210 lines
31 KiB
XML
210 lines
31 KiB
XML
<document id="393DB3160077AD13A099FE6402097072" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D509216FDF9F959CB0E2EB4" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius merdeka Telnov 2020, sp. nov." docType="treatment" docUuid="4CE118C9-AC74-47C5-93B3-82F7844A0000" docUuidSource="ZooBank" docVersion="10" lastPageNumber="78" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="77" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="09A52A35B7BC300FD07785281161B14B">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:roleTerm id="86D313D6111166DB8A22E048F0A2CD77">Author</mods:roleTerm>
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<mods:namePart id="F1B128F2914E531221EEB9F45772ED35">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="5624CBBAB684F0AE05C73E263DB192E8">text</mods:typeOfResource>
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<mods:title id="5A84197975BF3EE0A5D4EAEFE5093195">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="4AA0BB2F8F8D9965BEA17358540EA9C2">
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<mods:date id="9BFE473BA27F0623EE43E2B7CDB16007">2020</mods:date>
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<mods:number id="DF1F0C090F0745668AC1E050CD9125BA">2020-10-19</mods:number>
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<mods:detail id="6B1A6305BCF542D7E6BEA4174ADB13BB" type="volume">
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<mods:number id="765666BC372541B4D3449138EC4C9820">721</mods:number>
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<mods:classification id="0EA7C665B5277345E8E468A89E527736">journal article</mods:classification>
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<mods:identifier id="FE94FCC97E6974D4131BC6E55D401EEC" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D509216FDF9F959CB0E2EB4" ID-DOI="http://doi.org/10.5281/zenodo.4344341" ID-GBIF-Taxon="168648160" ID-Zenodo-Dep="4344341" LSID="urn:lsid:zoobank.org:act:4CE118C9-AC74-47C5-93B3-82F7844A0000" httpUri="http://treatment.plazi.org/id/1A3787F69D509216FDF9F959CB0E2EB4" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
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<subSubSection id="DA84656B9D509217FDF9F959CB9C2F60" box="[618,970,1767,1794]" pageId="76" pageNumber="77" type="nomenclature">
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<paragraph id="922136E09D509217FDF9F959CB9C2F60" blockId="76.[372,1215,1767,1870]" box="[618,970,1767,1794]" pageId="76" pageNumber="77">
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<heading id="C969818C9D509217FDF9F959CB9C2F60" box="[618,970,1767,1794]" centered="true" fontSize="11" level="2" pageId="76" pageNumber="77" reason="2">
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<taxonomicName id="559E4D639D509217FDF9F959CB312F60" ID-CoL="7X3X6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[618,871,1767,1794]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="merdeka" status="sp. nov.">
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<emphasis id="A0EAEAF29D509217FDF9F959CB312F60" bold="true" box="[618,871,1767,1794]" italics="true" pageId="76" pageNumber="77">Maechidius merdeka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D509217FCFCF956CB9C2F60" box="[879,970,1768,1794]" pageId="76" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DA84656B9D509217FEE7F8B3CC452F2F" pageId="76" pageNumber="77" type="description">
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<paragraph id="922136E09D509217FEE7F8B3CCE92F4A" blockId="76.[372,1215,1767,1870]" box="[372,1215,1805,1832]" pageId="76" pageNumber="77">
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<uri id="E60F3AE29D509217FEE7F8B3CCE92F4A" box="[372,1215,1805,1832]" pageId="76" pageNumber="77">
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urn:lsid:zoobank.org:act:
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<uuid id="E6380C359D509217FD06F8B3CCE92F4A" box="[661,1215,1805,1832]" pageId="76" pageNumber="77">4CE118C9-AC74-47C5-93B3-82F7844A0000</uuid>
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</uri>
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</paragraph>
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<paragraph id="922136E09D509217FDB2F88DCC452F2F" blockId="76.[372,1215,1767,1870]" box="[545,1043,1843,1870]" pageId="76" pageNumber="77">
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<figureCitation id="0AA52A659D509217FDB2F88DCA2D2F2F" box="[545,635,1843,1869]" captionStart="Figs 58–61" captionStartId="75.[189,241,1928,1954]" captionTargetBox="[336,1249,266,1914]" captionTargetId="figure-62@75.[333,1249,257,1923]" captionTargetPageId="75" captionText="Figs 58–61. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 58. M. maleo sp. nov., holotype, ♂ (IRSN). 59. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 60. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 61. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122152" httpUri="https://zenodo.org/record/4122152/files/figure.png" pageId="76" pageNumber="77">Figs 61</figureCitation>
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,
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<figureCitation id="0AA52A659D509217FD14F88DCAE32F2C" box="[647,693,1843,1870]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="76" pageNumber="77">145</figureCitation>
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,
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<figureCitation id="0AA52A659D509217FD52F88DCAA62F2F" box="[705,752,1843,1869]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="76" pageNumber="77">232</figureCitation>
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,
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<figureCitation id="0AA52A659D509217FD6FF88DCB7D2F2F" box="[764,811,1843,1869]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="76" pageNumber="77">323</figureCitation>
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, 396, 492, 664–666
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D509217FF2EF8C9CC532F8A" pageId="76" pageNumber="77" type="etymology">
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<paragraph id="922136E09D509217FF2EF8C9C9122FF3" blockId="76.[189,324,1911,1937]" box="[189,324,1911,1937]" pageId="76" pageNumber="77">
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||
<heading id="C969818C9D509217FF2EF8C9C9122FF3" bold="true" box="[189,324,1911,1937]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
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<emphasis id="A0EAEAF29D509217FF2EF8C9C9122FF3" bold="true" box="[189,324,1911,1937]" pageId="76" pageNumber="77">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D509217FF2EF816CC532F8A" blockId="76.[189,1399,1960,2025]" pageId="76" pageNumber="77">The name is derived from the Indonesian ‘merdeka’ (meaning ‘independent’, ‘free’) to celebrate the craving for spiritual freedom of the Papuan people. Noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D519216FF2EFEB7CCA22A62" pageId="77" pageNumber="78" type="materials_examined">
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||
<paragraph id="922136E09D519216FF2EFEB7C93A2940" blockId="77.[189,364,264,291]" box="[189,364,264,291]" pageId="77" pageNumber="78">
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||
<heading id="C969818C9D519216FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="77" pageNumber="78" reason="3">
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<emphasis id="A0EAEAF29D519216FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" pageId="77" pageNumber="78">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D519216FF79FE86C90A2930" blockId="77.[189,1398,312,410]" box="[234,348,312,338]" pageId="77" pageNumber="78">
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<emphasis id="A0EAEAF29D519216FF79FE86C90A2930" bold="true" box="[234,348,312,338]" pageId="77" pageNumber="78">
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<typeStatus id="4D2588429D519216FF79FE86C90A2930" box="[234,348,312,338]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="922136E09D519216FF2EFEE2C99F29F8" blockId="77.[189,1398,312,410]" pageId="77" pageNumber="78">
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<materialsCitation id="22F63CBD9D519216FF2EFEE2C99F29F8" ID-GBIF-Occurrence="2898878416" collectingDate="1921-11" collectionCode="BMNH" country="Guinea-Bissau" location="Island of New" pageId="77" pageNumber="78" specimenCount="1" typeStatus="holotype">
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COUNTRY UNKNOWN (
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<location id="9741603B9D519216FE6AFEE2CAC92914" LSID="urn:lsid:plazi:treatment:1A3787F69D509216FDF9F959CB0E2EB4:9741603B9D519216FE6AFEE2CAC92914" box="[505,671,348,374]" country="Guinea-Bissau" name="Island of New" pageId="77" pageNumber="78">Island of New</location>
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<collectingCountry id="EA8976709D519216FD34FEE2CAA82915" box="[679,766,348,375]" name="Guinea-Bissau" pageId="77" pageNumber="78">Guinea</collectingCountry>
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||
) •
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<specimenCount id="8498FD699D519216FC8EFEE2CB632914" box="[797,821,348,374]" pageId="77" pageNumber="78" type="male">♂</specimenCount>
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; “
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<collectingCountry id="EA8976709D519216FCDEFEE2CBB52914" box="[845,995,348,374]" pageId="77" pageNumber="78">New Guinea</collectingCountry>
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Doron
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<date id="E62010209D519216FBA8FEE2CCC52914" box="[1083,1171,348,374]" pageId="77" pageNumber="78" value="1921-11">
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||
<collectingDate id="F664E9C89D519216FBA8FEE2CCC52914" box="[1083,1171,348,374]" pageId="77" pageNumber="78" value="1921-11">xi.1921</collectingDate>
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||
</date>
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R.Neill Rothschild B.M.1939-1”;
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<collectionCode id="F48FAE259D519216FEFAFE3EC99329F8" box="[361,453,384,410]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="77" pageNumber="78">BMNH</collectionCode>
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||
.
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</materialsCitation>
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</paragraph>
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<paragraph id="922136E09D519216FF79FE7CC90A29BE" blockId="77.[189,1268,450,513]" box="[234,348,450,476]" pageId="77" pageNumber="78">
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<emphasis id="A0EAEAF29D519216FF79FE7CC90A29BE" bold="true" box="[234,348,450,476]" pageId="77" pageNumber="78">
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||
<typeStatus id="4D2588429D519216FF79FE7CC90A29BE" box="[234,348,450,476]" pageId="77" pageNumber="78" type="paratype">Paratype</typeStatus>
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</emphasis>
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||
</paragraph>
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<paragraph id="922136E09D519216FF2EFE58CCA22A62" blockId="77.[189,1268,450,513]" box="[189,1268,486,513]" pageId="77" pageNumber="78">
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||
<materialsCitation id="22F63CBD9D519216FF2EFE58CB512A63" ID-GBIF-Occurrence="2898878614" box="[189,775,486,513]" country="Guinea-Bissau" location="Island of New Guinea" pageId="77" pageNumber="78" specimenCount="1" typeStatus="paratype">
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COUNTRY UNKNOWN (
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<location id="9741603B9D519216FE6AFE58CB562A63" LSID="urn:lsid:plazi:treatment:1A3787F69D509216FDF9F959CB0E2EB4:9741603B9D519216FE6AFE58CB562A63" box="[505,768,486,513]" country="Guinea-Bissau" name="Island of New Guinea" pageId="77" pageNumber="78">Island of New Guinea</location>
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)
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</materialsCitation>
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•
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<materialsCitation id="22F63CBD9D519216FCB3FE58CCA62A62" ID-GBIF-Occurrence="2898878398" box="[800,1264,486,512]" collectingDate="2008-02-18" collectionCode="BMNH" collectorName="A. Skale & Coll. A. Skale Hof" country="Indonesia" elevation="260" latitude="1.3825" location="INDONESIA" pageId="77" pageNumber="78" specimenCount="1" specimenCount-male="1">
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<specimenCount id="8498FD699D519216FCB3FE58CB182A62" box="[800,846,486,512]" pageId="77" pageNumber="78" type="male">1 ♂</specimenCount>
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; same label as in holotype;
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<collectionCode id="F48FAE259D519216FB07FE58CCA62A62" box="[1172,1264,486,512]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="77" pageNumber="78">BMNH</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="DA84656B9D519216FF2EFD96C9FD2D91" pageId="77" pageNumber="78" type="description">
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||
<paragraph id="922136E09D519216FF2EFD96C9182A20" blockId="77.[189,334,552,578]" box="[189,334,552,578]" pageId="77" pageNumber="78">
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||
<emphasis id="A0EAEAF29D519216FF2EFD96C9182A20" bold="true" box="[189,334,552,578]" pageId="77" pageNumber="78">Description</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="922136E09D519216FF2EFDE6C9212AD8" blockId="77.[189,1399,599,698]" pageId="77" pageNumber="78">
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MEASUREMENTS. Male
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<typeStatus id="4D2588429D519216FE52FDE9CA7D2A13" box="[449,555,599,625]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
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||
, total body length
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||
<quantity id="55669B059D519216FC9AFDE9CB202A10" box="[777,886,599,626]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" pageId="77" pageNumber="78" unit="mm" value="7.05">7.05 mm</quantity>
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||
. Head
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||
<quantity id="55669B059D519216FC5AFDE9CC652A10" box="[969,1075,599,626]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="77" pageNumber="78" unit="mm" value="1.3">1.30 mm</quantity>
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long, across eyes
|
||
<quantity id="55669B059D519216FA9EFDE9CD212A10" box="[1293,1399,599,626]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="77" pageNumber="78" unit="mm" value="1.5">1.50 mm</quantity>
|
||
wide. Pronotum
|
||
<quantity id="55669B059D519216FEEDFDC5C9B02AF4" box="[382,486,635,662]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="77" pageNumber="78" unit="mm" value="1.45">1.45 mm</quantity>
|
||
long, maximum width
|
||
<quantity id="55669B059D519216FD66FDC5CB092AF4" box="[757,863,635,662]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="77" pageNumber="78" unit="mm" value="2.3">2.30 mm</quantity>
|
||
. Elytral length
|
||
<quantity id="55669B059D519216FB83FDC5CC2D2AF4" box="[1040,1147,635,662]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="77" pageNumber="78" unit="mm" value="4.3">4.30 mm</quantity>
|
||
, maximum combined width
|
||
<quantity id="55669B059D519216FE94FD21C9252AD8" box="[263,371,671,698]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="77" pageNumber="78" unit="mm" value="3.05">3.05 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="922136E09D519216FF2EFD5FCD092DE0" blockId="77.[189,1399,737,1410]" pageId="77" pageNumber="78">
|
||
With general features of
|
||
<taxonomicName id="559E4D639D519216FE45FD5CCA022A99" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[470,596,737,763]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="bintang" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D519216FE45FD5CCA022A99" box="[470,596,737,763]" italics="true" pageId="77" pageNumber="78">M. bintang</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D519216FDCAFD5CCAE22A9E" box="[601,692,738,764]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="559E4D639D519216FD2DFD5CCB1B2A99" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[702,845,738,763]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="crypticus" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D519216FD2DFD5CCB1B2A99" box="[702,845,738,763]" italics="true" pageId="77" pageNumber="78">M. crypticus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D519216FCC0FD5CCBF82A9E" box="[851,942,738,764]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="559E4D639D519216FC2BFD5CCC712A99" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[952,1063,737,763]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="lapsus" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D519216FC2BFD5CCC712A99" box="[952,1063,737,763]" italics="true" pageId="77" pageNumber="78">M. lapsus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D519216FBBEFD5CCCD32A9E" box="[1069,1157,738,764]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="559E4D639D519216FB29FD5CCD212A99" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1210,1399,737,763]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="owenstanleyi" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D519216FB29FD5CCD212A99" box="[1210,1399,737,763]" italics="true" pageId="77" pageNumber="78">M. owenstanleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D519216FF2EFCB8C9432B42" box="[189,277,774,800]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see descriptions above and below). Dorsal surface covered with microscopic velvety pubescence. Male labroclypeus (
|
||
<figureCitation id="0AA52A659D519216FE3EFC97CA432B26" box="[429,533,809,836]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="77" pageNumber="78">Fig. 145</figureCitation>
|
||
) straight on anterior margin. Anterolateral angles of male labroclypeus strongly protruding anteriad, acute. Lateral margins of male labroclypeus sinuous both in dorsal and lateral view. Head dorsal punctures deep, circular to ovoid. Antennae 9-segmented, club 3-lamellate. Head and pronotum covered with sparse, rather short suberect scale-like setae which are rounded apically. Pronotum constricted postmedially towards base. Obtuse median angulation on lateral margin of pronotum in dorsal view (
|
||
<figureCitation id="0AA52A659D519216FDB1FC63CADB2B95" box="[546,653,989,1015]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="77" pageNumber="78">Fig. 232</figureCitation>
|
||
). Pronotal punctures deep of different shape and size. Scales broader on sides of pronotal disc. With vague track of one longitudinal carina on each elytron; carina with multiple interruptions. Elytral punctures with ovoid to linear incision-shaped deep punctures (
|
||
<figureCitation id="0AA52A659D519216FA90FB9ACD3C2C5D" box="[1283,1386,1060,1087]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="77" pageNumber="78">Fig. 323</figureCitation>
|
||
). Intervening spaces microreticulate, subopaque, larger than punctures. Elytral scale-like setae strongly vary in size from minute (more abundant) to clavate and strongly surpassing length of corresponding punctures. Sutural carinae partly present, with interruptions. Setae scale-like, extremely minute (miniseta rising from anterior margin of each puncture) with sparse large scales. Male pygidium with deep ovoid punctures, intervening spaces covered with microscopical velvety pubescence (
|
||
<figureCitation id="0AA52A659D519216FB2BFB66CD772C90" box="[1208,1313,1240,1266]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="77" pageNumber="78">Fig. 492</figureCitation>
|
||
). Setae of pygidium scale-like, larger and clavate in median part (all pointing obliquely to the middle), minute and not surpassing length of corresponding punctures in most of anterior part of pygidium. Male protibia with two distal teeth on external margin (Fig. 396). Male metatibial terminal spurs distinctly shorter than length of basal metatarsomere. Spiculum gastrale rounded distally. Male aedeagus as in
|
||
<figureCitation id="0AA52A659D519216FB2AFAD6CD0A2DE0" box="[1209,1372,1384,1410]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="77" pageNumber="78">Figs 664–666</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="922136E09D519216FF2EFA14C9E62DA6" blockId="77.[189,432,1450,1476]" box="[189,432,1450,1476]" pageId="77" pageNumber="78">
|
||
<heading id="C969818C9D519216FF2EFA14C9E62DA6" bold="true" box="[189,432,1450,1476]" fontSize="11" level="3" pageId="77" pageNumber="78" reason="3">
|
||
<emphasis id="A0EAEAF29D519216FF2EFA14C9E62DA6" bold="true" box="[189,432,1450,1476]" pageId="77" pageNumber="78">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D519216FF2EFA67C9FD2D91" blockId="77.[189,427,1497,1523]" box="[189,427,1497,1523]" pageId="77" pageNumber="78">Female is unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D519216FF2EF9A2C96F2E07" box="[189,313,1564,1637]" pageId="77" pageNumber="78" type="biology_ecology">
|
||
<paragraph id="922136E09D519216FF2EF9A2C9762E54" blockId="77.[189,288,1564,1590]" box="[189,288,1564,1590]" pageId="77" pageNumber="78">
|
||
<heading id="C969818C9D519216FF2EF9A2C9762E54" bold="true" box="[189,288,1564,1590]" fontSize="11" level="3" pageId="77" pageNumber="78" reason="3">
|
||
<emphasis id="A0EAEAF29D519216FF2EF9A2C9762E54" bold="true" box="[189,288,1564,1590]" pageId="77" pageNumber="78">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D519216FF2EF9F5C96F2E07" blockId="77.[189,313,1611,1637]" box="[189,313,1611,1637]" pageId="77" pageNumber="78">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D519216FF2EF933CB0E2EB4" box="[189,856,1677,1751]" pageId="77" pageNumber="78" type="distribution">
|
||
<paragraph id="922136E09D519216FF2EF933C9002EC5" blockId="77.[189,342,1677,1703]" box="[189,342,1677,1703]" pageId="77" pageNumber="78">
|
||
<heading id="C969818C9D519216FF2EF933C9002EC5" bold="true" box="[189,342,1677,1703]" fontSize="11" level="3" pageId="77" pageNumber="78" reason="3">
|
||
<emphasis id="A0EAEAF29D519216FF2EF933C9002EC5" bold="true" box="[189,342,1677,1703]" pageId="77" pageNumber="78">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D519216FF2EF903CB0E2EB4" blockId="77.[189,856,1724,1751]" box="[189,856,1724,1751]" pageId="77" pageNumber="78">
|
||
New
|
||
<collectingCountry id="EA8976709D519216FF6FF902C9032EB4" box="[252,341,1724,1750]" name="Guinea-Bissau" pageId="77" pageNumber="78">Guinea</collectingCountry>
|
||
. Exact location of Doron cannot be tracked.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |