238 lines
25 KiB
XML
238 lines
25 KiB
XML
<document id="FE5E32C67AE1010A0A2FACA387EC5B58" ID-DOI="10.1080/00222930050020122" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747979" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620672552581" checkinUser="carolina" docAuthor="Gussmann S. M. & Scholtz, C. H." docDate="2000" docId="7B60F030FFCCFF9C01FAFC3CFC10DA03" docLanguage="en" docName="JNATHIST.34.1045-1123.pdf.imf" docOrigin="Journal of Natural History 34" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Bolboceratex holmi Gussmann S. M. & Scholtz 2000, spec. nov." docType="treatment" docVersion="4" lastPageNumber="1121" masterDocId="87598848FF81FFD20079FFE2FFB0DF5E" masterDocTitle="Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)" masterLastPageNumber="1123" masterPageNumber="1045" pageNumber="1120" updateTime="1698914725550" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="9879211C1CD46680BB38BEE8186017FD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3624DA28CA18FC9081894555A54AAD79">
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<mods:title id="22A3BA6CC30BA99B62B69666B7B2EBCB">Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2BF694A8D96C436F47EB2638A6481D97" type="personal">
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<mods:role id="9D897799AA38236CC8E5628295C1AB2A">
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<mods:roleTerm id="5AA9F42A35ED4B5ACD41102C92DE0881">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="50A0212F9D1434D5AE81F5CECF9E936B">Gussmann S. M.</mods:namePart>
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</mods:name>
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<mods:name id="E77777D8E8290D43A87E20660250F2DF" type="personal">
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<mods:role id="2FAA0A6ECB3A90CAB12699E04282BD01">
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<mods:roleTerm id="4A5C850A46D2EEB243B4FB6A8508A589">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F0CFDE60E118222B46CADF9905943A7A">Scholtz, C. H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0F2E9CC7F32EF264F81CDADC818F415B">text</mods:typeOfResource>
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<mods:relatedItem id="1DEF1A12302D5D65B3954ACB25481B70" type="host">
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<mods:titleInfo id="44613EF21C6489287F14FF3D718C338C">
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<mods:title id="561FC4ECA49B76D17F417A4A2D4B5C8A">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="6C862C1766969C08DD0E468CABDF338D">
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<mods:date id="34643835850D36DD2EDFAD5B4E0A0D7B">2000</mods:date>
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<mods:detail id="077CF6DC12D4A8AD2B78E8C628D3B36D" type="pubDate">
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<mods:number id="909B06B539F364F6481CC7892AC93601">2000-12-31</mods:number>
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</mods:detail>
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<mods:detail id="0853161944DD0910E6949B4DED5C6BBC" type="volume">
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<mods:number id="0F99557F99100237E2F133DE9D4D42F3">34</mods:number>
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</mods:detail>
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<mods:extent id="CCE2E89283BFBF65A4489C370A6CEBF3" unit="page">
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<mods:start id="377936DB725FCCDA4EC94882FC1D0447">1045</mods:start>
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<mods:end id="6C1A5B54D2395F5758D966795D615866">1123</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="7D29D8575B47C571901F0BB9B0578526">journal article</mods:classification>
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<mods:identifier id="6FFF23B7FC08ED798EA70FFC7E7EBEB3" type="DOI">10.1080/00222930050020122</mods:identifier>
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<mods:identifier id="001F94FAF525AEDEBB8D7B4A281746FB" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="505FBE03F4F2CE0835D392948A82A9B7" type="Zenodo-Dep">4747979</mods:identifier>
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<treatment id="7B60F030FFCCFF9C01FAFC3CFC10DA03" ID-DOI="http://doi.org/10.5281/zenodo.4747943" ID-Zenodo-Dep="4747943" LSID="urn:lsid:plazi:treatment:7B60F030FFCCFF9C01FAFC3CFC10DA03" httpUri="http://treatment.plazi.org/id/7B60F030FFCCFF9C01FAFC3CFC10DA03" lastPageId="78" lastPageNumber="1121" pageId="77" pageNumber="1120">
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<subSubSection id="BBD312ADFFCCFF9F01FAFC3CFCA0DB48" pageId="77" pageNumber="1120" type="nomenclature">
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<paragraph id="F3764126FFCCFF9F01FAFC3CFD67DCA8" blockId="77.[387,727,990,1014]" box="[387,727,990,1014]" pageId="77" pageNumber="1120">
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<heading id="A83EF64AFFCCFF9F01FAFC3CFD67DCA8" box="[387,727,990,1014]" centered="true" fontSize="10" level="2" pageId="77" pageNumber="1120" reason="2">
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<taxonomicName id="34C93AA5FFCCFF9F01FAFC3CFDEDDCA8" authority="Gussmann S. M. & Scholtz, 2000" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[387,605,990,1014]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="1120" phylum="Arthropoda" rank="species" species="holmi" status="sp. nov.">
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<emphasis id="C1BD9D34FFCCFF9F01FAFC3CFDEDDCA8" bold="true" box="[387,605,990,1014]" italics="true" pageId="77" pageNumber="1120">Bolboceratex holmi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="DA8E204FFFCCFF9F021CFC3DFD67DCA8" box="[613,727,991,1014]" pageId="77" pageNumber="1120" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
<paragraph id="F3764126FFCCFF9F0133FC1CFCA0DB48" blockId="77.[330,784,1022,1047]" box="[330,784,1022,1047]" pageId="77" pageNumber="1120">(®gures 24a±c, e, 27c, d, 29f±j, 33b, 35)</paragraph>
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</subSubSection>
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||
<subSubSection id="BBD312ADFFCCFF9C0036FBADFC10DA03" lastPageId="78" lastPageNumber="1121" pageId="77" pageNumber="1120" type="description">
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||
<paragraph id="F3764126FFCCFF9F0036FBADFF7DDB38" blockId="77.[79,1036,1103,1700]" box="[79,205,1103,1126]" pageId="77" pageNumber="1120">
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<emphasis id="C1BD9D34FFCCFF9F0036FBADFF7DDB38" box="[79,205,1103,1126]" italics="true" pageId="77" pageNumber="1120">Description</emphasis>
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||
</paragraph>
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<paragraph id="F3764126FFCCFF9F0001FB8CFE73DBD8" blockId="77.[79,1036,1103,1700]" box="[120,451,1134,1158]" pageId="77" pageNumber="1120">
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<emphasis id="C1BD9D34FFCCFF9F0001FB8CFF4DDBD8" box="[120,253,1134,1158]" italics="true" pageId="77" pageNumber="1120">Body length</emphasis>
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.
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<quantity id="3431ECC3FFCCFF9F0166FB8CFE0CDBD8" box="[287,444,1134,1158]" metricMagnitude="-2" metricUnit="m" metricValue="1.8350000000000002" metricValueMax="2.0100000000000002" metricValueMin="1.6600000000000001" pageId="77" pageNumber="1120" unit="mm" value="18.35" valueMax="20.1" valueMin="16.6">16.6±20.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="F3764126FFCCFF9F0001FB6CFEDFDBF8" blockId="77.[79,1036,1103,1700]" box="[120,367,1166,1190]" pageId="77" pageNumber="1120">
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<emphasis id="C1BD9D34FFCCFF9F0001FB6CFF73DBF8" box="[120,195,1166,1190]" italics="true" pageId="77" pageNumber="1120">Colour</emphasis>
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||
. Mid-brown.
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</paragraph>
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<paragraph id="F3764126FFCCFF9F0001FB4CFD1DD95A" blockId="77.[79,1036,1103,1700]" pageId="77" pageNumber="1120">
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<emphasis id="C1BD9D34FFCCFF9F0001FB4CFF03DB98" box="[120,179,1198,1222]" italics="true" pageId="77" pageNumber="1120">Head</emphasis>
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||
.
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<emphasis id="C1BD9D34FFCCFF9F00ACFB4CFEC6DB98" box="[213,374,1198,1222]" italics="true" pageId="77" pageNumber="1120">Dorsal surface</emphasis>
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asetose.
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<emphasis id="C1BD9D34FFCCFF9F01A5FB4CFD4ADB98" box="[476,762,1198,1222]" italics="true" pageId="77" pageNumber="1120">Outer margin of mandible</emphasis>
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(®gures 27c, 33b) rounded.
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<emphasis id="C1BD9D34FFCCFF9F00F4FB2CFF54DBB8" box="[141,228,1230,1254]" italics="true" pageId="77" pageNumber="1120">Labrum</emphasis>
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(®gure 27c) asetose; declivous in anterior third and posterior twothirds, transverse carina obsolete; anterior margin distinctly sinuate.
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<emphasis id="C1BD9D34FFCCFF9F03CAFB0CFBBBDA58" box="[947,1035,1262,1286]" italics="true" pageId="77" pageNumber="1120">Clypeus</emphasis>
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(®gure 27c) sloping upward to apex of feebly developed to strongly raised transverse anterior carina; lateral carina anteriorly feebly developed but raised and distinct above antennal insertion, continuing from there posteriorly in a straight and then arcuate line to almost posterior end of frons; antero-lateral angle of genae sharply raised; with posterior transverse bituberculate carina, carina U-shaped between tubercles; carina laterally terminating with tubercles, not joining lateral clypeal margin.
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<emphasis id="C1BD9D34FFCCFF9F00D4FA2CFF5CDABB" box="[173,236,1486,1509]" italics="true" pageId="77" pageNumber="1120">Frons</emphasis>
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with most of its surface slightly depressed.
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<emphasis id="C1BD9D34FFCCFF9F02AAFA2FFCD9DABB" box="[723,873,1485,1509]" italics="true" pageId="77" pageNumber="1120">Antennal club</emphasis>
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with glabrous area of basal segment about half of exposed surface.
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</paragraph>
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<paragraph id="F3764126FFCCFF9C0001F9ECFC85DF94" blockId="77.[79,1036,1103,1700]" lastBlockId="78.[79,1035,146,1373]" lastPageId="78" lastPageNumber="1121" pageId="77" pageNumber="1120">
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<emphasis id="C1BD9D34FFCCFF9F0001F9ECFF55D97B" box="[120,229,1550,1573]" italics="true" pageId="77" pageNumber="1120">Pronotum</emphasis>
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(®gures 29i, j, 33b). With large concavity occupying anterior median two-thirds of pronotum; concavity narrowed to posterior U- or V-shaped end (®gure 33b), sides steepest at posterior end; posteriorly with obsolete midline; each endpoint of the`U’ or`V’ feebly tuberculate; concavity with very ®ne punctures, interspersed with large punctures, the latter becoming increasingly more densely spacedÐbut not deeperÐand often conūent towards sides; concavity at its widest point about as wide as frons between eyes; lateral margin entire.
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</paragraph>
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<paragraph id="F3764126FFCFFF9C0001FF30FDEADE55" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
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<emphasis id="C1BD9D34FFCFFF9C0001FF30FF54DFB4" box="[120,228,210,234]" italics="true" pageId="78" pageNumber="1121">Scutellum</emphasis>
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(®gure 27d). Basal width about 1.5Ölength; with very ®ne punctures, interspersed with moderately sized punctures.
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</paragraph>
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<paragraph id="F3764126FFCFFF9C0001FEF6FCDDDE32" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
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<emphasis id="C1BD9D34FFCFFF9C0001FEF6FF7BDE72" box="[120,203,276,300]" italics="true" pageId="78" pageNumber="1121">Elytron</emphasis>
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. Striae moderately impressed, their punctures separated by two to three puncture diameters (®gure 27d); intervals with double punctation (with approximately four to ®ve ®ne and 20 minute punctures between two striae).
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</paragraph>
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<paragraph id="F3764126FFCFFF9C0001FE97FF6CDE90" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
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<emphasis id="C1BD9D34FFCFFF9C0001FE97FEBADED2" box="[120,266,373,396]" italics="true" pageId="78" pageNumber="1121">Metasternum</emphasis>
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. As in generic description but length of longitudinal suture variable, either reaching meso-metasternal suture (®gure 24a) or not reaching it (®gure 24d).
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||
</paragraph>
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<paragraph id="F3764126FFCFFF9C0001FE35FEDBDD11" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
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||
<emphasis id="C1BD9D34FFCFFF9C0001FE35FF1DDEB0" box="[120,173,471,494]" italics="true" pageId="78" pageNumber="1121">Legs</emphasis>
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.
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<emphasis id="C1BD9D34FFCFFF9C00B7FE34FE99DEB0" box="[206,297,470,494]" italics="true" pageId="78" pageNumber="1121">Protibia</emphasis>
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®ve-dentate.
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<emphasis id="C1BD9D34FFCFFF9C01AAFE34FDF0DEB0" box="[467,576,470,494]" italics="true" pageId="78" pageNumber="1121">Metatibia</emphasis>
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in lateral view elongate, only feebly broadening apically; with subapical carina distinctly bilobed; with toothlike projections arranged in two rows above subapical carina.
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<emphasis id="C1BD9D34FFCFFF9C02C3FDFAFC86DD71" box="[698,822,536,559]" italics="true" pageId="78" pageNumber="1121">Metatarsus</emphasis>
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lacking subapical setae on tarsomeres 2±4.
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</paragraph>
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<paragraph id="F3764126FFCFFF9C0001FDBAFC58DD2E" blockId="78.[79,1035,146,1373]" box="[120,1000,600,625]" pageId="78" pageNumber="1121">
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<emphasis id="C1BD9D34FFCFFF9C0001FDBAFEAADD2E" box="[120,282,600,624]" italics="true" pageId="78" pageNumber="1121">Male genitalia</emphasis>
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(®gure 29f±h). Aedeagus very large relative to the body size.
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</paragraph>
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<paragraph id="F3764126FFCFFF9C0001FD9BFC7BDDCE" blockId="78.[79,1035,146,1373]" box="[120,971,633,657]" pageId="78" pageNumber="1121">
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<emphasis id="C1BD9D34FFCFFF9C0001FD9BFF4CDDCF" box="[120,252,633,657]" italics="true" pageId="78" pageNumber="1121">Distribution</emphasis>
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(®gure 35). The species is known only from its
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<typeStatus id="2C72FF84FFCFFF9C034EFD9BFCD8DDCF" box="[823,872,633,657]" pageId="78" pageNumber="1121">type</typeStatus>
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locality.
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</paragraph>
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||
<paragraph id="F3764126FFCFFF9C0001FD78FE82DC6D" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
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||
<emphasis id="C1BD9D34FFCFFF9C0001FD78FE34DDEF" box="[120,388,665,689]" italics="true" pageId="78" pageNumber="1121">Type material examined</emphasis>
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.
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<materialsCitation id="43A14B7BFFCFFF9C01DFFD78FC37DD8C" collectingDate="1997-04-14" collectionCode="TMSA" collectorName="C. R. Owen'" country="South Africa" latitude="-30.27" location="Brandvlei" longLatPrecision="733" longitude="20.29" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="holotype">
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||
<typeStatus id="2C72FF84FFCFFF9C01DFFD78FDA9DDEF" box="[422,537,665,689]" pageId="78" pageNumber="1121" type="holotype">HOLOTYPE</typeStatus>
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||
: l`
|
||
<collectingCountry id="8BDE01B6FFCFFF9C0233FD7BFD6CDDEF" box="[586,732,665,689]" name="South Africa" pageId="78" pageNumber="1121">South Africa</collectingCountry>
|
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,
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||
<collectingRegion id="310D8FC4FFCFFF9C0295FD7BFBB2DDEF" box="[748,1026,665,689]" country="South Africa" name="Northern Cape" pageId="78" pageNumber="1121">Northern Cape Province</collectingRegion>
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,
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||
<location id="F61617FDFFCFFF9C0036FD58FF0CDD8C" LSID="urn:lsid:plazi:treatment:7B60F030FFCCFF9C01FAFC3CFC10DA03:F61617FDFFCFFF9C0036FD58FF0CDD8C" box="[79,188,698,722]" country="South Africa" latitude="-30.27" longLatPrecision="733" longitude="20.29" name="Brandvlei" pageId="78" pageNumber="1121" stateProvince="Northern Cape">Brandvlei</location>
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,
|
||
<geoCoordinate id="96FD27E1FFCFFF9C00B4FD58FEA8DD8C" box="[205,280,698,722]" degrees="30.27" direction="south" orientation="latitude" pageId="78" pageNumber="1121" precision="555" value="-30.27">30.27S</geoCoordinate>
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<geoCoordinate id="96FD27E1FFCFFF9C0158FD58FEDDDD8C" box="[289,365,698,722]" degrees="20.29" direction="east" orientation="longitude" pageId="78" pageNumber="1121" precision="555" value="20.29">20.29E</geoCoordinate>
|
||
, 12±
|
||
<date id="877767E6FFCFFF9C01D3FD58FDAADD8C" box="[426,538,698,722]" pageId="78" pageNumber="1121" value="1997-04-14">
|
||
<collectingDate id="97339E0EFFCFFF9C01D3FD58FDAADD8C" box="[426,538,698,722]" pageId="78" pageNumber="1121" value="1997-04-14">14.iv.1997</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="5E3C24F0FFCFFF9C0224FD58FD5BDD8C" box="[605,747,698,722]" pageId="78" pageNumber="1121">C. R. Owen’</collectorName>
|
||
(
|
||
<collectionCode id="95D8D9E3FFCFFF9C037BFD58FCE4DD8C" box="[770,852,698,722]" country="South Africa" name="Transvaal Museum" pageId="78" pageNumber="1121">TMSA</collectionCode>
|
||
); 13
|
||
</materialsCitation>
|
||
<materialsCitation id="43A14B7BFFCFFF9C03E8FD58FF6EDDAD" collectionCode="TMSA" country="South Africa" elevation="1" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="paratype">
|
||
<typeStatus id="2C72FF84FFCFFF9C03E8FD58FBB3DD8C" box="[913,1027,698,722]" pageId="78" pageNumber="1121" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<quantity id="3431ECC3FFCFFF9C0036FD39FFDBDDAC" box="[79,107,731,755]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">
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<elevation id="78E4A615FFCFFF9C0036FD39FFDBDDAC" box="[79,107,731,755]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">1m</elevation>
|
||
</quantity>
|
||
(
|
||
<collectionCode id="95D8D9E3FFCFFF9C00F8FD39FF66DDAD" box="[129,214,731,755]" country="South Africa" name="Transvaal Museum" pageId="78" pageNumber="1121">TMSA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
, 1, l
|
||
<materialsCitation id="43A14B7BFFCFFF9C0164FD39FE16DDAD" box="[285,422,731,755]" collectionCode="UPSA" country="South Africa" elevation="1" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="paratype">
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<quantity id="3431ECC3FFCFFF9C0164FD39FE89DDAC" box="[285,313,731,755]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">
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<elevation id="78E4A615FFCFFF9C0164FD39FE89DDAC" box="[285,313,731,755]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">1m</elevation>
|
||
</quantity>
|
||
(
|
||
<collectionCode id="95D8D9E3FFCFFF9C0136FD39FE2EDDAD" box="[335,414,731,755]" country="South Africa" name="University of Pretoria" pageId="78" pageNumber="1121">UPSA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
, 2 ll,
|
||
<quantity id="3431ECC3FFCFFF9C018EFD39FD91DDAC" box="[503,545,731,755]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="78" pageNumber="1121" unit="mm" value="2.0">2mm</quantity>
|
||
(COCS), 1, l
|
||
<materialsCitation id="43A14B7BFFCFFF9C02B5FD39FCD3DDAD" box="[716,867,731,755]" collectionCode="SMWN" country="South Africa" elevation="1" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="paratype">
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<quantity id="3431ECC3FFCFFF9C02B5FD39FD58DDAC" box="[716,744,731,755]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">
|
||
<elevation id="78E4A615FFCFFF9C02B5FD39FD58DDAC" box="[716,744,731,755]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">1m</elevation>
|
||
</quantity>
|
||
(
|
||
<collectionCode id="95D8D9E3FFCFFF9C0283FD39FCE5DDAD" box="[762,853,731,755]" country="Namibia" name="State Museum" pageId="78" pageNumber="1121">SMWN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="43A14B7BFFCFFF9C030DFD39FF48DC4D" collectingDate="1997-04-14" collectionCode="TMSA" collectorName="C. R. Owen'" country="South Africa" latitude="-30.27" location="Brandvlei" longLatPrecision="733" longitude="20.29" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="paratype">same data as holotype; 2 ll</materialsCitation>
|
||
,
|
||
<materialsCitation id="43A14B7BFFCFFF9C0175FD19FCA3DC4D" box="[268,787,763,787]" collectingDate="1997-04-14" collectionCode="AJCS" collectorName="A. Joubert" country="South Africa" elevation="1" latitude="-30.27" location="Brandvlei" longLatPrecision="733" longitude="20.29" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="paratype">
|
||
<quantity id="3431ECC3FFCFFF9C0175FD19FE97DC4C" box="[268,295,763,787]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">
|
||
<elevation id="78E4A615FFCFFF9C0175FD19FE97DC4C" box="[268,295,763,787]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="78" pageNumber="1121" unit="m" value="1.0">1m</elevation>
|
||
</quantity>
|
||
, same data but: leg.
|
||
<collectorName id="5E3C24F0FFCFFF9C0254FD19FD1CDC4D" box="[557,684,763,787]" pageId="78" pageNumber="1121">A. Joubert</collectorName>
|
||
(
|
||
<collectionCode id="95D8D9E3FFCFFF9C02BDFD19FCB6DC4D" box="[708,774,763,787]" pageId="78" pageNumber="1121">AJCS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="43A14B7BFFCFFF9C035FFD19FE9DDC6D" collectingDate="1997-03-30" collectionCode="COCS" collectorName="A. Joubert" country="South Africa" elevation="1" latitude="-30.27" location="Brandvlei" longLatPrecision="733" longitude="20.29" pageId="78" pageNumber="1121" specimenCount="1" stateProvince="Northern Cape" typeStatus="paratype">
|
||
1, l same data but:
|
||
<date id="877767E6FFCFFF9C0036FCF9FF73DC6D" box="[79,195,795,819]" pageId="78" pageNumber="1121" value="1997-03-30">
|
||
<collectingDate id="97339E0EFFCFFF9C0036FCF9FF73DC6D" box="[79,195,795,819]" pageId="78" pageNumber="1121" value="1997-03-30">30.iii.1997</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="95D8D9E3FFCFFF9C00ADFCF9FEAFDC6D" box="[212,287,795,819]" pageId="78" pageNumber="1121">COCS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F3764126FFCFFF9C0001FCDEFDAFDC2A" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
|
||
<emphasis id="C1BD9D34FFCFFF9C0001FCDEFF5FDC0A" box="[120,239,828,852]" italics="true" pageId="78" pageNumber="1121">Etymology</emphasis>
|
||
. The species is named after emeritus Prof. Erik Holm who did most of the habitus illustrations in this paper.
|
||
</paragraph>
|
||
<paragraph id="F3764126FFCFFF9C0001FC9FFD43DC88" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
|
||
<emphasis id="C1BD9D34FFCFFF9C0001FC9FFF5CDCCB" box="[120,236,893,917]" italics="true" pageId="78" pageNumber="1121">Comments</emphasis>
|
||
. In external morphology,
|
||
<taxonomicName id="34C93AA5FFCFFF9C0253FC9FFCB1DCCB" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[554,769,893,917]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="1121" phylum="Arthropoda" rank="species" species="holmi">
|
||
<emphasis id="C1BD9D34FFCFFF9C0253FC9FFCB1DCCB" box="[554,769,893,917]" italics="true" pageId="78" pageNumber="1121">Bolboceratex holmi</emphasis>
|
||
</taxonomicName>
|
||
agrees with the general generic characteristics and is thus justi®ably placed in
|
||
<taxonomicName id="34C93AA5FFCFFF9C036DFC7CFC15DCE8" box="[788,933,926,950]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="1121" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C1BD9D34FFCFFF9C036DFC7CFC15DCE8" box="[788,933,926,950]" italics="true" pageId="78" pageNumber="1121">Bolboceratex</emphasis>
|
||
</taxonomicName>
|
||
. Sexual dimorphism is absent and intraspeci®c variability minimal.
|
||
</paragraph>
|
||
<paragraph id="F3764126FFCFFF9C0001FC3CFC10DA03" blockId="78.[79,1035,146,1373]" pageId="78" pageNumber="1121">
|
||
The new species can only be confused with the sympatric
|
||
<taxonomicName id="34C93AA5FFCFFF9C036DFC3CFBB4DCA8" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[788,1028,990,1014]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="1121" phylum="Arthropoda" rank="species" species="jouberti">
|
||
<emphasis id="C1BD9D34FFCFFF9C036DFC3CFBB4DCA8" box="[788,1028,990,1014]" italics="true" pageId="78" pageNumber="1121">Bolboceratex jouberti</emphasis>
|
||
</taxonomicName>
|
||
. Reliable external morphological characters to distinguish the two species are the outer margin of the mandible, i.e. rounded in the latter but distinctly sinuate in the former species, the metatibia in lateral view which is elongate and only feebly broadening towards apex in the latter species but distinctly broadening in the former species. The metatibial armature also diOEers between the two species.
|
||
<figureCitation id="6BF25DA3FFCFFF9C031CFB60FC59DBC4" box="[869,1001,1154,1178]" captionStart="FIG" captionStartId="61.[79,95,1415,1436]" captionTargetBox="[272,841,148,1387]" captionTargetId="figure-0@61.[269,844,143,1389]" captionTargetPageId="61" captionText="FIG. 29. Bolboceratex jouberti: aedeagus, ventral (a), dorsal (b), lateral view (c); pronotum male, lateral (d) and frontal view (e). B. holmi: aedeagus, ventral (f), dorsal (g), lateral view (h); pronotum male, lateral (i) and frontal view (j)." figureDoi="http://doi.org/10.5281/zenodo.4748073" httpUri="https://zenodo.org/record/4748073/files/figure.png" pageId="78" pageNumber="1121">Figures 29a</figureCitation>
|
||
±c, 29f±h clearly illustrate the diOEerences of the aedeagi in size and structure. While
|
||
<taxonomicName id="34C93AA5FFCFFF9C0036FB21FEF9DB85" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[79,329,1219,1243]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="1121" phylum="Arthropoda" rank="species" species="jouberti">
|
||
<emphasis id="C1BD9D34FFCFFF9C0036FB21FEF9DB85" box="[79,329,1219,1243]" italics="true" pageId="78" pageNumber="1121">Bolboceratex jouberti</emphasis>
|
||
</taxonomicName>
|
||
’s genitalia have the more typical shape for the genus,
|
||
<taxonomicName id="34C93AA5FFCFFF9C0036FB01FE98DBA5" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[79,296,1251,1275]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="1121" phylum="Arthropoda" rank="species" species="holmi">
|
||
<emphasis id="C1BD9D34FFCFFF9C0036FB01FE98DBA5" box="[79,296,1251,1275]" italics="true" pageId="78" pageNumber="1121">Bolboceratex holmi</emphasis>
|
||
</taxonomicName>
|
||
has shortened parameres. The aedeagus of
|
||
<taxonomicName id="34C93AA5FFCFFF9C0363FB01FC42DBA5" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" box="[794,1010,1251,1275]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="1121" phylum="Arthropoda" rank="species" species="holmi">
|
||
<emphasis id="C1BD9D34FFCFFF9C0363FB01FC42DBA5" box="[794,1010,1251,1275]" italics="true" pageId="78" pageNumber="1121">Bolboceratex holmi</emphasis>
|
||
</taxonomicName>
|
||
is the most modi®ed within the genus but does share the strongly sclerotized parameres, armed with toothlike projections, with its congeneric species. A pair of strongly sclerotized, elongate projections as part of the internal sac is also present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |