111 lines
14 KiB
XML
111 lines
14 KiB
XML
<document id="9C62957A3F392FEF665B135F530077D1" ID-DOI="10.11646/zootaxa.3722.2.1" ID-GBIF-Dataset="f2c46514-f165-43d3-a574-47c55b89dcd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="218998" ID-ZooBank="29D77A16-096D-4FC1-A5B4-9EEDF2E761A7" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460690754500" checkinUser="plazi" docAuthor="Vallejo, Luís Fernando & Wolff, Martha" docDate="2013" docId="3009D414FFF2FFB939B6FC7DFC4AA431" docLanguage="en" docName="zt03722p142.pdf" docOrigin="Zootaxa 3722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllophaga (Phyllophaga) densata Moser 1918" docType="key" docVersion="8" lastPageNumber="112" masterDocId="CC30AC6CFFF9FFB23921FFF1FF91A242" masterDocTitle="The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains" masterLastPageNumber="142" masterPageNumber="101" pageNumber="112" updateTime="1698351836216" updateUser="plazi">
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<mods:titleInfo id="795CA06CFDDF1A65D30EDF5FDD354B79">
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<mods:title id="E14357CE419809B7E409CD44061FAFF5">The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains</mods:title>
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</mods:titleInfo>
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<mods:name id="1AB9C6A6BAAABE08B9F40762A13DA68C" type="personal">
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<mods:namePart id="566D96A485C8C91F2273BCD8C2FFB90D">Vallejo, Luís Fernando</mods:namePart>
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<mods:roleTerm id="BB1780E062C69444B6EDD95F88A7624C">Author</mods:roleTerm>
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<mods:namePart id="78023C2B6A218B1D9CB540092A9FB8AB">Wolff, Martha</mods:namePart>
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<mods:typeOfResource id="1595E59A3164540D4ED8DFA41E0F97A7">text</mods:typeOfResource>
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<mods:title id="CFB5A792A64AFC27F7AA4B24A9B6F56D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4E9C71A84CCBEC8DDF4C2A970146C00C">
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<mods:date id="5A4F0B831D3F7C4008F7B51318AC75B4">2013</mods:date>
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<mods:detail id="A8DC5EF3B2240D4F5BB083C4CA5C8C6E" type="volume">
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<mods:number id="89B4B407EE47192A91F7D50027BB9B97">3722</mods:number>
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<mods:number id="7B3AD110B5E01AC1428928D241AE3769">2</mods:number>
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<mods:classification id="00347461DD354AC041F062D40E24D24B">journal article</mods:classification>
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<mods:identifier id="16B745D64240960286C7D82E783F76D9" type="DOI">10.11646/zootaxa.3722.2.1</mods:identifier>
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<mods:identifier id="F32451944E205DD99E2817B6AC4E8052" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="359E75532BF42049CC3AD144487767ED" type="Zenodo-Dep">218998</mods:identifier>
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<treatment id="3009D414FFF2FFB939B6FC7DFC4AA431" ID-DOI="http://doi.org/10.5281/zenodo.6151923" ID-Zenodo-Dep="6151923" LSID="urn:lsid:plazi:treatment:3009D414FFF2FFB939B6FC7DFC4AA431" httpUri="http://treatment.plazi.org/id/3009D414FFF2FFB939B6FC7DFC4AA431" lastPageNumber="112" pageId="11" pageNumber="112">
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<subSubSection id="F0BA3689FFF2FFB939B6FC7DFAFAA735" pageId="11" pageNumber="112" type="key">
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<paragraph id="B81F6502FFF2FFB939B6FC7DFC84A1E4" blockId="11.[151,789,908,934]" box="[151,789,908,934]" pageId="11" pageNumber="112">
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<heading id="E357D26EFFF2FFB939B6FC7DFC84A1E4" bold="true" box="[151,789,908,934]" fontSize="11" level="1" pageId="11" pageNumber="112" reason="1">
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<keyLead id="0351C637FFF2FFB939B6FC7DFC84A1E4" box="[151,789,908,934]" pageId="11" pageNumber="112">
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<key id="CC313233FFF2FFB939B6FC7DFC84A1E4" box="[151,789,908,934]" pageId="11" pageNumber="112">
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<keyStep id="03547DA7FFF2FFB939B6FC7DFC84A1E4" box="[151,789,908,934]" pageId="11" pageNumber="112">
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<emphasis id="8AD4B910FFF2FFB939B6FC7DFC84A1E4" bold="true" box="[151,789,908,934]" pageId="11" pageNumber="112">
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5.
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<taxonomicName id="7FA01E81FFF2FFB93995FC7DFC84A1E4" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[180,789,908,934]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="densata" subGenus="Phyllophaga">
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<emphasis id="8AD4B910FFF2FFB93995FC7DFDF5A1E4" bold="true" box="[180,612,908,934]" italics="true" pageId="11" pageNumber="112">Phyllophaga (Phyllophaga) densata</emphasis>
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(Moser, 1918)
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</taxonomicName>
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</emphasis>
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</keyStep>
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</key>
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</keyLead>
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</heading>
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</paragraph>
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<paragraph id="B81F6502FFF2FFB939B6FC23FAFAA735" blockId="11.[151,1436,978,1651]" pageId="11" pageNumber="112">
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<emphasis id="8AD4B910FFF2FFB939B6FC23FED5A1A9" bold="true" box="[151,324,978,1003]" pageId="11" pageNumber="112">Redescription.</emphasis>
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Length,
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<quantity id="7F58C8E7FFF2FFB93894FC23FD9CA1A8" box="[437,525,978,1003]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="112" unit="mm" value="26.0">26 mm</quantity>
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. Body is elongated, oval, with
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<quantity id="7F58C8E7FFF2FFB93AAAFC22FC71A1A8" box="[907,992,979,1003]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="112" unit="mm" value="12.0">12 mm</quantity>
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humeral amplitude; dorsal surface is glabrous, bright black. Pronotum (
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<figureCitation id="209B7987FFF2FFB93B37FC07FDF6A64D" box="[534,615,1014,1039]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Fig. 22</figureCitation>
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) extended, wider than it is long, surface densely punctate, lateral margins crenulate. Clypeus bilobed, concave, with raised margins, surface densely punctate, and with a roughly punctate vertex. Antennae (
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<figureCitation id="209B7987FFF2FFB93845FBCFFE27A615" box="[356,438,1086,1111]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Fig. 23</figureCitation>
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) have 10 antennomeres and a wide and elongated antennal club with three antennomeres. Tarsal claws (
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<figureCitation id="209B7987FFF2FFB93815FB93FE23A639" box="[308,434,1122,1147]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Fig. 24, 26</figureCitation>
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) are cleft, with a short, dull tooth very close to the unguinal base. Protibia (
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<figureCitation id="209B7987FFF2FFB93C37FB93FAF8A639" box="[1302,1385,1122,1147]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Fig. 24</figureCitation>
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) are flattened with two pronounced anterior teeth and a third, smaller tooth; tibial spur well defined. Metatibia (
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<figureCitation id="209B7987FFF2FFB93C63FB77FA02A6DD" box="[1346,1427,1158,1183]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Fig. 25</figureCitation>
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) has sinuous, transversal carinae, more defined in the dorsolateral region, with two very sharp spurs, the inner shorter than the outer, both fused with tibia. The pygidial plate (
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<figureCitation id="209B7987FFF2FFB93A5BFB3FFC6DA6A5" box="[890,1020,1230,1255]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Figs 27–28</figureCitation>
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) is convex, with minute setae on its frontal and ventral extremes; anterodistal punctures are present in the ventral position. The male genital capsule (
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<figureCitation id="209B7987FFF2FFB939BEFAE7FEBBA76D" box="[159,298,1302,1327]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="11" pageNumber="112">Figs. 29–30</figureCitation>
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) is strongly sclerotized, compact, cylindrical, and curved. Parameres are long and wide, strongly curved at their distal extreme, with a very distinctive angular projection on each side; the aedeagus is membranous, its distal portion retracts to the internal side of each paramere forming a sort of an accordion; it is not adorned.
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</paragraph>
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</subSubSection>
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<subSubSection id="F0BA3689FFF2FFB939E6FA73FC4AA431" pageId="11" pageNumber="112" type="discussion">
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<paragraph id="B81F6502FFF2FFB939E6FA73FF66A468" blockId="11.[151,1436,978,1651]" pageId="11" pageNumber="112">
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<emphasis id="8AD4B910FFF2FFB939E6FA73FEDEA7D9" bold="true" box="[199,335,1410,1435]" pageId="11" pageNumber="112">Comments.</emphasis>
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The specimen collected in Parque Nacional Natural Tatamá apparently represents a glabrous form of the species, given that the population from southern
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<collectingCountry id="C0B72592FFF2FFB93A37FA57FCFDA7FD" box="[790,876,1446,1471]" name="Mexico" pageId="11" pageNumber="112">Mexico</collectingCountry>
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to the Canal Zone,
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<collectingCountry id="C0B72592FFF2FFB93D75FA56FB21A7FC" box="[1108,1200,1447,1470]" name="Panama" pageId="11" pageNumber="112">Panama</collectingCountry>
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, usually has a dense dorsal vestiture of microtrichia. Other diagnostic features are the shape of the tarsal claws, tibiae, and clypeus. The structure of the genitalia in the Colombian specimen is very similar to that in the specimens from
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<collectingCountry id="C0B72592FFF2FFB93DDEFA1EFAEAA445" box="[1279,1403,1518,1543]" name="Costa Rica" pageId="11" pageNumber="112">Costa Rica</collectingCountry>
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or
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<collectingCountry id="C0B72592FFF2FFB939B6F9E2FF65A468" box="[151,244,1555,1578]" name="Panama" pageId="11" pageNumber="112">Panama</collectingCountry>
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.
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</paragraph>
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<paragraph id="B81F6502FFF2FFB939E6F9C7FC4AA431" blockId="11.[151,1436,978,1651]" pageId="11" pageNumber="112">
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<emphasis id="8AD4B910FFF2FFB939E6F9C7FCDBA40D" bold="true" box="[199,842,1590,1615]" pageId="11" pageNumber="112">New geographic record. Departamento de Risaralda</emphasis>
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: Municipio de Santuario, Parque Nacional Natural Tatamá (Risaralda-Chocó) (
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<quantity id="7F58C8E7FFF2FFB938ECF9AAFDB5A431" box="[461,548,1627,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="112" unit="m" value="1100.0">1100 m</quantity>
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)
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<date id="CC1E43C2FFF2FFB93B12F9ABFD35A431" box="[563,676,1626,1651]" pageId="11" pageNumber="112" value="2000-07">July 2000</date>
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, Fernando Vallejo,
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<specimenCount id="AEA6AE8BFFF2FFB93A5CF9AAFC46A431" box="[893,983,1626,1651]" pageId="11" pageNumber="112" type="male">2 males</specimenCount>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |