140 lines
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140 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.148.48957" ID-GBIF-Dataset="731107ec-bcb7-48d7-961a-18a761f48483" ID-PMC="PMC7266832" ID-Pensoft-Pub="1314-2003-148-51" ID-Pensoft-UUID="B20B6DE0095C5D7CAA6A7CF6C003EF81" ID-PubMed="32523393" ModsDocID="1314-2003-148-51" checkinTime="1590560150783" checkinUser="pensoft" docAuthor="Ito, Takuro, Yu, Chih-Chieh, Yokota, Masatsugu & Kokubugata, Goro" docDate="2020" docId="37C5934724F5555D89A99F274E8F30F4" docLanguage="en" docName="PhytoKeys 148: 51-70" docOrigin="PhytoKeys 148" docSource="http://dx.doi.org/10.3897/phytokeys.148.48957" docTitle="Sedum formosanum subsp. miyakojimense Ito & Yu & Yokota & Kokubugata 2020, subsp. nov." docType="treatment" docVersion="4" id="B20B6DE0095C5D7CAA6A7CF6C003EF81" lastPageNumber="51" masterDocId="B20B6DE0095C5D7CAA6A7CF6C003EF81" masterDocTitle="Sedum formosanum subsp. miyakojimense (Crassulaceae), a new subspecies from Miyako-jima Island of the Ryukyu Islands, Japan" masterLastPageNumber="70" masterPageNumber="51" pageNumber="51" updateTime="1668139803971" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Sedum formosanum subsp. miyakojimense (Crassulaceae), a new subspecies from Miyako-jima Island of the Ryukyu Islands, Japan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ito, Takuro</mods:namePart>
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<mods:affiliation>Division of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kitashirakawa Oiwake-cho, Sakyo-ku, Kyoto, 606 - 8502, Japan & Department of Botany, National Museum of Nature and Science, Amakubo 4 - 1 - 1, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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<mods:nameIdentifier type="email">896wfbt5@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yu, Chih-Chieh</mods:namePart>
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<mods:affiliation>CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yokota, Masatsugu</mods:namePart>
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<mods:affiliation>Laboratory of Ecology and Systematics, Faculty of Science, University of the Ryukyus, Senbaru 1, Nishihara, Okinawa 903 - 0213, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4620-4140</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kokubugata, Goro</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Nature and Science, Amakubo 4 - 1 - 1, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>148</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>70</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.148.48957</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.148.48957</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-148-51</mods:identifier>
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<treatment ID-GBIF-Taxon="164360805" LSID="urn:lsid:plazi:treatment:37C5934724F5555D89A99F274E8F30F4" httpUri="http://treatment.plazi.org/id/37C5934724F5555D89A99F274E8F30F4" lastPageNumber="51" pageId="0" pageNumber="51">
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<subSubSection pageId="0" pageNumber="51" type="nomenclature">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName LSID="37C59347-24F5-555D-89A9-9F274E8F30F4" authority="Takuro Ito, Yokota & Kokub." authorityName="Ito & Yu & Yokota & Kokubugata" authorityYear="2020" class="Magnoliopsida" family="Crassulaceae" genus="Sedum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sedum formosanum subsp. miyakojimense" order="Saxifragales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="subSpecies" species="formosanum" status="subsp. nov." subSpecies="miyakojimense">Sedum formosanum N.E. Brown., subsp. miyakojimense Takuro Ito, Yokota & Kokub.</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="51">subsp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sedum formosanum subsp. miyakojimense (A-D T. Ito 2402, 2408, Miyako-jima Island of the Ryukyus, Japan) and S. formosanum subsp. formosanum (E Kume-jima Island of the Ryukyus, Japan. F, G NewTaipei City, Taiwan). A, B, E habit C, F flower D, G leaf. Scale bars: 1 mm (D-F)." figureDoi="10.3897/phytokeys.148.48957.figure3" httpUri="https://binary.pensoft.net/fig/413780" pageId="0" pageNumber="51">Figs 3A-D</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sedum formosanum subsp. miyakojimense. A flower B sepal C leaf, adaxial D leaf, abaxial E habit. Scale bars: 1 mm (A-D); 1 cm (E). Line drawings by Naomi Kizaki." figureDoi="10.3897/phytokeys.148.48957.figure4" httpUri="https://binary.pensoft.net/fig/413781" pageId="0" pageNumber="51">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="materials_examined">
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<paragraph pageId="0" pageNumber="51">Type.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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Japan. The Ryukyus: Miyako Islands, Miyako-jima Island, Gusukube, 5 April 2015,
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<emphasis italics="true" pageId="0" pageNumber="51">Takuro Ito 2402</emphasis>
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(holotype: TNS)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="diagnosis">
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<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName class="Magnoliopsida" family="Crassulaceae" genus="Sedum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sedum formosanum subsp. miyakojimense" order="Saxifragales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="subSpecies" species="formosanum" subSpecies="miyakojimense">Sedum formosanum subsp. miyakojimense</taxonomicName>
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differs from its close relative
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<taxonomicName authority="subsp. formosanum" lsidName="S. formosanum subsp. formosanum" pageId="0" pageNumber="51" rank="subSpecies" species="formosanum" subSpecies="formosanum">S. formosanum subsp. formosanum</taxonomicName>
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in being perennial, polycarpic, and having lateral branches arising from the leaf axils.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="description">
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<paragraph pageId="0" pageNumber="51">Description.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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Perennial herb, fleshy, glabrous. First year stem stout, erect, partly woody, 1-5 lateral branches in the leaf axils, 3-10 cm tall, with lax rosettes; rosettes 2.5-6 cm wide with 7-15 leaves. Flowering stems fleshy, 10-20 cm tall, base ca. 5 mm broad, yellowish green, erect or sprawling and creeping at base. Roots fibrous, sometimes adventitious at the leaf scar. Leaves alternate, occasionally verticillate, sessile, green or yellowish, flattish,
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<normalizedToken originalValue="±">+/-</normalizedToken>
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thick, spatulate to oblanceolate, 1.1-3.1 cm long, 0.3-1.0 cm wide, apex rounded, base long, attenuate, margins entire. Inflorescences terminal, cymes, basically trifurcate with 3 primary axes, sometimes with 2, 4, or 5 primary axes; primary axis 2-8 cm long, ascending, 1 to several times irregularly and often unequally forking, with a flower at each fork, ultimate branches 1-2 cm long, 3-7 flowered; bracts leaf-like, smaller than cauline leaves. Flowers 5 (rarely 6)-merous, 7-11 mm wide, sessile. Sepals 5, free, yellowish green, fleshy, flattish, unequal in size, obovate to oblanceolate, 1.8-4.5 mm long, 1.2-3.3 mm wide, apex round or obtuse, base spurred. Petals 5, bright yellow, lanceolate, 4.6-6 mm long, 1.3-1.6 mm wide, apex acuminate, base slightly connate. Stamens 10, shorter than petals, 4.2-5 mm long, erect at flowering, two-whorled arrangement; anthers oblong-lanceolate, ca. 0.5 mm long, deep yellow before dehiscence. Pistils 5, 5.2-6.3 mm long; carpels 5, free, connate at the base, gibbous ventrally. Fruits star-shaped, follicle, erect, 5.3-6.8 mm long. Flowering in April to June.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="taxonomic note">
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<paragraph pageId="0" pageNumber="51">Taxonomic note.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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This new subspecies is classified in the sect.
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<taxonomicName lsidName="" pageId="0" pageNumber="51" rank="section" section="Sedum">Sedum</taxonomicName>
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because of its adaxially gibbous carpels (
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<bibRefCitation author="Fu, KT" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="51" refId="B3" refString="Fu, KT, Ohba, H, 2001. Crassulaceae. In: Wu ZY, Raven PH (Eds) Flora of China 8. Missouri Botanical Garden Press, St. Louis, 244-401." title="Crassulaceae. In: Wu ZY, Raven PH (Eds) Flora of China 8. Missouri Botanical Garden Press, St. Louis, 244 - 401." year="2001">Fu and Ohba 2001</bibRefCitation>
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) (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sedum formosanum subsp. miyakojimense (A-D T. Ito 2402, 2408, Miyako-jima Island of the Ryukyus, Japan) and S. formosanum subsp. formosanum (E Kume-jima Island of the Ryukyus, Japan. F, G NewTaipei City, Taiwan). A, B, E habit C, F flower D, G leaf. Scale bars: 1 mm (D-F)." figureDoi="10.3897/phytokeys.148.48957.figure3" httpUri="https://binary.pensoft.net/fig/413780" pageId="0" pageNumber="51">3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="etymology">
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<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="51">The epithet refers to the Japanese name of the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="distribution">
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<paragraph pageId="0" pageNumber="51">Distribution and habitat.</paragraph>
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<paragraph pageId="0" pageNumber="51">Endemic to the southeastern portion of Miyako-jima Island (The Ryukyus), on sunny, coastal limestone.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="materials_examined">
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<paragraph pageId="0" pageNumber="51">Additional specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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Japan. The Ryukyus: the Miyakojima Islands, Miyako-jima Island, Gusukube, 5 April 2015,
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<emphasis italics="true" pageId="0" pageNumber="51">Takuro Ito 2403, 2408</emphasis>
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(isotype: TNS).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="conservation">
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<paragraph pageId="0" pageNumber="51">Conservation.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<emphasis bold="true" pageId="0" pageNumber="51">IUCN Red list category</emphasis>
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: Critically Endangered (CR). The distribution of
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<taxonomicName class="Magnoliopsida" family="Crassulaceae" genus="Sedum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sedum formosanum subsp. miyakojimense" order="Saxifragales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="subSpecies" species="formosanum" subSpecies="miyakojimense">Sedum formosanum subsp. miyakojimense</taxonomicName>
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is restricted to only one location ca. 0.15 km2 in Miyako-jima Island, the Ryukyu Islands. The population of the species contains fewer than 200 mature individuals. The plant occurs on limestone rocks scattered in a private golf course, therefore, it is not formally protected. In the future, the population could become threatened, given ongoing land development for tourism in the Ryukyus. Because of the small population size (≤ 250 mature individuals) and small area of occupancy (≤ 10 km2),
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<taxonomicName lsidName="S. formosanum subsp. miyakojimense" pageId="0" pageNumber="51" rank="subSpecies" species="formosanum" subSpecies="miyakojimense">S. formosanum subsp. miyakojimense</taxonomicName>
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is classified as CR (
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<bibRefCitation author="IUCN" journalOrPublisher="Series B, Botany" pageId="0" pageNumber="51" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B11" refString="IUCN, 2019. Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee. https://www.iucnredlist.org/resources/redlistguidelines" title="Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee." url="https://www.iucnredlist.org/resources/redlistguidelines" year="2019">IUCN 2019</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="japanese common name">
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<paragraph pageId="0" pageNumber="51">Japanese common name.</paragraph>
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<paragraph pageId="0" pageNumber="51">Miyako-hama-mannen-gusa (nov.).</paragraph>
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</subSubSection>
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</treatment>
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</document> |