenschaften und Kunste in alphabetischer Folge von genannten Schriftstellern bearbeitet. Theil 2 Aga - Aldus. J. F. Gleditsch, Leipzig, pp. 162 - 168. [Source for date of publication: Blomeke, E. &amp; Muller, J. (1890) Katalog der Bibliothek des Reichtags. Reichstag, Berlin, 744 pp.]" type="book chapter" year="1819">Merrem, 1819</bibRefCitation>
</taxonomicName>
”. This, however, threatens nomenclatural stability as the name 
<taxonomicName id="34D1C0A90914FF9C9A68A7E2C00661EE" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[552,772,187,213]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380914FF9C9A68A7E2C00661EE" box="[552,772,187,213]" italics="true" pageId="12" pageNumber="349">Bronchocela jubata</emphasis>
</taxonomicName>
has been used consistently since 
<bibRefCitation id="9740C6DB0914FF9C9C37A7E2C69A61EE" author="Dumeril, A. M. C. &amp; Bibron, G." box="[1143,1432,187,213]" pageId="12" pageNumber="349" refId="ref12679" refString="Dumeril, A. M. C. &amp; Bibron, G. (1837) Erpetologie generale ou histoire naturelle complete des Reptiles. Tome quatrieme. Contenant l'histoire de quarante-six genres et de cent quarante-six especes de la famille des iguaniens, de l'ordre des Sauriens. Vol. IV. Librairie Encyclopedique de Roret, Paris, ii + 572 pp." type="book" year="1837">Duméril &amp; Bibron (1837)</bibRefCitation>
. In a recent publication dealing with the taxonomy of 
<taxonomicName id="34D1C0A90914FF9C9B4CA786C62C61C2" authority=", Amarasinghe et al. (2022)" authorityName="Amarasinghe" authorityYear="2022" box="[780,1326,223,249]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380914FF9C9B4CA786C0EF61C2" box="[780,1005,223,249]" italics="true" pageId="12" pageNumber="349">Bronchocela jubata</emphasis>
, 
<bibRefCitation id="9740C6DB0914FF9C9BBDA786C62C61C2" author="Amarasinghe, A. A. T. &amp; Ineich, I. &amp; Riyanto, A. &amp; Hallermann, J. &amp; Andayani, N. &amp; Abinawanto, A. &amp; Supriatna, J." box="[1021,1326,223,249]" pageId="12" pageNumber="349" pagination="62 - 85" refId="ref11140" refString="Amarasinghe, A. A. T., Ineich, I., Riyanto, A., Hallermann, J., Andayani, N., Abinawanto, A. &amp; Supriatna, J. (2022) Taxonomy and distribution of a common arboreal lizard, Bronchocela jubata Dumeril &amp; Bibron, 1837 (Reptilia: Agamidae), with designation of its lectotype from Java, Indonesia. Zootaxa, 5150 (1), 62 - 85. https: // doi. org / 10.11646 / zootaxa. 5150.1.3" type="journal article" year="2022">
Amarasinghe 
<emphasis id="C1A567380914FF9C9CDDA786C7DB61C2" box="[1181,1241,223,249]" italics="true" pageId="12" pageNumber="349">et al.</emphasis>
(2022)
</bibRefCitation>
</taxonomicName>
followed 
<bibRefCitation id="9740C6DB0914FF9C98D7A65AC24D6026" author="Figueroa, A." box="[151,335,259,285]" pageId="12" pageNumber="349" pagination="397 - 400" refId="ref12915" refString="Figueroa, A. (2021) The type species of Bronchocela Kaup, 1827 and lectotype designations for Agama cristatella Kuhl, 1820 and Agama gutturosa Merrem, 1820 (Squamata: Iguania: Agamidae). Zootaxa, 4980 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4980.2.12" type="journal article" year="2021">Figueroa (2021)</bibRefCitation>
and treated 
<taxonomicName id="34D1C0A90914FF9C99A2A65AC0506026" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" box="[482,850,259,285]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380914FF9C99A2A65AC1A76027" box="[482,677,259,285]" italics="true" pageId="12" pageNumber="349">Agama gutturosa</emphasis>
<bibRefCitation id="9740C6DB0914FF9C9AEFA65AC0506026" author="Merrem, B." box="[687,850,259,285]" pageId="12" pageNumber="349" refId="ref14083" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis ampibiorum. Johann Christian Krieger, Marburg, xv + 191 pp., 1 pl. https: // doi. org / 10.5962 / bhl. title. 5037" type="book" year="1820">Merrem, 1820</bibRefCitation>
</taxonomicName>
as a junior objective synonym of, and conspecific with, 
<taxonomicName id="34D1C0A90914FF9C989AA67EC2E6607A" authorityName="Kuhl" authorityYear="1820" box="[218,484,295,321]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="cristatella">
<emphasis id="C1A567380914FF9C989AA67EC2E6607A" box="[218,484,295,321]" italics="true" pageId="12" pageNumber="349">Bronchocela cristatella</emphasis>
</taxonomicName>
in order “to uphold the taxonomic stability of a long standing nomen, 
<taxonomicName id="34D1C0A90914FF9C9D5FA67EC68B607A" box="[1311,1417,295,321]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380914FF9C9D5FA67EC632607B" box="[1311,1328,295,320]" italics="true" pageId="12" pageNumber="349">B</emphasis>
. 
<emphasis id="C1A567380914FF9C9D01A67EC68B607A" box="[1345,1417,295,321]" italics="true" pageId="12" pageNumber="349">jubata</emphasis>
</taxonomicName>
”. Our analysis has revealed that this assumption is incorrect, and the only way to stabilise nomenclature is to reverse precedence according to Art. 23.9 of the 
<emphasis id="C1A567380914FF9C9A19A636C19660B2" box="[601,660,367,393]" italics="true" pageId="12" pageNumber="349">Code</emphasis>
(
<bibRefCitation id="9740C6DB0914FF9C9AE1A636C02560B2" author="ICZN" box="[673,807,367,393]" pageId="12" pageNumber="349" refId="ref13461" refString="ICZN (1999) International Code of Zoological Nomenclature. International Commission on Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
). The last use of the older name, “ 
<taxonomicName id="34D1C0A90914FF9C9CDEA636C7F660B2" box="[1182,1268,367,393]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C9CDEA636C7F660B2" box="[1182,1268,367,393]" italics="true" pageId="12" pageNumber="349">Calotes</emphasis>
</taxonomicName>
(?) 
<taxonomicName id="34D1C0A90914FF9C9D5EA636C68A60B3" box="[1310,1416,367,392]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380914FF9C9D5EA636C68A60B3" box="[1310,1416,367,392]" italics="true" pageId="12" pageNumber="349">gutturosa</emphasis>
</taxonomicName>
”, as valid was by Fitzinger (1860). The species epithet 
<taxonomicName id="34D1C0A90914FF9C9B50A6CAC0786097" box="[784,890,403,428]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380914FF9C9B50A6CAC0786097" box="[784,890,403,428]" italics="true" pageId="12" pageNumber="349">gutturosa</emphasis>
</taxonomicName>
has not been in use after 
<quantity id="342916CF0914FF9C9CF4A6CAC60F6096" box="[1204,1293,403,429]" metricMagnitude="1" metricUnit="m" metricValue="4.82346" pageId="12" pageNumber="349" unit="in" value="1899.0">1899 in</quantity>
conjunction with either 
<taxonomicName id="34D1C0A90914FF9C995AA6EEC2AA60EA" authorityName="Kaup" authorityYear="1827" box="[282,424,439,465]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C995AA6EEC2AA60EA" box="[282,424,439,465]" italics="true" pageId="12" pageNumber="349">Bronchocela</emphasis>
</taxonomicName>
or 
<taxonomicName id="34D1C0A90914FF9C9991A6EEC12560EA" box="[465,551,439,465]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C9991A6EEC12560EA" box="[465,551,439,465]" italics="true" pageId="12" pageNumber="349">Calotes</emphasis>
</taxonomicName>
apart from the mere mentioning as 
<taxonomicName id="34D1C0A90914FF9C9B83A6EEC78760EB" box="[963,1157,439,465]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380914FF9C9B83A6EEC78760EB" box="[963,1157,439,465]" italics="true" pageId="12" pageNumber="349">Agama gutturosa</emphasis>
</taxonomicName>
in a non-taxonomic and non-nomenclatural compilation of earlier published names by 
<bibRefCitation id="9740C6DB0914FF9C9B18A682C71760CE" author="Sherborn, C. D." box="[856,1045,474,501]" pageId="12" pageNumber="349" refId="ref14482" refString="Sherborn, C. D. (1905) Index animalium; sive, Index nominum quae ab A. D. MDCCLVIII generibus et speciebus animalium imposita sunt. Cambridge University Press, Cambridge. [unknown pagination]" type="book" year="1905">Sherborn (1905)</bibRefCitation>
. We therefore consider the species epithet 
<taxonomicName id="34D1C0A90914FF9C98A9A6A6C2516323" box="[233,339,511,536]" class="Squamata" family="Agamidae" genus="Agama" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="gutturosa">
<emphasis id="C1A567380914FF9C98A9A6A6C2516323" box="[233,339,511,536]" italics="true" pageId="12" pageNumber="349">gutturosa</emphasis>
</taxonomicName>
/- 
<emphasis id="C1A567380914FF9C9925A6A6C27C6323" box="[357,382,511,536]" italics="true" pageId="12" pageNumber="349">us</emphasis>
in conjunction with either 
<taxonomicName id="34D1C0A90914FF9C9AE6A6A6C0366322" authorityName="Kaup" authorityYear="1827" box="[678,820,511,537]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C9AE6A6A6C0366322" box="[678,820,511,537]" italics="true" pageId="12" pageNumber="349">Bronchocela</emphasis>
</taxonomicName>
(or 
<taxonomicName id="34D1C0A90914FF9C9B20A6A6C0B46322" box="[864,950,511,537]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C9B20A6A6C0B46322" box="[864,950,511,537]" italics="true" pageId="12" pageNumber="349">Calotes</emphasis>
</taxonomicName>
) as a nomen oblitum and the species epithet 
<taxonomicName id="34D1C0A90914FF9C98D7A57AC3DD6306" box="[151,223,547,573]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380914FF9C98D7A57AC3DD6306" box="[151,223,547,573]" italics="true" pageId="12" pageNumber="349">jubata</emphasis>
</taxonomicName>
/- 
<emphasis id="C1A567380914FF9C98B0A57AC20B6307" box="[240,265,547,572]" italics="true" pageId="12" pageNumber="349">us</emphasis>
in conjunction with either 
<taxonomicName id="34D1C0A90914FF9C9A77A57AC1C76306" authorityName="Kaup" authorityYear="1827" box="[567,709,547,573]" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C9A77A57AC1C76306" box="[567,709,547,573]" italics="true" pageId="12" pageNumber="349">Bronchocela</emphasis>
</taxonomicName>
(or 
<taxonomicName id="34D1C0A90914FF9C9AB4A57AC0486306" box="[756,842,547,573]" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="C1A567380914FF9C9AB4A57AC0486306" box="[756,842,547,573]" italics="true" pageId="12" pageNumber="349">Calotes</emphasis>
</taxonomicName>
) as a nomen protectum. In support of our decision to reverse precedence we provide a list of publications in Appendix I in accordance with Art. 23.9 of the 
<emphasis id="C1A567380914FF9C9D4FA51EC648635A" box="[1295,1354,583,609]" italics="true" pageId="12" pageNumber="349">Code</emphasis>
(
<bibRefCitation id="9740C6DB0914FF9C9D19A51EC3D163BE" author="ICZN" pageId="12" pageNumber="349" refId="ref13461" refString="ICZN (1999) International Code of Zoological Nomenclature. International Commission on Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
). In agreement with currently accepted taxonomy, the valid name of the species remains 
<taxonomicName id="34D1C0A90914FF9C9CFFA532C2AD6392" authority="Dumeril &amp; Bibron, 1837" authorityName="Dumeril &amp; Bibron" authorityYear="1837" class="Squamata" family="Agamidae" genus="Bronchocela" kingdom="Animalia" pageId="12" pageNumber="349" phylum="Chordata" rank="species" species="jubata">
<emphasis id="C1A567380914FF9C9CFFA532C69F63BE" box="[1215,1437,619,645]" italics="true" pageId="12" pageNumber="349">Bronchocela jubata</emphasis>
<bibRefCitation id="9740C6DB0914FF9C98D7A5D6C2AD6392" author="Dumeril, A. M. C. &amp; Bibron, G." box="[151,431,655,681]" pageId="12" pageNumber="349" refId="ref12679" refString="Dumeril, A. M. C. &amp; Bibron, G. (1837) Erpetologie generale ou histoire naturelle complete des Reptiles. Tome quatrieme. Contenant l'histoire de quarante-six genres et de cent quarante-six especes de la famille des iguaniens, de l'ordre des Sauriens. Vol. IV. Librairie Encyclopedique de Roret, Paris, ii + 572 pp." type="book" year="1837">Duméril &amp; Bibron, 1837</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>