146 lines
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146 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.225.3721" ID-GBIF-Dataset="1301ded3-c86d-4e07-aae6-dd84e348cd25" ID-PMC="PMC3487652" ID-Pensoft-Pub="1313-2970-225-1" ID-PubMed="23166461" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-225-1" ModsDocOrigin="ZooKeys 225" ModsDocTitle="A revision of the genus Planinasus Cresson (Diptera, Periscelididae)" checkinTime="1451248711591" checkinUser="pensoft" docAuthor="Mathis, Wayne N., Rung, Alessandra & Kotrba, Marion" docDate="2012" docId="3F44678C60AB3BE59AD5E71F7D0F778D" docLanguage="en" docName="ZooKeys 225: 1-83" docOrigin="ZooKeys 225" docSource="http://dx.doi.org/10.3897/zookeys.225.3721" docTitle="Planinasus aenigmaticus Mathis, Rung & Kotrba, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="5C00FF9EFFDAFF805469564E9829FF97" masterDocTitle="A revision of the genus Planinasus Cresson (Diptera, Periscelididae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="11" updateTime="1668154510379" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the genus Planinasus Cresson (Diptera, Periscelididae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rung, Alessandra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kotrba, Marion</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>225</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.225.3721</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.225.3721</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-225-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152037215" LSID="urn:lsid:zoobank.org:act:707380A1-90E5-417C-88E3-C30704091CF1" httpUri="http://treatment.plazi.org/id/3F44678C60AB3BE59AD5E71F7D0F778D" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<subSubSection pageId="10" pageNumber="11" type="nomenclature">
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<paragraph pageId="10" pageNumber="11">
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1.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:707380A1-90E5-417C-88E3-C30704091CF1" class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus aenigmaticus" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aenigmaticus">Planinasus aenigmaticus</taxonomicName>
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<taxonomicNameLabel pageId="10" pageNumber="11">sp. n.</taxonomicNameLabel>
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Figures 1-5
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
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<paragraph pageId="10" pageNumber="11">Description.</paragraph>
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<paragraph pageId="10" pageNumber="11">Moderately small flies, body length 2.70 mm (holotype).</paragraph>
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<paragraph lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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Head: Frons mostly bare, shiny, except for more densely microtomentose, anterolateral angles; frons wider than long, frontal ratio 0.50-0.58; interfrontal seta moderately short, about 2/3 length of lateral vertical seta. Antenna unicolorous,
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<pageBreakToken pageId="11" pageNumber="12" start="start">blackish</pageBreakToken>
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brown; pedicel with ventral projection short, not extended anteriorly much beyond dorsal margin; basal flagellomere short, about as long as high, mostly yellow (faintly black around base of arista); arista bearing 8-9 dorsal rays, 3-4 ventral rays. Face comparatively narrow; facial ratio 0.24-0.39; dorsal half of face shield-like, mostly bare, roughly pentagonal, with yellowish to slightly bluish reflections; large
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<pageBreakToken pageId="12" pageNumber="13" start="start">facial</pageBreakToken>
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setae arranged in 2-3 irregular, transverse rows, dorsal row with 4 setae, 2 dorsoclinate, 2 ventroclinate; ventral facial row with 4 ventroclinate setae. Clypeus and palpus blackish brown.
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</paragraph>
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<paragraph pageId="12" pageNumber="13">Thorax: Generally dark brown but with some paler, mostly yellowish areas along margins of sclerites; mesonotum moderately invested with whitish gray microtomentum, appearing dull medially, becoming subshiny, less microtomentose laterally; postpronotum yellowish; area from postpronotum and through notopleuron mostly bare, shiny; anepisternum moderately invested with very fine microtomentum, mostly appearing dull; other pleural areas less densely invested. Wing mostly hyaline to very faintly infumate, without pattern. Coxae whitish yellow; femora whitish yellow on basal 2/3, only distal 1/3 dark brown; forefemur bearing 1-3 subequal, posteroventral setae at apical 1/3; lacking a preapical annulus; tibiae with base and apex yellow, middle 2/3 brownish black; basitarsomere yellowish, apical 3-4 tarsomeres darker; forefemur bearing 2 setae at apical 1/3 along posteroventral surface.</paragraph>
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<paragraph lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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Abdomen: Uniformly blackish brown, mostly subshiny, moderately invested with microtomentum. Male abdomen: Tergites 1+2-6 well developed, lengths of tergites 3-6 subequal; sternite 3 only slightly wider than long, posterior margin sinuous with a medial, broad, rounded projection; width of sternite 4 over twice length, sclerotized portion moderately deeply emarginate with broad, V-shaped membranous area posteriorly; width of sternite 5 over twice length, becoming slightly wider pos
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<pageBreakToken pageId="13" pageNumber="14" start="start">teriorly</pageBreakToken>
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, posterior margin shallowly emarginate, bearing 1-2 setulae at posterolateral corners; sternite 6 apparently absent; sternite 7 narrow, forming an annulus with tergite 7. Male terminalia (Figs 1-4): Epandrium in lateral view (Fig. 1) triangular to trapezoidal, dorsal margin narrowly angulate; surstylar length slightly less than half length of epandrium, surstylus extended from ventral margin of epandrium in nearly vertical alignment with anterior margin of it, in lateral and ventral views (Figs 1-2) digitiform, apex bluntly rounded to truncate, length approximately twice width, bearing 4 strong posteromedial setulae and a few smaller setulae along posterior and apical portions; hypandrium in ventral view (Fig. 2) U-shaped with anterior portion of U with a short, trapezoidal projection; pregonite in ventral view (Fig. 2) approximately triangular, with anterior margin receded; postgonite in ventral view convoluted (Fig. 2), apex bearing narrow subrectangular projection, with robustly developed, clavate lobe bearing more than 20 setulae, in lateral view (Fig. 4) with lobe moderately expanded, broadly rounded apically, bearing numerous setulae; phallus large, in ventral and lateral views (Figs 2, 4) complex, mostly sclerotized; phallapodeme in lateral and ventral views (Figs 2, 4) elongate, tubular, truncate apically; ejaculatory apodeme in lateral view (Fig. 3) large, about 1/3 longer than length of phallapodeme, apical half greatly expanded, fan-like, basal half narrow, stem-like but slightly expanded toward base.
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</paragraph>
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<caption pageId="13" pageNumber="14">
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<paragraph pageId="13" pageNumber="14">
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Figures 1-4 Illustrations of
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus aenigmaticus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigmaticus">Planinasus aenigmaticus</taxonomicName>
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sp. n. (male). 1 epandrium, surstylus, hypandrium, lateral view 2 epandrium, hypandrium, and internal structures of male terminalia, ventral view 3 ejaculatory apodeme, lateral view 4 internal structures of male terminalia, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="13" pageNumber="14">
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<paragraph pageId="13" pageNumber="14">
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Figure 5. Distribution of
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus aenigmaticus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigmaticus">Planinasus aenigmaticus</taxonomicName>
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sp. n. (squares) and
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus shannoni" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="shannoni">Planinasus shannoni</taxonomicName>
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(Malloch) (dots).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="type material">
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<paragraph pageId="13" pageNumber="14">Type material.</paragraph>
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<paragraph pageId="13" pageNumber="14">
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The holotype male is labeled "W of Bogota COLOMBIA/M R Wheeler collector/USNM ENT 00118284 [plastic bar code label]/HOLOTYPE ♂
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus aenigmaticus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigmaticus">Planinasus aenigmaticus</taxonomicName>
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Mathis & Rung USNM [red]." The holotype is double mounted (glued to a paper point), is in good condition (abdomen removed, dissected, stored in an attached microvial), and is deposited in the USNM. A male paratype (CAS) is as follows: COLOMBIA. Valle del Cauca:
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Raposo (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="3.7166667">03°43'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-77.13333">77°08'W</geoCoordinate>
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), Oct 1964, V. H. Lee.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="type locality">
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<paragraph pageId="13" pageNumber="14">Type locality.</paragraph>
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<paragraph pageId="13" pageNumber="14">
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Colombia. Bogota: Bogota (west;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="4.596667">04°35.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-74.14667">74°08.8'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="distribution">
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<paragraph pageId="13" pageNumber="14">Distribution</paragraph>
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<paragraph pageId="13" pageNumber="14">
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(Fig. 5).
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<taxonomicName genus="Neotropical" lsidName="Neotropical" pageId="13" pageNumber="14" rank="genus">Neotropical</taxonomicName>
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: Colombia (Bogota, Valle del Cauca).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="etymology">
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<paragraph pageId="13" pageNumber="14">Etymology.</paragraph>
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<paragraph pageId="13" pageNumber="14">The specific epithet, aenigmaticus, is of Greek derivation, meaning something baffling or puzzling, and refers to the puzzling nature of this species, as well as the genus.</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="remarks">
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<paragraph pageId="13" pageNumber="14">Remarks.</paragraph>
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<paragraph pageId="13" pageNumber="14">
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This species, represented by two males only, is known only from Colombia and is the only species of
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Planinasus</taxonomicName>
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known to occur in that country. It is very similar to
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus neotropicus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neotropicus">Planinasus neotropicus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus ambiguous" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ambiguous">Planinasus ambiguous</taxonomicName>
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, and analysis of the male genitalia is generally necessary to distinguish among the three species. Characters of the male terminalia of
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus ambiguous" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ambiguous">Planinasus ambiguous</taxonomicName>
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are very distinct, particularly the shape of the epandrium (Fig. 10), the length and shape of the surstylus (Fig. 9), and the uniform, U-shaped hypandrium. From
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus neotropicus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neotropicus">Planinasus neotropicus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus aenigmaticus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigmaticus">Planinasus aenigmaticus</taxonomicName>
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can be easily distinguished by the surstylus, digitiform over its entire length and with about four posteromedial setulae (Fig. 1), and the U-shaped hypandrium with the anterior margin bearing a short, trapezoidal projection (Fig. 2). In
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<taxonomicName class="Insecta" family="Periscelididae" genus="Planinasus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planinasus neotropicus" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neotropicus">Planinasus neotropicus</taxonomicName>
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, the surstylus is bulged basally, there is only one medial seta on it (Fig. 15), and the hypandrium is V-shaped (Fig. 16).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |