158 lines
19 KiB
XML
158 lines
19 KiB
XML
<document id="D0EC320704DE9C051408A9EC99D26550" ID-CLB-Dataset="21316" ID-DOI="10.11646/zootaxa.4753.1.1" ID-GBIF-Dataset="f7974bcf-11e7-4d1b-b09d-530dc2dbd54f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3983782" ID-ZooBank="AA9F66B3-EF8C-4F6B-8F35-0BCBEE5122ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597351967001" checkinUser="felipe" docAuthor="Shear, William A., Richart, Casey H. & Wong, Victoria L." docDate="2020" docId="131D87EFFFBAFF8BFFDC5CB0FC98F921" docLanguage="en" docName="zootaxa.4753.1.1.pdf" docOrigin="Zootaxa 4753 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calityla essigi Shear & Richart & Wong 2020, new species" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="EF24FF97FFA9FF98FF4B5F01FFFFFF9E" masterDocTitle="The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="20" updateTime="1698844398316" updateUser="ExternalLinkService">
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<mods:mods id="79A51975469771E0A6F6A119C4DEF2FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BA05D374E266B1F8E598D12C885F231F">
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<mods:title id="1457B07DC0434F4AED8FF084141DEED6">The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="3E4FFC746FA988D550B39194CF2726E8" type="personal">
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<mods:namePart id="C8B0A0859C6AD3F0E5C2F6AF58C0DC51">Shear, William A.</mods:namePart>
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<mods:roleTerm id="F475161A1425E055D31099010F1942A5">Author</mods:roleTerm>
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<mods:namePart id="CFB0A13D3773BC0825C657007B487244">Richart, Casey H.</mods:namePart>
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<mods:name id="8872983D8AFCD16E33FFA6EC05882DAA" type="personal">
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<mods:roleTerm id="575B384784006F7978FC6F6D959042FB">Author</mods:roleTerm>
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<mods:namePart id="3D00F8D5147401859F0210F842C513E3">Wong, Victoria L.</mods:namePart>
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<mods:typeOfResource id="805142B140A60FA42D3C5C779D30F05D">text</mods:typeOfResource>
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<mods:title id="901499FE263CE81CD0909D45FFF90679">Zootaxa</mods:title>
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<mods:part id="13B5A0F2754C38F3473ABF436CB7ACC3">
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<mods:date id="FA32AC6627AAE0456A1A1051FD2CEFB0">2020</mods:date>
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<mods:number id="F22B8A2BA56C825D9BE636226B1AF5A0">2020-03-19</mods:number>
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<mods:detail id="A4FCC5DFC15F5A57A39573DCE6162F7F" type="volume">
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<mods:number id="C39F594B0B33D306E2462B5905059C35">4753</mods:number>
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<mods:classification id="68B7AFBA43475BB8387E5F7FA68117AB">journal article</mods:classification>
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<mods:identifier id="97272EDFE4CFAC160796F62F176C90A9" type="CLB-Dataset">21316</mods:identifier>
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<mods:identifier id="A4FD447F1560E9F22C06F943172D8336" type="DOI">10.11646/zootaxa.4753.1.1</mods:identifier>
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<mods:identifier id="8519E424DD3EA5C7C86D95A83DEC54FC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2423FCE7815FE021DAF2857520668EDC" type="Zenodo-Dep">3983782</mods:identifier>
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<treatment id="131D87EFFFBAFF8BFFDC5CB0FC98F921" ID-DOI="http://doi.org/10.5281/zenodo.4341634" ID-GBIF-Taxon="166240468" ID-Zenodo-Dep="4341634" LSID="urn:lsid:plazi:treatment:131D87EFFFBAFF8BFFDC5CB0FC98F921" httpUri="http://treatment.plazi.org/id/131D87EFFFBAFF8BFFDC5CB0FC98F921" lastPageNumber="20" pageId="19" pageNumber="20">
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<subSubSection id="D3AE6572FFBAFF8BFFDC5CB0FE21FC55" box="[151,478,945,972]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="9B0B36F9FFBAFF8BFFDC5CB0FE21FC55" blockId="19.[151,478,945,1008]" box="[151,478,945,972]" pageId="19" pageNumber="20">
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<heading id="C0438195FFBAFF8BFFDC5CB0FE21FC55" bold="true" box="[151,478,945,972]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="4">
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||
<emphasis id="A9C0EAEBFFBAFF8BFFDC5CB0FE21FC55" bold="true" box="[151,478,945,972]" pageId="19" pageNumber="20">
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<emphasis id="A9C0EAEBFFBAFF8BFFDC5CB0FEB8FC55" bold="true" box="[151,327,945,972]" italics="true" pageId="19" pageNumber="20">
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<taxonomicName id="5CB44D7AFFBAFF8BFFDC5CB0FEBEFC55" ID-CoL="c3433450-52e6-4953-b25a-f9427fc42aec" authority="Shear & Richart & Wong, 2020" authorityName="Shear & Richart & Wong" authorityYear="2020" box="[151,321,945,972]" class="Diplopoda" family="Conotylidae" genus="Calityla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="essigi" status="sp. nov.">Calityla essigi</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="B2F35790FFBAFF8BFE045CB3FE21FC55" box="[335,478,945,972]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3AE6572FFBAFF8BFFDC5CD4FDB7FBE1" pageId="19" pageNumber="20" type="description">
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<paragraph id="9B0B36F9FFBAFF8BFFDC5CD4FE90FC71" blockId="19.[151,478,945,1008]" box="[151,367,981,1008]" pageId="19" pageNumber="20">
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<figureCitation id="038F2A7CFFBAFF8BFFDC5CD4FEE1FC6E" box="[151,286,981,1008]" captionStart="FIGS" captionStartId="18.[151,212,1910,1935]" captionTargetBox="[193,1395,181,1884]" captionTargetId="figure@18.[191,1395,181,1884]" captionTargetPageId="18" captionText="FIGS. 45–50. Calityla spp. Figs. 45–47. Calityla siskiyou. Fig. 45. Left anterior gonopod, posterior view. Fig. 46. Tip of left posterior gonopod coxite, posterior view. Fig. 47. Right posterior gonopod coxite, posterior view. Fig. 48–50. Calityla essigi. Fig. 48. Right anterior gonopod, posterior view. Fig. 49. Tip of right posterior gonopod, posterior view. Fig. 50. Tip of left posterior gonopod coxite, mesoposterior view." figureDoi="http://doi.org/10.5281/zenodo.3983794" httpUri="https://zenodo.org/record/3983794/files/figure.png" pageId="19" pageNumber="20">Figs. 48–50</figureCitation>
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, 67–74
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</paragraph>
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<paragraph id="9B0B36F9FFBAFF8BFFDC5B1FFDB7FBE1" blockId="19.[151,1437,1053,1728]" pageId="19" pageNumber="20">
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<emphasis id="A9C0EAEBFFBAFF8BFFDC5B1FFF23FBA6" bold="true" box="[151,220,1054,1080]" pageId="19" pageNumber="20">Types</emphasis>
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:
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<materialsCitation id="2BDC3CA4FFBAFF8BFFA65B1CFDBBFBE1" ID-GBIF-Occurrence="2831262392" collectingDate="1955-01-22" collectionCode="EMEC" collectorName="D. Burdick & M. Wasbauer" county="Santa Cruz Co." latitude="40.56" location="Ben Lomond" longLatPrecision="697" longitude="-123.42" municipality="Male" pageId="19" pageNumber="20" specimenCount="2" stateProvince="California" typeStatus="holotype">
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<collectingMunicipality id="7B6FAC83FFBAFF8BFFA65B1CFED8FBA9" box="[237,295,1053,1079]" pageId="19" pageNumber="20">Male</collectingMunicipality>
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<typeStatus id="440F885BFFBAFF8BFE655B1CFE6FFBA9" box="[302,400,1053,1079]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
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and four male
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<typeStatus id="440F885BFFBAFF8BFD755B1FFD56FBA6" box="[574,681,1054,1080]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
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from
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<collectingRegion id="5970F81BFFBAFF8BFDBB5B1CFC61FBA9" box="[752,926,1053,1079]" country="United States of America" name="California" pageId="19" pageNumber="20">CALIFORNIA</collectingRegion>
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:
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<emphasis id="A9C0EAEBFFBAFF8BFCE15B1CFBA2FBA9" bold="true" box="[938,1117,1053,1079]" italics="true" pageId="19" pageNumber="20">
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<collectingCounty id="726A4E75FFBAFF8BFCE15B1CFBA2FBA9" box="[938,1117,1053,1079]" pageId="19" pageNumber="20">Santa Cruz Co.</collectingCounty>
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</emphasis>
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,
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<location id="9E6B6022FFBAFF8BFB275B1FFAFCFBA9" LSID="urn:lsid:plazi:treatment:131D87EFFFBAFF8BFFDC5CB0FC98F921:9E6B6022FFBAFF8BFB275B1FFAFCFBA9" box="[1132,1283,1053,1080]" county="Santa Cruz Co." latitude="40.56" longLatPrecision="697" longitude="-123.42" municipality="Male" name="Ben Lomond" pageId="19" pageNumber="20" stateProvince="California">Ben Lomond</location>
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, ca.
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<geoCoordinate id="FE80503EFFBAFF8BFA7C5B1CFA67FBA6" box="[1335,1432,1053,1080]" degrees="40.56" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="40.56">40.56°N</geoCoordinate>
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, -
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<geoCoordinate id="FE80503EFFBAFF8BFFEA5B40FEECFBC5" box="[161,275,1089,1115]" degrees="123.42" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="-123.42">123.42°W</geoCoordinate>
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(estimated coordinates taken from
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<location id="9E6B6022FFBAFF8BFDD65B40FBA1FBC2" LSID="urn:lsid:plazi:treatment:131D87EFFFBAFF8BFFDC5CB0FC98F921:9E6B6022FFBAFF8BFDD65B40FBA1FBC2" box="[669,1118,1089,1116]" county="Santa Cruz Co." latitude="40.56" longLatPrecision="697" longitude="-123.42" municipality="Male" name="Highlands County Park in Ben Lomond" pageId="19" pageNumber="20" stateProvince="California">Highlands County Park in Ben Lomond</location>
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), collected
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<date id="EF0A1039FFBAFF8BFB965B40FA67FBC2" box="[1245,1432,1089,1116]" pageId="19" pageNumber="20" value="1955-01-22">
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<collectingDate id="FF4EE9D1FFBAFF8BFB965B40FA67FBC2" box="[1245,1432,1089,1116]" pageId="19" pageNumber="20" value="1955-01-22">22 January 1955</collectingDate>
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</date>
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, by
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<collectorName id="3641532FFFBAFF8BFFF15B67FEC6FBE1" box="[186,313,1125,1152]" pageId="19" pageNumber="20">D. Burdick</collectorName>
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,
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<collectorName id="3641532FFFBAFF8BFE0E5B67FE26FBE1" box="[325,473,1125,1152]" pageId="19" pageNumber="20">M. Wasbauer</collectorName>
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(
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<collectionCode id="FDA5AE3CFFBAFF8BFEA35B64FDC3FBE1" box="[488,572,1125,1151]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="19" pageNumber="20" type="University or college">EMEC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3AE6572FFBAFF8BFF8C5B88FA8DFB59" pageId="19" pageNumber="20" type="diagnosis">
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<paragraph id="9B0B36F9FFBAFF8BFF8C5B88FA8DFB59" blockId="19.[151,1437,1053,1728]" pageId="19" pageNumber="20">
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<emphasis id="A9C0EAEBFFBAFF8BFF8C5B88FEC5FB3A" bold="true" box="[199,314,1161,1188]" pageId="19" pageNumber="20">Diagnosis</emphasis>
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: This species resembles the preceding one, but is distinct in the more massive fimbriate branch of the posterior gonopod coxites and their more robust terminal process, with the subterminal tooth displaced basally.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3AE6572FFBAFF8BFF8C5BD3FF3AFAAD" pageId="19" pageNumber="20" type="etymology">
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<paragraph id="9B0B36F9FFBAFF8BFF8C5BD3FF3AFAAD" blockId="19.[151,1437,1053,1728]" pageId="19" pageNumber="20">
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<emphasis id="A9C0EAEBFFBAFF8BFF8C5BD3FEB7FB72" bold="true" box="[199,328,1234,1260]" pageId="19" pageNumber="20">Etymology</emphasis>
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: The species epithet honors the late Edward O. Essig, founder of the Essig Museum of Entomology at the University of
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<collectingRegion id="5970F81BFFBAFF8BFEED5BF4FDE5FA91" box="[422,538,1269,1295]" country="United States of America" name="California" pageId="19" pageNumber="20">California</collectingRegion>
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, Berkeley. Dr. Essig shared with WAS an interest in the horticulture of the genus
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<emphasis id="A9C0EAEBFFBAFF8BFFDC5A1BFF3AFAAD" box="[151,197,1306,1331]" italics="true" pageId="19" pageNumber="20">
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||
<taxonomicName id="5CB44D7AFFBAFF8BFFDC5A1BFF3EFAAD" box="[151,193,1306,1331]" class="Insecta" family="Tarachodidae" genus="Iris" kingdom="Animalia" order="Mantodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Iris</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3AE6572FFBAFF8BFF8C5A3CFC85F9E9" pageId="19" pageNumber="20" type="description">
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<paragraph id="9B0B36F9FFBAFF8BFF8C5A3CFC85F9E9" blockId="19.[151,1437,1053,1728]" pageId="19" pageNumber="20">
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<emphasis id="A9C0EAEBFFBAFF8BFF8C5A3CFEAEFAC6" bold="true" box="[199,337,1341,1368]" pageId="19" pageNumber="20">Description</emphasis>
|
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:
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<emphasis id="A9C0EAEBFFBAFF8BFE2B5A3FFD25FAC9" box="[352,730,1341,1368]" italics="true" pageId="19" pageNumber="20">
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<typeStatus id="440F885BFFBAFF8BFE2B5A3FFE38FAC9" box="[352,455,1342,1367]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
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male from Ben Lomond:
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</emphasis>
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Length, 11.0 mm. 22 ocelli in triangular eyepatch. Metazonites with prominent angular shoulders on rings 2–27. Color chestnut brown marked darker purplish brown. Legpairs one and two reduced, three to seven enlarged, with mesal knobs on femora of legpairs three to seven; hardly discernible on femora 3, gradually enlarged on more posterior femora, large, squarish and angular on femora seven (
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<figureCitation id="038F2A7CFFBAFF8BFA2F5AA8FF1DFA79" captionStart="FIGS" captionStartId="21.[151,212,1760,1785]" captionTargetBox="[201,1387,193,1721]" captionTargetId="figure@21.[189,1398,181,1734]" captionTargetPageId="21" captionText="FIGS. 68–84. Calityla spp. Figs. 68–74. Calityla essigi. Fig. 68. Left posterior gonopod coxite, posterior view. Fig. 69. Tip of left posterior gonopod coxite, posterior view. Figs. 70–74. Male legs 3–7, respectively. Figs. 75–85. Calityla humboldtensis. Fig. 75. Anterior gonopods, posterior view. Fig. 76. Posterior gonopods, posterior view. Fig. 77. Tip of left posterior gonopod coxite, posterior view. Figs. 78–82. Male legs 3–7, respectively. Fig. 83. Male leg 10. Fig. 84. Male leg 11. Fig. 85. Femoral knob of male leg 7." figureDoi="http://doi.org/10.5281/zenodo.3983798" httpUri="https://zenodo.org/record/3983798/files/figure.png" pageId="19" pageNumber="20">Figs. 70–74</figureCitation>
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). Anterior gonopods (
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<figureCitation id="038F2A7CFFBAFF8BFE995ACCFDA7FA79" box="[466,600,1485,1511]" captionStart="FIGS" captionStartId="18.[151,212,1910,1935]" captionTargetBox="[193,1395,181,1884]" captionTargetId="figure@18.[191,1395,181,1884]" captionTargetPageId="18" captionText="FIGS. 45–50. Calityla spp. Figs. 45–47. Calityla siskiyou. Fig. 45. Left anterior gonopod, posterior view. Fig. 46. Tip of left posterior gonopod coxite, posterior view. Fig. 47. Right posterior gonopod coxite, posterior view. Fig. 48–50. Calityla essigi. Fig. 48. Right anterior gonopod, posterior view. Fig. 49. Tip of right posterior gonopod, posterior view. Fig. 50. Tip of left posterior gonopod coxite, mesoposterior view." figureDoi="http://doi.org/10.5281/zenodo.3983794" httpUri="https://zenodo.org/record/3983794/files/figure.png" pageId="19" pageNumber="20">Figs. 48, 49</figureCitation>
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, 67) with prominent mesobasal, single seta on each side, simple, anteroposteriorly flattened, with sinuate posterior ridge, small serrations on lateral sides near tips (
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<figureCitation id="038F2A7CFFBAFF8BFBCF5AF0FB27F995" box="[1156,1240,1521,1547]" captionStart="FIGS" captionStartId="18.[151,212,1910,1935]" captionTargetBox="[193,1395,181,1884]" captionTargetId="figure@18.[191,1395,181,1884]" captionTargetPageId="18" captionText="FIGS. 45–50. Calityla spp. Figs. 45–47. Calityla siskiyou. Fig. 45. Left anterior gonopod, posterior view. Fig. 46. Tip of left posterior gonopod coxite, posterior view. Fig. 47. Right posterior gonopod coxite, posterior view. Fig. 48–50. Calityla essigi. Fig. 48. Right anterior gonopod, posterior view. Fig. 49. Tip of right posterior gonopod, posterior view. Fig. 50. Tip of left posterior gonopod coxite, mesoposterior view." figureDoi="http://doi.org/10.5281/zenodo.3983794" httpUri="https://zenodo.org/record/3983794/files/figure.png" pageId="19" pageNumber="20">Fig. 49</figureCitation>
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), slightly hooked apically. Posterior gonopod coxites (
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<figureCitation id="038F2A7CFFBAFF8BFD795914FD6FF9AE" box="[562,656,1557,1584]" captionStart="FIGS" captionStartId="18.[151,212,1910,1935]" captionTargetBox="[193,1395,181,1884]" captionTargetId="figure@18.[191,1395,181,1884]" captionTargetPageId="18" captionText="FIGS. 45–50. Calityla spp. Figs. 45–47. Calityla siskiyou. Fig. 45. Left anterior gonopod, posterior view. Fig. 46. Tip of left posterior gonopod coxite, posterior view. Fig. 47. Right posterior gonopod coxite, posterior view. Fig. 48–50. Calityla essigi. Fig. 48. Right anterior gonopod, posterior view. Fig. 49. Tip of right posterior gonopod, posterior view. Fig. 50. Tip of left posterior gonopod coxite, mesoposterior view." figureDoi="http://doi.org/10.5281/zenodo.3983794" httpUri="https://zenodo.org/record/3983794/files/figure.png" pageId="19" pageNumber="20">Figs. 50</figureCitation>
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,
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<figureCitation id="038F2A7CFFBAFF8BFDD65914FD1BF9AE" box="[669,740,1557,1584]" captionStart="FIGS" captionStartId="21.[151,212,1760,1785]" captionTargetBox="[201,1387,193,1721]" captionTargetId="figure@21.[189,1398,181,1734]" captionTargetPageId="21" captionText="FIGS. 68–84. Calityla spp. Figs. 68–74. Calityla essigi. Fig. 68. Left posterior gonopod coxite, posterior view. Fig. 69. Tip of left posterior gonopod coxite, posterior view. Figs. 70–74. Male legs 3–7, respectively. Figs. 75–85. Calityla humboldtensis. Fig. 75. Anterior gonopods, posterior view. Fig. 76. Posterior gonopods, posterior view. Fig. 77. Tip of left posterior gonopod coxite, posterior view. Figs. 78–82. Male legs 3–7, respectively. Fig. 83. Male leg 10. Fig. 84. Male leg 11. Fig. 85. Femoral knob of male leg 7." figureDoi="http://doi.org/10.5281/zenodo.3983798" httpUri="https://zenodo.org/record/3983798/files/figure.png" pageId="19" pageNumber="20">68, 69</figureCitation>
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||
) bearing large, complex fimbriate branch distal of midlength, terminating in large bifurcate process that bears a tooth near its base. Legpair 10 coxae of normal size, with small glands, legpair 11 femora with long, thin, dorsally directed knobs.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3AE6572FFBAFF8BFF8C5980FC98F921" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="9B0B36F9FFBAFF8BFF8C5980FC98F921" blockId="19.[151,1437,1053,1728]" pageId="19" pageNumber="20">
|
||
<emphasis id="A9C0EAEBFFBAFF8BFF8C5980FEA6F902" bold="true" box="[199,345,1665,1692]" pageId="19" pageNumber="20">Distribution</emphasis>
|
||
: Known only from the
|
||
<typeStatus id="440F885BFFBAFF8BFD145983FD70F902" box="[607,655,1666,1692]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality. Ben Lomond is a town; the same name is applied to a nearby mountain, and both are close to Big Bend Redwoods State Park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |