749 lines
96 KiB
XML
749 lines
96 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.947.53906" ID-GBIF-Dataset="20958e7a-dd5a-4363-afc3-b1cd8186cf46" ID-PMC="PMC7363738" ID-Pensoft-Pub="1313-2970-947-39" ID-Pensoft-UUID="E27A1F2F6D9150EB83BE9E7552BD32CA" ID-PubMed="32733128" ID-ZooBank="1A80132DD67949C090D1069770305473" ModsDocID="1313-2970-947-39" checkinTime="1594285666152" checkinUser="pensoft" docAuthor="Sidabalok, Conni M., Wong, Helen P. - S. & Ng, Peter K. L." docDate="2020" docId="2C1A752EBD93579E8A09338DED303E6C" docLanguage="en" docName="ZooKeys 947: 39-52" docOrigin="ZooKeys 947" docSource="http://dx.doi.org/10.3897/zookeys.947.53906" docTitle="Bathynomus raksasa Sidabalok & Wong & Ng 2020, sp. nov." docType="treatment" docUuid="84D71359-90FB-4CC6-856F-B96402F23211" docUuidSource="ZooBank" docVersion="4" id="E27A1F2F6D9150EB83BE9E7552BD32CA" lastPageNumber="39" masterDocId="E27A1F2F6D9150EB83BE9E7552BD32CA" masterDocTitle="Description of the supergiant isopod Bathynomus raksasa sp. nov. (Crustacea, Isopoda, Cirolanidae) from southern Java, the first record of the genus from Indonesia" masterLastPageNumber="52" masterPageNumber="39" pageNumber="39" updateTime="1668169071850" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of the supergiant isopod Bathynomus raksasa sp. nov. (Crustacea, Isopoda, Cirolanidae) from southern Java, the first record of the genus from Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sidabalok, Conni M.</mods:namePart>
|
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<mods:affiliation>Division of Zoology, Research Center for Biology, Indonesian Institute of Sciences (LIPI), Gedung Widyasatwaloka, Cibinong Science Center, Jl Raya Jakarta-Bogor Km 46, Cibinong 16911, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wong, Helen P. - S.</mods:namePart>
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<mods:affiliation>St. John's Island National Marine Laboratory, Tropical Marine Science Institute, National University of Singapore (NUS), 18 Kent Ridge Road, 119227, Singapore</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart>Ng, Peter K. L.</mods:namePart>
|
||
<mods:affiliation>Lee Kong Chian Natural History Museum (LKCNHM), 2 Conservatory Drive, National University of Singapore, Singapore 117377, Singapore</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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||
<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>947</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>52</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.947.53906</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.947.53906</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-947-39</mods:identifier>
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<mods:identifier type="ZooBank">1A80132DD67949C090D1069770305473</mods:identifier>
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<mods:identifier type="Pensoft-UUID">E27A1F2F6D9150EB83BE9E7552BD32CA</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164986667" LSID="urn:lsid:zoobank.org:act:84D71359-90FB-4CC6-856F-B96402F23211" httpUri="http://treatment.plazi.org/id/2C1A752EBD93579E8A09338DED303E6C" lastPageNumber="39" pageId="0" pageNumber="39">
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<subSubSection pageId="0" pageNumber="39" type="nomenclature">
|
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<paragraph pageId="0" pageNumber="39">
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||
<taxonomicName LSID="2C1A752E-BD93-579E-8A09-338DED303E6C" authority="Sidabalok & Wong & Ng, 2020" authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa" status="sp. nov.">Bathynomus raksasa</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="39">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">Figs 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">, 4</figureCitation>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">, 5</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Holotype</emphasis>
|
||
, male, 363 mm; Indonesia, Sunda Strait (between Sumatra and Java);
|
||
<geoCoordinate degrees="6" direction="south" minutes="00.828" orientation="latitude" precision="1" value="-6.0138">6°00.828'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="49.428" orientation="longitude" precision="1" value="104.8238">104°49.428'E</geoCoordinate>
|
||
; 26 Mar. 2018; SJADES exped.; station CP 13, beam trawl 1259 m; MZB Cru.Iso 097.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Paratype</emphasis>
|
||
, female, 298 mm; Indonesia, Indian Ocean (East of Tinjil Island);
|
||
<geoCoordinate degrees="6" direction="south" minutes="59.778" orientation="latitude" precision="1" value="-6.9962997">6°59.778'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="105" direction="east" minutes="55.224" orientation="longitude" precision="1" value="105.9204">105°55.224'E</geoCoordinate>
|
||
; 28 Mar. 2018; SJADES exped.; station CP 28, beam trawl 957 m; ZRC 2020.0015.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="39">Comparative material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<taxonomicName class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus giganteus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus giganteus</emphasis>
|
||
</taxonomicName>
|
||
A. Milne-Edwards, 1879 - 1 male, 354 mm; U.S.A., Virginia, 100 miles off Virginia Beach; 36.483N, 74.8W; 30 May 1962; 73 m depth; ZRC 2014.0837.
|
||
<taxonomicName authorityName="Ortmann" authorityYear="1894" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus doederleini" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus doederleini</emphasis>
|
||
</taxonomicName>
|
||
Ortmann, 1894 - 6 males, 100, 120, 120, 128, 136, 145 mm; 7 females, 88, 90, 94, 130, 130, 138, 145 mm; 3 juveniles; Taiwan; AM P68684. 1 male, 125 mm; 1 female, 85 mm; 4 juveniles; Taiwan, Tashi port; 1990s; P. K. L. Ng leg.; deep-water; ZRC 1998.417.
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. - 1 subadult, not sexually mature, pereopod 7 not fully developed, 107 mm; Indonesia, Indian Ocean (East of Tinjil Island);
|
||
<geoCoordinate degrees="6" direction="south" minutes="56.664" orientation="latitude" precision="1" value="-6.9444">6°56.664'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="105" direction="east" minutes="55.315" orientation="longitude" precision="1" value="105.92191">105°55.315'E</geoCoordinate>
|
||
; 28 Mar. 2018; SJADES exped.; station CP 26, beam trawl 517 m; MZB Cru.Iso 098. 1 juvenile; Indonesia, Sunda Strait (between Tabuan Island and Sumatra);
|
||
<geoCoordinate degrees="5" direction="south" minutes="45.126" orientation="latitude" precision="1" value="-5.7521">5°45.126'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="51.080" orientation="longitude" precision="1" value="104.85133">104°51.080'E</geoCoordinate>
|
||
; 25 Mar. 2018; SJADES exped.; station CP 08, beam trawl 442 m; ZRC 2020.0016. 2 juveniles, 60, 63 mm; Indonesia, Indian Ocean (Pelabuhan Ratu Bay);
|
||
<geoCoordinate degrees="7" direction="south" minutes="01.116" orientation="latitude" precision="1" value="-7.0186">7°01.116'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="106" direction="east" minutes="26.421" orientation="longitude" precision="1" value="106.44035">106°26.421'E</geoCoordinate>
|
||
; 3 Apr. 2018; SJADES exped.; station CP 55, beam trawl 379 m; ZRC 2020.0017.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="type-locality">
|
||
<paragraph pageId="0" pageNumber="39">Type-locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Indonesia, Sunda Strait: between Sumatra and Java,
|
||
<geoCoordinate degrees="06" direction="south" minutes="00.828" orientation="latitude" precision="1" value="-6.0138">06°00.828'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="49.428" orientation="longitude" precision="1" value="104.8238">104°49.428'E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Narrowly rounded clypeus apex (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2C</figureCitation>
|
||
); prominent longitudinal carina on clypeus (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2C</figureCitation>
|
||
); convex lateral margins of uropodal exopod and endopod (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3D, E</figureCitation>
|
||
); produced distolateral corners of uropodal exopod and endopod with acute tips (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3D, E</figureCitation>
|
||
); uropodal exopod with medium-length setal fringe (69%) (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3D, E</figureCitation>
|
||
); pleotelson 1.6 times wider than long with posterior margin medially concave (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2D</figureCitation>
|
||
); 11-13 spines on pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<emphasis bold="true" pageId="0" pageNumber="39">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (363 mm) (MZB Cru.Iso 097), Indonesia
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
cephalon, anterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
clypeal region
|
||
<emphasis bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
pleotelson
|
||
<emphasis bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
body, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">F</emphasis>
|
||
pereon, lateral view. Scale bars: 5 cm (
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A, D, E</emphasis>
|
||
);1 cm (
|
||
<emphasis bold="true" pageId="0" pageNumber="39">B, C, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="description">
|
||
<paragraph pageId="0" pageNumber="39">Description of holotype male.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Body (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2A</figureCitation>
|
||
) 363 mm long, 155 mm wide at pereonite 5, length 2.3 times width. Head (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2B</figureCitation>
|
||
) with ridge above eyes discontinuous; clypeus (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2C</figureCitation>
|
||
) with prominent longitudinal carina, distal margins slightly concave, apex narrowly rounded.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Antenna 2 (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2A, E</figureCitation>
|
||
) flagellum extending to end of pleonite 2.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Pereopod 1 (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3A</figureCitation>
|
||
) ischium with 2 posteroproximal robust setae, 2 robust setae on posterodistal margin; merus with 4 short robust setae on anterodistal angle, posterior margin with 4 robust setae in proximal row and 2 robust setae in distal row; propodus length 2.3 times width, with 5 robust setae on posterior margin. Pereopod 2 (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3B, C</figureCitation>
|
||
) ischium with 3 robust setae on posterior margin and 2 robust setae on posterodistal margin; merus with 7 short robust setae on anterodistal angle, posteromedial margin with 3 robust setae in proximal row and 2 robust setae in distal row; propodus with 4 robust setae on posterior margin. Pereopod 7 coxa (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2F</figureCitation>
|
||
) distally attenuated, curved posteriorly.
|
||
</paragraph>
|
||
<caption doi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<emphasis bold="true" pageId="0" pageNumber="39">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (363 mm) (MZB Cru.Iso 097), Indonesia
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
pereopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
pereopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
pereopod 2 merus, posterolateral margin
|
||
<emphasis bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
uropod, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
uropod, dorsal view. Scale bars: 1 cm (
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A, B, D, E</emphasis>
|
||
); 0.5 cm (
|
||
<emphasis bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Pleonite 3 (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2F</figureCitation>
|
||
) not extending beyond pleonite 5.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Uropod (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3D, E</figureCitation>
|
||
) not extending beyond pleotelson; peduncle with 3 robust setae; exopod and endopod with smooth lateral and distal margins; exopodal lateral margin convex with 10 robust setae along margin, setal fringe medium to continuous in length (69%), medial margin straight, distomedial corner rounded, distal margin convex with 5 robust setae, distolateral corner slightly produced, acute; endopodal lateral margin convex, distally sinuate, with 4 robust setae; medial margin straight; distomedial corner rounded; distal margin straight with 11 robust setae; distolateral corner produced, acute.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A dorsal view B cephalon, anterior view C clypeal region D pleotelson E body, lateral view F pereon, lateral view. Scale bars: 5 cm (A, D, E); 1 cm (B, C, F)." figureDoi="/10.3897/zookeys.947.53906.figure2" httpUri="https://binary.pensoft.net/fig/430114" pageId="0" pageNumber="39">2D</figureCitation>
|
||
) broader than long, 1.6 times as wide as length, posterior margin medially concave, smooth (minute pores), conspicuous longitudinal carina on dorsal surface, with 11 distal and 2 lateral straight acute prominent spines along distal margin, without setae between spines, central distal spine simple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="female">
|
||
<paragraph pageId="0" pageNumber="39">Female.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">Similar to male.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="variation">
|
||
<paragraph pageId="0" pageNumber="39">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">Robust setae count on female as follows: exopodal lateral margin with 7-10 robust setae, distal margin with 4 or 5, endopodal lateral margin with 3-5 and distal margin with 8-10; pleotelson with 9 distal and 2 lateral straight acute prominent spines along distal margin.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="etymology">
|
||
<paragraph pageId="0" pageNumber="39">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
The epithet is the Indonesian word "
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">raksasa</taxonomicName>
|
||
" for giant, alluding to its enormous size and the significance of the find. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="remarks">
|
||
<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be readily identified by its large size (330 mm on average), narrowly rounded clypeus apex, produced and acute distolateral corners of uropodal rami, wider rather than long pleotelson with medially concave posterior margin and the presence of 11-13 pleotelson spines.
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is the sixth
|
||
<normalizedToken originalValue="“supergiant”">"supergiant"</normalizedToken>
|
||
species from the Indo-West Pacific and is one of the largest known members of the genus.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
In general appearance,
|
||
<taxonomicName lsidName="B. raksasa" pageId="0" pageNumber="39" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="B. lowryi" pageId="0" pageNumber="39" rank="species" species="lowryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. lowryi</emphasis>
|
||
</taxonomicName>
|
||
. All three are large, averaging 300 mm in length, possess a prominent longitudinal carina on the dorsal surface of the pleotelson and have acute spines on the distal margin of the pleotelson. The new species is closest to
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
, sharing the relatively medium length of antenna 2 (reaching to between the posterior of pereonite 2 and anterior of pereonite 3), lateral and posterior shape of the uropodal exopod and endopod, and the pleotelson spine count.
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., however, differs markedly from
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
by its more conspicuous longitudinal carina on the clypeus ventral surface (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">4A</figureCitation>
|
||
) (vs. less conspicuous in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">4B</figureCitation>
|
||
), absence of a transverse carina on the anterior of the head (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">4C</figureCitation>
|
||
) (vs. carina present in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">4D</figureCitation>
|
||
), the relatively shorter uropodal endopod (0.12 total body length, Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">4E</figureCitation>
|
||
) (vs. relatively longer, 0.15 body length in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A, C, E Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B clypeus ventral surface C, D anterior of head E, F body, ventral view." figureDoi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39">4F</figureCitation>
|
||
), the body surface, including that of the pleotelson, being covered with small low granules and smooth to the touch (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5A</figureCitation>
|
||
) (vs. granules more prominent and the surfaces distinctly rough in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5B</figureCitation>
|
||
), the almost flat posterior ventral surface of the pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5C</figureCitation>
|
||
) (vs. surface distinctly concave in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5D</figureCitation>
|
||
), the straight spines of pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5E</figureCitation>
|
||
) (vs. gently curved upwards in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5F</figureCitation>
|
||
), the pleotelson is broader than long (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5A</figureCitation>
|
||
) (vs. as long as broad in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5B</figureCitation>
|
||
), and the posterior margin of the pleotelson is broad and medially concave (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5A</figureCitation>
|
||
) (vs. broadly rounded in
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5B</figureCitation>
|
||
).
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can easily be distinguished from
|
||
<taxonomicName lsidName="B. lowryi" pageId="0" pageNumber="39" rank="species" species="lowryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. lowryi</emphasis>
|
||
</taxonomicName>
|
||
in possessing a relatively longer antenna 2 which reaches to the ends of pereonite 2 (vs. shorter antenna 2 which reaches only to the anterior part of pereonite 2 in
|
||
<taxonomicName lsidName="B. lowryi" pageId="0" pageNumber="39" rank="species" species="lowryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. lowryi</emphasis>
|
||
</taxonomicName>
|
||
), the narrowly rounded clypeus apex (vs. apex truncate in
|
||
<taxonomicName lsidName="B. lowryi" pageId="0" pageNumber="39" rank="species" species="lowryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. lowryi</emphasis>
|
||
</taxonomicName>
|
||
), straight pleotelson spines (vs. spines upwardly curved in
|
||
<taxonomicName lsidName="B. lowryi" pageId="0" pageNumber="39" rank="species" species="lowryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. lowryi</emphasis>
|
||
</taxonomicName>
|
||
) and the larger number (13) of robust setae on the pleotelson (vs. 9 in
|
||
<taxonomicName lsidName="B. lowryi" pageId="0" pageNumber="39" rank="species" species="lowryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. lowryi</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Bruce, NL" journalOrPublisher="Phuket Marine Biological Center, Research Bulletin" pageId="0" pageNumber="39" pagination="1 - 8" refId="B4" refString="Bruce, NL, Bussarawit, S, 2004. Bathynomus lowryi sp. nov. (Crustacea: Isopoda: Cirolanidae), the first record of the 'giant' marine isopod genus from Thailand waters. Phuket Marine Biological Center, Research Bulletin 65: 1 - 8" title="Bathynomus lowryi sp. nov. (Crustacea: Isopoda: Cirolanidae), the first record of the ' giant' marine isopod genus from Thailand waters." volume="65" year="2004">Bruce and Bussarawit 2004</bibRefCitation>
|
||
: figs 1, 6).
|
||
</paragraph>
|
||
<caption doi="/10.3897/zookeys.947.53906.figure4" httpUri="https://binary.pensoft.net/fig/430116" pageId="0" pageNumber="39" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<emphasis bold="true" pageId="0" pageNumber="39">Figure 4.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A, C, E</emphasis>
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype male (363 mm) (MZB Cru.Iso 097), Indonesia
|
||
<emphasis bold="true" pageId="0" pageNumber="39">B, D, F</emphasis>
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
male (354 mm) (ZRC 2014.0837), Caribbean
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A, B</emphasis>
|
||
clypeus ventral surface
|
||
<emphasis bold="true" pageId="0" pageNumber="39">C, D</emphasis>
|
||
anterior of head
|
||
<emphasis bold="true" pageId="0" pageNumber="39">E, F</emphasis>
|
||
body, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. shares the same general uropodal exopod and endopod shape as
|
||
<taxonomicName lsidName="B. crosnieri" pageId="0" pageNumber="39" rank="species" species="crosnieri">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. crosnieri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="B. kensleyi" pageId="0" pageNumber="39" rank="species" species="kensleyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. kensleyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="B. richeri" pageId="0" pageNumber="39" rank="species" species="richeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. richeri</emphasis>
|
||
</taxonomicName>
|
||
but can easily be distinguished from them in its possession of a conspicuous longitudinal carina on the dorsal surface of the pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5A</figureCitation>
|
||
). Although the number of spines on the margin of the pleotelson (at least 11) is similar to those of
|
||
<taxonomicName lsidName="B. crosnieri" pageId="0" pageNumber="39" rank="species" species="crosnieri">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. crosnieri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="B. richeri" pageId="0" pageNumber="39" rank="species" species="richeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. richeri</emphasis>
|
||
</taxonomicName>
|
||
, the presence of the longitudinal ridge on the pleotelson easily separates
|
||
<taxonomicName lsidName="B. raksasa" pageId="0" pageNumber="39" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from these species.
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. also has the same number of spines on the margin of the pleotelson but can easily be distinguished from
|
||
<taxonomicName lsidName="B. keablei" pageId="0" pageNumber="39" rank="species" species="keablei">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. keablei</emphasis>
|
||
</taxonomicName>
|
||
in having the distolateral corners of the uropodal exopod and endopod distinctly produced (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bathynomus raksasa sp. nov., holotype male (363 mm) (MZB Cru. Iso 097), Indonesia A pereopod 1 B pereopod 2 C pereopod 2 merus, posterolateral margin D uropod, ventral view E uropod, dorsal view. Scale bars: 1 cm (A, B, D, E); 0.5 cm (C)." figureDoi="/10.3897/zookeys.947.53906.figure3" httpUri="https://binary.pensoft.net/fig/430115" pageId="0" pageNumber="39">3D, E</figureCitation>
|
||
) (vs. rounded and not produced in
|
||
<taxonomicName lsidName="B. keablei" pageId="0" pageNumber="39" rank="species" species="keablei">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. keablei</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 17).
|
||
</paragraph>
|
||
<caption doi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<emphasis bold="true" pageId="0" pageNumber="39">Figure 5.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A, C, E, G</emphasis>
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype male (363 mm) (MZB Cru.Iso 097), Indonesia
|
||
<emphasis bold="true" pageId="0" pageNumber="39">B, D, F, H</emphasis>
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
male (354 mm) (ZRC 2014.0837), Caribbean
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A, B</emphasis>
|
||
pleotelson dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">C, D</emphasis>
|
||
pleotelson posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">E, F</emphasis>
|
||
pleotelson lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">G, H</emphasis>
|
||
pleopod 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
The appendix masculina is absent on the holotype male of
|
||
<taxonomicName lsidName="B. raksasa" pageId="0" pageNumber="39" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5G</figureCitation>
|
||
) but this is almost certainly not a species-character. It is known to be sometimes absent in
|
||
<taxonomicName lsidName="B. doederleini" pageId="0" pageNumber="39" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. doederleini</emphasis>
|
||
</taxonomicName>
|
||
from Taiwan (present study); with five out of seven males below the size of 130 mm lacking it. The largest males of
|
||
<taxonomicName lsidName="B. doederleini" pageId="0" pageNumber="39" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. doederleini</emphasis>
|
||
</taxonomicName>
|
||
(136-145 mm) possess appendix masculina. The absence or presence of appendix masculina has been previously used by
|
||
<bibRefCitation DOI="https://doi.org/10.2307/1549056" author="Soong, K" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="39" pagination="72 - 79" refId="B17" refString="Soong, K, Mok, H-K, 1994. Size and maturity stage observations of the deep-sea isopod Bathynomus doederleini Ortmann, 1894 (Flabellifera: Cirolanidae), in eastern Taiwan. Journal of Crustacean Biology 14: 72 - 79, DOI: https://doi.org/10.2307/1549056" title="Size and maturity stage observations of the deep-sea isopod Bathynomus doederleini Ortmann, 1894 (Flabellifera: Cirolanidae), in eastern Taiwan." url="https://doi.org/10.2307/1549056" volume="14" year="1994">Soong and Mok (1994)</bibRefCitation>
|
||
to determine the maturity of males of
|
||
<taxonomicName authorityName="Ortmann" authorityYear="1894" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus doederleini" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus doederleini</emphasis>
|
||
</taxonomicName>
|
||
; "mature males" were males with appendix masculina and "maturing males" were those without appendix masculina and testes.
|
||
<bibRefCitation DOI="https://doi.org/10.1016/S0967-0637(03)00036-0" author="Barradas-Ortiz, C" journalOrPublisher="Deep Sea Research Part I: Oceanographic Research Papers" pageId="0" pageNumber="39" pagination="495 - 513" refId="B1" refString="Barradas-Ortiz, C, Briones-Fourzan, P, Lozano-Alvarez, E, 2003. Seasonal reproduction and feeding ecology of giant isopods Bathynomus giganteus from the continental slope of the Yucatan peninsula. Deep Sea Research Part I: Oceanographic Research Papers 50: 495 - 513, DOI: https://doi.org/10.1016/S0967-0637(03)00036-0" title="Seasonal reproduction and feeding ecology of giant isopods Bathynomus giganteus from the continental slope of the Yucatan peninsula." url="https://doi.org/10.1016/S0967-0637(03)00036-0" volume="50" year="2003">Barradas-Ortiz et al. (2003)</bibRefCitation>
|
||
reported that some adult males of
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
from Brazil (mostly smaller specimens below 290 mm) lacked appendix masculina, especially in summer. They suggested that these smaller males might have been less reproductively active in summer and/or the appendix masculina may be a non-permanent organ which is lost or regrown when the animals moult (
|
||
<bibRefCitation DOI="https://doi.org/10.1016/S0967-0637(03)00036-0" author="Barradas-Ortiz, C" journalOrPublisher="Deep Sea Research Part I: Oceanographic Research Papers" pageId="0" pageNumber="39" pagination="495 - 513" refId="B1" refString="Barradas-Ortiz, C, Briones-Fourzan, P, Lozano-Alvarez, E, 2003. Seasonal reproduction and feeding ecology of giant isopods Bathynomus giganteus from the continental slope of the Yucatan peninsula. Deep Sea Research Part I: Oceanographic Research Papers 50: 495 - 513, DOI: https://doi.org/10.1016/S0967-0637(03)00036-0" title="Seasonal reproduction and feeding ecology of giant isopods Bathynomus giganteus from the continental slope of the Yucatan peninsula." url="https://doi.org/10.1016/S0967-0637(03)00036-0" volume="50" year="2003">Barradas-Ortiz et al. 2003</bibRefCitation>
|
||
).
|
||
<bibRefCitation DOI="https://doi.org/10.1016/S0967-0637(03)00036-0" author="Barradas-Ortiz, C" journalOrPublisher="Deep Sea Research Part I: Oceanographic Research Papers" pageId="0" pageNumber="39" pagination="495 - 513" refId="B1" refString="Barradas-Ortiz, C, Briones-Fourzan, P, Lozano-Alvarez, E, 2003. Seasonal reproduction and feeding ecology of giant isopods Bathynomus giganteus from the continental slope of the Yucatan peninsula. Deep Sea Research Part I: Oceanographic Research Papers 50: 495 - 513, DOI: https://doi.org/10.1016/S0967-0637(03)00036-0" title="Seasonal reproduction and feeding ecology of giant isopods Bathynomus giganteus from the continental slope of the Yucatan peninsula." url="https://doi.org/10.1016/S0967-0637(03)00036-0" volume="50" year="2003">Barradas-Ortiz et al. (2003)</bibRefCitation>
|
||
also noted that larger male specimens of
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
tend to keep the organ for longer periods than smaller ones, although even large individuals (310 mm) sometimes do not possess the structure. We cannot be certain that either of the patterns above apply to
|
||
<taxonomicName lsidName="B. raksasa" pageId="0" pageNumber="39" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. as only one male was collected. The appendix masculina (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, C, E, G Bathynomus raksasa sp. nov. holotype male (363 mm) (MZB Cru. Iso 097), Indonesia B, D, F, H B. giganteus male (354 mm) (ZRC 2014.0837), Caribbean A, B pleotelson dorsal view C, D pleotelson posterior view E, F pleotelson lateral view G, H pleopod 2." figureDoi="/10.3897/zookeys.947.53906.figure5" httpUri="https://binary.pensoft.net/fig/430117" pageId="0" pageNumber="39">5H</figureCitation>
|
||
) is present on the large male American specimen of
|
||
<taxonomicName lsidName="B. giganteus" pageId="0" pageNumber="39" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. giganteus</emphasis>
|
||
</taxonomicName>
|
||
(ZRC 2014.0837) examined here.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
The SJADES cruise also obtained four juvenile and subadult specimens from southern Java (here identified as
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp.) (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6</figureCitation>
|
||
) which we are unable to identify to the species level, especially as the diagnostic characters may not be developed. They are clearly not
|
||
<taxonomicName authorityName="Sidabalok & Wong & Ng" authorityYear="2020" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus raksasa" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="raksasa">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus raksasa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. with a different pleotelson spination, shapes of pleotelson and uropodal rami. The largest specimen in the lot (107 mm) has an almost fully-developed pereopod 7 which indicates that the adult would not be too much larger in size. This, along with the presence of setae between the pleotelson spines, suggest that this species belongs to the
|
||
<normalizedToken originalValue="“giant”">"giant"</normalizedToken>
|
||
group. The number of spines on the posterior margin of the pleotelson ranges between 5+2, 7+2 and 9+2.
|
||
<bibRefCitation DOI="https://doi.org/10.2307/1549056" author="Soong, K" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="39" pagination="72 - 79" refId="B17" refString="Soong, K, Mok, H-K, 1994. Size and maturity stage observations of the deep-sea isopod Bathynomus doederleini Ortmann, 1894 (Flabellifera: Cirolanidae), in eastern Taiwan. Journal of Crustacean Biology 14: 72 - 79, DOI: https://doi.org/10.2307/1549056" title="Size and maturity stage observations of the deep-sea isopod Bathynomus doederleini Ortmann, 1894 (Flabellifera: Cirolanidae), in eastern Taiwan." url="https://doi.org/10.2307/1549056" volume="14" year="1994">Soong and Mok (1994)</bibRefCitation>
|
||
used the development of pereopod 7 as one of the characters to classify the development stages of
|
||
<taxonomicName authorityName="Ortmann" authorityYear="1894" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus doederleini" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus doederleini</emphasis>
|
||
</taxonomicName>
|
||
. According to
|
||
<bibRefCitation DOI="https://doi.org/10.2307/1549056" author="Soong, K" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="39" pagination="72 - 79" refId="B17" refString="Soong, K, Mok, H-K, 1994. Size and maturity stage observations of the deep-sea isopod Bathynomus doederleini Ortmann, 1894 (Flabellifera: Cirolanidae), in eastern Taiwan. Journal of Crustacean Biology 14: 72 - 79, DOI: https://doi.org/10.2307/1549056" title="Size and maturity stage observations of the deep-sea isopod Bathynomus doederleini Ortmann, 1894 (Flabellifera: Cirolanidae), in eastern Taiwan." url="https://doi.org/10.2307/1549056" volume="14" year="1994">Soong and Mok (1994)</bibRefCitation>
|
||
, individuals with "small, white" pereopod 7 and lacking either oostegites or penes and/or appendix masculina were categorised as "subadult I" which equals to stage 2 of five development stages they proposed. However, we will not apply this approach to
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. because of the limited specimen number.
|
||
</paragraph>
|
||
<caption doi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<emphasis bold="true" pageId="0" pageNumber="39">Figure 6.</emphasis>
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. (107 mm) (MZB Cru.Iso 098), Indonesia
|
||
<emphasis bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
uropod, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
uropod, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
pleotelson central spine
|
||
<emphasis bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
pleotelson
|
||
<emphasis bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
length of antenna 2
|
||
<emphasis bold="true" pageId="0" pageNumber="39">F</emphasis>
|
||
cephalon, anterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">G</emphasis>
|
||
clypeal region
|
||
<emphasis bold="true" pageId="0" pageNumber="39">H</emphasis>
|
||
pereopod 7, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="39">I</emphasis>
|
||
pereopod 7 coxa.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. superficially resembles the poorly known
|
||
<taxonomicName authorityName="Richardson" authorityYear="1910" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus affinis" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus affinis</emphasis>
|
||
</taxonomicName>
|
||
Richardson, 1910, described from the Philippines from one specimen. There is, however, a problem with what has been identified as "
|
||
<taxonomicName authorityName="Richardson" authorityYear="1910" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus affinis" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus affinis</emphasis>
|
||
</taxonomicName>
|
||
" by
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey (2006</bibRefCitation>
|
||
: 169, figs 2, 3), who listed among their material, the type from the Philippines as well as two females from the Arafura Sea, providing figures of the latter.
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce (1986</bibRefCitation>
|
||
: fig. 87A-E) had earlier figured the uropods, and pereopods 1 and 3 of the type specimen (sex not specified). The problem is that the distolateral corners of uropodal rami of the holotype from the Philippines is distinctly acute and curved (
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 87A-C) whereas that of
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey (2006</bibRefCitation>
|
||
: fig. 3D, E) from the Arafura Sea is distinctly wider and not produced. Significantly,
|
||
<bibRefCitation author="Richardson, H" journalOrPublisher="Bureau of Fisheries Document" pageId="0" pageNumber="39" pagination="1 - 44" refId="B15" refString="Richardson, H, 1910. Marine isopods collected in the Philippines by the U.S. Fisheries steamer Albatross in 1907-08. Bureau of Fisheries Document 736: 1 - 44" title="Marine isopods collected in the Philippines by the U. S. Fisheries steamer Albatross in 1907 - 08." volume="736" year="1910">
|
||
<normalizedToken originalValue="Richardson’s">Richardson's</normalizedToken>
|
||
(1910
|
||
</bibRefCitation>
|
||
: fig. 1) figures of the uropods are the same as those by
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce (1986)</bibRefCitation>
|
||
. The material from Arafura Sea are thus unlikely to be
|
||
<taxonomicName lsidName="B. affinis" pageId="0" pageNumber="39" rank="species" species="affinis">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. affinis</emphasis>
|
||
</taxonomicName>
|
||
s. str.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Our material of
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. from Java resembles the "
|
||
<taxonomicName lsidName="B. affinis" pageId="0" pageNumber="39" rank="species" species="affinis">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. affinis</emphasis>
|
||
</taxonomicName>
|
||
" of
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey (2006)</bibRefCitation>
|
||
in possessing the same relative length of antenna 2 (reaching between pereonites 3 and 4), straight clypeus distal margins, the setal fringe on the uropodal exopod is long and continuous (
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
90%), and similar pleotelson spine count (5+2, 7+2 and 9+2). The marked difference in the form of the uropodal endopod distolateral corner, however, indicates they are not conspecific. In addition, the uropod of
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. reaches to the end of the pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6D</figureCitation>
|
||
) (vs. slightly extended beyond the pleotelson;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 2F) and the pleotelson central spine is weakly bifid (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6C</figureCitation>
|
||
) (vs. simple;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 2F). The uropods of our material from Java agree very well with the figures by
|
||
<bibRefCitation author="Richardson, H" journalOrPublisher="Bureau of Fisheries Document" pageId="0" pageNumber="39" pagination="1 - 44" refId="B15" refString="Richardson, H, 1910. Marine isopods collected in the Philippines by the U.S. Fisheries steamer Albatross in 1907-08. Bureau of Fisheries Document 736: 1 - 44" title="Marine isopods collected in the Philippines by the U. S. Fisheries steamer Albatross in 1907 - 08." volume="736" year="1910">Richardson (1910)</bibRefCitation>
|
||
and
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce (1986)</bibRefCitation>
|
||
, but until a complete redescription of the holotype of
|
||
<taxonomicName lsidName="B. affinis" pageId="0" pageNumber="39" rank="species" species="affinis">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. affinis</emphasis>
|
||
</taxonomicName>
|
||
is done and more character states are known, we are not certain if they are actually conspecific.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. differs from
|
||
<taxonomicName lsidName="B. pelor" pageId="0" pageNumber="39" rank="species" species="pelor">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. pelor</emphasis>
|
||
</taxonomicName>
|
||
Bruce, 1986 (from northwestern Australia) in having a longer antenna 2 that reaches to the middle of pereonite 4 (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6E</figureCitation>
|
||
) (vs. middle of pereonite 2;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 91A), weakly bifid pleotelson central spine (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6C</figureCitation>
|
||
) (vs. strongly bifid;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 91B), and the conspicuous longitudinal carina on the pleotelson (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6D</figureCitation>
|
||
) (vs. inconspicuous;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 91C). Both species share similar shape of uropodal rami with more acute and curved distolateral corner on the endopod of
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6A, B</figureCitation>
|
||
) (vs. less acute and curved;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 91D). It differs from
|
||
<taxonomicName lsidName="B. immanis" pageId="0" pageNumber="39" rank="species" species="immanis">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. immanis</emphasis>
|
||
</taxonomicName>
|
||
Bruce, 1986, in the slightly concave lateral of uropodal exopod (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6A, B</figureCitation>
|
||
) (vs. strongly concave;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 90C, D), greater length of fringing setae (
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
80%) on the lateral uropod exopod (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6A, B</figureCitation>
|
||
) (vs. 66%;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 90C, D) and the weakly bifid pleotelson central spine (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6C</figureCitation>
|
||
) (vs. simple;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.6.1986.98" author="Bruce, NL" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="39" pagination="1 - 239" refId="B3" refString="Bruce, NL, 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239, DOI: https://doi.org/10.3853/j.0812-7387.6.1986.98" title="Cirolanidae (Crustacea: Isopoda) of Australia." url="https://doi.org/10.3853/j.0812-7387.6.1986.98" volume="6" year="1986">Bruce 1986</bibRefCitation>
|
||
: fig. 89 D). The two species together with
|
||
<taxonomicName lsidName="B. doederleini" pageId="0" pageNumber="39" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. doederleini</emphasis>
|
||
</taxonomicName>
|
||
share similar uropodal endopod shapes (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. shares with
|
||
<taxonomicName lsidName="B. kapala" pageId="0" pageNumber="39" rank="species" species="kapala">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. kapala</emphasis>
|
||
</taxonomicName>
|
||
Griffin, 1975 (from Australia) a similar bifid central pleotelson spine but can easily be distinguished by its relatively longer antenna 2 (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6E</figureCitation>
|
||
) (middle of pereonite 4 vs. within pereonite 3;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 14 C), the straight head ridge (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6F</figureCitation>
|
||
) (vs. curved;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 14 D), a narrowly rounded clypeus apex (6G) (vs. broadly rounded;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 14 E), with only one row of fringing setae on the anterior margin of the basis of pereopod 7 (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6H</figureCitation>
|
||
) (vs. with two rows;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 23 F) and the uropodal endopod distolateral margin is subacute and only slightly produced (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6A, B</figureCitation>
|
||
) (vs. not produced;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 15 D, E).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">
|
||
Compared to
|
||
<taxonomicName lsidName="B. doederleini" pageId="0" pageNumber="39" rank="species" species="doederleini">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">B. doederleini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1879" class="Malacostraca" family="Cirolanidae" genus="Bathynomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynomus" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="39">Bathynomus</emphasis>
|
||
</taxonomicName>
|
||
sp. has pereopod 7 coxa more slender (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6I</figureCitation>
|
||
) (vs. relatively broader;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 10B), there is one row of fringing setae on the anterior margin of the basis of pereopod 7 (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6H</figureCitation>
|
||
) (vs. with two rows;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 23D), and the lengths of the pleotelson spines are similarly sized (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bathynomus sp. (107 mm) (MZB Cru. Iso 098), Indonesia A uropod, ventral view B uropod, dorsal view C pleotelson central spine D pleotelson E length of antenna 2 F cephalon, anterior view G clypeal region H pereopod 7, ventral view I pereopod 7 coxa." figureDoi="/10.3897/zookeys.947.53906.figure6" httpUri="https://binary.pensoft.net/fig/430118" pageId="0" pageNumber="39">6D</figureCitation>
|
||
) (vs. uneven;
|
||
<bibRefCitation author="Lowry, JK" editor="Richer De Forges, B" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="39" pagination="163 - 192" refId="B11" refString="Lowry, JK, Dempsey, K, 2006. The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific. In: Richer De Forges, B, Justine, J-L, Eds., Tropical Deep-Sea Benthos, volume 24. Memoires du Museum national d'Histoire naturelle 193: 163 - 192" title="The giant deep-sea scavenger genus Bathynomus (Crustacea, Isopoda, Cirolanidae) in the Indo-West Pacific." volume="193" volumeTitle="Tropical Deep-Sea Benthos, volume 24." year="2006">Lowry and Dempsey 2006</bibRefCitation>
|
||
: fig. 10F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="39" type="distribution">
|
||
<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="39">Sunda Strait and Indian Ocean, South Java, Indonesia; at depths of 957-1259 m.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |