262 lines
24 KiB
XML
262 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.85399" ID-Pensoft-Pub="1313-2970-1110-39" ID-Pensoft-UUID="766F9A94793D5793A9BE1E70F575798B" ID-ZooBank="4681CC6DE5F347C6B1D052DEA78BE7C3" ModsDocID="1313-2970-1110-39" checkinTime="1657006630715" checkinUser="pensoft" docAuthor="Sian Man, Nem, Siriboon, Thanit, Lin, Aung, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="CD2F41B7771E529F8BCA4B98EF88C87F" docLanguage="en" docName="ZooKeys 1110: 39-102" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.85399" docTitle="Haploptychius thebawi" docType="treatment" docVersion="2" id="766F9A94793D5793A9BE1E70F575798B" lastPageNumber="39" masterDocId="766F9A94793D5793A9BE1E70F575798B" masterDocTitle="Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species" masterLastPageNumber="102" masterPageNumber="39" pageNumber="39" updateTime="1657006933044" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sian Man, Nem</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4453-734X</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Siriboon, Thanit</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok, 10110, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lin, Aung</mods:namePart>
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<mods:affiliation>Fauna and Flora International, No. 35, 3</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1110</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1110.85399</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.85399</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1110-39</mods:identifier>
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<mods:identifier type="ZooBank">4681CC6DE5F347C6B1D052DEA78BE7C3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">766F9A94793D5793A9BE1E70F575798B</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F" httpUri="http://treatment.plazi.org/id/CD2F41B7771E529F8BCA4B98EF88C87F" lastPageNumber="39" pageId="0" pageNumber="39">
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<subSubSection pageId="0" pageNumber="39" type="nomenclature">
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName LSID="CD2F41B7-771E-529F-8BCA-4B98EF88C87F" authority="(Godwin-Austin, 1888)" baseAuthorityName="Godwin-Austin" baseAuthorityYear="1888" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius thebawi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="thebawi">Haploptychius thebawi (Godwin-Austin, 1888)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="description">
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<paragraph pageId="0" pageNumber="39">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Approximate sampling locations used in this study. The location name and coordinates are listed in Table 1. Historical type localities of a broad scale, such as Tavoy, Burma, Moulmein, Mergui, and Tenasserim, are not indicated on this map." figureDoi="10.3897/zookeys.1110.85399.figure1" httpUri="https://binary.pensoft.net/fig/710466" pageId="0" pageNumber="39">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Haploptychius thebawi A-C specimens CUMZ 13008 from Pyinyaung Village, Meiktila, Mandalay Region, with apertural dentition D, E specimens CUMZ 13010 from Aik Kham Cave, Taunggyi, Shan State." figureDoi="10.3897/zookeys.1110.85399.figure12" httpUri="https://binary.pensoft.net/fig/710477" pageId="0" pageNumber="39">, 12</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="reference_group">
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1888" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis thebawi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="thebawi">Streptaxis thebawi</taxonomicName>
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Godwin-Austin, 1888: 243. Type locality: Pingoung, Shan Hills [Pinlaung Township, Taunggyi District, Shan State, Myanmar].
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<bibRefCitation author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" pagination="201 - 275" refId="B32" refString="Gude, GK, 1903. A synopsis of the genus Streptaxis and its allies. Proceedings of the Malacological Society of London 5: 201 - 275" title="A synopsis of the genus Streptaxis and its allies." volume="5" year="1903">Gude 1903</bibRefCitation>
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: 212, 322, 325, pl. 12, figs 11-13.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName baseAuthorityName="Godwin-Austin" baseAuthorityYear="1888" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius thebawi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="thebawi">Haploptychius thebawi</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Journal of the Society for Bibliography of Natural History" pageId="0" pageNumber="39" refId="B37" refString="Kobelt, W, 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1(12b) (2): 1-211 pls42-71 [pp. 1-96 pls 42-59 (1905) ; 97-211, pls 60-71 (1906)]." title="Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (12 b) (2): 1 - 211 pls 42 - 71 [pp. 1 - 96 pls 42 - 59 (1905); 97 - 211, pls 60 - 71 (1906)]." year="1905">Kobelt 1906</bibRefCitation>
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: 145, 146: pl. 62, figs 11-13.
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<bibRefCitation author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
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: 222.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="materials_examined">
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<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Myanmar" county="Taunggyi District" latitude="20.818611" location="Limestone" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" specimenCount="1" stateProvince="Mandalay Region">
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:BD1594F6D84966DF8736A0F5EBF8F4DC" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Limestone" stateProvince="Mandalay Region">Limestone</location>
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hills (Apache Cement Factory),
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:C3370F1B5056BA526AA3BC9F7604A13D" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Pyinyaung Village" stateProvince="Mandalay Region">Pyinyaung Village</location>
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,
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:AC3C1BC5426C5EF34A3F2D18D3952DA2" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Thazi Township" stateProvince="Mandalay Region">Thazi Township</location>
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,
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:165EA5B01027473F746B3BC853ABD229" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Meiktila District" stateProvince="Mandalay Region">Meiktila District</location>
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,
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<collectingRegion country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
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,
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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(
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<geoCoordinate degrees="20" direction="north" minutes="49" orientation="latitude" precision="1" seconds="39.1" value="20.827528">20°49'39.1"N</geoCoordinate>
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,
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<geoCoordinate degrees="96" direction="east" minutes="23" orientation="longitude" precision="1" seconds="35.1" value="96.39308">96°23'35.1"E</geoCoordinate>
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): CUMZ 13008 (4 shells; Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Haploptychius thebawi A-C specimens CUMZ 13008 from Pyinyaung Village, Meiktila, Mandalay Region, with apertural dentition D, E specimens CUMZ 13010 from Aik Kham Cave, Taunggyi, Shan State." figureDoi="10.3897/zookeys.1110.85399.figure12" httpUri="https://binary.pensoft.net/fig/710477" pageId="0" pageNumber="39">12A-C</figureCitation>
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).
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:EDAF250AB6BD64DC9DB49111E7971ABF" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Ywangan Village" stateProvince="Mandalay Region">Ywangan Village</location>
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, near Lin Way Monastery,
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:DC9C587B48EE6B65E5230A8E67F086D6" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Kalaw Township" stateProvince="Mandalay Region">Kalaw Township</location>
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,
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:131AE2319E8B100EA55933EE796A49A2" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Taunggyi District" stateProvince="Mandalay Region">Taunggyi District</location>
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,
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
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,
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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(
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<geoCoordinate degrees="21" direction="north" minutes="13" orientation="latitude" precision="1" seconds="43.3" value="21.228695">21°13'43.3"N</geoCoordinate>
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,
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<geoCoordinate degrees="96" direction="east" minutes="33" orientation="longitude" precision="1" seconds="19.2" value="96.555336">96°33'19.2"E</geoCoordinate>
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): CUMZ 13009 (7 shells).
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<location LSID="urn:lsid:plazi:treatment:CD2F41B7771E529F8BCA4B98EF88C87F:F9446B82D1187B4EF14954B6502CC8A4" country="Myanmar" county="Taunggyi District" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Taunggyi Township" name="Aik Kham Cave" stateProvince="Mandalay Region">Aik Kham Cave</location>
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,
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<collectingMunicipality>Taunggyi Township</collectingMunicipality>
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,
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<collectingCounty>Taunggyi District</collectingCounty>
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,
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
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,
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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(
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<geoCoordinate degrees="20" direction="north" minutes="49" orientation="latitude" precision="1" seconds="07.0" value="20.818611">20°49'07.0"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="13" orientation="longitude" precision="1" seconds="42.0" value="97.22833">97°13'42.0"E</geoCoordinate>
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): CUMZ 13010 (3 shells; Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Haploptychius thebawi A-C specimens CUMZ 13008 from Pyinyaung Village, Meiktila, Mandalay Region, with apertural dentition D, E specimens CUMZ 13010 from Aik Kham Cave, Taunggyi, Shan State." figureDoi="10.3897/zookeys.1110.85399.figure12" httpUri="https://binary.pensoft.net/fig/710477" pageId="0" pageNumber="39">12D, E</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.1110.85399.figure12" httpUri="https://binary.pensoft.net/fig/710477" pageId="0" pageNumber="39" start="Figure 12" startId="F12">
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<paragraph pageId="0" pageNumber="39">
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<emphasis bold="true" pageId="0" pageNumber="39">Figure 12.</emphasis>
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<taxonomicName baseAuthorityName="Godwin-Austin" baseAuthorityYear="1888" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius thebawi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius thebawi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="39">A-C</emphasis>
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specimens CUMZ13008 from Pyinyaung Village, Meiktila,
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<collectingRegion country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
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, with apertural dentition
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<emphasis bold="true" pageId="0" pageNumber="39">D, E</emphasis>
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specimens CUMZ 13010 from Aik Kham Cave, Taunggyi,
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="diagnosis">
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<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName baseAuthorityName="Godwin-Austin" baseAuthorityYear="1888" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius thebawi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius thebawi</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName genus="H." lsidName="H. solidulus" pageId="0" pageNumber="39" rank="species" species="solidulus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. solidulus</emphasis>
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</taxonomicName>
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in having a high conical spire and globosely ovate shell. The former species can be discriminated by its smoother shell surface and narrower aperture, more rounded and extended penultimate whorl, and less inflated last whorl. Additionally,
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName genus="H." lsidName="H. porrectus" pageId="0" pageNumber="39" rank="species" species="porrectus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. porrectus</emphasis>
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</taxonomicName>
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(Pfeiffer, 1863) from Laos in having strong radial ridges, conical spire, and an inflated and deflected last whorl, whereas
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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exhibits a larger shell, less depressed and deflected last whorl, fine radial ridges, and higher spire.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="description">
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<paragraph pageId="0" pageNumber="39">Description.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Shell oblique-heliciform, white and translucent; whorls
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<normalizedToken originalValue="5½-6½">51/2-61/2</normalizedToken>
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; spire conical with distinct suture. Shell surface glossy with transverse ridges that diminish below periphery. Embryonic shell ~
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<normalizedToken originalValue="2½">21/2</normalizedToken>
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whorls with smooth surface; following whorls regularly coiled. Shell periphery rounded; last whorl axially deflected. Aperture subcircular; peristome discontinuous, thickened, expanded, and slightly reflected. Apertural dentition with one strong parietal lamella. Umbilicus open and deep (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Haploptychius thebawi A-C specimens CUMZ 13008 from Pyinyaung Village, Meiktila, Mandalay Region, with apertural dentition D, E specimens CUMZ 13010 from Aik Kham Cave, Taunggyi, Shan State." figureDoi="10.3897/zookeys.1110.85399.figure12" httpUri="https://binary.pensoft.net/fig/710477" pageId="0" pageNumber="39">12</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="distribution">
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<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="39">This species occurs in central-eastern Myanmar, based on the type locality and two localities in Shan State, one locality in Mandalay Region, and it is relatively low in population density.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="remarks">
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<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Originally,
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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was described based on specimens from Pingoung, Shan Hills, which is probably now referred to as Pinlaung Township, Taunggyi District, Shan State. Unfortunately, the type specimens of
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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could not be located in the NHM, London collection. However, the specimens examined herein match well with the original description and the syntype illustrated in
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<bibRefCitation author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" pagination="201 - 275" refId="B32" refString="Gude, GK, 1903. A synopsis of the genus Streptaxis and its allies. Proceedings of the Malacological Society of London 5: 201 - 275" title="A synopsis of the genus Streptaxis and its allies." volume="5" year="1903">Gude (1903</bibRefCitation>
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: figs 11-13). Despite that, the specimens from Mandalay Region tend to have a more ovate shell and elongated last whorl, and their smooth shell surface, narrow umbilicus, and conical spire make them most similar to this nominal species. Therefore, we treat this population as an intraspecific variation. Examination of genitalia of specimens from this population is necessary to clarify their taxonomic status.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="39" refId="B10" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen (1908)</bibRefCitation>
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treated this nominal species as a junior synonym of
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<taxonomicName genus="H." lsidName="H. burmanicus" pageId="0" pageNumber="39" rank="species" species="burmanicus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. burmanicus</emphasis>
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</taxonomicName>
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|
. However, the recently collected specimens of
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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|
can be distinguished from
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<taxonomicName genus="H." lsidName="H. burmanicus" pageId="0" pageNumber="39" rank="species" species="burmanicus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. burmanicus</emphasis>
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|
</taxonomicName>
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|
and
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<taxonomicName genus="H." lsidName="H. blanfordi" pageId="0" pageNumber="39" rank="species" species="blanfordi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. blanfordi</emphasis>
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</taxonomicName>
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by having an oblique-ovate to oblique-heliciform shell, elevated and conical spire, nearly smooth shell surface, penultimate whorl extended beyond last whorl, only one parietal lamella, and narrow umbilicus. Conversely, the two latter species have sub-oblique heliciform shells, depressed spire, prominent radial ridges on the shell surface, penultimate whorl slightly extended beyond the last whorl, with one parietal and one palatal lamella, and a wide umbilicus. Therefore, we consider
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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to stand as a separate species.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Among three populations of
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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|
(Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Shell measurements of streptaxid species from Myanmar recognized in this study. Numbers listed before collection locality correspond to those in the map in Fig. 1." httpUri="http://table.plazi.org/id/96B9BDC5D55CC6E7CCCBB1D55A5AA162" pageId="0" pageNumber="39" tableUuid="96B9BDC5D55CC6E7CCCBB1D55A5AA162">1</tableCitation>
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), Aik Kham Cave specimens have the smallest size with a more depressed shell, followed by Ywangan Village; Apache Cement Factory specimens have the largest shell with more conical spire and rounded last whorl.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |