844 lines
95 KiB
XML
844 lines
95 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4662071" ID-GBIF-Dataset="3fb65f60-0e30-4201-b360-72058e04f055" ID-GBIF-Taxon="180873596" ID-ISSN="0067-1975" ID-Zenodo-Dep="4662071" checkinTime="1617227073522" checkinUser="felipe" docAuthor="Shea, Glenn M." docDate="1995" docId="6E1687E8E121FFDD2A6611A36889F8D4" docLanguage="en" docName="RecAustMus.47.1.83-115.pdf.imf" docOrigin="Records of the Australian Museum 47 (1)" docSource="https://journals.australian.museum/shea-1995-rec-aust-mus-471-83115/" docStyle="DocumentStyle:D3147822FB22E0797CFA40BA2FBFB595.2:RecAustMus.1986-1997.journal_article" docStyleId="D3147822FB22E0797CFA40BA2FBFB595" docStyleName="RecAustMus.1986-1997.journal_article" docStyleVersion="2" docTitle="Cyclodomorphus casuarinae" docType="treatment" docVersion="5" lastPageId="13" lastPageNumber="96" masterDocId="922FFF90E127FFD02F4316166F06FFCB" masterDocTitle="A taxonomic revision of the Cyclodomorphus casuarinae complex (Squamata: Scincidae)" masterLastPageNumber="115" masterPageNumber="83" pageId="6" pageNumber="89" updateTime="1634708750209" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
|
||
<mods:title>A taxonomic revision of the Cyclodomorphus casuarinae complex (Squamata: Scincidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart>Shea, Glenn M.</mods:namePart>
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</mods:name>
|
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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<mods:titleInfo>
|
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<mods:title>Records of the Australian Museum</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>1995</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>1995-05-10</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>47</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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||
<mods:start>83</mods:start>
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<mods:end>115</mods:end>
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||
</mods:extent>
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||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>https://journals.australian.museum/shea-1995-rec-aust-mus-471-83115/</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.3853/j.0067-1975.47.1995.1</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">3fb65f60-0e30-4201-b360-72058e04f055</mods:identifier>
|
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<mods:identifier type="ISSN">0067-1975</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">4654787</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4662071" ID-GBIF-Taxon="180873596" ID-Zenodo-Dep="4662071" LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4" httpUri="http://treatment.plazi.org/id/6E1687E8E121FFDD2A6611A36889F8D4" lastPageId="13" lastPageNumber="96" pageId="6" pageNumber="89">
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<subSubSection pageId="6" pageNumber="89" type="nomenclature">
|
||
<paragraph blockId="6.[1317,2251,1973,2061]" pageId="6" pageNumber="89">
|
||
<taxonomicName authority="TODO" class="Reptilia" family="Scincidae" genus="Cyclodomorphus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="89" phylum="Chordata" rank="species" species="casuarinae">
|
||
<heading box="[1317,1846,1973,2012]" fontSize="10" level="2" pageId="6" pageNumber="89" reason="2">
|
||
<emphasis box="[1317,1620,1973,2012]" italics="true" pageId="6" pageNumber="89">Cyclodomorphus</emphasis>
|
||
TODO
|
||
</heading>
|
||
Bibron,
|
||
<heading box="[1737,1835,2019,2061]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="89" reason="3">TODO</heading>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="89" type="reference_group">
|
||
<paragraph blockId="6.[1290,2276,2124,2368]" pageId="6" pageNumber="89">
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<treatmentCitationGroup pageId="6" pageNumber="89">
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||
<treatmentCitation author="Dumeril, AM & G. Bibron" box="[1294,2097,2124,2160]" page="749" pageId="6" pageNumber="89" year="1839">
|
||
<taxonomicName authority="Casuarinae Dumeril & Bibron, 1839: 749" authorityName="Casuarinae Dumeril & Bibron" authorityPageNumber="749" authorityYear="1839" box="[1294,2097,2124,2160]" class="Reptilia" family="Scincidae" genus="Cyclodus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="89" phylum="Chordata" rank="genus" species="casuarinae">
|
||
<emphasis box="[1294,1622,2124,2160]" italics="true" pageId="6" pageNumber="89">Cyclodus Casuarinae</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3068164301" box="[1634,2097,2124,2160]" collectorName="Peron & Lesueur" location="Bruny Island" pageId="6" pageNumber="89" specimenCount="1" stateProvince="Tasmania" typeStatus="lectotype">
|
||
<bibRefCitation author="Dumeril, AM & G. Bibron" box="[1634,2097,2124,2160]" pageId="6" pageNumber="89" refId="ref19199" refString="Dumeril, AM. C. & G. Bibron, 1839. Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Vol. 5. Librairie Encyclopedique de Roret, Paris, viii + 854 pp." type="book" year="1839">Dumeril & Bibron, 1839: 749</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
<typeStatus box="[2115,2270,2124,2160]" pageId="6" pageNumber="89">Lectotype</typeStatus>
|
||
: MNHP 7131,
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E121FFD629611E6369F5F752" box="[1570,1779,2165,2201]" name="Bruny Island" pageId="6" pageNumber="89" stateProvince="Tasmania">Bruny Island</location>
|
||
,
|
||
<collectingRegion box="[1804,1949,2165,2201]" country="Australia" name="Tasmania" pageId="6" pageNumber="89">Tasmania</collectingRegion>
|
||
(
|
||
<collectorName box="[1978,2068,2165,2201]" pageId="6" pageNumber="89">Peron</collectorName>
|
||
&
|
||
<collectorName box="[2133,2260,2165,2201]" pageId="6" pageNumber="89">Lesueur</collectorName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="6.[1290,2276,2124,2368]" pageId="6" pageNumber="89">
|
||
<treatmentCitationGroup pageId="6" pageNumber="89">
|
||
<treatmentCitation author="Peters, W." box="[1293,1898,2208,2243]" page="621" pageId="6" pageNumber="89" year="1875">
|
||
<taxonomicName authority="Peters, 1875: 621" authorityName="Peters" authorityPageNumber="621" authorityYear="1875" box="[1293,1898,2208,2243]" family="Scincidae" genus="Cyclodus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="89" rank="species" species="nigricans">
|
||
<emphasis box="[1293,1595,2208,2243]" italics="true" pageId="6" pageNumber="89">Cyclodus nigricans</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3068164316" box="[1614,1898,2208,2243]" country="Australia" location="Australia" pageId="6" pageNumber="89" specimenCount="1" typeStatus="holotype">
|
||
<bibRefCitation author="Peters, W." box="[1614,1898,2208,2243]" pageId="6" pageNumber="89" pagination="616 - 624" refId="ref20295" refString="Peters, W., 1875. Uber neue Amphibien (Gymnopis, Siphonops, Polypedates, Rhacophorus, Hyla, Cyciodus, Euprepes, Clemmys). Monatsberichte der koniglich preussischen Akademie der Wissenschaften zu Berlin 1874: 616 - 624, I-II pIs." type="journal article" year="1875">Peters, 1875: 621</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
<typeStatus box="[1923,2068,2208,2243]" pageId="6" pageNumber="89">Holotype</typeStatus>
|
||
: 2MB 8193,
|
||
<collectingCountry box="[1339,1479,2249,2285]" name="Australia" pageId="6" pageNumber="89">Australia</collectingCountry>
|
||
(Flower).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="6.[1290,2276,2124,2368]" pageId="6" pageNumber="89">
|
||
<treatmentCitationGroup pageId="6" pageNumber="89">
|
||
<treatmentCitation author="Frost, C. & AH & S. Lucas" box="[1290,2271,2291,2326]" page="227" pageId="6" pageNumber="89" year="1894">
|
||
<taxonomicName authority="Frost & Lucas, 1894: 227" authorityName="Frost & Lucas" authorityPageNumber="227" authorityYear="1894" box="[1290,2271,2291,2326]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="89" phylum="Chordata" rank="species" species="tasmanicum">
|
||
<emphasis box="[1290,1776,2291,2326]" italics="true" pageId="6" pageNumber="89">Hemisphaeriodon tasmanicum</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3068164323" box="[1806,2271,2291,2326]" location="Tasmania" pageId="6" pageNumber="89" specimenCount="1" stateProvince="Tasmania" typeStatus="lectotype">
|
||
<bibRefCitation author="Frost, C. & AH & S. Lucas" box="[1806,2271,2291,2326]" pageId="6" pageNumber="89" pagination="227 - 228" refId="ref19368" refString="Frost, C. & AH. S. Lucas, 1894. On a new skink lizard from Tasmania. Proceedings of the Linnean Society of New South Wales (2) 8 (2): 227 - 228." type="journal article" year="1894">Frost & Lucas, 1894: 227</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
<typeStatus box="[1340,1496,2331,2368]" pageId="6" pageNumber="89">Lectotype</typeStatus>
|
||
: MV D2087,
|
||
<collectingRegion box="[1744,1889,2331,2368]" country="Australia" name="Tasmania" pageId="6" pageNumber="89">Tasmania</collectingRegion>
|
||
(c. Frost).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="89" type="diagnosis">
|
||
<paragraph blockId="6.[1292,2278,2433,2794]" pageId="6" pageNumber="89">
|
||
<emphasis bold="true" box="[1293,1473,2433,2473]" pageId="6" pageNumber="89">Diagnosis.</emphasis>
|
||
A moderately large
|
||
<taxonomicName box="[1862,2146,2433,2473]" class="Reptilia" family="Scincidae" genus="Cyclodomorphus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="89" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1862,2146,2433,2473]" italics="true" pageId="6" pageNumber="89">Cyclodomorphus</emphasis>
|
||
</taxonomicName>
|
||
(maximum SVL
|
||
<quantity box="[1508,1661,2479,2519]" metricMagnitude="-1" metricUnit="m" metricValue="1.7399999999999998" pageId="6" pageNumber="89" unit="mm" value="174.0">174 mm</quantity>
|
||
), differing from all other species in the genus in the combination of prefrontals usually contacting, postnarial groove absent, postmental usually contacting two infralabials on each side, subcaudal scales 68-84, and dorsal colour pattern often present and complex in adults (dark edges to scales and dark streaks basally and centrally on scales).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="8" lastPageNumber="91" pageId="6" pageNumber="89" type="description">
|
||
<paragraph blockId="6.[1292,2279,2860,3221]" lastBlockId="7.[226,1216,414,2200]" lastPageId="7" lastPageNumber="90" pageId="6" pageNumber="89">
|
||
<emphasis bold="true" box="[1293,1508,2860,2899]" pageId="6" pageNumber="89">Description.</emphasis>
|
||
Nasals usually broadly to narrowly separated (55.6%, n = 117), less commonly in point to narrow contact (21.4%) or moderate to broad contact (20.5%), rarely a median internasal present (2.6%); prefrontals usually in moderate to broad contact (55.2%, n = 116), less commonly in narrow contact (31.0%), rarely in point contact (1.7%), narrowly to broadly separated (10.3%), or separated by a small median scale (1.7%); transversely enlarged nuchals 0-6 on each side (x = 2.4, sd = 1.08, n = 225), usually three (48.0%) or two (26.2%); loreals two bilaterally; supraoculars usually three bilaterally, rostral two in contact with frontal, second largest (98.3%, n = 116), rarely two unilaterally (1.7%), reduction due to fusion of first and second supraoculars (n = 1) or second and third supraoculars (n = 1); supraciliaries 5-7 (x = 6.0, sd = 0.29, n = 232), usually six (91.8%); presuboculars 1- 3, usually two (82.2%, n = 231), rarely one (0.4%); postsuboculars 2-5 (x = 3.6, sd = 0.56, n = 231), usually four (55.4%); upper palpebrals 7-12 (x = 8.2, sd = 1.00, n = 69); lower palpebrals 7-12 (x = 9.3, sd = 0.95, n = 68); secondary temporals usually in a-configuration bilaterally, rarely in {3-configuration unilaterally (n = 2) or bilaterally (n = 1); supralabials 6-8 (x = 7.0, sd = 0.30, n = 232), usually seven (90.9%), third-last below centre of eye, separating pre- and postsuboculars; infralabials 6-9 (x = 7.4, sd = 0.56, n = 227), usually seven (56.4%) or eight (40.1%); usually first two infralabials contacting postmental (n = 110), rarely one only unilaterally (n = 4) or bilaterally (n = 1); ear small, usually with a single small lobule along rostral margin (81.2%, n = 218), rarely two (6.4%) or lobules absent (12.4%).
|
||
</paragraph>
|
||
<paragraph blockId="7.[226,1216,414,2200]" pageId="7" pageNumber="90">Body scales in 22-26 (x = 23.3, sd = 1.00, n = 108) longitudinal rows at midbody; scales in paravertebral rows not or only slightly broader than adjacent scales, 61-73 (x = 66.7, sd = 2.87, n = 107); subcaudal scales 68-84 (x = 76.9, sd = 3.89, n = 69); lamellae below fourth toe 9-14 (x = 11.5, sd = 1.03, n = 201).</paragraph>
|
||
<paragraph blockId="7.[226,1216,414,2200]" pageId="7" pageNumber="90">
|
||
SVL
|
||
<quantity box="[373,609,1840,1880]" metricMagnitude="-1" metricUnit="m" metricValue="1.0775000000000001" metricValueMax="1.7399999999999998" metricValueMin="0.41500000000000004" pageId="7" pageNumber="90" unit="mm" value="107.75" valueMax="174.0" valueMin="41.5">41.5-174 mm</quantity>
|
||
(n = 114); AGL/SVL 51.8-69.1% (x = 62.0%, n = 112); TL/SVL 63.6-133.6% (x = 103.9%, n = 70); FLL/SVL 12.7-21.4% (x = 15.8%, n = 114); HLL/SVL 17.5-26.2% (x = 21.5%, n = 114); FLL/HLL 64.2-85.7% (x = 73.6%, n = 115); HL/SVL 13.6-24.6% (x = 17.2%, n = 113); HW/HL 57.9-75.2% (x = 67.9%, n = 113); HD/HL 40.4--61.1% (x = 50.8%, n = 112).
|
||
</paragraph>
|
||
<paragraph blockId="7.[222,1210,2266,3179]" box="[227,1208,2266,2306]" pageId="7" pageNumber="90">Coloration (in preservative). Adult coloration variable.</paragraph>
|
||
<paragraph blockId="7.[222,1210,2266,3179]" pageId="7" pageNumber="90">Dorsal ground colour olive-grey to green, rarely reddish. Rarely immaculate, usually with narrow to broad dark brown or black lateral margins to most dorsal body and tail scales, giving solid to broken narrow dark stripes on body and at least tail base. Many individuals also with multiple fine mid brown-grey or russet streaks basally and centrally on most scales, which may in extreme development obscure the dark stripes and ground colour. Some individuals with a few dorsal body and tail scales also dark brown to black edged apically, in rare extreme development leading to irregularly defined narrow dark bands across back and tail.</paragraph>
|
||
<paragraph blockId="7.[222,1210,2266,3179]" pageId="7" pageNumber="90">Head dorsum usually immaculate, but in some strongly patterned individuals with dark flecks or spots along margins of some head shields.</paragraph>
|
||
<paragraph blockId="7.[222,1210,2266,3179]" pageId="7" pageNumber="90">Laterally, body and tail with dorsal ground colour and predominantly striped pattern grading evenly into ventral colour and predominantly banded pattern.</paragraph>
|
||
<paragraph blockId="7.[222,1210,2266,3179]" lastBlockId="7.[1284,2266,416,1926]" pageId="7" pageNumber="90">Face olive-grey or green, sutures black edged, especially subocular supralabials and less commonly other circumocular scalation, giving a dark mask about eyes.</paragraph>
|
||
<paragraph blockId="7.[1284,2266,416,1926]" pageId="7" pageNumber="90">Venter olive-green to blue-grey, rarely immaculate, usually with scattered scales black, especially along apical and lateral margins, sometimes with cream bases, aligned to give a series of narrow dark bands, usually in the form of irregular vermiculations on the body, more regular and on alternate scale rows on tail. Throat variably patterned, from immaculate through a few scattered dark flecks or spots, to three or four solid narrow dark bands cranial to level of forelimbs.</paragraph>
|
||
<paragraph blockId="7.[1284,2266,416,1926]" pageId="7" pageNumber="90">Limbs above with dorsal ground, ventrally with ventral ground, with varying development of dark markings corresponding to dorsal and ventral patterns.</paragraph>
|
||
<paragraph blockId="7.[1284,2266,416,1926]" pageId="7" pageNumber="90">Soles and palms yellow, occasionally with light brown calli or low tubercles.</paragraph>
|
||
<paragraph blockId="7.[1284,2266,416,1926]" pageId="7" pageNumber="90">
|
||
Rare individuals entirely melanistic, or in one case (MV
|
||
<accessionNumber box="[1391,1525,1197,1237]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D11218" pageId="7" pageNumber="90">D11218</accessionNumber>
|
||
) with dorsal pattern and ground colour largely obscured by broad, black, nearly confluent bands.
|
||
</paragraph>
|
||
<paragraph blockId="7.[1284,2266,416,1926]" pageId="7" pageNumber="90">
|
||
Juveniles with red to olive-green dorsal ground, body dorsum with dark scales and/or scale margins tending to align transversely to give narrow, closely spaced dark bands on body and at least tail base. Nape with two similar but broader and more pronounced dark bands, the more cranial extending rostroventrolaterally towards ears, continuing below ears as one or two narrower bars. Head dorsum of neonates often with a dark brown wash, especially over parietal shields. Subocular supralabial and usually some adjacent circumocular shields solid black, giving a black mask over eyes. Coloration otherwise as adults. Dark nape bands are the most longlasting element of juvenile coloration, but rarely persist as solid elements beyond SVL
|
||
<quantity box="[1868,2001,1887,1926]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="90" unit="mm" value="50.0">50 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[1282,2268,1992,2306]" pageId="7" pageNumber="90">
|
||
Coloration (in life). Three lizards (AM
|
||
<accessionNumber box="[1984,2126,1992,2033]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R65206" pageId="7" pageNumber="90">R65206</accessionNumber>
|
||
-08) had a faint lavender tint to the sides of the body, the smallest individual having the throat and body and tail venter yellowish-brown (A. Greer field notes). An individual from Mount Wellington (
|
||
<figureCitation box="[1738,1849,2175,2216]" captionStart="Fig" captionStartId="8.[400,459,1830,1868]" captionTargetBox="[320,2245,485,1774]" captionTargetPageId="8" captionText="Fig. 4. A live C. casuarinae from Mt Wellington, Tasmania." figureDoi="http://doi.org/10.5281/zenodo.4654789" httpUri="https://zenodo.org/record/4654789/files/figure.png" pageId="7" pageNumber="90">Fig. 4</figureCitation>
|
||
) had the pale parts of the body venter and flanks yellow, the iris mid-brown and the tongue dark blue-black.
|
||
</paragraph>
|
||
<paragraph blockId="7.[1281,2268,2372,2596]" pageId="7" pageNumber="90">
|
||
Allometry (
|
||
<tableCitation box="[1499,1638,2372,2412]" captionStart="Table 3" captionStartId="30.[303,395,550,590]" captionTargetBox="[420,1866,776,1193]" captionTargetPageId="30" captionText="Table 3. Allometric equations and calculated values for cranial and somatic proportions in C. casuarinae. Values a and b solve the equation y = bxa, r = correlation coefficient, s.e. = standard error of a, C42, C1D3 and Cl74 are calculated percentage proportions (y/x) at SVL = 42, 103 and 174 mm (minimum, minimum mature female and maximum size). Sample sizes are as for ratios." httpUri="http://table.plazi.org/id/B2C06676E139FFCE2E6C143068E1FD13" pageId="7" pageNumber="90" tableUuid="B2C06676E139FFCE2E6C143068E1FD13">Table 3</tableCitation>
|
||
). With respect to SVL, AGL and TL show positive allometry, while HL and limb lengths show negative allometry. With respect to HLL, FLL shows negative allometry, while HD shows negative allometry with respect to HL.
|
||
</paragraph>
|
||
<paragraph blockId="7.[1281,2267,2655,3200]" pageId="7" pageNumber="90">Sexual dimorphism. No significant differences were detected in the degree of separation/contact of nasals or prefrontals, or in mean number of nuchals, presuboculars, postsuboculars, supraciliaries, lower palpebrals, supralabials, infralabials, midbody scales, subcaudal scales or subdigital lamellae (t-tests). Significant differences were detected between males and females in mean number of upper palpebrals (males: x = 7.9, sd = 0.60, n = 25; females: x = 8.4, sd = 1.10, n = 42; t65 = 2.06') and paravertebral scales (males: x = 65.4, sd = 2.51, n = 33; females: x = 67.5, sd = 2.90, n = 57; t88 = 3.44'*').</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4654789" ID-Zenodo-Dep="4654789" box="[400,1470,1830,1868]" httpUri="https://zenodo.org/record/4654789/files/figure.png" pageId="8" pageNumber="91" startId="8.[400,459,1830,1868]" targetBox="[320,2245,485,1774]" targetPageId="8">
|
||
<paragraph blockId="8.[400,1470,1830,1868]" box="[400,1470,1830,1868]" pageId="8" pageNumber="91">
|
||
<emphasis bold="true" box="[400,518,1830,1868]" pageId="8" pageNumber="91">Fig. 4.</emphasis>
|
||
A live
|
||
<materialsCitation ID-GBIF-Occurrence="3068164302" box="[675,1466,1832,1867]" location="Mt Wellington" pageId="8" pageNumber="91" specimenCount="1" stateProvince="Tasmania">
|
||
C.
|
||
<taxonomicName baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1839" box="[734,909,1832,1867]" class="Reptilia" family="Scincidae" genus="Cyclodomorphus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="casuarinae">
|
||
<emphasis box="[734,909,1832,1867]" italics="true" pageId="8" pageNumber="91">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12FFFD82B49113E6A06F880" box="[1034,1280,1832,1867]" name="Mt Wellington" pageId="8" pageNumber="91" stateProvince="Tasmania">Mt Wellington</location>
|
||
,
|
||
<collectingRegion box="[1311,1466,1832,1867]" country="Australia" name="Tasmania" pageId="8" pageNumber="91">Tasmania</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="8.[242,1228,1979,2340]" pageId="8" pageNumber="91">
|
||
Mature females (SVL
|
||
<quantity box="[690,919,1979,2020]" metricMagnitude="-1" metricUnit="m" metricValue="1.385" metricValueMax="1.7399999999999998" metricValueMin="1.03" pageId="8" pageNumber="91" unit="mm" value="138.5" valueMax="174.0" valueMin="103.0">103-174 mm</quantity>
|
||
; x =
|
||
<quantity box="[1043,1221,1979,2020]" metricMagnitude="-1" metricUnit="m" metricValue="1.276" pageId="8" pageNumber="91" unit="mm" value="127.6">127.6 mm</quantity>
|
||
, sd ::: 16.97, n ::: 48) were much larger than mature males (
|
||
<quantity box="[256,463,2072,2112]" metricMagnitude="-1" metricUnit="m" metricValue="1.055" metricValueMax="1.26" metricValueMin="0.85" pageId="8" pageNumber="91" unit="mm" value="105.5" valueMax="126.0" valueMin="85.0">85-126 mm</quantity>
|
||
, x :::
|
||
<quantity box="[567,739,2072,2112]" metricMagnitude="-1" metricUnit="m" metricValue="1.0779999999999998" pageId="8" pageNumber="91" unit="mm" value="107.8">107.8 mm</quantity>
|
||
, sd ::: 12.48, n = 21; MannWhitney U test, Z = 4.416***).
|
||
</paragraph>
|
||
<paragraph blockId="8.[242,1228,1979,2340]" pageId="8" pageNumber="91">
|
||
Females have significantly longer bodies, shorter tails and legs and shorter but broader heads than males, although the differences in proportions are slight in most characters (
|
||
<tableCitation box="[451,595,2300,2340]" captionStart="Table 4" captionStartId="30.[484,573,1398,1438]" captionTargetBox="[481,1747,1713,3136]" captionTargetPageId="30" captionText="Table 4. Sexual dimorphism III cranial and somatic proportions in C." httpUri="http://table.plazi.org/id/B2C06676E139FFCE2EA713606DC3F94F" pageId="8" pageNumber="91" tableUuid="B2C06676E139FFCE2EA713606DC3F94F">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="91" type="distribution">
|
||
<paragraph blockId="8.[240,1228,2400,2717]" pageId="8" pageNumber="91">
|
||
<emphasis bold="true" box="[241,472,2400,2440]" pageId="8" pageNumber="91">Distribution.</emphasis>
|
||
Northern, central and eastern
|
||
<collectingRegion box="[1058,1221,2400,2440]" country="Australia" name="Tasmania" pageId="8" pageNumber="91">Tasmania</collectingRegion>
|
||
, from sea level to the central plateau (
|
||
<figureCitation box="[894,1002,2446,2486]" captionStart="Fig" captionStartId="2.[387,444,2361,2398]" captionTargetBox="[333,2084,521,2303]" captionTargetPageId="2" captionText="Fig. 2. Distribution of C. casuarinae in Tasmania. Dashed lines and numbers indicate populations used for analysing geographic variation, as follows: 1, north-east; 2, north-west; 3, south-east; 4, central. Solid lines represent 500 m contour." figureDoi="http://doi.org/10.5281/zenodo.4660401" httpUri="https://zenodo.org/record/4660401/files/figure.png" pageId="8" pageNumber="91">Fig. 2</figureCitation>
|
||
). Also Betsy (
|
||
<bibRefCitation author="Green, R. H. & J. L. Rainbird" box="[252,708,2492,2532]" pageId="8" pageNumber="91" refId="ref19509" refString="Green, R. H. & J. L. Rainbird, 1993. Reptiles from the islands of Tasmania. Queen Victoria Museum and Art Gallery Technical Report 1993 / 1 (unpaginated)." type="book" year="1993">Green & Rainbird, 1993</bibRefCitation>
|
||
), Brony, Maria and Tasman Islands on the east coast. Although there are no specimen-based records from the south-west of
|
||
<collectingRegion country="Australia" name="Tasmania" pageId="8" pageNumber="91">Tasmania</collectingRegion>
|
||
, sight records exist for Mount Anne and Mount Melaleuca (A.
|
||
<collectingRegion box="[522,644,2675,2717]" country="United Kingdom" name="Dudley" pageId="8" pageNumber="91">Dudley</collectingRegion>
|
||
, M. Hutchinson, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="11" lastPageNumber="94" pageId="8" pageNumber="91" type="discussion">
|
||
<paragraph blockId="8.[237,1231,2774,3183]" lastBlockId="8.[1297,2286,1979,3180]" pageId="8" pageNumber="91">
|
||
<emphasis bold="true" box="[238,515,2774,2816]" pageId="8" pageNumber="91">Type material.</emphasis>
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[532,902,2774,2816]" genus="Cyclodus" kingdom="Animalia" pageId="8" pageNumber="91" rank="species" species="casuarinae">
|
||
<emphasis box="[532,902,2774,2816]" italics="true" pageId="8" pageNumber="91">Cyclodus casuarinae</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation author="Dumeril, AM & G. Bibron" box="[239,699,2820,2862]" pageId="8" pageNumber="91" refId="ref19199" refString="Dumeril, AM. C. & G. Bibron, 1839. Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Vol. 5. Librairie Encyclopedique de Roret, Paris, viii + 854 pp." type="book" year="1839">Dumeril & Bibron (1839)</bibRefCitation>
|
||
mostly from a single MNHP specimen from NouveIle Hollande. A second specimen, in the collection of the Zoological Society of London, is mentioned in their description of coloration. Dumeril and Bibron did not specifically designate a holotype. However, the Paris specimen (
|
||
<collectionCode box="[1101,1227,3050,3091]" country="USA" name="Princeton University" pageId="8" pageNumber="91">MNHP</collectionCode>
|
||
7131) has consistently been considered the
|
||
<typeStatus box="[1075,1226,3097,3137]" pageId="8" pageNumber="91">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation author="Dumeril, M. C. & A Dumeril" box="[250,768,3142,3183]" pageId="8" pageNumber="91" refId="ref19237" refString="Dumeril, M. C. & M. A Dumeril, 1851. Catalogue methodique de la collection des reptiles. Gide et Baudry, Paris, iv + 224 pp." type="book" year="1851">Dumeril & Dumeril, 1851</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Guibe, J." box="[807,1047,3142,3183]" pageId="8" pageNumber="91" refId="ref19621" refString="Guibe, J., 1954. Catalogue des types de lezards du Museum National d'Histoire Naturelle. Colas imp., Bayeux, 119 pp." type="book" year="1954">Guibe, 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Brygoo, E. - R." box="[1087,1385,1979,3183]" pageId="8" pageNumber="91" pagination="1 - 126" refId="ref18900" refString="Brygoo, E. - R., 1985. Les types des Scincides (Reptiles, Sauriens) du Museum national d'Histoire naturelle Catalogue critique. Bulletin du Museum national d'Histoire naturelle (4) Section A (Zoologie, Biologie et Ecologie animales) 7 (3) (suppl.): 1 - 126." type="journal article" year="1985">Brygoo, 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Cogger, H. G. & E. E. Cameron & H. M. Cogger" box="[1418,1812,1979,2020]" pageId="8" pageNumber="91" refId="ref19088" refString="Cogger, H. G., E. E. Cameron & H. M. Cogger, 1983. Zoological Catalogue of Australia Vol. 1 Amphibia and Reptilia. Australian Government Publishing Service, Canberra, 313 pp." type="book" year="1983">
|
||
Cogger
|
||
<emphasis box="[1579,1689,1979,2020]" italics="true" pageId="8" pageNumber="91">et al.,</emphasis>
|
||
1983
|
||
</bibRefCitation>
|
||
), and must therefore be considered to be lectotype, nominated by assumption of holotype status (Article 74b of the Code of Zoological Nomenclature). The lectotype has been subsequently identified as collected by Peron and Lesueur from Bruny Island during the Baudin Expedition of 1801-04 (
|
||
<bibRefCitation author="Dumeril, M. C. & A Dumeril" box="[1656,2140,2255,2296]" pageId="8" pageNumber="91" refId="ref19237" refString="Dumeril, M. C. & M. A Dumeril, 1851. Catalogue methodique de la collection des reptiles. Gide et Baudry, Paris, iv + 224 pp." type="book" year="1851">Dumeril & Dumeril, 1851</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Guibe, J." pageId="8" pageNumber="91" refId="ref19621" refString="Guibe, J., 1954. Catalogue des types de lezards du Museum National d'Histoire Naturelle. Colas imp., Bayeux, 119 pp." type="book" year="1954">Guibe, 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Brygoo, E. - R." box="[1424,1685,2301,2341]" pageId="8" pageNumber="91" pagination="1 - 126" refId="ref18900" refString="Brygoo, E. - R., 1985. Les types des Scincides (Reptiles, Sauriens) du Museum national d'Histoire naturelle Catalogue critique. Bulletin du Museum national d'Histoire naturelle (4) Section A (Zoologie, Biologie et Ecologie animales) 7 (3) (suppl.): 1 - 126." type="journal article" year="1985">Brygoo, 1985</bibRefCitation>
|
||
). This specimen is presumably one of the "quelques beaux Iezards analogues aux Scinques, differant toutefois essentiellement des animaux de celle famille par l'elegance des formes et le rapport des proportions" observed by
|
||
<bibRefCitation author="Peron, E" pageId="8" pageNumber="91" refId="ref20194" refString="Peron, E, 1807. Voyage de Decouvertes Aux Terres Australes, execute par ordre de Sa Majeste l'Empereur et Roi, Sur les Corvettes le Geographe, le Naturaliste, et la Goelette le Casuarina, Pendant les Annees 1800, 1801, 1802, 1803 et 1804. Vol. I. Imprimerie Imperiale, Paris, 496 pp." type="book" year="1807">Peron (1807)</bibRefCitation>
|
||
on Bruny Island between mid January and early
|
||
<date box="[1405,1669,2577,2617]" pageId="8" pageNumber="91" value="1802-02">February 1802</date>
|
||
(see also
|
||
<bibRefCitation author="MacFarlane, C. W. & A Triebel" pageId="8" pageNumber="91" refId="ref19877" refString="MacFarlane, C. W. & L. A Triebel, 1937. French explorers in Tasmania and in southern seas. Australasian Publishing Co., Sydney, 120 pp." type="book" year="1937">MacFarlane & Triebel, 1937</bibRefCitation>
|
||
, for a reprint of Peron's account). The whereabouts of the Zoological Society of London paralectotype are unknown. That collection was largely dispersed to other national and provincial collections in 1852, and the remaining specimens disposed of in 1856 (
|
||
<bibRefCitation author="Sclater, P. L." box="[1310,1567,2852,2894]" pageId="8" pageNumber="91" refId="ref20567" refString="Sclater, P. L., 1901. A record of the progress of the Zoological Society of London during the Nineteenth Century. William Clowes & Sons, London, 248 pp." type="book" year="1901">Sclater, 1901</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="8.[1297,2286,1979,3180]" lastBlockId="10.[214,1208,415,3166]" lastPageId="10" lastPageNumber="93" pageId="8" pageNumber="91">
|
||
The lectotype of C.
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1758,1950,2910,2950]" genus="Cyclodus" kingdom="Animalia" pageId="8" pageNumber="91" rank="species" species="casuarinae">
|
||
<emphasis box="[1758,1950,2910,2950]" italics="true" pageId="8" pageNumber="91">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[1990,2110,2910,2950]" captionStart="Fig" captionStartId="9.[422,480,1759,1795]" captionTargetBox="[344,2222,447,1711]" captionTargetPageId="9" captionText="Fig. 5. Lectotype of Cyclodus casuarinae Dumeril & Bibron, 1839 (MNHP 7131)." figureDoi="http://doi.org/10.5281/zenodo.4654791" httpUri="https://zenodo.org/record/4654791/files/figure.png" pageId="8" pageNumber="91">Fig. 5</figureCitation>
|
||
) has the following combination of character states: nasals narrowly separated; prefrontals in narrow contact; supraoculars three; presuboculars two; postsuboculars three; supraciIiaries six; supralabials seven; infralabials 7/8, first two contacting postmental; nuchals three;
|
||
<materialsCitation ID-GBIF-Occurrence="3068164321" collectionCode="SVL, AGL" pageId="10" pageNumber="93" specimenCount="1">
|
||
temporals in a-configuration; midbody scales 24; paravertebral scales 68; tail regenerated from 12th subcaudal; subdigital lamellae
|
||
<emphasis box="[808,912,507,546]" italics="true" pageId="10" pageNumber="93">14/13;</emphasis>
|
||
SVL
|
||
<quantity box="[1046,1200,507,546]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="10" pageNumber="93" unit="mm" value="145.0">145 mm</quantity>
|
||
; AGL
|
||
<quantity box="[335,499,553,593]" metricMagnitude="-2" metricUnit="m" metricValue="9.55" pageId="10" pageNumber="93" unit="mm" value="95.5">95.5 mm</quantity>
|
||
; TL
|
||
<quantity box="[597,753,553,593]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="10" pageNumber="93" unit="mm" value="25.5">25.5 mm</quantity>
|
||
(original part) +
|
||
<quantity box="[1083,1208,553,593]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="93" unit="mm" value="28.0">28 mm</quantity>
|
||
(regenerate); FLL
|
||
<quantity box="[563,689,599,639]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="93" unit="mm" value="20.0">20 mm</quantity>
|
||
; HLL
|
||
<quantity box="[813,940,599,639]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="93" unit="mm" value="26.0">26 mm</quantity>
|
||
; HL
|
||
<quantity box="[1040,1200,599,639]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="10" pageNumber="93" unit="mm" value="21.2">21.2 mm</quantity>
|
||
; HW
|
||
<quantity box="[321,486,646,685]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="10" pageNumber="93" unit="mm" value="15.2">15.2 mm</quantity>
|
||
; HD
|
||
<quantity box="[600,759,646,685]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="10" pageNumber="93" unit="mm" value="11.1">11.1 mm</quantity>
|
||
. Although the specimen has been eviscerated, the large size suggests that it was female. Most of the measurements and scalational characters are in close agreement with those given in the
|
||
<typeStatus box="[300,373,829,869]" pageId="10" pageNumber="93">type</typeStatus>
|
||
description, only the tail length (
|
||
<quantity box="[1011,1141,829,869]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="93" unit="mm" value="62.0">62 mm</quantity>
|
||
<emphasis box="[1173,1208,829,869]" italics="true" pageId="10" pageNumber="93">vs</emphasis>
|
||
<quantity box="[226,397,875,915]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" pageId="10" pageNumber="93" unit="mm" value="53.5">53.5 mm</quantity>
|
||
) being noticeably different
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4654791" ID-Zenodo-Dep="4654791" box="[422,1870,1759,1797]" httpUri="https://zenodo.org/record/4654791/files/figure.png" pageId="9" pageNumber="92" startId="9.[422,480,1759,1795]" targetBox="[344,2222,447,1711]" targetPageId="9">
|
||
<paragraph box="[422,1870,1759,1797]" pageId="9" pageNumber="92">
|
||
<emphasis bold="true" box="[422,537,1759,1795]" pageId="9" pageNumber="92">Fig. 5.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName authority="Dumeril & Bibron, 1839" authorityName="Dumeril & Bibron" authorityYear="1839" box="[799,1597,1760,1797]" genus="Cyclodus" kingdom="Animalia" pageId="9" pageNumber="92" rank="species" species="casuarinae">
|
||
<emphasis box="[799,1145,1760,1797]" italics="true" pageId="9" pageNumber="92">Cyclodus casuarinae</emphasis>
|
||
<bibRefCitation author="Dumeril, AM & G. Bibron" box="[1170,1597,1760,1797]" pageId="9" pageNumber="92" refId="ref19199" refString="Dumeril, AM. C. & G. Bibron, 1839. Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Vol. 5. Librairie Encyclopedique de Roret, Paris, viii + 854 pp." type="book" year="1839">Dumeril & Bibron, 1839</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode box="[1633,1746,1760,1797]" country="USA" name="Princeton University" pageId="9" pageNumber="92">MNHP</collectionCode>
|
||
7131).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4654793" ID-Zenodo-Dep="4654793" box="[413,1585,3101,3139]" httpUri="https://zenodo.org/record/4654793/files/figure.png" pageId="9" pageNumber="92" startId="9.[413,473,3101,3137]" targetBox="[425,2112,2390,2971]" targetPageId="9">
|
||
<paragraph blockId="9.[413,1585,3101,3139]" box="[413,1585,3101,3139]" pageId="9" pageNumber="92">
|
||
<emphasis bold="true" box="[413,531,3101,3137]" pageId="9" pageNumber="92">Fig. 6.</emphasis>
|
||
Holotype of
|
||
<taxonomicName authority="Peters, 1875" authorityName="Peters" authorityYear="1875" box="[783,1336,3102,3139]" genus="Cyclodus" kingdom="Animalia" pageId="9" pageNumber="92" rank="species" species="nigricans">
|
||
<emphasis box="[783,1105,3102,3139]" italics="true" pageId="9" pageNumber="92">Cyclodus nigricans</emphasis>
|
||
<bibRefCitation author="Peters, W." box="[1130,1336,3102,3139]" pageId="9" pageNumber="92" pagination="616 - 624" refId="ref20295" refString="Peters, W., 1875. Uber neue Amphibien (Gymnopis, Siphonops, Polypedates, Rhacophorus, Hyla, Cyciodus, Euprepes, Clemmys). Monatsberichte der koniglich preussischen Akademie der Wissenschaften zu Berlin 1874: 616 - 624, I-II pIs." type="journal article" year="1875">Peters, 1875</bibRefCitation>
|
||
</taxonomicName>
|
||
(ZMB 8193).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="10.[214,1208,415,3166]" pageId="10" pageNumber="93">
|
||
While the type description does not give a precise locality for the lectotype and the regenerated tail precludes use of subcaudal counts to accurately assign it to a population, the more precise locality given by
|
||
<bibRefCitation author="Dumeril, M. C. & A Dumeril" box="[224,737,1105,1145]" pageId="10" pageNumber="93" refId="ref19237" refString="Dumeril, M. C. & M. A Dumeril, 1851. Catalogue methodique de la collection des reptiles. Gide et Baudry, Paris, iv + 224 pp." type="book" year="1851">Dumeril & Dumeril (1851)</bibRefCitation>
|
||
and subsequent authors, based on MNHP catalogue data, is sufficient to assign the name to the Tasmanian taxon.
|
||
</paragraph>
|
||
<paragraph blockId="10.[214,1208,415,3166]" pageId="10" pageNumber="93">
|
||
The description of
|
||
<taxonomicName authorityName="Peters" authorityYear="1875" box="[621,1207,1243,1282]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="nigricans" subGenus="Homolepida">
|
||
<emphasis box="[621,1207,1243,1282]" italics="true" pageId="10" pageNumber="93">Cyclodus (Homolepida) nigricans</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Peters, W." box="[293,535,1288,1329]" pageId="10" pageNumber="93" pagination="616 - 624" refId="ref20295" refString="Peters, W., 1875. Uber neue Amphibien (Gymnopis, Siphonops, Polypedates, Rhacophorus, Hyla, Cyciodus, Euprepes, Clemmys). Monatsberichte der koniglich preussischen Akademie der Wissenschaften zu Berlin 1874: 616 - 624, I-II pIs." type="journal article" year="1875">Peters (1875)</bibRefCitation>
|
||
is brief, mentioning only the dark coloration,
|
||
<emphasis box="[437,489,1335,1374]" italics="true" pageId="10" pageNumber="93">7/6</emphasis>
|
||
supralabials, two loreals longer than high, 25 midbody scales, and 70 scales along the body between lower jaw and vent. Peters considered that the combination of these features and a long snout differentiated his species from C.
|
||
<emphasis box="[774,968,1519,1558]" italics="true" pageId="10" pageNumber="93">
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[774,962,1519,1558]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="casuarinae">casuarinae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
although he gave no comparative data for the latter species.
|
||
<taxonomicName authorityName="Peters" authorityYear="1875" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="nigricans">
|
||
<emphasis italics="true" pageId="10" pageNumber="93">Cyclodus nigricans</emphasis>
|
||
</taxonomicName>
|
||
was placed in the synonymy of C.
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1022,1207,1611,1650]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="casuarinae">
|
||
<emphasis box="[1022,1207,1611,1650]" italics="true" pageId="10" pageNumber="93">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Boulenger, G." box="[287,602,1657,1697]" pageId="10" pageNumber="93" refId="ref18844" refString="Boulenger, G. A, 1887. Catalogue of the Lizards in the British Museum (Natural History). Vol. Ill. Lacertidae, Gerrhosauridae, Scincidae, Anelytropidae, Dibamidae, Chamaeleontidae. Trustees of the British Museum (Natural History), London, 575 pp., xl pIs." type="book" year="1887">Boulenger (1887)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="10.[214,1208,415,3166]" lastBlockId="10.[1277,2265,415,3166]" pageId="10" pageNumber="93">
|
||
Peters did not give any locality for his species, nor did he explicitly indicate the extent of his type series or its provenance or repository. Although he states at the end of the description that he found additional specimens of the species in collections sent to him by Prof. Flower of the Royal College of Surgeons in London, it is clear from the single set of scale counts and the asymmetry reported in supralabial scales (seven left, six right) that he described his species from only a single specimen. A single specimen (ZMB 8193;
|
||
<collectingCountry box="[217,377,2162,2202]" name="Australia" pageId="10" pageNumber="93">Australia</collectingCountry>
|
||
; pre: Flower) is identified as the type in Berlin, and in coloration and the direction of asymmetry of the supralabial shields agrees with Peters' description, although Peters counts one fewer supralabial and 25 midbody scales rather than the 24 that I count. The
|
||
<typeStatus box="[217,358,2392,2432]" pageId="10" pageNumber="93">holotype</typeStatus>
|
||
of C.
|
||
<taxonomicName authorityName="Peters" authorityYear="1875" box="[501,659,2392,2432]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="nigricans">
|
||
<emphasis box="[501,659,2392,2432]" italics="true" pageId="10" pageNumber="93">nigricans</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[692,809,2392,2432]" captionStart="Fig" captionStartId="9.[413,473,3101,3137]" captionTargetBox="[425,2112,2390,2971]" captionTargetPageId="9" captionText="Fig. 6. Holotype of Cyclodus nigricans Peters, 1875 (ZMB 8193)." figureDoi="http://doi.org/10.5281/zenodo.4654795" httpUri="https://zenodo.org/record/4654795/files/figure.png" pageId="10" pageNumber="93">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation box="[839,860,2392,2432]" captionStart="Fig" captionStartId="11.[378,436,2613,2648]" captionTargetBox="[464,1961,481,2564]" captionTargetPageId="11" captionText="Fig. 7. Dorsal and lateral views of head shields of holotype of Cyclodus nigricans." figureDoi="http://doi.org/10.5281/zenodo.4654797" httpUri="https://zenodo.org/record/4654797/files/figure.png" pageId="10" pageNumber="93">7</figureCitation>
|
||
) has the following combination of character states: nasals in moderate contact; prefrontals in moderate contact; supraoculars three; presuboculars two; postsuboculars
|
||
<emphasis box="[909,971,2530,2570]" italics="true" pageId="10" pageNumber="93">3/4;</emphasis>
|
||
supraciliaries 617; supralabials 817; infralabials
|
||
<emphasis box="[817,879,2576,2615]" italics="true" pageId="10" pageNumber="93">8/9,</emphasis>
|
||
first two contacting postmental; rostral ear lobules one; nuchals three; temporals in a-configuration; upper and lower palpebrals nine; midbody scales 24; paravertebral scales 67; tail regenerated from 34th subcaudal; subdigital lamellae 13; SVL
|
||
<quantity box="[314,459,2805,2844]" metricMagnitude="-1" metricUnit="m" metricValue="1.48" pageId="10" pageNumber="93" unit="mm" value="148.0">148 mm</quantity>
|
||
; AGL
|
||
<quantity box="[585,710,2805,2844]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="93" unit="mm" value="97.0">97 mm</quantity>
|
||
; FLL
|
||
<quantity box="[825,950,2805,2844]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="93" unit="mm" value="21.0">21 mm</quantity>
|
||
; HLL
|
||
<quantity box="[1072,1197,2805,2844]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="93" unit="mm" value="30.0">30 mm</quantity>
|
||
; HL
|
||
<quantity box="[304,479,2851,2890]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="10" pageNumber="93" unit="mm" value="22.9">22.9 mm</quantity>
|
||
; HW
|
||
<quantity box="[624,794,2851,2890]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="10" pageNumber="93" unit="mm" value="16.3">16.3 mm</quantity>
|
||
; HD
|
||
<quantity box="[931,1101,2851,2890]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="10" pageNumber="93" unit="mm" value="11.7">11.7 mm</quantity>
|
||
. The coloration is uniformly dark browniblack dorsally and dark but with evidence of darker macules centrally on scales ventrally. Although the gonads have not been examined to confirm the sex, the large size of the specimen suggests that it is female. As with C.
|
||
<emphasis box="[1010,1204,3080,3120]" italics="true" pageId="10" pageNumber="93">
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1010,1198,3080,3120]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="casuarinae">casuarinae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the regenerated tail of this specimen precludes use of the sole completely diagnostic character to assign the name to this species. However, the canonical variates analysis (
|
||
<figureCitation box="[1460,1580,507,548]" captionStart="Fig" captionStartId="5.[381,436,2475,2511]" captionTargetBox="[471,1951,528,2389]" captionTargetPageId="5" captionText="Fig. 3. Ordination of members of the C. casuarinae complex on the first two discriminant functions. Dots are C. michaeli, triangles are C. casuarinae, squares are C. praealtus." figureDoi="http://doi.org/10.5281/zenodo.4660405" httpUri="https://zenodo.org/record/4660405/files/figure.png" pageId="10" pageNumber="93">Fig. 3</figureCitation>
|
||
) unequivocally (98.9% probability) identifies the type as Tasmanian (67.6% probability of being from south-east
|
||
<collectingRegion box="[1688,1854,599,639]" country="Australia" name="Tasmania" pageId="10" pageNumber="93">Tasmania</collectingRegion>
|
||
). Further, I am aware of two other individuals with uniformly dark coloration, both from
|
||
<collectingRegion box="[1481,1640,690,731]" country="Australia" name="Tasmania" pageId="10" pageNumber="93">Tasmania</collectingRegion>
|
||
(A. Dudley, pers. comm.).
|
||
</paragraph>
|
||
<paragraph blockId="10.[1277,2265,415,3166]" pageId="10" pageNumber="93">
|
||
<taxonomicName authorityName="Frost & Lucas" authorityYear="1894" box="[1324,1841,737,777]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="species" species="tasmanicum">
|
||
<emphasis box="[1324,1841,737,777]" italics="true" pageId="10" pageNumber="93">Hemisphaeriodon tasmanicum</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation author="Frost, C. & AH & S. Lucas" pageId="10" pageNumber="93" pagination="227 - 228" refId="ref19368" refString="Frost, C. & AH. S. Lucas, 1894. On a new skink lizard from Tasmania. Proceedings of the Linnean Society of New South Wales (2) 8 (2): 227 - 228." type="journal article" year="1894">Frost & Lucas (1894)</bibRefCitation>
|
||
from material collected by Baldwin Spencer from Lake St Clair. They initially only compared their species with
|
||
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1845" box="[1706,2155,875,915]" class="Reptilia" family="Scincidae" genus="Cyclodomorphus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="species" species="gerrardii">
|
||
<emphasis box="[1706,2155,875,915]" italics="true" pageId="10" pageNumber="93">Cyclodomorphus gerrardii</emphasis>
|
||
</taxonomicName>
|
||
(then in the monotypic genus
|
||
<emphasis box="[1749,2071,921,961]" italics="true" pageId="10" pageNumber="93">
|
||
<taxonomicName authorityName="Peters" authorityYear="1867" box="[1749,2054,921,961]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="genus">Hemisphaeriodon</taxonomicName>
|
||
),
|
||
</emphasis>
|
||
but later (Lucas & Frost, 1896), after examining additional material, recognised its affinities with C.
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1999,2182,1014,1053]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="casuarinae">
|
||
<emphasis box="[1999,2182,1014,1053]" italics="true" pageId="10" pageNumber="93">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
and placed their species in its synonymy.
|
||
</paragraph>
|
||
<paragraph blockId="10.[1277,2265,415,3166]" lastBlockId="11.[221,1208,2732,3186]" lastPageId="11" lastPageNumber="94" pageId="10" pageNumber="93">
|
||
Although Frost and Lucas did not indicate the number of specimens on which their description was based, it is clear from the variation expressed ("one to four pairs of nuchals", ventral surface "greyish or brownish") that more than one individual was involved. This is further borne out by their later (Lucas & Frost, 1896) mention of "specimens" from Lake St Clair. No indication was given of the repository of their material, and no types were located by
|
||
<bibRefCitation author="Cogger, H. G. & E. E. Cameron & H. M. Cogger" box="[1591,1994,1473,1513]" pageId="10" pageNumber="93" refId="ref19088" refString="Cogger, H. G., E. E. Cameron & H. M. Cogger, 1983. Zoological Catalogue of Australia Vol. 1 Amphibia and Reptilia. Australian Government Publishing Service, Canberra, 313 pp." type="book" year="1983">
|
||
Cogger
|
||
<emphasis box="[1755,1849,1473,1513]" italics="true" pageId="10" pageNumber="93">et al.</emphasis>
|
||
(1983)
|
||
</bibRefCitation>
|
||
. Amongst the material examined for this paper are five MV specimens from Frost's collection (D2087-90, D2092,
|
||
<collectingRegion box="[2089,2252,1565,1605]" country="Australia" name="Tasmania" pageId="10" pageNumber="93">Tasmania</collectingRegion>
|
||
, received 12 October, 1915, but not registered until 23 June, 1943; A.J.
|
||
<collectingRegion box="[1597,1752,1657,1697]" country="United Kingdom" name="Coventry" pageId="10" pageNumber="93">Coventry</collectingRegion>
|
||
, pers. comm.) and a single AM specimen (R4142) from Lake St Clair, collected by Spencer and donated by Lucas. The AM animal forms part of a collection from Lucas that includes a
|
||
<typeStatus box="[2130,2259,1795,1835]" pageId="10" pageNumber="93">syntype</typeStatus>
|
||
of
|
||
<emphasis box="[1324,1756,1841,1881]" italics="true" pageId="10" pageNumber="93">
|
||
<taxonomicName authorityName="Lucas & Frost" authorityYear="1896" box="[1324,1750,1841,1881]" class="Reptilia" family="Scincidae" genus="Ablepharus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="species" species="rhodonoides">Ablepharus rhodonoides</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
described by Lucas & Frost (1896) in the same paper that synonymised
|
||
<emphasis italics="true" pageId="10" pageNumber="93">
|
||
<taxonomicName authorityName="Frost & Lucas" authorityYear="1894" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="species" species="tasmanicum">H. tasmanicum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation author="Frost, C. & AH & S. Lucas" box="[1510,1898,1933,1973]" pageId="10" pageNumber="93" pagination="227 - 228" refId="ref19368" refString="Frost, C. & AH. S. Lucas, 1894. On a new skink lizard from Tasmania. Proceedings of the Linnean Society of New South Wales (2) 8 (2): 227 - 228." type="journal article" year="1894">Frost & Lucas (1894)</bibRefCitation>
|
||
placed
|
||
<taxonomicName authorityName="Frost & Lucas" authorityYear="1894" box="[2060,2259,1933,1973]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="species" species="tasmanicum">
|
||
<emphasis box="[2060,2259,1933,1973]" italics="true" pageId="10" pageNumber="93">tasmanicum</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName authorityName="Peters" authorityYear="1867" box="[1326,1627,1979,2019]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="93" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1326,1627,1979,2019]" italics="true" pageId="10" pageNumber="93">Hemisphaeriodon</emphasis>
|
||
</taxonomicName>
|
||
on the basis of enlarged maxillary teeth, while Lucas & Frost (1896) synonymised it partly on palatal osteology. The mouths of both R4142 and D2087 have been opened subsequent to preservation, in the former case by fracturing the mandibles, in the latter by transecting the adductor musculature, and the palatal mucosa has been stripped back, allowing access to the palatal elements. Both specimens are similar in coloration and in position and condition of preservation. The measurements and scalation of D2087 are very close to those presented by
|
||
<bibRefCitation author="Frost, C. & AH & S. Lucas" box="[1655,2070,2437,2478]" pageId="10" pageNumber="93" pagination="227 - 228" refId="ref19368" refString="Frost, C. & AH. S. Lucas, 1894. On a new skink lizard from Tasmania. Proceedings of the Linnean Society of New South Wales (2) 8 (2): 227 - 228." type="journal article" year="1894">Frost & Lucas (1894)</bibRefCitation>
|
||
, only the midbody count (24) being outside the variation given in the description (26). However, counts of 26 midbody scales are very rare in C.
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1749,1933,2576,2616]" genus="Cyclodus" kingdom="Animalia" pageId="10" pageNumber="93" rank="species" species="casuarinae">
|
||
<emphasis box="[1749,1933,2576,2616]" italics="true" pageId="10" pageNumber="93">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
(only seen in two of
|
||
<specimenCount box="[1334,1587,2622,2662]" pageId="10" pageNumber="93" type="generic">108 specimens</specimenCount>
|
||
examined) and it is possible that the count provided by Frost and Lucas is erroneous. On the basis of collection data and the dissection of the oral cavity, I believe that AM R4142 is certainly a
|
||
<typeStatus box="[2123,2256,2759,2799]" pageId="10" pageNumber="93">syntype</typeStatus>
|
||
, and on the basis of the similarity between this specimen and D2087, and between the latter and the type description, that MV D2087 is also a
|
||
<typeStatus box="[1986,2117,2898,2937]" pageId="10" pageNumber="93">syntype</typeStatus>
|
||
. Of the other four MY specimens from Frost's collection, three (D2088-89, D2092) are strongly patterned, and cannot be
|
||
<typeStatus box="[1331,1475,3035,3075]" pageId="10" pageNumber="93">syntypes</typeStatus>
|
||
(Frost and Lucas emphasised the unpatterned dorsum of their species). The other specimen, D2090, while unpatterned, has been more neatly set, is in a much better state of preservation, and has clearly been treated very differently to the two identifiable
|
||
<typeStatus box="[1052,1201,2778,2817]" pageId="11" pageNumber="94">syntypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4654797" ID-Zenodo-Dep="4654797" box="[378,1862,2613,2648]" httpUri="https://zenodo.org/record/4654797/files/figure.png" pageId="11" pageNumber="94" startId="11.[378,436,2613,2648]" targetBox="[464,1961,481,2564]" targetPageId="11">
|
||
<paragraph blockId="11.[378,1494,2613,2648]" box="[378,1862,2613,2648]" lastBlockId="11.[1529,1862,2613,2648]" pageId="11" pageNumber="94">
|
||
Fig. 7. Dorsal and lateral views of head shields of holotype of
|
||
<heading box="[1529,1862,2613,2648]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="94" reason="5">
|
||
<emphasis box="[1529,1862,2613,2648]" italics="true" pageId="11" pageNumber="94">
|
||
<taxonomicName authorityName="Peters" authorityYear="1875" box="[1529,1857,2613,2648]" genus="Cyclodus" kingdom="Animalia" pageId="11" pageNumber="94" rank="species" species="nigricans">Cyclodus nigricans</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[221,1208,2732,3186]" pageId="11" pageNumber="94">
|
||
Because of its closer correspondence to the measurements provided by
|
||
<bibRefCitation author="Frost, C. & AH & S. Lucas" box="[583,983,2869,2910]" pageId="11" pageNumber="94" pagination="227 - 228" refId="ref19368" refString="Frost, C. & AH. S. Lucas, 1894. On a new skink lizard from Tasmania. Proceedings of the Linnean Society of New South Wales (2) 8 (2): 227 - 228." type="journal article" year="1894">Frost & Lucas (1894)</bibRefCitation>
|
||
, I nominate MV D2087 as
|
||
<typeStatus box="[505,661,2916,2955]" pageId="11" pageNumber="94">lectotype</typeStatus>
|
||
, leaving AM R4142 the only identifiable
|
||
<typeStatus box="[441,670,2962,3001]" pageId="11" pageNumber="94">paralectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="94" type="description">
|
||
<paragraph blockId="11.[221,1208,2732,3186]" lastBlockId="11.[1279,2266,2731,3184]" pageId="11" pageNumber="94">
|
||
The
|
||
<typeStatus box="[353,501,3008,3047]" pageId="11" pageNumber="94">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Frost & Lucas" authorityYear="1894" box="[582,846,3008,3047]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="94" phylum="Chordata" rank="species" species="tasmanicum">
|
||
<emphasis box="[582,846,3008,3047]" italics="true" pageId="11" pageNumber="94">H. tasmanicum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[879,996,3008,3047]" captionStart="Fig" captionStartId="13.[390,448,1428,1463]" captionTargetBox="[332,2110,471,1345]" captionTargetPageId="13" captionText="Fig. 8. Lectotype of Hemisphaeriodon tasmanicum Frost & Lucas, 1894 (MV D2087)." figureDoi="http://doi.org/10.5281/zenodo.4654799" httpUri="https://zenodo.org/record/4654799/files/figure.png" pageId="11" pageNumber="94">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation box="[1027,1048,3008,3047]" captionStart="Fig" captionStartId="14.[366,424,2686,2722]" captionTargetBox="[452,2026,524,2621]" captionTargetPageId="14" captionText="Fig. 9. Dorsal and lateral views of head shields of lectotype of Hemisphaeriodon tasmanicum." figureDoi="http://doi.org/10.5281/zenodo.4654801" httpUri="https://zenodo.org/record/4654801/files/figure.png" pageId="11" pageNumber="94">9</figureCitation>
|
||
) has the following combination of character states: nasals separated; prefrontals in narrow contact; supraoculars three; presuboculars two; postsuboculars four; supraciliaries six; supralabials six; infralabials seven, first two contacting postmental; nuchals three; upper palpebrals eight; lower palpebrals nine; temporals in a-configuration; midbody scales 24; paravertebral scales 62; subcaudal scales 71; subdigital lamellae 11/10; SVL
|
||
<quantity box="[2016,2157,2916,2955]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="11" pageNumber="94" unit="mm" value="101.0">101 mm</quantity>
|
||
; AGL
|
||
<quantity box="[1280,1406,2962,3000]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="11" pageNumber="94" unit="mm" value="63.0">63 mm</quantity>
|
||
; TL
|
||
<quantity box="[1504,1651,2962,3000]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="11" pageNumber="94" unit="mm" value="101.0">101 mm</quantity>
|
||
; FLL
|
||
<quantity box="[1775,1901,2962,3000]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="94" unit="mm" value="17.0">17 mm</quantity>
|
||
; HLL
|
||
<quantity box="[2027,2184,2962,3000]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="94" unit="mm" value="22.5">22.5 mm</quantity>
|
||
; HL
|
||
<quantity box="[1282,1442,3007,3046]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="11" pageNumber="94" unit="mm" value="15.9">15.9 mm</quantity>
|
||
; HW
|
||
<quantity box="[1568,1729,3007,3046]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="11" pageNumber="94" unit="mm" value="11.9">11.9 mm</quantity>
|
||
; HD
|
||
<quantity box="[1846,1988,3007,3046]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="11" pageNumber="94" unit="mm" value="7.6">7.6 mm</quantity>
|
||
. Both the type locality (Lake St Clair) and the morphology of the two type specimens clearly identify the name as belonging to the Tasmanian taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="95" type="biology_ecology">
|
||
<paragraph blockId="12.[215,1207,422,3172]" pageId="12" pageNumber="95">
|
||
<emphasis bold="true" box="[227,609,422,462]" pageId="12" pageNumber="95">Habitat and habits.</emphasis>
|
||
<bibRefCitation author="Rawlinson, P." box="[640,944,422,462]" pageId="12" pageNumber="95" pagination="291 - 338" refId="ref20413" refString="Rawlinson, P. A, 1974. Biogeography and ecology of the reptiles of Tasmania and the Bass Strait area. Pp. 291 - 338. In W. D. Williams (ed.). Biogeography and Ecology in Tasmania. W. Junk, The Hague, 498 pp." type="book chapter" year="1974">Rawlinson (1974)</bibRefCitation>
|
||
describes C.
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[227,412,468,507]" genus="Cyclodus" kingdom="Animalia" pageId="12" pageNumber="95" rank="species" species="casuarinae">
|
||
<emphasis box="[227,412,468,507]" italics="true" pageId="12" pageNumber="95">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
as terrestrial, commonest in clearings bordered or surrounded by dense arboreal vegetation, and using exposed patches of low vegetation or litter for basking sites and microenvironments under fallen logs and in deep litter for periods of inactivity. In his tabulation of habitat preferences (
|
||
<bibRefCitation author="Rawlinson, P." box="[811,1101,697,737]" pageId="12" pageNumber="95" pagination="291 - 338" refId="ref20413" refString="Rawlinson, P. A, 1974. Biogeography and ecology of the reptiles of Tasmania and the Bass Strait area. Pp. 291 - 338. In W. D. Williams (ed.). Biogeography and Ecology in Tasmania. W. Junk, The Hague, 498 pp." type="book chapter" year="1974">Rawlinson, 1974</bibRefCitation>
|
||
, table 11.5) he records the species from savanna woodland, dry sclerophyll forest and wet sclerophyll forest.
|
||
</paragraph>
|
||
<paragraph blockId="12.[215,1207,422,3172]" pageId="12" pageNumber="95">
|
||
Three more general publications give probably composite accounts of the ecology of members of the C.
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[289,484,928,967]" genus="Cyclodus" kingdom="Animalia" pageId="12" pageNumber="95" rank="species" species="casuarinae">
|
||
<emphasis box="[289,484,928,967]" italics="true" pageId="12" pageNumber="95">casuarinae</emphasis>
|
||
</taxonomicName>
|
||
complex, although the authors of all three have had some experience with the Tasmanian species.
|
||
<bibRefCitation author="Wilson, S. & D. G. Knowles" box="[383,872,1020,1060]" pageId="12" pageNumber="95" refId="ref21543" refString="Wilson, S. K & D. G. Knowles, 1988. Australia's Reptiles. A Photographic Reference to the Terrestrial Reptiles of Australia. William Collins, Sydney, 447 pp." type="book" year="1988">Wilson & Knowles (1988)</bibRefCitation>
|
||
describe the species as crepuscular to nocturnal, sheltering in grass tussocks and beneath leaf-litter, logs and surface debris in dry sclerophyll forest, woodlands, heathlands and swamplands, particularly where ground cover is dominated by tussock grasses.
|
||
<bibRefCitation author="Ehmann, H." box="[786,1059,1250,1290]" pageId="12" pageNumber="95" refId="ref19293" refString="Ehmann, H., 1992. Encyclopedia of Australian Animals. Reptiles. Collins Angus & Robertson, Pymble, 495 pp." type="book" year="1992">Ehmann (1992)</bibRefCitation>
|
||
records the species from "coastal plains, dunes, river flats, valleys and ranges. Vegetation of forests, woodlands, heathlands and tussock grasslands. In relatively closed vegetation structures, the species inhabits clearings, edges and other natural canopy openings that are invaded by dense low ground cover, especially spreading tussock grass." The species "shelters under embedded fallen logs, deep litter, stones and the procumbent spreading apron around the base of large tussock grasses and low bushes. It forages during the day and on warm nights, stalking and ambushing insects, snails, grubs and small lizards under partial cover."
|
||
<bibRefCitation author="Cogger, H. G." box="[363,622,1847,1887]" pageId="12" pageNumber="95" refId="ref19060" refString="Cogger, H. G., 1986. Reptiles and Amphibians of Australia. 4 th edn. Reed Books, Frenchs Forest, 688 pp." type="book" year="1986">Cogger (1986)</bibRefCitation>
|
||
reports the species from a wide variety of habitats, from "coastal heaths and sand dunes to the dry sclerophyll forests of the ranges. Common in coastal grazing lands. Normally crepuscular to nocturnal, it is usually found during the day in leaf-litter or under fallen timber."
|
||
</paragraph>
|
||
<paragraph blockId="12.[215,1207,422,3172]" pageId="12" pageNumber="95">
|
||
Specific data are available for few specimens.
|
||
<bibRefCitation author="Green, R. H." box="[219,466,2169,2209]" pageId="12" pageNumber="95" pagination="1 - 40" refId="ref19477" refString="Green, R. H., 1977. The vertebrate fauna of Maggs Mountain, Tasmania. Records of the Queen Victoria Museum (58): 1 - 40." type="journal article" year="1977">Green (1977)</bibRefCitation>
|
||
reported the species to be uncommon at his Maggs Mountain study site, with only three records. One adult was collected by a roadside in wet sclerophyll forest. Among the material examined for this study, habitat and microhabitat data are available for AM
|
||
<accessionNumber box="[372,518,2397,2438]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R65206" pageId="12" pageNumber="95">R65206</accessionNumber>
|
||
-08 (under pieces of thin wood/timber which were very dry underneath, on a north-east facing grassy slope with lots of timber and large dead trees scattered about the paddocks; A. Greer field notes), AM
|
||
<accessionNumber box="[477,622,2581,2622]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R65209" pageId="12" pageNumber="95">R65209</accessionNumber>
|
||
-1O (under dry rubbish in a cleared south-facing sloping paddock with rock outcrops, boards, rubbish and tree pieces scattered throughout, below a eucalypt forest; A. Greer field notes) and AM
|
||
<accessionNumber box="[310,457,2765,2805]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70069" pageId="12" pageNumber="95">R70069</accessionNumber>
|
||
-72 (under sheet iron in grassland bordered by dry sclerophyll forest; G. Stephenson field notes).
|
||
</paragraph>
|
||
<paragraph blockId="12.[215,1207,422,3172]" pageId="12" pageNumber="95">
|
||
For a series of six lizards,
|
||
<bibRefCitation author="Rawlinson, P." box="[741,1048,2903,2943]" pageId="12" pageNumber="95" pagination="291 - 338" refId="ref20413" refString="Rawlinson, P. A, 1974. Biogeography and ecology of the reptiles of Tasmania and the Bass Strait area. Pp. 291 - 338. In W. D. Williams (ed.). Biogeography and Ecology in Tasmania. W. Junk, The Hague, 498 pp." type="book chapter" year="1974">Rawlinson (1974)</bibRefCitation>
|
||
reported a voluntary thermal minimum of 27.1°C, a voluntary thermal maximum of 39°C and a mean active temperature of 32.6°C.
|
||
</paragraph>
|
||
<paragraph blockId="12.[215,1207,422,3172]" pageId="12" pageNumber="95">
|
||
<bibRefCitation author="Fleay, D." box="[264,501,3086,3126]" pageId="12" pageNumber="95" pagination="275 - 280" refId="ref19317" refString="Fleay, D., 1952. The Tasmanian or Marsupial Devil Its Habits and Family Life. Australian Museum Magazine 10 (9): 275 - 280." type="journal article" year="1952">Fleay (1952)</bibRefCitation>
|
||
reports predation by the Tasmanian Devil,
|
||
<emphasis box="[345,708,3133,3172]" italics="true" pageId="12" pageNumber="95">
|
||
<taxonomicName baseAuthorityName="Boitard" baseAuthorityYear="1841" box="[345,703,3133,3172]" class="Mammalia" family="Dasyuridae" genus="Sarcophilus" kingdom="Animalia" order="Dasyuromorphia" pageId="12" pageNumber="95" phylum="Chordata" rank="species" species="harrisii">Sarcophilus harrisii</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="12.[1277,2262,422,2713]" pageId="12" pageNumber="95">
|
||
<emphasis bold="true" box="[1278,1540,422,461]" pageId="12" pageNumber="95">Reproduction.</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3068164311" accessionNumber="D39139" collectingDate="1972-03-17" collectingDateMax="1984-03-13" collectingDateMin="1972-03-17" collectionCode="TM, OVM, MV, SVL" collectorName="However, I" county="March" location="Adult" municipality="January" pageId="12" pageNumber="95" specimenCode="C127, D39151-56" specimenCount="1">
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC295B17B0697CFE06" box="[1560,1658,422,461]" county="March" municipality="January" name="Adult" pageId="12" pageNumber="95">Adult</location>
|
||
females are available only from
|
||
<locationDeviation box="[1278,1790,467,507]" firstLocation="October" pageId="12" pageNumber="95" secondLocation="March">between October and March</locationDeviation>
|
||
(
|
||
<figureCitation box="[1822,1958,467,507]" captionStart="Fig" captionStartId="15.[420,475,1741,1777]" captionTargetBox="[469,2019,541,1647]" captionTargetPageId="15" captionText="Fig. 10. Seasonal variation in size of largest ovarian follicle, and occurrence of oviducal embryos (ov) i1 C. casuarinae." figureDoi="http://doi.org/10.5281/zenodo.4660407" httpUri="https://zenodo.org/record/4660407/files/figure.png" pageId="12" pageNumber="95">Fig. 10</figureCitation>
|
||
).
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC288C17C568F7FE30" box="[1999,2033,467,507]" county="March" municipality="January" name="In" pageId="12" pageNumber="95">In</location>
|
||
this period, females with grossly enlarged ovarian follicles were only present between late
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC29A814396873FD9C" box="[1771,1909,559,599]" county="March" municipality="January" name="October" pageId="12" pageNumber="95">October</location>
|
||
and early
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC270A14396A31FD4E" county="March" municipality="January" name="December" pageId="12" pageNumber="95">December</location>
|
||
, while oviducal yolks and embryos were present between early
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC2946149A69BCFD78" box="[1541,1722,652,691]" county="March" municipality="January" name="November" pageId="12" pageNumber="95">November</location>
|
||
and
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC285B149A688AFD78" box="[1816,1932,652,691]" county="March" municipality="January" name="March" pageId="12" pageNumber="95">March</location>
|
||
. Between January and
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC2A1A14AF6ACDFD2A" box="[1369,1483,697,737]" county="March" municipality="January" name="March" pageId="12" pageNumber="95">March</location>
|
||
, non-gravid females with small ovarian follicles were also collected.
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC286714F16863FCC5" box="[1828,1893,743,782]" county="March" municipality="January" name="The" pageId="12" pageNumber="95">The</location>
|
||
data, being pooled over many years, are insufficient to determine whether the occurrence of both gravid and non-gravid material in summer reflects non-annual reproduction or variation in the timing of parturition.
|
||
<collectorName box="[1910,2110,927,967]" pageId="12" pageNumber="95">However, I</collectorName>
|
||
suspect from the occurrence of enlarged ovarian follicles over almost a two month period, the occurrence of fully scaled and pigmented embryos in females collected as early as 19
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC2A9912416966FBB4" box="[1498,1632,1111,1151]" county="March" municipality="January" name="January" pageId="12" pageNumber="95">January</location>
|
||
(
|
||
<collectionCode box="[1664,1730,1111,1151]" country="Germany" name="Teylers Museum, Paleontologische" pageId="12" pageNumber="95">TM</collectionCode>
|
||
<specimenCode box="[1750,1843,1111,1151]" collectionCode="TM" country="Germany" name="Teylers Museum, Paleontologische" pageId="12" pageNumber="95">C127</specimenCode>
|
||
) and as late as
|
||
<location LSID="urn:lsid:plazi:treatment:6E1687E8E121FFDD2A6611A36889F8D4:E3606025E12BFFDC2720124167D4FBB4" box="[2147,2258,1111,1151]" county="March" municipality="January" name="March" pageId="12" pageNumber="95">March</location>
|
||
(
|
||
<collectionCode box="[1290,1390,1157,1198]" pageId="12" pageNumber="95">OVM</collectionCode>
|
||
<date box="[1424,1601,1157,1198]" pageId="12" pageNumber="95" value="1972-03-17">
|
||
<collectingDate box="[1424,1601,1157,1198]" pageId="12" pageNumber="95" value="1972-03-17">1972.3.17</collectingDate>
|
||
</date>
|
||
b), and the existence of neonates born
|
||
<collectingMunicipality box="[1373,1506,1204,1243]" pageId="12" pageNumber="95">January</collectingMunicipality>
|
||
and
|
||
<collectingCounty box="[1604,1715,1204,1243]" pageId="12" pageNumber="95">March</collectingCounty>
|
||
(
|
||
<collectionCode box="[1745,1815,1204,1243]" country="0" name="University of Montana Museum" pageId="12" pageNumber="95">MV</collectionCode>
|
||
<specimenCode box="[1832,2038,1204,1243]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="12" pageNumber="95">D39151-56</specimenCode>
|
||
,
|
||
<collectionCode box="[2065,2145,1204,1243]" pageId="12" pageNumber="95">SVL</collectionCode>
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="4.45" metricValueMin="4.15" pageId="12" pageNumber="95" unit="mm" value="43.0" valueMax="44.5" valueMin="41.5">41.5- 44.5 mm</quantity>
|
||
, born to
|
||
<accessionNumber box="[1604,1746,1250,1288]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39139" pageId="12" pageNumber="95">D39139</accessionNumber>
|
||
, collected 31 January, 1967; OVM
|
||
<date box="[1401,1580,1295,1335]" pageId="12" pageNumber="95" value="1984-03-13">
|
||
<collectingDate box="[1401,1580,1295,1335]" pageId="12" pageNumber="95" value="1984-03-13">1984.3.13</collectingDate>
|
||
</date>
|
||
b-g,
|
||
<collectionCode box="[1672,1751,1295,1335]" pageId="12" pageNumber="95">SVL</collectionCode>
|
||
<quantity box="[1773,2005,1295,1335]" metricMagnitude="-2" metricUnit="m" metricValue="4.375" metricValueMax="4.5" metricValueMin="4.25" pageId="12" pageNumber="95" unit="mm" value="43.75" valueMax="45.0" valueMin="42.5">42.5-45 mm</quantity>
|
||
, born after 4 weeks captivity to
|
||
<date box="[1644,1818,1341,1381]" pageId="12" pageNumber="95" value="1984-03-13">
|
||
<collectingDate box="[1644,1818,1341,1381]" pageId="12" pageNumber="95" value="1984-03-13">1984.3.13</collectingDate>
|
||
</date>
|
||
a, collected 9 February, 1984; SAM R8784-8785,
|
||
<collectionCode box="[1802,1882,1387,1426]" pageId="12" pageNumber="95">SVL</collectionCode>
|
||
<quantity box="[1915,2060,1387,1426]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="95" unit="mm" value="42.0">42 mm</quantity>
|
||
; collected January, 1967) that the latter is the case.
|
||
<bibRefCitation author="Rawlinson, P." pageId="12" pageNumber="95" pagination="291 - 338" refId="ref20413" refString="Rawlinson, P. A, 1974. Biogeography and ecology of the reptiles of Tasmania and the Bass Strait area. Pp. 291 - 338. In W. D. Williams (ed.). Biogeography and Ecology in Tasmania. W. Junk, The Hague, 498 pp." type="book chapter" year="1974">Rawlinson (1974)</bibRefCitation>
|
||
reported parturition in mid to late February
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="95" type="description">
|
||
<paragraph blockId="12.[1277,2262,422,2713]" pageId="12" pageNumber="95">
|
||
Mature males have turgid testes throughout the year (
|
||
<figureCitation box="[1291,1438,1571,1612]" captionStart="Fig" captionStartId="15.[414,471,3032,3067]" captionTargetBox="[450,2011,2016,2916]" captionTargetPageId="15" captionText="Fig. 11. Seasonal variation in testis length in C. casuarinae. Dots are turgid rounded testes, triangles are flaccid or flattened testes." figureDoi="http://doi.org/10.5281/zenodo.4660409" httpUri="https://zenodo.org/record/4660409/files/figure.png" pageId="12" pageNumber="95">Fig. 11</figureCitation>
|
||
). The largest testes were seen in males collected 27 May, 30 September and December. The latter two dates correspond to the inferred timing of ovulation, but the May date is well before this period.
|
||
<bibRefCitation author="Rawlinson, P." box="[1278,1608,1755,1795]" pageId="12" pageNumber="95" pagination="291 - 338" refId="ref20413" refString="Rawlinson, P. A, 1974. Biogeography and ecology of the reptiles of Tasmania and the Bass Strait area. Pp. 291 - 338. In W. D. Williams (ed.). Biogeography and Ecology in Tasmania. W. Junk, The Hague, 498 pp." type="book chapter" year="1974">Rawlinson (1974)</bibRefCitation>
|
||
stated that copulation occurs in Spring.
|
||
</paragraph>
|
||
<paragraph blockId="12.[1277,2262,422,2713]" pageId="12" pageNumber="95">
|
||
Females (SVL
|
||
<quantity box="[1597,1829,1847,1887]" metricMagnitude="-1" metricUnit="m" metricValue="1.385" metricValueMax="1.7399999999999998" metricValueMin="1.03" pageId="12" pageNumber="95" unit="mm" value="138.5" valueMax="174.0" valueMin="103.0">103-174 mm</quantity>
|
||
, x = 127.6, sd = 16.90, n = 25) carried from 4-14 (x = 7.6, sd = 3.06, n = 26) enlarged yolking ovarian follicles or oviducal yolks or embryos. There was a significant positive correlation between litter size and maternal SVL (litter = 0.117SVL - 7.219; r = 0.655*'*).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="95" type="discussion">
|
||
<paragraph blockId="12.[1277,2262,422,2713]" pageId="12" pageNumber="95">
|
||
There are several literature reports of litter size for C.
|
||
<emphasis box="[1338,1540,2169,2208]" italics="true" pageId="12" pageNumber="95">
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1338,1533,2169,2208]" genus="Cyclodus" kingdom="Animalia" pageId="12" pageNumber="95" rank="species" species="casuarinae">casuarinae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
although in the absence of associated locality data most could refer to any of the three species here recognised.
|
||
<bibRefCitation author="Worrell, E." box="[1715,1975,2260,2300]" pageId="12" pageNumber="95" refId="ref21578" refString="Worrell, E., 1963. Reptiles of Australia. Angus & Robertson, Sydney, 207 pp." type="book" year="1963">Worrell (1963)</bibRefCitation>
|
||
reported "about six young",
|
||
<bibRefCitation author="Frauca, H." box="[1498,1746,2306,2346]" pageId="12" pageNumber="95" refId="ref19347" refString="Frauca, H., 1966. Harry Frauca's Book of Reptiles. Jacaranda Press, Brisbane, 101 pp." type="book" year="1966">Frauca (1966)</bibRefCitation>
|
||
a litter of six, Peters (1967) 6-8 young,
|
||
<bibRefCitation author="Bustard, H. R." box="[1512,1787,2352,2392]" pageId="12" pageNumber="95" refId="ref18959" refString="Bustard, H. R., 1970. Australian Lizards. William Collins (Australia), Sydney, 162 pp." type="book" year="1970">Bustard (1970)</bibRefCitation>
|
||
a litter of 19,
|
||
<bibRefCitation author="Swanson, S." pageId="12" pageNumber="95" refId="ref21114" refString="Swanson, S., 1976. Lizards of Australia. Angus & Robertson, Sydney, 80 pp., 119 pIs." type="book" year="1976">Swanson (1976)</bibRefCitation>
|
||
around five young,
|
||
<bibRefCitation author="Wilson, S. & D. G. Knowles" box="[1779,2261,2397,2438]" pageId="12" pageNumber="95" refId="ref21543" refString="Wilson, S. K & D. G. Knowles, 1988. Australia's Reptiles. A Photographic Reference to the Terrestrial Reptiles of Australia. William Collins, Sydney, 447 pp." type="book" year="1988">Wilson & Knowles (1988)</bibRefCitation>
|
||
2-17, usually about four, and
|
||
<bibRefCitation author="Ehmann, H." box="[1858,2140,2443,2483]" pageId="12" pageNumber="95" refId="ref19293" refString="Ehmann, H., 1992. Encyclopedia of Australian Animals. Reptiles. Collins Angus & Robertson, Pymble, 495 pp." type="book" year="1992">Ehmann (1992)</bibRefCitation>
|
||
up to 19, usually about seven. More precise figures are given by
|
||
<bibRefCitation author="Rawlinson, P." box="[1462,1789,2535,2576]" pageId="12" pageNumber="95" pagination="291 - 338" refId="ref20413" refString="Rawlinson, P. A, 1974. Biogeography and ecology of the reptiles of Tasmania and the Bass Strait area. Pp. 291 - 338. In W. D. Williams (ed.). Biogeography and Ecology in Tasmania. W. Junk, The Hague, 498 pp." type="book chapter" year="1974">Rawlinson (1974)</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Greer, AE" box="[1821,2056,2535,2576]" pageId="12" pageNumber="95" refId="ref19593" refString="Greer, AE., 1989. The Biology and Evolution of Australian Lizards. Surrey Beatty & Sons, Chipping Norton, xvi + 264 pp." type="book" year="1989">Greer (1989)</bibRefCitation>
|
||
and Shine & Greer (1991), who give 2-7 (x = 4.1, n = 15), 4-14 (x = 6.8, n = 8) and 4-14 (x = 6.87, n = 8) respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="95" type="description">
|
||
<paragraph blockId="12.[1278,2264,2778,3095]" pageId="12" pageNumber="95">
|
||
Sex
|
||
<emphasis bold="true" box="[1362,1461,2778,2817]" pageId="12" pageNumber="95">ratio.</emphasis>
|
||
Overall, the ratio of mature males:females was 21:49, significantly different to 1:1 (XI = 10.41*"), although animals were only available between September and May. Seasonally, there was a significant difference in sex ratio between the gestation period and other months (male:female; November-March, 9:29; April-October, 6:3; Xl = 6.05*).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4654799" ID-Zenodo-Dep="4654799" box="[390,1900,1427,1464]" httpUri="https://zenodo.org/record/4654799/files/figure.png" pageId="13" pageNumber="96" startId="13.[390,448,1428,1463]" targetBox="[332,2110,471,1345]" targetPageId="13">
|
||
<paragraph blockId="13.[390,1900,1427,1464]" box="[390,1900,1427,1464]" pageId="13" pageNumber="96">
|
||
Fig. 8. Lectotype of
|
||
<taxonomicName authority="Frost & Lucas, 1894" authorityName="Frost & Lucas" authorityYear="1894" box="[765,1650,1427,1464]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="96" phylum="Chordata" rank="species" species="tasmanicum">
|
||
<emphasis box="[765,1263,1427,1464]" italics="true" pageId="13" pageNumber="96">Hemisphaeriodon tasmanicum</emphasis>
|
||
<bibRefCitation author="Frost, C. & AH & S. Lucas" box="[1288,1650,1427,1464]" pageId="13" pageNumber="96" pagination="227 - 228" refId="ref19368" refString="Frost, C. & AH. S. Lucas, 1894. On a new skink lizard from Tasmania. Proceedings of the Linnean Society of New South Wales (2) 8 (2): 227 - 228." type="journal article" year="1894">Frost & Lucas, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
(MV D2087).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="13" pageNumber="96" type="materials_examined">
|
||
<paragraph blockId="13.[229,1218,1578,3201]" lastBlockId="13.[1289,2273,1578,1823]" pageId="13" pageNumber="96">
|
||
Specimens examined. 1. NORTH-EAST
|
||
<collectingRegion box="[844,1012,1578,1613]" country="Australia" name="Tasmania" pageId="13" pageNumber="96">TASMANIA</collectingRegion>
|
||
: MV
|
||
<emphasis box="[1105,1216,1578,1613]" italics="true" pageId="13" pageNumber="96">D1051,</emphasis>
|
||
Scottsdale;
|
||
<accessionNumber box="[430,555,1620,1655]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39139" pageId="13" pageNumber="96">D39139</accessionNumber>
|
||
,
|
||
<accessionNumber box="[590,719,1620,1655]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39151" pageId="13" pageNumber="96">D39151</accessionNumber>
|
||
-56, 16 km SSW Scottsdale; QVM
|
||
<emphasis box="[356,440,1662,1697]" italics="true" pageId="13" pageNumber="96">1006,</emphasis>
|
||
Launceston; 1940.302, Montana; 1942.220, Deloraine; 1943.142, Hill Street, Launceston; 1944.76, Lebrina;
|
||
<date box="[237,408,1746,1780]" pageId="13" pageNumber="96" value="1962-03-28">1962.3.28</date>
|
||
a-b,
|
||
<date box="[505,683,1746,1780]" pageId="13" pageNumber="96" value="1963-03-12">1963.3.12</date>
|
||
-13, Winnaleah;
|
||
<date box="[994,1147,1746,1780]" pageId="13" pageNumber="96" value="1969-03-08">1969.3.8</date>
|
||
.a-b, Sideling, Launceston-Scottsdale road; 1972.3.188, south-east slope Ben Lomond; 1981.3.96, North Lilydale;
|
||
<date box="[1031,1189,1829,1864]" pageId="13" pageNumber="96" value="1984-03-13">1984.3.13</date>
|
||
ag, Liffey; 1987.3.74, Dairy Plains; TM C48, St Columba Falls, Pyengana. 2. NORTH-WEST
|
||
<collectingRegion box="[648,814,1913,1948]" country="Australia" name="Tasmania" pageId="13" pageNumber="96">TASMANIA</collectingRegion>
|
||
: AM
|
||
<accessionNumber box="[900,1029,1913,1948]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R37702" pageId="13" pageNumber="96">R37702</accessionNumber>
|
||
-03, Zeehan;
|
||
<accessionNumber box="[233,363,1954,1990]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R37704" pageId="13" pageNumber="96">R37704</accessionNumber>
|
||
-06,
|
||
<quantity box="[436,554,1954,1990]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="13" pageNumber="96" unit="mi" value="4.0">4 miles</quantity>
|
||
west Queenstown; MV D915, Emu Bay;
|
||
<accessionNumber box="[232,357,1996,2031]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39138" pageId="13" pageNumber="96">D39138</accessionNumber>
|
||
, 24 km east Marrawah;
|
||
<accessionNumber box="[781,905,1996,2031]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39140" pageId="13" pageNumber="96">D39140</accessionNumber>
|
||
, 25 km south-east Zeehan;
|
||
<accessionNumber box="[376,501,2038,2073]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39142" pageId="13" pageNumber="96">D39142</accessionNumber>
|
||
, 17 km south-west Smithton;
|
||
<accessionNumber box="[1015,1139,2038,2073]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39143" pageId="13" pageNumber="96">D39143</accessionNumber>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="96" unit="km" value="6.4">6.4 km</quantity>
|
||
east Queenstown;
|
||
<accessionNumber box="[582,705,2080,2115]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39144" pageId="13" pageNumber="96">D39144</accessionNumber>
|
||
, Collingwood River,
|
||
<quantity box="[1044,1143,2080,2115]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="96" unit="km" value="44.0">44 km</quantity>
|
||
east Queenstown; QM
|
||
<accessionNumber box="[539,650,2122,2157]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/J41561" pageId="13" pageNumber="96">J41561</accessionNumber>
|
||
, Franklin River, below Goodwins Peak;
|
||
<accessionNumber box="[330,443,2163,2199]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/J41562" pageId="13" pageNumber="96">J41562</accessionNumber>
|
||
, south side Macquarie Harbour; QVM
|
||
<date box="[1075,1208,2163,2199]" pageId="13" pageNumber="96" value="1958-03-07">1958.3.7</date>
|
||
, Burnie;
|
||
<date box="[374,527,2206,2240]" pageId="13" pageNumber="96" value="1969-03-13">1969.3.13</date>
|
||
, near Devonport;
|
||
<date box="[852,1004,2206,2240]" pageId="13" pageNumber="96" value="1972-03-19">1972.3.19</date>
|
||
, Ulverstone; 1972.3.121, Renison Bell. 3. CENTRAL
|
||
<collectingRegion box="[850,1019,2247,2282]" country="Australia" name="Tasmania" pageId="13" pageNumber="96">TASMANIA</collectingRegion>
|
||
: AM R4142 (
|
||
<typeStatus box="[242,445,2288,2325]" pageId="13" pageNumber="96">paralectotype</typeStatus>
|
||
of
|
||
<emphasis box="[514,769,2288,2325]" italics="true" pageId="13" pageNumber="96">
|
||
<taxonomicName authorityName="Frost & Lucas" authorityYear="1894" box="[514,754,2288,2325]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="96" phylum="Chordata" rank="species" species="tasmanicum">H. tasmanicum</taxonomicName>
|
||
),
|
||
</emphasis>
|
||
Lake St Clair;
|
||
<accessionNumber box="[1029,1159,2288,2325]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R65206" pageId="13" pageNumber="96">R65206</accessionNumber>
|
||
-08,
|
||
<quantity box="[234,360,2330,2366]" metricMagnitude="4" metricUnit="m" metricValue="1.7399999999999998" pageId="13" pageNumber="96" unit="km" value="17.4">17.4 km</quantity>
|
||
north Breona via Highway 5;
|
||
<emphasis box="[860,1045,2330,2366]" italics="true" pageId="13" pageNumber="96">
|
||
<accessionNumber box="[860,989,2330,2366]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R65209" pageId="13" pageNumber="96">R65209</accessionNumber>
|
||
-1O,
|
||
</emphasis>
|
||
north-east side Bronte Lagoon, just north Lyell Highway; MV
|
||
<accessionNumber box="[1073,1205,2372,2407]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D56347" pageId="13" pageNumber="96">D56347</accessionNumber>
|
||
- 49, Mount Field;
|
||
<accessionNumber box="[510,635,2414,2449]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D56658" pageId="13" pageNumber="96">D56658</accessionNumber>
|
||
,
|
||
<quantity box="[657,764,2414,2449]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="96" unit="km" value="2.6">2.6 km</quantity>
|
||
south Bronte Lagoon; NTM R9292, Cradle Mountain; QVM 1962.3.40, Erriba;
|
||
<date box="[1074,1207,2455,2491]" pageId="13" pageNumber="96" value="1964-03-03">1964.3.3</date>
|
||
, Waratah Hellyer Spur road;
|
||
<date box="[774,931,2498,2533]" pageId="13" pageNumber="96" value="1972-03-17">1972.3.17</date>
|
||
a-b, Great Lake;
|
||
<date box="[233,384,2539,2575]" pageId="13" pageNumber="96" value="1976-03-22">1976.3.22</date>
|
||
, Maggs Mountain; 1979.3.37, 1979.3.39, Maggs Mountain Hut; SAM R8784-85, south extreme, Lake Sinclair [St Clair]; R8798, Lake Sinclair [St Clair]. 4. SOUTH-EAST
|
||
<collectingRegion box="[230,400,2665,2700]" country="Australia" name="Tasmania" pageId="13" pageNumber="96">TASMANIA</collectingRegion>
|
||
: AM 4785, Ouse River;
|
||
<emphasis box="[876,1006,2665,2700]" italics="true" pageId="13" pageNumber="96">
|
||
<accessionNumber box="[876,1000,2665,2700]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R10053" pageId="13" pageNumber="96">R10053</accessionNumber>
|
||
,
|
||
</emphasis>
|
||
Catamaran;
|
||
<accessionNumber box="[231,351,2706,2741]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R68001" pageId="13" pageNumber="96">R68001</accessionNumber>
|
||
, Russell Falls National Park;
|
||
<accessionNumber box="[889,1018,2706,2741]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70069" pageId="13" pageNumber="96">R70069</accessionNumber>
|
||
-72, Russell River,
|
||
<quantity box="[353,451,2748,2783]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="96" unit="km" value="5.0">5 km</quantity>
|
||
north Judbury in Huon Valley;
|
||
<emphasis box="[1062,1213,2748,2783]" italics="true" pageId="13" pageNumber="96">R107594,</emphasis>
|
||
Eaglehawk Neck; ANWC R3071, Huon River,
|
||
<quantity box="[1027,1133,2790,2825]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="96" unit="km" value="14.0">14 km</quantity>
|
||
east Judbury; MNHP 7131, Bruny Island (type of C.
|
||
<emphasis box="[1021,1212,2831,2867]" italics="true" pageId="13" pageNumber="96">
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1839" box="[1021,1195,2831,2867]" genus="Cyclodus" kingdom="Animalia" pageId="13" pageNumber="96" rank="species" species="casuarinae">casuarinae</taxonomicName>
|
||
);
|
||
</emphasis>
|
||
MV D29, Port Arthur; D2593, Port Esperance; D5733-34, Tasman Island; D7919-20, Kingston; D7991, Ridgeway; D1l218, Huon Valley;
|
||
<accessionNumber box="[625,748,2957,2992]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39128" pageId="13" pageNumber="96">D39128</accessionNumber>
|
||
,
|
||
<quantity box="[779,897,2957,2992]" metricMagnitude="3" metricUnit="m" metricValue="9.6" pageId="13" pageNumber="96" unit="km" value="9.6">9.6 km</quantity>
|
||
west Geavestown;
|
||
<accessionNumber box="[230,352,2998,3033]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39133" pageId="13" pageNumber="96">D39133</accessionNumber>
|
||
, Coal Mines;
|
||
<accessionNumber box="[619,742,2998,3033]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39141" pageId="13" pageNumber="96">D39141</accessionNumber>
|
||
,
|
||
<quantity box="[779,914,2998,3033]" metricMagnitude="4" metricUnit="m" metricValue="1.13" pageId="13" pageNumber="96" unit="km" value="11.3">11.3 km</quantity>
|
||
south Huonville;
|
||
<accessionNumber box="[230,352,3040,3076]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/D39145" pageId="13" pageNumber="96">D39145</accessionNumber>
|
||
,
|
||
<quantity box="[384,503,3040,3076]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="13" pageNumber="96" unit="km" value="5.6">5.6 km</quantity>
|
||
east-south-east Longley; QVM
|
||
<date box="[1054,1206,3040,3076]" pageId="13" pageNumber="96" value="1972-03-16">1972.3.16</date>
|
||
, Antill Ponds; TM C39, Kettering; C1l4, Lunawanna, South Bruny Island; C126-27, Lower Longley; C258, Dover; C267ab, Cygnet; C273, Maria Island; C305, West Moonah; C318, Counsel Creek, Maria Island; C997, Wellesley Street, South Hobart. UNLOCALISED: AM R2917, no data;
|
||
<accessionNumber box="[2042,2164,1620,1655]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R14402" pageId="13" pageNumber="96">R14402</accessionNumber>
|
||
, north
|
||
<collectingRegion box="[1289,1437,1662,1698]" country="Australia" name="Tasmania" pageId="13" pageNumber="96">Tasmania</collectingRegion>
|
||
; MV D161l, D2087 (
|
||
<typeStatus box="[1798,1936,1662,1698]" pageId="13" pageNumber="96">lectotype</typeStatus>
|
||
of
|
||
<emphasis box="[2009,2271,1662,1698]" italics="true" pageId="13" pageNumber="96">
|
||
<taxonomicName authorityName="Frost & Lucas" authorityYear="1894" box="[2009,2255,1662,1698]" class="Reptilia" family="Scincidae" genus="Hemisphaeriodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="96" phylum="Chordata" rank="species" species="tasmanicum">H. tasmanicum</taxonomicName>
|
||
),
|
||
</emphasis>
|
||
D2088-90, D2092, D4919,
|
||
<accessionNumber box="[1727,1911,1703,1739]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/QM113774" pageId="13" pageNumber="96">QM 113774</accessionNumber>
|
||
, QVM
|
||
<date box="[2036,2170,1703,1739]" pageId="13" pageNumber="96" value="1950-03-01">1950.3.1</date>
|
||
, SAM R2231, R6131,
|
||
<collectingRegion box="[1538,1687,1745,1780]" country="Australia" name="Tasmania" pageId="13" pageNumber="96">Tasmania</collectingRegion>
|
||
; SAM R59, "
|
||
<collectingRegion box="[1900,2159,1745,1780]" country="Australia" name="South Australia" pageId="13" pageNumber="96">South Australia</collectingRegion>
|
||
"; 2MB 8193,
|
||
<collectingCountry box="[1393,1529,1787,1823]" name="Australia" pageId="13" pageNumber="96">Australia</collectingCountry>
|
||
(type of C.
|
||
<emphasis box="[1772,1935,1787,1823]" italics="true" pageId="13" pageNumber="96">
|
||
<taxonomicName authorityName="Peters" authorityYear="1875" box="[1772,1920,1787,1823]" genus="Cyclodus" kingdom="Animalia" pageId="13" pageNumber="96" rank="species" species="nigricans">nigricans</taxonomicName>
|
||
).
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |