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<document id="7134647E1C7B34FD587609EE83E9C838" ID-CLB-Dataset="9327" ID-DOI="10.11646/zootaxa.4894.3.4" ID-GBIF-Dataset="74322c33-5f04-4474-a9ff-0dbc181e7bfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4315887" ID-ZooBank="451A3DBD-A70C-426B-8455-8A554DE99144" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607645634773" checkinUser="plazi" docAuthor="Oanh, Le Thi Kieu &amp; Boyko, Christopher B." docDate="2020" docId="34489947DA70CC00FF7039235603B25D" docLanguage="en" docName="zootaxa.4894.3.4.pdf" docOrigin="Zootaxa 4894 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cancrion khanhensis Oanh &amp; Boyko 2020, sp. nov." docType="treatment" docUuid="5DACACB1-3B6D-4EB9-916A-27F4C6DC6490" docUuidSource="ZooBank" docVersion="7" lastPageNumber="375" masterDocId="C871E13FDA72CC09FFE73868546FB428" masterDocTitle="Cancrion khanhensis sp. nov. (Crustacea: Isopoda: Entoniscidae) infesting Monomia haanii (Stimpson, 1858) (Crustacea: Brachyura: Portunidae) from Nha Trang Bay, Khanh Hoa, Vietnam, with remarks on larval stages of entoniscids and description of a new family, genus and two new species of hyperparasites" masterLastPageNumber="386" masterPageNumber="366" pageNumber="368" updateTime="1698847412080" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="18EB16D235158DC831DCCD8B1AB0D6AA">Cancrion khanhensis sp. nov. (Crustacea: Isopoda: Entoniscidae) infesting Monomia haanii (Stimpson, 1858) (Crustacea: Brachyura: Portunidae) from Nha Trang Bay, Khanh Hoa, Vietnam, with remarks on larval stages of entoniscids and description of a new family, genus and two new species of hyperparasites</mods:title>
</mods:titleInfo>
<mods:name id="892E0EF71FF675E264921370E48F8F2A" type="personal">
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</mods:role>
<mods:namePart id="37F16AFC2B82BF1AD51BE89F0BFF8501">Oanh, Le Thi Kieu</mods:namePart>
<mods:affiliation id="902612055E39B7AA0E904B42CF91D8BB">Vietnam-Russia Tropical Centre, Nha Trang city, Khanh Hoa province, Vietnam.</mods:affiliation>
</mods:name>
<mods:name id="43F3A99A6981EAB3495A301DCF234C6B" type="personal">
<mods:role id="B0D787DC0A356DF8CE43E46E39869118">
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<mods:namePart id="16240B8BDFAC4E5013CC5DD5057BB59E">Boyko, Christopher B.</mods:namePart>
<mods:affiliation id="74A3089BC28FE63D96C15D4D5BEF84E3">Department of Biology, Hofstra University, Hempstead, NY 11549, USA.</mods:affiliation>
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<mods:title id="AD609D0B47FB1014480E064C367746D6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B06B842750C3F421716511D01C303084">
<mods:date id="76B0E458D33C8570131F249D06D66608">2020</mods:date>
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<mods:number id="ED1135CFFDAF34961E157EFA2A03FB20">2020-12-10</mods:number>
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<mods:identifier id="0F2967E7BB47753C0AD2316624AE8993" type="CLB-Dataset">9327</mods:identifier>
<mods:identifier id="F73B5C32197027A0D355DB8F9DB15222" type="DOI">10.11646/zootaxa.4894.3.4</mods:identifier>
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<treatment id="34489947DA70CC00FF7039235603B25D" ID-DOI="http://doi.org/10.5281/zenodo.4329024" ID-GBIF-Taxon="176084951" ID-Zenodo-Dep="4329024" LSID="urn:lsid:zoobank.org:act:5DACACB1-3B6D-4EB9-916A-27F4C6DC6490" httpUri="http://treatment.plazi.org/id/34489947DA70CC00FF7039235603B25D" lastPageId="9" lastPageNumber="375" pageId="2" pageNumber="368">
<subSubSection id="F4FB7BDADA70CC0BFF703923566FB54D" box="[151,512,330,357]" pageId="2" pageNumber="368" type="nomenclature">
<paragraph id="BC5E2851DA70CC0BFF703923566FB54D" blockId="2.[151,512,330,392]" box="[151,512,330,357]" pageId="2" pageNumber="368">
<heading id="E7169F3DDA70CC0BFF703923566FB54D" bold="true" box="[151,512,330,357]" fontSize="11" level="1" pageId="2" pageNumber="368" reason="1">
<emphasis id="8E95F443DA70CC0BFF703923566FB54D" bold="true" box="[151,512,330,357]" pageId="2" pageNumber="368">
<taxonomicName id="7BE153D2DA70CC0BFF70392355F3B54D" ID-CoL="84765" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[151,412,330,357]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA70CC0BFF70392355F3B54D" bold="true" box="[151,412,330,357]" italics="true" pageId="2" pageNumber="368">Cancrion khanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFE443923566FB54D" box="[419,512,331,357]" pageId="2" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA70CC0BFF70390657A1B5E3" pageId="2" pageNumber="368" type="description">
<paragraph id="BC5E2851DA70CC0BFF70390655DEB5A0" blockId="2.[151,512,330,392]" box="[151,433,366,392]" pageId="2" pageNumber="368">
<heading id="E7169F3DDA70CC0BFF70390655DEB5A0" bold="true" box="[151,433,366,392]" fontSize="11" level="1" pageId="2" pageNumber="368" reason="1">
<emphasis id="8E95F443DA70CC0BFF70390655DEB5A0" bold="true" box="[151,433,366,392]" pageId="2" pageNumber="368">
(
<figureCitation id="24DA34D4DA70CC0BFF783906556AB5A0" box="[159,261,366,392]" captionStart-0="FIGURE" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,168,1766,1792]" captionStartId-1="4.[151,250,1920,1944]" captionStartId-2="6.[151,250,1740,1764]" captionStartId-3="8.[151,250,1943,1967]" captionTargetBox-0="[158,1425,187,1734]" captionTargetBox-1="[158,1424,189,1892]" captionTargetBox-2="[209,1380,658,1710]" captionTargetBox-3="[209,1378,189,1911]" captionTargetId-0="figure-25@3.[151,1436,179,1741]" captionTargetId-1="figure-17@4.[151,1436,181,1896]" captionTargetId-2="figure-227@6.[201,1387,651,1716]" captionTargetId-3="figure-17@8.[200,1387,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." captionText-1="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." captionText-2="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." captionText-3="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4315889" figureDoi-1="http://doi.org/10.5281/zenodo.4315891" figureDoi-2="http://doi.org/10.5281/zenodo.4315893" figureDoi-3="http://doi.org/10.5281/zenodo.4315895" httpUri-0="https://zenodo.org/record/4315889/files/figure.png" httpUri-1="https://zenodo.org/record/4315891/files/figure.png" httpUri-2="https://zenodo.org/record/4315893/files/figure.png" httpUri-3="https://zenodo.org/record/4315895/files/figure.png" pageId="2" pageNumber="368">Figs 14</figureCitation>
,
<figureCitation id="24DA34D4DA70CC0BFEF639065571B5A0" box="[273,286,366,392]" captionStart="FIGURE 8" captionStartId="14.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure-17@14.[151,1436,181,1786]" captionTargetPageId="14" captionText="FIGURE 8. A, Cancrion khanhensis sp. nov. mature ovigerous female; B, C. khanhensis sp. nov. mature female showing dorsolateral ovarian processes (arrows); C, C. khanhensis sp. nov, immature female (stage 8); D, C. khanhensis sp. nov. mature female infested by Stellatoniscus tentaculus gen. nov., sp. nov. mature female; E, S. tentaculus gen. nov., sp. nov. immature females (stage III); F, S. tentaculus gen. nov., sp. nov. mature female (stage IV). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4315903" httpUri="https://zenodo.org/record/4315903/files/figure.png" pageId="2" pageNumber="368">8</figureCitation>
AC, 9C, D)
</emphasis>
</heading>
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF7039DB57A1B5E3" blockId="2.[151,974,435,459]" box="[151,974,435,459]" pageId="2" pageNumber="368">
LSID
<uri id="C8702453DA70CC0BFF3139DB57A1B5E3" box="[214,974,435,459]" pageId="2" pageNumber="368">
urn:lsid:zoobank.org:act:
<uuid id="C8471284DA70CC0BFE2839DB57A1B5E3" box="[463,974,435,459]" pageId="2" pageNumber="368">5DACACB1-3B6D-4EB9-916A-27F4C6DC6490</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA70CC0CFF7039915748B039" lastPageId="5" lastPageNumber="371" pageId="2" pageNumber="368" type="materials_examined">
<paragraph id="BC5E2851DA70CC0BFF70399156B0B674" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF703991552AB63B" bold="true" box="[151,325,505,531]" pageId="2" pageNumber="368">Type material.</emphasis>
<materialsCitation id="0C89220CDA70CC0BFEAC399156B0B674" ID-GBIF-Occurrence="2995018305" collectingDate="2019-04-25" collectionCode="NOMV-E" collectorName="L. T. K. Oanh" country="Vietnam" county="Cu Hin" latitude="12.15" location="Nha Trang bay" longLatPrecision="776" longitude="109.23" municipality="Ke Ga" pageId="2" pageNumber="368" specimenCode="NOMV-E.56943" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<emphasis id="8E95F443DA70CC0BFEAC399155D4B63B" box="[331,443,505,532]" italics="true" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFEAC399155DCB63C" bold="true" box="[331,435,505,532]" pageId="2" pageNumber="368">
<typeStatus id="635A96F3DA70CC0BFEAC399155DCB63C" box="[331,435,505,532]" pageId="2" pageNumber="368" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
Female (
<quantity id="7B1985B4DA70CC0BFDC5399156E7B63C" box="[546,648,505,532]" metricMagnitude="-2" metricUnit="m" metricValue="2.44" pageId="2" pageNumber="368" unit="mm" value="24.4">24.4 mm</quantity>
TL, stage 12),
<location id="B93E7E8ADA70CC0BFCD6399157B7B63B" LSID="urn:lsid:plazi:treatment:34489947DA70CC00FF7039235603B25D:B93E7E8ADA70CC0BFCD6399157B7B63B" box="[817,984,505,532]" country="Vietnam" county="Cu Hin" latitude="12.15" longLatPrecision="776" longitude="109.23" municipality="Ke Ga" name="Nha Trang bay" pageId="2" pageNumber="368" stateProvince="Khanh Hoa">Nha Trang bay</location>
,
<collectingRegion id="7E25E6B3DA70CC0BFC043991500AB63C" box="[995,1125,505,532]" country="Vietnam" name="Khanh Hoa" pageId="2" pageNumber="368">Khanh Hoa</collectingRegion>
,
<collectingCountry id="C4F668C1DA70CC0BFB97399150BDB63B" box="[1136,1234,505,531]" name="Vietnam" pageId="2" pageNumber="368">Vietnam</collectingCountry>
, from Ke Ga cape (
<geoCoordinate id="D9D54E96DA70CC0BFF783A75556EB61F" box="[159,257,541,567]" degrees="12.30" direction="north" orientation="latitude" pageId="2" pageNumber="368" precision="555" value="12.3">12.30°N</geoCoordinate>
,
<geoCoordinate id="D9D54E96DA70CC0BFEEB3A755519B61F" box="[268,374,541,567]" degrees="109.24" direction="east" orientation="longitude" pageId="2" pageNumber="368" precision="555" value="109.24">109.24°E</geoCoordinate>
) to Cu Hin cape (12.15°N, 109.23°E), ex male
<taxonomicName id="7BE153D2DA70CC0BFC993A765092B610" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[894,1277,541,568]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFC993A765058B61F" box="[894,1079,541,567]" italics="true" pageId="2" pageNumber="368">Monomia haanii</emphasis>
(Stimpson, 1858)
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFAEC3A755135B610" box="[1291,1370,541,568]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="368" unit="mm" value="82.0">82 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFF703A295549B673" box="[151,294,577,604]" pageId="2" pageNumber="368" value="2019-04-25">
<collectingDate id="D81BF779DA70CC0BFF703A295549B673" box="[151,294,577,604]" pageId="2" pageNumber="368" value="2019-04-25">25 Apr 2019</collectingDate>
</date>
, coll.
<collectorName id="11144D87DA70CC0BFE8E3A2A566BB673" box="[361,516,577,604]" pageId="2" pageNumber="368">L. T. K. Oanh</collectorName>
(
<specimenCode id="EC47802ADA70CC0BFDF53A2956BBB674" box="[530,724,577,604]" pageId="2" pageNumber="368">NOMV-E.56943</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF203A0D5503B68C" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<materialsCitation id="0C89220CDA70CC0BFF203A0D5503B68C" ID-GBIF-Occurrence="2995018306" collectingDate="2019-02-23" collectionCode="NOMV-E" pageId="2" pageNumber="368" specimenCount="2" typeStatus="allotype">
<emphasis id="8E95F443DA70CC0BFF203A0D5549B6A8" bold="true" box="[199,294,613,640]" italics="true" pageId="2" pageNumber="368">
<typeStatus id="635A96F3DA70CC0BFF203A0D5549B6A8" box="[199,294,613,640]" pageId="2" pageNumber="368" type="allotype">Allotype</typeStatus>
</emphasis>
. Male (
<quantity id="7B1985B4DA70CC0BFE633A0D558CB6A8" box="[388,483,613,640]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="368" unit="mm" value="4.9">4.9 mm</quantity>
), same locality and collector data as holotype, ex female
<taxonomicName id="7BE153D2DA70CC0BFB603A0E5097B657" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1159,1272,613,640]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFB603A0E50F1B657" box="[1159,1182,614,639]" italics="true" pageId="2" pageNumber="368">M</emphasis>
.
<emphasis id="8E95F443DA70CC0BFB573A0D5097B657" box="[1200,1272,613,639]" italics="true" pageId="2" pageNumber="368">haanii</emphasis>
</taxonomicName>
,
<date id="C85F0E91DA70CC0BFAEE3A0D51F3B657" box="[1289,1436,613,639]" pageId="2" pageNumber="368" value="2019-02-23">
<collectingDate id="D81BF779DA70CC0BFAEE3A0D51F3B657" box="[1289,1436,613,639]" pageId="2" pageNumber="368" value="2019-02-23">23 Feb 2019</collectingDate>
</date>
(
<collectionCode id="DAF0B094DA70CC0BFF783AE1557AB68C" box="[159,277,649,676]" pageId="2" pageNumber="368">NOMV-E</collectionCode>
.56945).
</materialsCitation>
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF203AC657EAB788" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF203AC6552FB6EF" box="[199,320,686,712]" italics="true" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF203AC65556B6E0" bold="true" box="[199,313,686,712]" italics="true" pageId="2" pageNumber="368">
<typeStatus id="635A96F3DA70CC0BFF203AC65556B6E0" box="[199,313,686,712]" pageId="2" pageNumber="368" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
All same locality and collector data as
<typeStatus id="635A96F3DA70CC0BFCE63AC55708B6EF" box="[769,871,685,711]" pageId="2" pageNumber="368" type="holotype">holotype</typeStatus>
.
<emphasis id="8E95F443DA70CC0BFC953AC55073B6EF" bold="true" box="[882,1052,685,711]" pageId="2" pageNumber="368">
<date id="C85F0E91DA70CC0BFC953AC55078B6EF" box="[882,1047,685,711]" pageId="2" pageNumber="368" value="2019-01">January 2019</date>
.
</emphasis>
Mature female (
<quantity id="7B1985B4DA70CC0BFB3E3AC55144B6E0" box="[1241,1323,685,712]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="368" unit="mm" value="26.0">26 mm</quantity>
TL, stage 12), ex female
<taxonomicName id="7BE153D2DA70CC0BFEA63ABA55C0B6C3" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[321,431,721,747]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFEA63ABA55C0B6C3" box="[321,431,721,747]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFE593AB95660B6C4" box="[446,527,721,748]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="368" unit="mm" value="57.0">57 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFDBC3AB956BAB6C3" box="[603,725,721,748]" pageId="2" pageNumber="368" value="2019-01-02">2 Jan 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFD033AB9575CB6C3" box="[740,819,721,747]" pageId="2" pageNumber="368">CNVB</collectionCode>
10002); Mature femae (
<quantity id="7B1985B4DA70CC0BFBA03AB950F8B6C4" box="[1095,1175,721,748]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="368" unit="mm" value="24.0">24 mm</quantity>
TL, stage 12), ex male
<taxonomicName id="7BE153D2DA70CC0BFF703A9E556AB727" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[151,261,757,783]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFF703A9E556AB727" box="[151,261,757,783]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFEF43A9D550BB738" box="[275,356,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="368" unit="mm" value="92.0">92 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFE483A9D5646B727" box="[431,553,757,784]" pageId="2" pageNumber="368" value="2019-01-02">2 Jan 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFDDF3A9D56E8B727" box="[568,647,757,783]" pageId="2" pageNumber="368">CNVB</collectionCode>
10003); Two mature females (
<quantity id="7B1985B4DA70CC0BFC073A9D5035B738" box="[992,1114,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="2" pageNumber="368" unit="mm" value="30.5" valueMax="35.0" valueMin="26.0">2635 mm</quantity>
TL, stage 12, larger infested with
<taxonomicName id="7BE153D2DA70CC0BFF363B715588B71B" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[209,487,793,819]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFF363B715588B71B" box="[209,487,793,819]" italics="true" pageId="2" pageNumber="368">Stellatoniscus tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFE083B7156ADB71B" box="[495,706,793,820]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFE083B715636B71B" bold="true" box="[495,601,793,819]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFD8F3B7156ADB71B" bold="true" box="[616,706,793,819]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA70CC0BFCC63B715713B71C" box="[801,892,793,820]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="368" unit="mm" value="3.1">3.1 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFBE13B72501BB71B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1030,1140,793,819]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFBE13B72501BB71B" box="[1030,1140,793,819]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB633B7150BAB71C" box="[1156,1237,793,820]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="368" unit="mm" value="70.0">70 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFAC63B7151F3B71B" box="[1313,1436,793,820]" pageId="2" pageNumber="368" value="2019-01-02">2 Jan 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFF783B555481B77F" box="[159,238,829,855]" pageId="2" pageNumber="368">CNVB</collectionCode>
10004); Mature female (
<quantity id="7B1985B4DA70CC0BFDE23B55563AB770" box="[517,597,829,856]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="368" unit="mm" value="29.0">29 mm</quantity>
TL, stage 12), male (
<quantity id="7B1985B4DA70CC0BFCA43B5557F1B770" box="[835,926,829,856]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="368" unit="mm" value="4.3">4.3 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFBE93B565014B77F" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1038,1147,829,855]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFBE93B565014B77F" box="[1038,1147,829,855]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB6D3B5550B6B770" box="[1162,1241,829,856]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="368" unit="mm" value="61.0">61 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFAC33B5551F3B77F" box="[1316,1436,829,856]" pageId="2" pageNumber="368" value="2019-01-08">8 Jan 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFF783B095481B753" box="[159,238,865,891]" pageId="2" pageNumber="368">CNVB</collectionCode>
10005); Mature female (
<quantity id="7B1985B4DA70CC0BFDE83B09560EB754" box="[527,609,865,892]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="368" unit="mm" value="32.0">32 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFC413B0A5058B753" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[934,1079,865,892]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFC413B0A57DBB754" box="[934,948,866,892]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFC233B095058B753" box="[964,1079,865,891]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFBA73B095178B753" box="[1088,1303,865,892]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFBA73B0950C3B753" bold="true" box="[1088,1196,865,891]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFB5B3B095178B753" bold="true" box="[1212,1303,865,891]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA70CC0BFA9F3B0954A8B788" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="368" unit="mm" value="4.3">4.3 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFEDE3BEE55C9B7B7" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[313,422,901,927]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFEDE3BEE55C9B7B7" box="[313,422,901,927]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFE523BED566AB788" box="[437,517,901,928]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="368" unit="mm" value="89.0">89 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFDB73BED56A6B7B7" box="[592,713,901,928]" pageId="2" pageNumber="368" value="2019-01-08">8 Jan 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFD3F3BED5748B7B7" box="[728,807,901,927]" pageId="2" pageNumber="368">CNVB</collectionCode>
10006).
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF203BC156EDB12F" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF203BC155EEB7EB" bold="true" box="[199,385,937,963]" pageId="2" pageNumber="368">
<date id="C85F0E91DA70CC0BFF203BC15513B7EB" box="[199,380,937,963]" pageId="2" pageNumber="368" value="2019-02">February 2019</date>
.
</emphasis>
Mature female (
<quantity id="7B1985B4DA70CC0BFDA43BC156FAB7EC" box="[579,661,937,964]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="368" unit="mm" value="26.0">26 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFC3A3BC1509AB7EB" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[989,1269,937,963]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFC3A3BC1509AB7EB" box="[989,1269,937,963]" italics="true" pageId="2" pageNumber="368">Stellatoniscus tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFB183BC154A7B7CF" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFB183BC15105B7EB" bold="true" box="[1279,1386,937,963]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFA9C3BC154A7B7CF" bold="true" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFEBF3BA655A8B7CF" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[344,455,973,999]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFEBF3BA655A8B7CF" box="[344,455,973,999]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFE303BA55646B7C0" box="[471,553,973,1000]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="368" unit="mm" value="69.0">69 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFD913BA5576AB7CF" box="[630,773,973,999]" pageId="2" pageNumber="368" value="2019-02-23">23 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFCF23BA5570BB7CF" box="[789,868,973,999]" pageId="2" pageNumber="368">CNVB</collectionCode>
10007); Juvenile female (
<quantity id="7B1985B4DA70CC0BFB693BA5508FB7C0" box="[1166,1248,973,1000]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="368" unit="mm" value="22.0">22 mm</quantity>
TL, stage 8), ex male
<taxonomicName id="7BE153D2DA70CC0BFF323B9A552CB023" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[213,323,1009,1035]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFF323B9A552CB023" box="[213,323,1009,1035]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFEB43B9955CBB024" box="[339,420,1009,1036]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="368" unit="mm" value="65.0">65 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFE163B995611B023" box="[497,638,1009,1035]" pageId="2" pageNumber="368" value="2019-02-23">23 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFD693B9956B2B023" box="[654,733,1009,1035]" pageId="2" pageNumber="368">CNVB</collectionCode>
10008); Mature female (
<quantity id="7B1985B4DA70CC0BFC1E3B995026B024" box="[1017,1097,1009,1036]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="368" unit="mm" value="31.0">31 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFA603B9A5565B007" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFA603B9A51FAB024" box="[1415,1429,1010,1036]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFF703C7D5565B007" box="[151,266,1045,1071]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFEF53C7D5588B007" box="[274,487,1045,1072]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFEF53C7D5513B007" bold="true" box="[274,380,1045,1071]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFE6C3C7D5588B007" bold="true" box="[395,487,1045,1071]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="8E95F443DA70CC0BFE003C7D5580B007" bold="true" box="[487,495,1045,1071]" pageId="2" pageNumber="368">)</emphasis>
, male (
<quantity id="7B1985B4DA70CC0BFDA33C7D56CEB018" box="[580,673,1045,1072]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="368" unit="mm" value="4.5">4.5 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFCF23C7E57EBB007" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[789,900,1045,1071]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFCF23C7E57EBB007" box="[789,900,1045,1071]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFC743C7D578AB018" box="[915,997,1045,1072]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="368" unit="mm" value="65.0">65 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFBD63C7D50D0B007" box="[1073,1215,1045,1071]" pageId="2" pageNumber="368" value="2019-02-28">28 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFB283C7D512AB018" box="[1231,1349,1045,1072]" pageId="2" pageNumber="368">NOMV-E</collectionCode>
.56944); Two mature females (
<quantity id="7B1985B4DA70CC0BFE773C515662B07C" box="[400,525,1081,1108]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="2" pageNumber="368" unit="mm" value="30.5" valueMax="32.0" valueMin="29.0">2932 mm</quantity>
TL, stage 12) male (
<quantity id="7B1985B4DA70CC0BFCE73C515732B07C" box="[768,861,1081,1108]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="368" unit="mm" value="3.9">3.9 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFC0C3C525035B07B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1003,1114,1081,1107]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFC0C3C525035B07B" box="[1003,1114,1081,1107]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB8C3C5150D2B07C" box="[1131,1213,1081,1108]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="368" unit="mm" value="65.0">65 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFAEB3C5151F3B07B" box="[1292,1436,1081,1107]" pageId="2" pageNumber="368" value="2019-02-28">28 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFF783C355481B05F" box="[159,238,1117,1143]" pageId="2" pageNumber="368">CNVB</collectionCode>
10009); Juvenile female (
<quantity id="7B1985B4DA70CC0BFDC73C35561CB050" box="[544,627,1117,1144]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="368" unit="mm" value="26.0">26 mm</quantity>
TL, stage 8), ex male
<taxonomicName id="7BE153D2DA70CC0BFC663C36579DB05F" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[897,1010,1117,1143]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFC663C36579DB05F" box="[897,1010,1117,1143]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFBE33C355038B050" box="[1028,1111,1117,1144]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="368" unit="mm" value="64.0">64 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFB4E3C355154B05F" box="[1193,1339,1117,1143]" pageId="2" pageNumber="368" value="2019-02-28">28 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFAAA3C3551F3B05F" box="[1357,1436,1117,1143]" pageId="2" pageNumber="368">CNVB</collectionCode>
10010); Two mature females (
<quantity id="7B1985B4DA70CC0BFE0D3CE9560BB0B4" box="[490,612,1153,1180]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="2" pageNumber="368" unit="mm" value="29.5" valueMax="31.0" valueMin="28.0">2831 mm</quantity>
TL, stage 12, largest infested with
<taxonomicName id="7BE153D2DA70CC0BFC153CEA50EEB0B3" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1010,1153,1153,1180]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFC153CEA506FB0B4" box="[1010,1024,1154,1180]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFBE93CE950EEB0B3" box="[1038,1153,1153,1179]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFB6E3CE95134B0B3" box="[1161,1371,1153,1180]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFB6E3CE9509DB0B3" bold="true" box="[1161,1266,1153,1179]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFAE63CE95134B0B3" bold="true" box="[1281,1371,1153,1179]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), two males (
<quantity id="7B1985B4DA70CC0BFF003CCD551CB0E8" box="[231,371,1189,1216]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.8" metricValueMin="3.4" pageId="2" pageNumber="368" unit="mm" value="4.1" valueMax="4.8" valueMin="3.4">3.44.8 mm</quantity>
), ex immature male
<taxonomicName id="7BE153D2DA70CC0BFDB13CCE56ACB097" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[598,707,1189,1215]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFDB13CCE56ACB097" box="[598,707,1189,1215]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFD353CCD574DB0E8" box="[722,802,1189,1216]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="368" unit="mm" value="55.0">55 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFC8B3CCD5797B097" box="[876,1016,1189,1215]" pageId="2" pageNumber="368" value="2019-02-28">28 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFBE13CCD503AB097" box="[1030,1109,1189,1215]" pageId="2" pageNumber="368">CNVB</collectionCode>
10011); Two mature females (
<quantity id="7B1985B4DA70CC0BFF783CA15575B0CC" box="[159,282,1225,1252]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.4" metricValueMin="2.4" pageId="2" pageNumber="368" unit="mm" value="29.0" valueMax="34.0" valueMin="24.0">2434 mm</quantity>
TL, stage 12, both infested with
<taxonomicName id="7BE153D2DA70CC0BFD743CA2574CB0CB" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[659,803,1225,1252]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFD743CA256CEB0CC" box="[659,673,1226,1252]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFD573CA1574CB0CB" box="[688,803,1225,1251]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFCCC3CA15790B0CB" box="[811,1023,1225,1252]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFCCC3CA157FAB0CB" bold="true" box="[811,917,1225,1251]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFC433CA15790B0CB" bold="true" box="[932,1023,1225,1251]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA70CC0BFBBA3CA150D5B0CC" box="[1117,1210,1225,1252]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="368" unit="mm" value="4.2">4.2 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFAC93CA251F2B0CB" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1326,1437,1225,1251]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFAC93CA251F2B0CB" box="[1326,1437,1225,1251]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFF473C85549FB120" box="[160,240,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="368" unit="mm" value="61.0">61 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFEDC3C8555A8B12F" box="[315,455,1261,1287]" pageId="2" pageNumber="368" value="2019-02-28">28 Feb 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFE323C85564BB12F" box="[469,548,1261,1287]" pageId="2" pageNumber="368">CNVB</collectionCode>
10012).
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF203D79558BB20F" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF203D795531B103" bold="true" box="[199,350,1297,1323]" pageId="2" pageNumber="368">
<date id="C85F0E91DA70CC0BFF203D795536B103" box="[199,345,1297,1323]" pageId="2" pageNumber="368" value="2019-03">March 2019</date>
.
</emphasis>
Juvenile female (
<quantity id="7B1985B4DA70CC0BFDC03D795618B104" box="[551,631,1297,1324]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="368" unit="mm" value="21.0">21 mm</quantity>
TL, stage 8), ex immature female
<taxonomicName id="7BE153D2DA70CC0BFC193D7A5003B103" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1022,1132,1297,1323]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFC193D7A5003B103" box="[1022,1132,1297,1323]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB9C3D7950A3B104" box="[1147,1228,1297,1324]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="368" unit="mm" value="47.0">47 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFAFF3D7951F3B103" box="[1304,1436,1297,1324]" pageId="2" pageNumber="368" value="2019-03-05">5 Mar 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFF783D5D5481B167" box="[159,238,1333,1359]" pageId="2" pageNumber="368">CNVB</collectionCode>
10013); Mature female (
<quantity id="7B1985B4DA70CC0BFDEB3D5D5632B178" box="[524,605,1333,1360]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="368" unit="mm" value="31.0">31 mm</quantity>
TL, stage 12), male (
<quantity id="7B1985B4DA70CC0BFCB53D5D57C0B178" box="[850,943,1333,1360]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="368" unit="mm" value="4.2">4.2 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFBDD3D5E50C6B167" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1082,1193,1333,1359]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFBDD3D5E50C6B167" box="[1082,1193,1333,1359]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB5E3D5D5165B178" box="[1209,1290,1333,1360]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="368" unit="mm" value="73.0">73 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFAB03D5D54A0B15B" pageId="2" pageNumber="368" value="2019-03-05">5 Mar 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFF3A3D315543B15B" box="[221,300,1369,1395]" pageId="2" pageNumber="368">CNVB</collectionCode>
10014); Two mature females (
<quantity id="7B1985B4DA70CC0BFD673D315696B15C" box="[640,761,1369,1396]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="2" pageNumber="368" unit="mm" value="29.5" valueMax="34.0" valueMin="25.0">2534 mm</quantity>
TL, stage 12), two males (
<quantity id="7B1985B4DA70CC0BFBC73D3150C4B15C" box="[1056,1195,1369,1396]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.7" metricValueMin="3.7" pageId="2" pageNumber="368" unit="mm" value="4.2" valueMax="4.7" valueMin="3.7">3.74.7 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFAD73D3251F2B15B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1328,1437,1369,1395]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFAD73D3251F2B15B" box="[1328,1437,1369,1395]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFF783D15549EB1B0" box="[159,241,1405,1432]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="368" unit="mm" value="70.0">70 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFED93D1555ACB1BF" box="[318,451,1405,1432]" pageId="2" pageNumber="368" value="2019-03-05">5 Mar 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFE343D15564DB1BF" box="[467,546,1405,1431]" pageId="2" pageNumber="368">CNVB</collectionCode>
10015); Mature female (
<quantity id="7B1985B4DA70CC0BFCA73D1557FDB1B0" box="[832,914,1405,1432]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="368" unit="mm" value="27.0">27 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFB343D16510CB1BF" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1235,1379,1405,1432]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFB343D16508EB1B0" box="[1235,1249,1406,1432]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFB173D15510CB1BF" box="[1264,1379,1405,1431]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFA8C3D15555FB193" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFA8C3D1554A7B193" bold="true" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFF313DC9555FB193" bold="true" box="[214,304,1441,1467]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (4.0 mm), ex female
<taxonomicName id="7BE153D2DA70CC0BFD883DCA56B3B193" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[623,732,1441,1467]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFD883DCA56B3B193" box="[623,732,1441,1467]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFD0C3DC95754B194" box="[747,827,1441,1468]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="368" unit="mm" value="66.0">66 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFC613DC95066B193" box="[902,1033,1441,1468]" pageId="2" pageNumber="368" value="2019-03-05">5 Mar 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFBF03DC95009B193" box="[1047,1126,1441,1467]" pageId="2" pageNumber="368">CNVB</collectionCode>
10016); Mature female (
<quantity id="7B1985B4DA70CC0BFA983DC954ACB1C8" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="368" unit="mm" value="32.0">32 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFE193DAE56E2B1F7" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[510,653,1477,1504]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFE193DAE5663B1C8" box="[510,524,1478,1504]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFDFD3DAD56E2B1F7" box="[538,653,1477,1503]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFD723DAD5709B1F7" box="[661,870,1477,1504]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFD723DAD5691B1F7" bold="true" box="[661,766,1477,1503]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFCEB3DAD5709B1F7" bold="true" box="[780,870,1477,1503]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFC143DAE500FB1F7" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1011,1120,1477,1503]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFC143DAE500FB1F7" box="[1011,1120,1477,1503]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB883DAD50AFB1C8" box="[1135,1216,1477,1504]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="368" unit="mm" value="58.0">58 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFAEC3DAD51F3B1F7" box="[1291,1436,1477,1504]" pageId="2" pageNumber="368" value="2019-03-21">21 Mar 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFF783D815481B22B" box="[159,238,1513,1539]" pageId="2" pageNumber="368">CNVB</collectionCode>
10017); Two immature females (
<quantity id="7B1985B4DA70CC0BFD833D8156B1B22C" box="[612,734,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="2" pageNumber="368" unit="mm" value="22.0" valueMax="23.0" valueMin="21.0">2123 mm</quantity>
TL, stage 8), male (
<quantity id="7B1985B4DA70CC0BFC583D815075B22C" box="[959,1050,1513,1540]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="368" unit="mm" value="3.6">3.6 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFB6B3D825096B22B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1164,1273,1513,1539]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFB6B3D825096B22B" box="[1164,1273,1513,1539]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFAEF3D815137B22C" box="[1288,1368,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="368" unit="mm" value="73.0">73 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFF703E655547B20F" box="[151,296,1549,1576]" pageId="2" pageNumber="368" value="2019-03-21">21 Mar 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFED13E6555EAB20F" box="[310,389,1549,1575]" pageId="2" pageNumber="368">CNVB</collectionCode>
10018).
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF203E59517DB2BC" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF203E595520B263" bold="true" box="[199,335,1585,1611]" pageId="2" pageNumber="368">
<date id="C85F0E91DA70CC0BFF203E595525B263" box="[199,330,1585,1611]" pageId="2" pageNumber="368" value="2019-04">April 2019</date>
.
</emphasis>
Mature female (
<quantity id="7B1985B4DA70CC0BFDE83E59560EB264" box="[527,609,1585,1612]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="368" unit="mm" value="26.0">26 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFC403E5A5057B263" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[935,1080,1585,1612]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFC403E5A57DAB264" box="[935,949,1586,1612]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFC223E595057B263" box="[965,1080,1585,1611]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFBA63E595178B263" box="[1089,1303,1585,1612]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFBA63E5950C3B263" bold="true" box="[1089,1196,1585,1611]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFB5B3E595178B263" bold="true" box="[1212,1303,1585,1611]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA70CC0BFA9F3E5954A8B258" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="368" unit="mm" value="3.6">3.6 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFEDE3E3E55C9B247" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[313,422,1621,1647]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFEDE3E3E55C9B247" box="[313,422,1621,1647]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFE523E3D566AB258" box="[437,517,1621,1648]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="368" unit="mm" value="74.0">74 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFDB73E3D56A1B247" box="[592,718,1621,1648]" pageId="2" pageNumber="368" value="2019-04-05">5 Apr 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFD3A3E3D5743B247" box="[733,812,1621,1647]" pageId="2" pageNumber="368">CNVB</collectionCode>
10019); Immature female (
<quantity id="7B1985B4DA70CC0BFB863E3D50DEB258" box="[1121,1201,1621,1648]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="368" unit="mm" value="21.0">21 mm</quantity>
TL, stage 8, infested with
<taxonomicName id="7BE153D2DA70CC0BFF373E125530B2BB" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[208,351,1657,1684]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFF373E1254B1B2BC" box="[208,222,1658,1684]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFF0B3E115530B2BB" box="[236,351,1657,1683]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFE813E115658B2BB" box="[358,567,1657,1684]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFE813E1155A0B2BB" bold="true" box="[358,463,1657,1683]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFE3A3E115658B2BB" bold="true" box="[477,567,1657,1683]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFD4A3E125775B2BB" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[685,794,1657,1683]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFD4A3E125775B2BB" box="[685,794,1657,1683]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFCCE3E115716B2BC" box="[809,889,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="368" unit="mm" value="82.0">82 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFC233E11503FB2BB" box="[964,1104,1657,1684]" pageId="2" pageNumber="368" value="2019-04-25">25 Apr 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFBB83E1150C1B2BB" box="[1119,1198,1657,1683]" pageId="2" pageNumber="368">CNVB</collectionCode>
10020).
</paragraph>
<paragraph id="BC5E2851DA70CC0BFF203EF555AAB3F0" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="368">
<emphasis id="8E95F443DA70CC0BFF203EF5552BB29F" bold="true" box="[199,324,1693,1719]" pageId="2" pageNumber="368">
<date id="C85F0E91DA70CC0BFF203EF55550B29F" box="[199,319,1693,1719]" pageId="2" pageNumber="368" value="2019-05">May 2019</date>
.
</emphasis>
Mature male (
<quantity id="7B1985B4DA70CC0BFE0F3EF55657B290" box="[488,568,1693,1720]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="368" unit="mm" value="30.0">30 mm</quantity>
TL, stage 12), ex male
<taxonomicName id="7BE153D2DA70CC0BFCA63EF657C0B29F" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[833,943,1693,1719]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFCA63EF657C0B29F" box="[833,943,1693,1719]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFC5A3EF55061B290" box="[957,1038,1693,1720]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="368" unit="mm" value="75.0">75 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFBBE3EF5508FB29F" box="[1113,1248,1693,1720]" pageId="2" pageNumber="368" value="2019-05-05">5 May 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFB083EF55151B29F" box="[1263,1342,1693,1719]" pageId="2" pageNumber="368">CNVB</collectionCode>
10021); Mature female (
<quantity id="7B1985B4DA70CC0BFEAA3EA955F1B2F4" box="[333,414,1729,1756]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="368" unit="mm" value="29.0">29 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFD383EAA571FB2F3" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[735,880,1729,1756]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFD383EAA5682B2F4" box="[735,749,1730,1756]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFD1A3EA9571FB2F3" box="[765,880,1729,1755]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFC9F3EA95023B2F3" box="[888,1100,1729,1756]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFC9F3EA9578DB2F3" bold="true" box="[888,994,1729,1755]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFC153EA95023B2F3" bold="true" box="[1010,1100,1729,1755]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA70CC0BFB4B3EA95167B2F4" box="[1196,1288,1729,1756]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="368" unit="mm" value="3.9">3.9 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFA993EAA54B0B2D7" baseAuthorityName="Stimpson" baseAuthorityYear="1858" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFA993EAA54B0B2D7" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFF083E8D552FB328" box="[239,320,1765,1792]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="368" unit="mm" value="65.0">65 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFE6A3E8D5678B2D7" box="[397,535,1765,1792]" pageId="2" pageNumber="368" value="2019-05-05">5 May 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFDC03E8D5619B2D7" box="[551,630,1765,1791]" pageId="2" pageNumber="368">CNVB</collectionCode>
10022); Immature female (
<quantity id="7B1985B4DA70CC0BFC483E8D579DB328" box="[943,1010,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="368" unit="mm" value="5.0">5 mm</quantity>
TL, stage 2), mature female (
<quantity id="7B1985B4DA70CC0BFAA33E8D51F7B328" box="[1348,1432,1765,1792]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="368" unit="mm" value="24.0">24 mm</quantity>
, stage 12, infested with
<taxonomicName id="7BE153D2DA70CC0BFE453F62565CB30B" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[418,563,1801,1828]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFE453F6255DFB30C" box="[418,432,1802,1828]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFE273F61565CB30B" box="[448,563,1801,1827]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFDDB3F61577CB30B" box="[572,787,1801,1828]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFDDB3F6156C7B30B" bold="true" box="[572,680,1801,1827]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFD5F3F61577CB30B" bold="true" box="[696,787,1801,1827]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFC683F625790B30B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[911,1023,1801,1827]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFC683F625790B30B" box="[911,1023,1801,1827]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFBE83F61500DB30C" box="[1039,1122,1801,1828]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="368" unit="mm" value="59.0">59 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFB563F615153B30B" box="[1201,1340,1801,1828]" pageId="2" pageNumber="368" value="2019-05-05">5 May 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFAAA3F6151F3B30B" box="[1357,1436,1801,1827]" pageId="2" pageNumber="368">CNVB</collectionCode>
10023); Two mature females (
<quantity id="7B1985B4DA70CC0BFE083F455604B360" box="[495,619,1837,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="2" pageNumber="368" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">2630 mm</quantity>
TL, stage 12, both infested with
<taxonomicName id="7BE153D2DA70CC0BFC0D3F465015B36F" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1002,1146,1837,1864]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFC0D3F465797B360" box="[1002,1016,1838,1864]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFBE03F455015B36F" box="[1031,1146,1837,1863]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFB643F455136B36F" box="[1155,1369,1837,1864]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFB643F455081B36F" bold="true" box="[1155,1262,1837,1863]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFB193F455136B36F" bold="true" box="[1278,1369,1837,1863]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), two males (
<quantity id="7B1985B4DA70CC0BFF003F39551CB344" box="[231,371,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.7" metricValueMin="3.2" pageId="2" pageNumber="368" unit="mm" value="3.95" valueMax="4.7" valueMin="3.2">3.24.7 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA70CC0BFE043F3A563FB343" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[483,592,1873,1899]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFE043F3A563FB343" box="[483,592,1873,1899]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFDB83F3956C1B344" box="[607,686,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="368" unit="mm" value="71.0">71 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFD1E3F395710B343" box="[761,895,1873,1900]" pageId="2" pageNumber="368" value="2019-05-05">5 May 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFC6A3F3957B3B343" box="[909,988,1873,1899]" pageId="2" pageNumber="368">CNVB</collectionCode>
10024); Mature male (
<quantity id="7B1985B4DA70CC0BFB3A3F395142B344" box="[1245,1325,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="368" unit="mm" value="32.0">32 mm</quantity>
TL, stage 12), ex female
<taxonomicName id="7BE153D2DA70CC0BFEA53F1E55DFB3A7" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[322,432,1909,1935]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFEA53F1E55DFB3A7" box="[322,432,1909,1935]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFE583F1D567EB3B8" box="[447,529,1909,1936]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="368" unit="mm" value="57.0">57 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFDBA3F1D569BB3A7" box="[605,756,1909,1936]" pageId="2" pageNumber="368" value="2019-05-16">16 May 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFCE33F1D573CB3A7" box="[772,851,1909,1935]" pageId="2" pageNumber="368">CNVB</collectionCode>
10025); Two mature females (
<quantity id="7B1985B4DA70CC0BFB483F1D5144B3B8" box="[1199,1323,1909,1936]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="2" pageNumber="368" unit="mm" value="27.5" valueMax="29.0" valueMin="26.0">2629 mm</quantity>
TL, stage 12, smaller infested with
<taxonomicName id="7BE153D2DA70CC0BFE513FF25629B39B" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[438,582,1945,1972]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA70CC0BFE513FF255ABB39C" box="[438,452,1946,1972]" italics="true" pageId="2" pageNumber="368">S</emphasis>
.
<emphasis id="8E95F443DA70CC0BFE343FF15629B39B" box="[467,582,1945,1971]" italics="true" pageId="2" pageNumber="368">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA70CC0BFDA93FF1574EB39B" box="[590,801,1945,1972]" pageId="2" pageNumber="368" rank="species">
<emphasis id="8E95F443DA70CC0BFDA93FF156D7B39B" bold="true" box="[590,696,1945,1971]" pageId="2" pageNumber="368">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA70CC0BFD213FF1574EB39B" bold="true" box="[710,801,1945,1971]" pageId="2" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA70CC0BFC983FF157B4B39C" box="[895,987,1945,1972]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="368" unit="mm" value="4.1">4.1 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA70CC0BFB823FF250BCB39B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1125,1235,1945,1971]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA70CC0BFB823FF250BCB39B" box="[1125,1235,1945,1971]" italics="true" pageId="2" pageNumber="368">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA70CC0BFB043FF1515BB39C" box="[1251,1332,1945,1972]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="368" unit="mm" value="57.0">57 mm</quantity>
CW),
<date id="C85F0E91DA70CC0BFA673FF15565B3FF" pageId="2" pageNumber="368" value="2019-05-16">16 May 2019</date>
(
<collectionCode id="DAF0B094DA70CC0BFEFF3FD55508B3FF" box="[280,359,1981,2007]" pageId="2" pageNumber="368">CNVB</collectionCode>
10026).
</paragraph>
<caption id="E89E78D9DA71CC0AFF703E8E5156B3FF" ID-DOI="http://doi.org/10.5281/zenodo.4315889" ID-Zenodo-Dep="4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="3" pageNumber="369" startId="3.[151,168,1766,1792]" targetBox="[158,1425,187,1734]" targetPageId="3">
<paragraph id="BC5E2851DA71CC0AFF703E8E5156B3FF" blockId="3.[151,1436,1766,2008]" pageId="3" pageNumber="369">
<emphasis id="8E95F443DA71CC0AFF703E8E5561B328" bold="true" box="[151,270,1766,1792]" pageId="3" pageNumber="369">FIGURE 1</emphasis>
. Female of
<taxonomicName id="7BE153D2DA71CC0AFE633E805634B328" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[388,603,1768,1792]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA71CC0AFE633E805634B328" box="[388,603,1768,1792]" italics="true" pageId="3" pageNumber="369">Cancrion khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA71CC0AFD873E8056C0B328" bold="true" box="[608,687,1768,1792]" pageId="3" pageNumber="369">
<taxonomicNameLabel id="95A64938DA71CC0AFD873E8056C0B328" box="[608,687,1768,1792]" pageId="3" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find
<bibRefCitation id="D87055A0DA71CC0AFD8D3FA85747B3F0" author="Kuris, A. M. &amp; Poinar, G. O. &amp; Hess, R. T." box="[618,808,1984,2008]" pageId="3" pageNumber="369" pagination="211 - 232" refId="ref13241" refString="Kuris, A. M., Poinar, G. O. &amp; Hess, R. T. (1980) Post-larval mortality of the endoparasitic castrator Portunion conformis (Epicaridea: Entoniscidae) in the shore crab, Hemigrapsus oregonensis, with a description of the host response. Parasitology, 80, 211 - 232, pls. 1 - 4. https: // doi. org / 10.1017 / S 003118200000069 X" type="journal article" year="1980">
Kuris
<emphasis id="8E95F443DA71CC0AFD4F3FA856BAB3F0" box="[680,725,1984,2008]" italics="true" pageId="3" pageNumber="369">et al</emphasis>
. (1980)
</bibRefCitation>
. Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm.
</paragraph>
</caption>
<caption id="E89E78D9DA76CC0DFF703FE85659B3C8" ID-DOI="http://doi.org/10.5281/zenodo.4315891" ID-Zenodo-Dep="4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="4" pageNumber="370" startId="4.[151,250,1920,1944]" targetBox="[158,1424,189,1892]" targetPageId="4">
<paragraph id="BC5E2851DA76CC0DFF703FE85659B3C8" blockId="4.[151,1436,1920,2016]" pageId="4" pageNumber="370">
<emphasis id="8E95F443DA76CC0DFF703FE85563B3B0" bold="true" box="[151,268,1920,1944]" pageId="4" pageNumber="370">FIGURE 2</emphasis>
. Mature male of
<taxonomicName id="7BE153D2DA76CC0DFE5E3FE956E0B3B0" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[441,655,1920,1944]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA76CC0DFE5E3FE956E0B3B0" box="[441,655,1920,1944]" italics="true" pageId="4" pageNumber="370">Cancrion khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA76CC0DFD733FE8568BB3B0" bold="true" box="[660,740,1920,1944]" pageId="4" pageNumber="370">
<taxonomicNameLabel id="95A64938DA76CC0DFD733FE8568BB3B0" box="[660,740,1920,1944]" pageId="4" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm.
</paragraph>
</caption>
<paragraph id="BC5E2851DA77CC0CFF2038FF55D6B54D" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF2038FF552AB498" bold="true" box="[199,325,150,177]" pageId="5" pageNumber="371">
<date id="C85F0E91DA77CC0CFF2038FF552FB498" box="[199,320,150,177]" pageId="5" pageNumber="371" value="2019-06">June 2019</date>
.
</emphasis>
Mature female (
<quantity id="7B1985B4DA77CC0CFE1B38FF5624B499" box="[508,587,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="371" unit="mm" value="28.0">28 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA77CC0CFC9B38FF5065B499" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[892,1034,151,177]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA77CC0CFC9B38FF57E5B499" box="[892,906,151,177]" italics="true" pageId="5" pageNumber="371">S</emphasis>
.
<emphasis id="8E95F443DA77CC0CFC7038FF5065B499" box="[919,1034,151,177]" italics="true" pageId="5" pageNumber="371">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA77CC0CFBF738FF50B3B499" box="[1040,1244,151,177]" pageId="5" pageNumber="371" rank="species">
<emphasis id="8E95F443DA77CC0CFBF738FF5018B499" bold="true" box="[1040,1143,151,177]" pageId="5" pageNumber="371">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA77CC0CFB6338FF50B3B499" bold="true" box="[1156,1244,151,177]" pageId="5" pageNumber="371">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA77CC0CFAD138FF51FFB499" box="[1334,1424,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="371" unit="mm" value="4.1">4.1 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA77CC0CFEE838D35511B4FD" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[271,382,187,213]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFEE838D35511B4FD" box="[271,382,187,213]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFE6838D3558EB4FD" box="[399,481,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="371" unit="mm" value="66.0">66 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFDD738D356C1B4FD" box="[560,686,187,213]" pageId="5" pageNumber="371" value="2019-06-09">9 Jun 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFD5938D35762B4FD" box="[702,781,187,213]" pageId="5" pageNumber="371">CNVB</collectionCode>
10027); Mature female (
<quantity id="7B1985B4DA77CC0CFBC938D35010B4FD" box="[1070,1151,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="371" unit="mm" value="33.0">33 mm</quantity>
TL, stage 12), male (
<quantity id="7B1985B4DA77CC0CFA9F38D354A8B4D1" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="371" unit="mm" value="4.4">4.4 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA77CC0CFEA838B755D2B4D1" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[335,445,223,249]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFEA838B755D2B4D1" box="[335,445,223,249]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFE2B38B65673B4D1" box="[460,540,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="371" unit="mm" value="65.0">65 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFD8F38B7569EB4D1" box="[616,753,223,249]" pageId="5" pageNumber="371" value="2019-06-30">30 Jun 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFCE738B75720B4D1" box="[768,847,223,249]" pageId="5" pageNumber="371">CNVB</collectionCode>
10028); Immature female (
<quantity id="7B1985B4DA77CC0CFB6238B750BAB4D1" box="[1157,1237,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="371" unit="mm" value="18.0">18 mm</quantity>
TL, stage 5), two cryptoniscus larvae (partially damaged), ex female
<taxonomicName id="7BE153D2DA77CC0CFD3A396B5724B535" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[733,843,259,285]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFD3A396B5724B535" box="[733,843,259,285]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFCBC396B57C3B535" box="[859,940,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="371" unit="mm" value="69.0">69 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFC1D396B50EAB535" box="[1018,1157,259,285]" pageId="5" pageNumber="371" value="2019-06-30">30 Jun 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFB72396B508BB535" box="[1173,1252,259,285]" pageId="5" pageNumber="371">CNVB</collectionCode>
10029); Mature female (
<quantity id="7B1985B4DA77CC0CFF14394F552BB569" box="[243,324,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="371" unit="mm" value="33.0">33 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA77CC0CFD65394F577DB569" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[642,786,295,321]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA77CC0CFD65394F56FFB569" box="[642,656,295,321]" italics="true" pageId="5" pageNumber="371">S</emphasis>
.
<emphasis id="8E95F443DA77CC0CFD78394F577DB569" box="[671,786,295,321]" italics="true" pageId="5" pageNumber="371">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA77CC0CFCFD394F5782B569" box="[794,1005,295,321]" pageId="5" pageNumber="371" rank="species">
<emphasis id="8E95F443DA77CC0CFCFD394F57EBB569" bold="true" box="[794,900,295,321]" pageId="5" pageNumber="371">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA77CC0CFC75394F5782B569" bold="true" box="[914,1005,295,321]" pageId="5" pageNumber="371">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex male
<taxonomicName id="7BE153D2DA77CC0CFB82394F50BCB569" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1125,1235,295,321]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFB82394F50BCB569" box="[1125,1235,295,321]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFB04394F515BB569" box="[1251,1332,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="5" pageNumber="371" unit="mm" value="82.0">82 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFA67394F5492B54D" pageId="5" pageNumber="371" value="2019-06-30">30 Jun 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFEEB39235534B54D" box="[268,347,331,357]" pageId="5" pageNumber="371">CNVB</collectionCode>
10030).
</paragraph>
<paragraph id="BC5E2851DA77CC0CFF20390757ADB615" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203907552EB5A0" bold="true" box="[199,321,366,393]" pageId="5" pageNumber="371">
<date id="C85F0E91DA77CC0CFF2039075553B5A0" box="[199,316,366,393]" pageId="5" pageNumber="371" value="2019-07">July 2019</date>
.
</emphasis>
Immature female (
<quantity id="7B1985B4DA77CC0CFDFD3906561DB5A1" box="[538,626,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="371" unit="mm" value="3.5">3.5 mm</quantity>
TL, stage 1), mature female (
<quantity id="7B1985B4DA77CC0CFC233907507AB5A1" box="[964,1045,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="371" unit="mm" value="26.0">26 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA77CC0CFAB339075487B585" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA77CC0CFAB33907510DB5A1" box="[1364,1378,367,393]" italics="true" pageId="5" pageNumber="371">S</emphasis>
.
<emphasis id="8E95F443DA77CC0CFA9639075487B585" italics="true" pageId="5" pageNumber="371">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA77CC0CFF0939FB55D4B585" box="[238,443,403,429]" pageId="5" pageNumber="371" rank="species">
<emphasis id="8E95F443DA77CC0CFF0939FB5539B585" bold="true" box="[238,342,403,429]" pageId="5" pageNumber="371">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA77CC0CFE8539FB55D4B585" bold="true" box="[354,443,403,429]" pageId="5" pageNumber="371">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex ovigerous female
<taxonomicName id="7BE153D2DA77CC0CFD5039FB574CB585" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[695,803,403,429]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFD5039FB574CB585" box="[695,803,403,429]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFCD739FB5710B585" box="[816,895,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="371" unit="mm" value="73.0">73 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFC2F39FB5055B585" box="[968,1082,403,429]" pageId="5" pageNumber="371" value="2019-07-06">6 Jul 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFBA039FB50F9B585" box="[1095,1174,403,429]" pageId="5" pageNumber="371">CNVB</collectionCode>
10031); Mature female (
<quantity id="7B1985B4DA77CC0CFF7839DF5480B5F9" box="[159,239,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="371" unit="mm" value="29.0">29 mm</quantity>
TL, stage 12), male (
<quantity id="7B1985B4DA77CC0CFE3839DF5655B5F9" box="[479,570,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="371" unit="mm" value="4.1">4.1 mm</quantity>
), ex immature male
<taxonomicName id="7BE153D2DA77CC0CFCFA39DF57E5B5F9" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[797,906,439,465]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFCFA39DF57E5B5F9" box="[797,906,439,465]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFC7E39DE5786B5F9" box="[921,1001,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="371" unit="mm" value="57.0">57 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFBD339DF50C7B5F9" box="[1076,1192,439,465]" pageId="5" pageNumber="371" value="2019-07-06">6 Jul 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFB5039DF5169B5F9" box="[1207,1286,439,465]" pageId="5" pageNumber="371">CNVB</collectionCode>
10032); Two mature females (
<quantity id="7B1985B4DA77CC0CFEB739B355A5B5DD" box="[336,458,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="5" pageNumber="371" unit="mm" value="29.0" valueMax="32.0" valueMin="26.0">2632 mm</quantity>
TL, stage 12, largest infested with
<taxonomicName id="7BE153D2DA77CC0CFCA939B357B3B5DD" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[846,988,475,501]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA77CC0CFCA939B35733B5DD" box="[846,860,475,501]" italics="true" pageId="5" pageNumber="371">S</emphasis>
.
<emphasis id="8E95F443DA77CC0CFC8E39B357B3B5DD" box="[873,988,475,501]" italics="true" pageId="5" pageNumber="371">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA77CC0CFC0439B350DEB5DD" box="[995,1201,475,501]" pageId="5" pageNumber="371" rank="species">
<emphasis id="8E95F443DA77CC0CFC0439B35024B5DD" bold="true" box="[995,1099,475,501]" pageId="5" pageNumber="371">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA77CC0CFBBF39B350DEB5DD" bold="true" box="[1112,1201,475,501]" pageId="5" pageNumber="371">sp. nov.</emphasis>
</taxonomicNameLabel>
), two males (
<quantity id="7B1985B4DA77CC0CFAA039B354A8B631" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="5" pageNumber="371" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.84.1 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA77CC0CFEB6399755D1B631" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[337,446,511,537]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFEB6399755D1B631" box="[337,446,511,537]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFE2939975670B631" box="[462,543,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="5" pageNumber="371" unit="mm" value="71.0">71 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFD8C39975680B631" box="[619,751,511,537]" pageId="5" pageNumber="371" value="2019-07-21">21 Jul 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFD1939975722B631" box="[766,845,511,537]" pageId="5" pageNumber="371">CNVB</collectionCode>
10033); Mature female (
<quantity id="7B1985B4DA77CC0CFB8F399750D6B631" box="[1128,1209,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="371" unit="mm" value="34.0">34 mm</quantity>
TL, stage 12), male (
<quantity id="7B1985B4DA77CC0CFF783A4B5494B615" box="[159,251,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="371" unit="mm" value="4.8">4.8 mm</quantity>
), ex male
<taxonomicName id="7BE153D2DA77CC0CFE8B3A4B55B5B615" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[364,474,547,573]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFE8B3A4B55B5B615" box="[364,474,547,573]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFE0F3A4B5657B615" box="[488,568,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="371" unit="mm" value="64.0">64 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFD643A4B5769B615" box="[643,774,547,573]" pageId="5" pageNumber="371" value="2019-07-21">21 Jul 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFCF23A4B570BB615" box="[789,868,547,573]" pageId="5" pageNumber="371">CNVB</collectionCode>
10034).
</paragraph>
<paragraph id="BC5E2851DA77CC0CFF203A2E50C0B6E5" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203A2E550BB648" bold="true" box="[199,356,582,608]" pageId="5" pageNumber="371">
<date id="C85F0E91DA77CC0CFF203A2E5530B648" box="[199,351,582,608]" pageId="5" pageNumber="371" value="2019-08">August 2019</date>
.
</emphasis>
Mature female (
<quantity id="7B1985B4DA77CC0CFDC63A2F561EB649" box="[545,625,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="371" unit="mm" value="31.0">31 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA77CC0CFC483A2F5050B649" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[943,1087,583,609]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA77CC0CFC483A2F57D2B649" box="[943,957,583,609]" italics="true" pageId="5" pageNumber="371">S</emphasis>
.
<emphasis id="8E95F443DA77CC0CFC2B3A2F5050B649" box="[972,1087,583,609]" italics="true" pageId="5" pageNumber="371">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA77CC0CFBA03A2F5175B649" box="[1095,1306,583,609]" pageId="5" pageNumber="371" rank="species">
<emphasis id="8E95F443DA77CC0CFBA03A2F50DEB649" bold="true" box="[1095,1201,583,609]" pageId="5" pageNumber="371">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA77CC0CFB583A2F5175B649" bold="true" box="[1215,1306,583,609]" pageId="5" pageNumber="371">sp. nov.</emphasis>
</taxonomicNameLabel>
), male (
<quantity id="7B1985B4DA77CC0CFA9F3A2F54A8B6AD" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="371" unit="mm" value="4.8">4.8 mm</quantity>
), ex female
<taxonomicName id="7BE153D2DA77CC0CFEAA3A0355D5B6AD" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[333,442,619,645]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFEAA3A0355D5B6AD" box="[333,442,619,645]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFE2F3A035678B6AD" box="[456,535,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="5" pageNumber="371" unit="mm" value="82.0">82 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFD863A03568DB6AD" box="[609,738,619,645]" pageId="5" pageNumber="371" value="2019-08-07">7 Aug 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFD173A035750B6AD" box="[752,831,619,645]" pageId="5" pageNumber="371">CNVB</collectionCode>
10035); Mature female (
<quantity id="7B1985B4DA77CC0CFBB13A0350CAB6AD" box="[1110,1189,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="371" unit="mm" value="39.0">39 mm</quantity>
TL, stage 12, infested with
<taxonomicName id="7BE153D2DA77CC0CFF363AE7550FB681" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[209,352,655,681]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA77CC0CFF363AE754B0B681" box="[209,223,655,681]" italics="true" pageId="5" pageNumber="371">S</emphasis>
.
<emphasis id="8E95F443DA77CC0CFF0A3AE7550FB681" box="[237,352,655,681]" italics="true" pageId="5" pageNumber="371">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA77CC0CFE8F3AE75654B681" box="[360,571,655,681]" pageId="5" pageNumber="371" rank="species">
<emphasis id="8E95F443DA77CC0CFE8F3AE755BDB681" bold="true" box="[360,466,655,681]" pageId="5" pageNumber="371">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA77CC0CFE073AE75654B681" bold="true" box="[480,571,655,681]" pageId="5" pageNumber="371">sp. nov.</emphasis>
</taxonomicNameLabel>
), ex male
<taxonomicName id="7BE153D2DA77CC0CFD543AE7574EB681" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[691,801,655,681]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFD543AE7574EB681" box="[691,801,655,681]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFCD73AE657EEB681" box="[816,897,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="371" unit="mm" value="58.0">58 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFC2A3AE75030B681" box="[973,1119,655,681]" pageId="5" pageNumber="371" value="2019-08-13">13 Aug 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFB883AE750D1B681" box="[1135,1214,655,681]" pageId="5" pageNumber="371">CNVB</collectionCode>
10036); Mature female (
<quantity id="7B1985B4DA77CC0CFF3A3ADB5542B6E5" box="[221,301,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="371" unit="mm" value="39.0">39 mm</quantity>
TL, stage 12), ex female
<taxonomicName id="7BE153D2DA77CC0CFDAB3ADB56D6B6E5" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[588,697,691,717]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA77CC0CFDAB3ADB56D6B6E5" box="[588,697,691,717]" italics="true" pageId="5" pageNumber="371">M. haanii</emphasis>
</taxonomicName>
(
<quantity id="7B1985B4DA77CC0CFD203ADB5777B6E5" box="[711,792,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="5" pageNumber="371" unit="mm" value="76.0">76 mm</quantity>
CW),
<date id="C85F0E91DA77CC0CFC843ADB579CB6E5" box="[867,1011,691,717]" pageId="5" pageNumber="371" value="2019-08-24">24 Aug 2019</date>
(
<collectionCode id="DAF0B094DA77CC0CFBE53ADB503EB6E5" box="[1026,1105,691,717]" pageId="5" pageNumber="371">CNVB</collectionCode>
10037).
</paragraph>
<paragraph id="BC5E2851DA77CC0CFF203ABE5748B039" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203ABE5639B6D8" bold="true" box="[199,598,726,752]" pageId="5" pageNumber="371">Comparative material examined.</emphasis>
<taxonomicName id="7BE153D2DA77CC0CFDB83ABF5015B6D9" authority="Shields &amp; Earley, 1993" authorityName="Shields &amp; Earley" authorityYear="1993" box="[607,1146,727,753]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA77CC0CFDB83ABF570DB6D9" box="[607,866,727,753]" italics="true" pageId="5" pageNumber="371">Cancrion australiensis</emphasis>
<materialsCitation id="0C89220CDA77CC0CFC8C3ABF5015B6D9" ID-GBIF-Occurrence="2995018304" accessionNumber="W17079" box="[875,1146,727,753]" collectingDate="1990-11-20" collectingDateMax="1990-11-23" collectingDateMin="1990-11-20" collectionCode="QM" collectorName="J. D. Shields" country="Australia" county="Brisbane River" elevation="6" latitude="-27.0" location="Moreton bay" longLatPrecision="1239" longitude="153.0" municipality="Cabbage Tree Creek" pageId="5" pageNumber="371" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<bibRefCitation id="D87055A0DA77CC0CFC8C3ABF5015B6D9" author="Shields, J. D. &amp; Earley, C. G." box="[875,1146,727,753]" pageId="5" pageNumber="371" pagination="601 - 608" refId="ref14016" refString="Shields, J. D. &amp; Earley, C. G. (1993) Cancrion australiensis new species (Isopoda: Entoniscidae) found in Thalamita sima (Brachyura: Portunidae) from Australia. International Journal for Parasitology, 23, 601 - 608. https: // doi. org / 10.1016 / 0020 - 7519 (93) 90166 - V" type="journal article" year="1993">Shields &amp; Earley, 1993</bibRefCitation>
</materialsCitation>
</taxonomicName>
: mature
<typeStatus id="635A96F3DA77CC0CFB393ABF512FB6D9" box="[1246,1344,727,753]" pageId="5" pageNumber="371" type="holotype">holotype</typeStatus>
female,
<quantity id="7B1985B4DA77CC0CFF703A92556EB73D" box="[151,257,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="5" pageNumber="371" unit="mm" value="15.1">15.1 mm</quantity>
,
<location id="B93E7E8ADA77CC0CFEF63A9355C9B73D" LSID="urn:lsid:plazi:treatment:34489947DA70CC00FF7039235603B25D:B93E7E8ADA77CC0CFEF63A9355C9B73D" box="[273,422,763,789]" country="Australia" county="Brisbane River" latitude="-27.0" longLatPrecision="1239" longitude="153.0" municipality="Cabbage Tree Creek" name="Moreton bay" pageId="5" pageNumber="371" stateProvince="Queensland">Moreton bay</location>
,
<geoCoordinate id="D9D54E96DA77CC0CFE523A93567CB73D" box="[437,531,763,789]" degrees="27" direction="south" minutes="00" orientation="latitude" pageId="5" pageNumber="371" precision="925" value="-27.0">27°00S</geoCoordinate>
,
<geoCoordinate id="D9D54E96DA77CC0CFDC43A9256FDB73D" box="[547,658,762,789]" degrees="153" direction="east" minutes="00" orientation="longitude" pageId="5" pageNumber="371" precision="925" value="153.0">153°00E</geoCoordinate>
, between the mouth of
<collectingMunicipality id="5C3AB22BDA77CC0CFC4B3A9350F6B73D" box="[940,1177,763,789]" pageId="5" pageNumber="371">Cabbage Tree Creek</collectingMunicipality>
and the mouth of the
<collectingCounty id="553F50DDDA77CC0CFF703B77552BB711" box="[151,324,799,825]" pageId="5" pageNumber="371">Brisbane River</collectingCounty>
,
<collectingRegion id="7E25E6B3DA77CC0CFEB63B7755B7B711" box="[337,472,799,825]" country="Australia" name="Queensland" pageId="5" pageNumber="371">Queensland</collectingRegion>
,
<collectingCountry id="C4F668C1DA77CC0CFE033B775621B711" box="[484,590,799,825]" name="Australia" pageId="5" pageNumber="371">Australia</collectingCountry>
,
<elevation id="37CCCF62DA77CC0CFDBC3B765680B711" box="[603,751,798,825]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="371" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">
<quantity id="7B1985B4DA77CC0CFDBC3B7656CBB711" box="[603,676,798,825]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="371" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">57 m</quantity>
depth
</elevation>
, ex
<taxonomicName id="7BE153D2DA77CC0CFCC73B775097B711" authority="H. Milne Edwards, 1834" authorityName="H. Milne Edwards" authorityYear="1834" box="[800,1272,799,825]" class="Malacostraca" family="Portunidae" genus="Thalamita" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="sima">
<emphasis id="8E95F443DA77CC0CFCC73B7757BEB710" box="[800,977,799,825]" italics="true" pageId="5" pageNumber="371">Thalamita sima</emphasis>
H. Milne Edwards, 1834
</taxonomicName>
,
<date id="C85F0E91DA77CC0CFAE23B7751F3B711" box="[1285,1436,799,825]" pageId="5" pageNumber="371" value="1990-11-20">
<collectingDate id="D81BF779DA77CC0CFAE23B7751F3B711" box="[1285,1436,799,825]" pageId="5" pageNumber="371" value="1990-11-20">20 Nov 1990</collectingDate>
</date>
[in publication;
<date id="C85F0E91DA77CC0CFEAA3B2B558EB775" box="[333,481,835,861]" pageId="5" pageNumber="371" value="1990-11-23">
<collectingDate id="D81BF779DA77CC0CFEAA3B2B558EB775" box="[333,481,835,861]" pageId="5" pageNumber="371" value="1990-11-23">23 Nov 1990</collectingDate>
</date>
on label in vial], coll.
<collectorName id="11144D87DA77CC0CFD013B2B5719B775" box="[742,886,835,861]" pageId="5" pageNumber="371">J. D. Shields</collectorName>
(
<collectionCode id="DAF0B094DA77CC0CFC603B2B57D9B775" box="[903,950,835,861]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="5" pageNumber="371" type="Museum">QM</collectionCode>
W17079);
<materialsCitation id="0C89220CDA77CC0CFBD03B2B5141B78D" ID-GBIF-Occurrence="2995018303" accessionNumber="W17079" collectingDate="1990-10-20" collectingDateMax="1991-02-13" collectingDateMin="1990-10-20" collectionCode="USNM" collectorName="Sta. &amp; J. D. Shields" country="Australia" county="Mud island" elevation="6" latitude="-27.25" location="Mature" longLatPrecision="1237" longitude="153.25" municipality="Moreton bay" pageId="5" pageNumber="371" specimenCode="USNM 284172" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<location id="B93E7E8ADA77CC0CFBD03B2B50E7B775" LSID="urn:lsid:plazi:treatment:34489947DA70CC00FF7039235603B25D:B93E7E8ADA77CC0CFBD03B2B50E7B775" box="[1079,1160,835,861]" country="Australia" county="Mud island" latitude="-27.25" longLatPrecision="1237" longitude="153.25" municipality="Moreton bay" name="Mature" pageId="5" pageNumber="371" stateProvince="Queensland">Mature</location>
<typeStatus id="635A96F3DA77CC0CFB773B2B509FB775" box="[1168,1264,835,861]" pageId="5" pageNumber="371" type="paratype">paratype</typeStatus>
male,
<quantity id="7B1985B4DA77CC0CFAD93B2B51F7B775" box="[1342,1432,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="371" unit="mm" value="2.3">2.3 mm</quantity>
, same data as holotype except
<date id="C85F0E91DA77CC0CFE043B0F5603B7A9" box="[483,620,871,897]" pageId="5" pageNumber="371" value="1990-10-20">
<collectingDate id="D81BF779DA77CC0CFE043B0F5603B7A9" box="[483,620,871,897]" pageId="5" pageNumber="371" value="1990-10-20">20 Oct 1990</collectingDate>
</date>
collection date; 1 mature female,
<collectorName id="11144D87DA77CC0CFC003B0F507DB7A9" box="[999,1042,871,897]" pageId="5" pageNumber="371">Sta.</collectorName>
36A-25,
<collectingMunicipality id="5C3AB22BDA77CC0CFB9B3B0F5163B7A9" box="[1148,1292,871,897]" pageId="5" pageNumber="371">Moreton bay</collectingMunicipality>
,
<collectingCounty id="553F50DDDA77CC0CFAF03B0F51F7B7A9" box="[1303,1432,871,897]" pageId="5" pageNumber="371">Mud island</collectingCounty>
,
<collectingRegion id="7E25E6B3DA77CC0CFF703BE35571B78D" box="[151,286,907,933]" country="Australia" name="Queensland" pageId="5" pageNumber="371">Queensland</collectingRegion>
,
<collectingCountry id="C4F668C1DA77CC0CFECF3BE355FDB78D" box="[296,402,907,933]" name="Australia" pageId="5" pageNumber="371">Australia</collectingCountry>
,
<geoCoordinate id="D9D54E96DA77CC0CFE7A3BE25594B78D" box="[413,507,906,933]" degrees="27" direction="south" minutes="15" orientation="latitude" pageId="5" pageNumber="371" precision="925" value="-27.25">27°15S</geoCoordinate>
,
<geoCoordinate id="D9D54E96DA77CC0CFDE03BE2561AB78D" box="[519,629,906,933]" degrees="153" direction="east" minutes="15" orientation="longitude" pageId="5" pageNumber="371" precision="925" value="153.25">153°15E</geoCoordinate>
, ex
<taxonomicName id="7BE153D2DA77CC0CFD443BE3569AB78C" authorityName="H. Milne Edwards" authorityYear="1834" box="[675,757,907,933]" class="Malacostraca" family="Portunidae" genus="Thalamita" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="sima">
<emphasis id="8E95F443DA77CC0CFD443BE356DCB78C" box="[675,691,907,932]" italics="true" pageId="5" pageNumber="371">T</emphasis>
.
<emphasis id="8E95F443DA77CC0CFD273BE3569AB78C" box="[704,757,907,932]" italics="true" pageId="5" pageNumber="371">sima</emphasis>
</taxonomicName>
,
<date id="C85F0E91DA77CC0CFCE33BE357FDB78D" box="[772,914,907,933]" pageId="5" pageNumber="371" value="1991-02-13">
<collectingDate id="D81BF779DA77CC0CFCE33BE357FDB78D" box="[772,914,907,933]" pageId="5" pageNumber="371" value="1991-02-13">13 Feb 1991</collectingDate>
</date>
, coll.
<collectorName id="11144D87DA77CC0CFC313BE3500BB78D" box="[982,1124,907,933]" pageId="5" pageNumber="371">J. D. Shields</collectorName>
(
<specimenCode id="EC47802ADA77CC0CFB943BE35149B78D" box="[1139,1318,907,933]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="371" type="Museum">USNM 284172</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="0C89220CDA77CC0CFADE3BE355DBB7C5" ID-GBIF-Occurrence="2995018301" collectingDate="1990-12-04" collectionCode="USNM" collectorName="J. D. Shields" country="Australia" latitude="-27.25" location="Sta." longLatPrecision="1237" longitude="153.25" municipality="Moreton bay" pageId="5" pageNumber="371" specimenCode="USNM 284173" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 mature female,
<location id="B93E7E8ADA77CC0CFF133BC75570B7E1" LSID="urn:lsid:plazi:treatment:34489947DA70CC00FF7039235603B25D:B93E7E8ADA77CC0CFF133BC75570B7E1" box="[244,287,943,969]" country="Australia" latitude="-27.25" longLatPrecision="1237" longitude="153.25" municipality="Moreton bay" name="Sta." pageId="5" pageNumber="371" stateProvince="Queensland">Sta.</location>
31-A,
<collectingMunicipality id="5C3AB22BDA77CC0CFE943BC75669B7E1" box="[371,518,943,969]" pageId="5" pageNumber="371">Moreton bay</collectingMunicipality>
,
<collectingRegion id="7E25E6B3DA77CC0CFDF23BC756F3B7E1" box="[533,668,943,969]" country="Australia" name="Queensland" pageId="5" pageNumber="371">Queensland</collectingRegion>
,
<collectingCountry id="C4F668C1DA77CC0CFD4F3BC7577DB7E1" box="[680,786,943,969]" name="Australia" pageId="5" pageNumber="371">Australia</collectingCountry>
,
<geoCoordinate id="D9D54E96DA77CC0CFCF83BC65712B7E1" box="[799,893,942,969]" degrees="27" direction="south" minutes="15" orientation="latitude" pageId="5" pageNumber="371" precision="925" value="-27.25">27°15S</geoCoordinate>
,
<geoCoordinate id="D9D54E96DA77CC0CFC6B3BC65795B7E1" box="[908,1018,942,969]" degrees="153" direction="east" minutes="15" orientation="longitude" pageId="5" pageNumber="371" precision="925" value="153.25">153°15E</geoCoordinate>
, ex
<taxonomicName id="7BE153D2DA77CC0CFBCA3BC750EDB7E0" authorityName="H. Milne Edwards" authorityYear="1834" box="[1069,1154,943,969]" class="Malacostraca" family="Portunidae" genus="Thalamita" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="sima">
<emphasis id="8E95F443DA77CC0CFBCA3BC75052B7E0" box="[1069,1085,943,968]" italics="true" pageId="5" pageNumber="371">T</emphasis>
.
<emphasis id="8E95F443DA77CC0CFBAA3BC750EDB7E0" box="[1101,1154,943,968]" italics="true" pageId="5" pageNumber="371">sima</emphasis>
</taxonomicName>
,
<date id="C85F0E91DA77CC0CFB743BC75172B7E1" box="[1171,1309,943,969]" pageId="5" pageNumber="371" value="1990-12-04">
<collectingDate id="D81BF779DA77CC0CFB743BC75172B7E1" box="[1171,1309,943,969]" pageId="5" pageNumber="371" value="1990-12-04">4 Dec 1990</collectingDate>
</date>
, coll.
<collectorName id="11144D87DA77CC0CFA823BC75486B7C5" pageId="5" pageNumber="371">J. D. Shields</collectorName>
(
<specimenCode id="EC47802ADA77CC0CFF1E3BBB55C3B7C5" box="[249,428,979,1005]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="371" type="Museum">USNM 284173</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="0C89220CDA77CC0CFE273BBB574BB039" ID-GBIF-Occurrence="2995018307" collectingDate="1990-12-20" collectionCode="USNM" collectorName="J. D. Shields" country="Australia" latitude="-27.25" location="Sta." longLatPrecision="1237" longitude="153.25" municipality="Moreton bay" pageId="5" pageNumber="371" specimenCode="USNM 284174" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
1 mature female,
<location id="B93E7E8ADA77CC0CFD623BBB56DFB7C5" LSID="urn:lsid:plazi:treatment:34489947DA70CC00FF7039235603B25D:B93E7E8ADA77CC0CFD623BBB56DFB7C5" box="[645,688,979,1005]" country="Australia" latitude="-27.25" longLatPrecision="1237" longitude="153.25" municipality="Moreton bay" name="Sta." pageId="5" pageNumber="371" stateProvince="Queensland">Sta.</location>
29-2,
<collectingMunicipality id="5C3AB22BDA77CC0CFD1E3BBB57E5B7C5" box="[761,906,979,1005]" pageId="5" pageNumber="371">Moreton bay</collectingMunicipality>
,
<collectingRegion id="7E25E6B3DA77CC0CFC703BBB5071B7C5" box="[919,1054,979,1005]" country="Australia" name="Queensland" pageId="5" pageNumber="371">Queensland</collectingRegion>
,
<collectingCountry id="C4F668C1DA77CC0CFBCF3BBB50FDB7C5" box="[1064,1170,979,1005]" name="Australia" pageId="5" pageNumber="371">Australia</collectingCountry>
,
<geoCoordinate id="D9D54E96DA77CC0CFB7A3BBA5094B7C5" box="[1181,1275,978,1005]" degrees="27" direction="south" minutes="15" orientation="latitude" pageId="5" pageNumber="371" precision="925" value="-27.25">27°15S</geoCoordinate>
,
<geoCoordinate id="D9D54E96DA77CC0CFAEF3BBA511AB7C5" box="[1288,1397,978,1005]" degrees="153" direction="east" minutes="15" orientation="longitude" pageId="5" pageNumber="371" precision="925" value="153.25">153°15E</geoCoordinate>
, ex
<taxonomicName id="7BE153D2DA77CC0CFF703B9F5485B038" authorityName="H. Milne Edwards" authorityYear="1834" box="[151,234,1015,1041]" class="Malacostraca" family="Portunidae" genus="Thalamita" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="sima">
<emphasis id="8E95F443DA77CC0CFF703B9F54C8B038" box="[151,167,1015,1040]" italics="true" pageId="5" pageNumber="371">T</emphasis>
.
<emphasis id="8E95F443DA77CC0CFF523B9F5485B038" box="[181,234,1015,1040]" italics="true" pageId="5" pageNumber="371">sima</emphasis>
</taxonomicName>
,
<date id="C85F0E91DA77CC0CFF1F3B9F55E5B039" box="[248,394,1015,1041]" pageId="5" pageNumber="371" value="1990-12-20">
<collectingDate id="D81BF779DA77CC0CFF1F3B9F55E5B039" box="[248,394,1015,1041]" pageId="5" pageNumber="371" value="1990-12-20">20 Dec 1990</collectingDate>
</date>
, coll.
<collectorName id="11144D87DA77CC0CFE2A3B9F5635B039" box="[461,602,1015,1041]" pageId="5" pageNumber="371">J. D. Shields</collectorName>
(
<specimenCode id="EC47802ADA77CC0CFD8E3B9F5773B039" box="[617,796,1015,1041]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="371" type="Museum">USNM 284174</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA77CC0CFF203C73510DB01D" box="[199,1378,1051,1077]" pageId="5" pageNumber="371" type="etymology">
<paragraph id="BC5E2851DA77CC0CFF203C73510DB01D" blockId="5.[151,1437,150,2013]" box="[199,1378,1051,1077]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203C735521B01D" bold="true" box="[199,334,1051,1077]" pageId="5" pageNumber="371">Etymology.</emphasis>
The name refers to the province of
<collectingCountry id="C4F668C1DA77CC0CFD393C73572FB01D" box="[734,832,1051,1077]" name="Vietnam" pageId="5" pageNumber="371">Vietnam</collectingCountry>
,
<collectingRegion id="7E25E6B3DA77CC0CFCAC3C7357A1B01D" box="[843,974,1051,1077]" country="Vietnam" name="Khanh Hoa" pageId="5" pageNumber="371">Khanh Hoa</collectingRegion>
, in which the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA77CC0EFF203C565644B089" lastPageId="7" lastPageNumber="373" pageId="5" pageNumber="371" type="description">
<paragraph id="BC5E2851DA77CC0CFF203C5656F7B11A" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203C565660B071" bold="true" box="[199,527,1086,1113]" pageId="5" pageNumber="371">
Description.
<emphasis id="8E95F443DA77CC0CFE853C575660B071" bold="true" box="[354,527,1086,1113]" italics="true" pageId="5" pageNumber="371">Mature female</emphasis>
</emphasis>
(stage 12, see Remarks) (
<figureCitation id="24DA34D4DA77CC0CFCDA3C5757EDB071" box="[829,898,1087,1113]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Figs 1</figureCitation>
AD)
<emphasis id="8E95F443DA77CC0CFC373C575069B071" box="[976,1030,1087,1113]" italics="true" pageId="5" pageNumber="371">body</emphasis>
occupying most of host hemocoel; total length of cephalon, thorax, and abdomen
<quantity id="7B1985B4DA77CC0CFD403C0A574BB055" box="[679,804,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="5" pageNumber="371" unit="mm" value="29.5" valueMax="35.0" valueMin="24.0">2435 mm</quantity>
. Sheath formed by host surrounds entire parasite except for perforation at posterior end (exit pore not observed). Body dorsally recurved, V-shaped from cephalon to posterior end of pleon.
<emphasis id="8E95F443DA77CC0CFE833CC355BEB0ED" box="[356,465,1195,1221]" italics="true" pageId="5" pageNumber="371">Cephalon</emphasis>
rounded, 3.0
<quantity id="7B1985B4DA77CC0CFD923CC256A2B0ED" box="[629,717,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="371" unit="mm" value="4.5">4.5 mm</quantity>
, dorsally distinctly divided into 2 bulbous lobes (
<figureCitation id="24DA34D4DA77CC0CFB1B3CC3513BB0ED" box="[1276,1364,1195,1221]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1B</figureCitation>
), with 2 pairs of antennae and pair of maxillipeds. Both pairs of
<emphasis id="8E95F443DA77CC0CFCC43CA757E5B0C0" box="[803,906,1231,1256]" italics="true" pageId="5" pageNumber="371">antennae</emphasis>
as slender lobes, subequal in length, inner pair slightly broader (
<figureCitation id="24DA34D4DA77CC0CFEBF3C9B55DFB125" box="[344,432,1267,1293]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1B</figureCitation>
).
<emphasis id="8E95F443DA77CC0CFE213C9B5650B125" box="[454,575,1267,1293]" italics="true" pageId="5" pageNumber="371">Maxilliped</emphasis>
with subquadrate coxopodite, folded concave, lamellar exopodite (
<figureCitation id="24DA34D4DA77CC0CFAD23C9B51E2B125" box="[1333,1421,1267,1293]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1B</figureCitation>
); endopodite absent or vestigial (not observed).
</paragraph>
<paragraph id="BC5E2851DA77CC0CFF203D535695B25D" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203D535578B17C" box="[199,279,1339,1364]" italics="true" pageId="5" pageNumber="371">Pereon</emphasis>
compact, 8.010.0 mm, ovaries cream to orange, four pairs of large dorsolateral ovarian processes, first pair slightly smaller than others, covering most or all of dorsal, lateral, and ventral margins of pereon (
<figureCitation id="24DA34D4DA77CC0CFAEA3D37510BB151" box="[1293,1380,1375,1401]" captionStart="FIGURE 8" captionStartId="14.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure-17@14.[151,1436,181,1786]" captionTargetPageId="14" captionText="FIGURE 8. A, Cancrion khanhensis sp. nov. mature ovigerous female; B, C. khanhensis sp. nov. mature female showing dorsolateral ovarian processes (arrows); C, C. khanhensis sp. nov, immature female (stage 8); D, C. khanhensis sp. nov. mature female infested by Stellatoniscus tentaculus gen. nov., sp. nov. mature female; E, S. tentaculus gen. nov., sp. nov. immature females (stage III); F, S. tentaculus gen. nov., sp. nov. mature female (stage IV). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4315903" httpUri="https://zenodo.org/record/4315903/files/figure.png" pageId="5" pageNumber="371">Fig. 8B</figureCitation>
),
<emphasis id="8E95F443DA77CC0CFA9F3D375495B1B5" italics="true" pageId="5" pageNumber="371">pereopod 1</emphasis>
absent,
<emphasis id="8E95F443DA77CC0CFEB23DEB5596B1B5" box="[341,505,1411,1437]" italics="true" pageId="5" pageNumber="371">pereopods 27</emphasis>
rudimentary; pereopods 25 present as small projections arising from dorsal side at median of respective oostegites, pereopods 6, 7 located on angle between pereon and pleon (
<figureCitation id="24DA34D4DA77CC0CFB463DCF5095B1E9" box="[1185,1274,1447,1473]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1H</figureCitation>
). Five pairs of oostegites forming brood pouch, margins entire. Oostegite 1 with ascendant lobe projecting anteriorly and dorsally, 2 simple transverse lobes, one recurrent lobe extending entire length of pereon with complex folding flap on lateral and flower-shaped fold on dorsal surface (
<figureCitation id="24DA34D4DA77CC0CFD943E7B56A4B205" box="[627,715,1555,1581]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1C</figureCitation>
). Oostegites 25 delicate, scarcely overlapping, fused in mature females; oostegite 2 covering base of oostegite 1 and ventral surface of cephalon (
<figureCitation id="24DA34D4DA77CC0CFBD03E5F50FFB279" box="[1079,1168,1591,1617]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1A</figureCitation>
); oostegites 35 arising from lateral margins of dorsolateral ovarian processes.
</paragraph>
<paragraph id="BC5E2851DA77CC0CFF203E1756B1B32D" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203E175567B2B1" box="[199,264,1663,1689]" italics="true" pageId="5" pageNumber="371">Pleon</emphasis>
white, 12.017.0 mm, composed of 5 segments. Pleomeres 15 each with distinct uniramous flattened pair of pleopods that overlay one another (
<figureCitation id="24DA34D4DA77CC0CFD923ECB56A0B295" box="[629,719,1699,1725]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1D</figureCitation>
). Two pairs of complex pleural lamellae positioned medially on pleomeres 1 and 2 (
<figureCitation id="24DA34D4DA77CC0CFE883EAF5586B2C9" box="[367,489,1735,1761]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1A, D</figureCitation>
). Oval, bulbous heart in pleomere 3 (
<figureCitation id="24DA34D4DA77CC0CFC663EAF5793B2C9" box="[897,1020,1735,1761]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1A, H</figureCitation>
). Distal end of pleon with two minute finger-like projections (possibly uropods) (
<figureCitation id="24DA34D4DA77CC0CFD9F3E8356BEB32D" box="[632,721,1771,1797]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="BC5E2851DA77CC0CFF203F675782B3F5" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="371">
<emphasis id="8E95F443DA77CC0CFF203F6755F4B301" bold="true" box="[199,411,1806,1833]" italics="true" pageId="5" pageNumber="371">Immature females</emphasis>
(
<figureCitation id="24DA34D4DA77CC0CFE4D3F675583B301" box="[426,492,1807,1833]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Figs 1</figureCitation>
EH, 8C) immature newly molted from cryptoniscus females found lying along the median shaft of host gill; vermiform, cylindrical, without appendages (
<figureCitation id="24DA34D4DA77CC0CFC543F5B5067B365" box="[947,1032,1843,1869]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1E</figureCitation>
), immature females found in hemocoel ranging from vermiform, recurved, without appendages (stage 2,
<figureCitation id="24DA34D4DA77CC0CFC783F3F5780B359" box="[927,1007,1879,1905]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1F</figureCitation>
) to V-shaped with rudimentary pleural lamellae (stage 5,
<figureCitation id="24DA34D4DA77CC0CFE843F1355D6B3BD" box="[355,441,1915,1941]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1G</figureCitation>
) to V-shaped with developing oostegites plus rudimentary pleural lamellae (stage 8,
<figureCitation id="24DA34D4DA77CC0CFA973F1354D3B391" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Fig. 1H</figureCitation>
); latter stage with ovaries as two cream-colored wavy ridges lying along lateral margins of pereon (
<figureCitation id="24DA34D4DA77CC0CFAE33FF75130B391" box="[1284,1375,1951,1977]" captionStart="FIGURE" captionStartId="3.[151,168,1766,1792]" captionTargetBox="[158,1425,187,1734]" captionTargetId="figure-25@3.[151,1436,179,1741]" captionTargetPageId="3" captionText="FIGURE 1. Female of Cancrion khanhensis sp. nov. A, mature female (stage 12) in host sheath, lateral view (1, bulbous heart); B, mature female (stage 12), cephalon (1, outer antenna; 2, inner antenna; 3, coxopodite; 4, exopodite of maxilliped); C, mature female (stage 12), oostegite 1; D, mature female (stage 12) abdomen with pleural lamellae (1, distal end of pleon; 2, pleural lamella 2; 3, pleural lamella 1); E, pre-asticot female (immediately post-molt from cryptoniscus); F, pre-asticot female, more developed (stage 2); G, immature female (stage 5) with rudimentary pleural lamellae; H, immature female (stage 8) with moderately developed pleural lamellae (1, anterior lobe of oostegite 1 extending anterodorsal to cephalon; 2, oostegite 2; 3, pereopod 3; 4, pereopods 6 and 7; 5, heart). Stages find Kuris et al. (1980). Scale bars: A, C, D, G, H = 5 mm; B, E, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315889" httpUri="https://zenodo.org/record/4315889/files/figure.png" pageId="5" pageNumber="371">Figs 1H</figureCitation>
,
<figureCitation id="24DA34D4DA77CC0CFA8D3FF751E1B391" box="[1386,1422,1951,1977]" captionStart="FIGURE 8" captionStartId="14.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure-17@14.[151,1436,181,1786]" captionTargetPageId="14" captionText="FIGURE 8. A, Cancrion khanhensis sp. nov. mature ovigerous female; B, C. khanhensis sp. nov. mature female showing dorsolateral ovarian processes (arrows); C, C. khanhensis sp. nov, immature female (stage 8); D, C. khanhensis sp. nov. mature female infested by Stellatoniscus tentaculus gen. nov., sp. nov. mature female; E, S. tentaculus gen. nov., sp. nov. immature females (stage III); F, S. tentaculus gen. nov., sp. nov. mature female (stage IV). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4315903" httpUri="https://zenodo.org/record/4315903/files/figure.png" pageId="5" pageNumber="371">8C</figureCitation>
); mature ovigerous females (stage 12,
<figureCitation id="24DA34D4DA77CC0CFDD43FAB56C5B3F5" box="[563,682,1987,2013]" captionStart="FIGURE 8" captionStartId="14.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure-17@14.[151,1436,181,1786]" captionTargetPageId="14" captionText="FIGURE 8. A, Cancrion khanhensis sp. nov. mature ovigerous female; B, C. khanhensis sp. nov. mature female showing dorsolateral ovarian processes (arrows); C, C. khanhensis sp. nov, immature female (stage 8); D, C. khanhensis sp. nov. mature female infested by Stellatoniscus tentaculus gen. nov., sp. nov. mature female; E, S. tentaculus gen. nov., sp. nov. immature females (stage III); F, S. tentaculus gen. nov., sp. nov. mature female (stage IV). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4315903" httpUri="https://zenodo.org/record/4315903/files/figure.png" pageId="5" pageNumber="371">Fig. 8A, B</figureCitation>
) with orange gonadal tissue.
</paragraph>
<paragraph id="BC5E2851DA74CC0FFF2038FE515EB531" blockId="6.[151,1437,150,628]" pageId="6" pageNumber="372">
<emphasis id="8E95F443DA74CC0FFF2038FE556DB499" bold="true" box="[199,258,150,177]" italics="true" pageId="6" pageNumber="372">Male</emphasis>
(
<figureCitation id="24DA34D4DA74CC0FFEF538FF5539B499" box="[274,342,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2</figureCitation>
) body elongated, white, markedly curved ventrally,
<quantity id="7B1985B4DA74CC0FFC4338FF5044B499" box="[932,1067,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.9" metricValueMin="3.1" pageId="6" pageNumber="372" unit="mm" value="4.0" valueMax="4.9" valueMin="3.1">3.14.9 mm</quantity>
long,
<quantity id="7B1985B4DA74CC0FFB9438FE5166B499" box="[1139,1289,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="7.749999999999999" metricValueMax="8.5" metricValueMin="7.0" pageId="6" pageNumber="372" unit="mm" value="0.7749999999999999" valueMax="0.85" valueMin="0.7">0.70.85 mm</quantity>
wide, all pereomeres clearly separated dorsally, pereomere 1 fused with rounded cephalon (
<figureCitation id="24DA34D4DA74CC0FFBF038D15035B4FB" box="[1047,1114,185,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2</figureCitation>
AC). Cephalon with pair of minute dorsolateral eyes (
<figureCitation id="24DA34D4DA74CC0FFE5238B45662B4DE" box="[437,525,220,246]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2C</figureCitation>
). Antennulae as large ovate lobes fused medially, with group of short setae anteroventrally (
<figureCitation id="24DA34D4DA74CC0FFEED389755EBB531" box="[266,388,255,281]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2A, B</figureCitation>
); antennae absent. Mandibles inside oral cone; no vestiges of maxillipeds (
<figureCitation id="24DA34D4DA74CC0FFB2C3897514CB531" box="[1227,1315,255,281]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="BC5E2851DA74CC0FFF20394A57F8B5C1" blockId="6.[151,1437,150,628]" pageId="6" pageNumber="372">
<emphasis id="8E95F443DA74CC0FFF20394A5578B513" box="[199,279,290,315]" italics="true" pageId="6" pageNumber="372">Pereon</emphasis>
maximal width at pereomeres 46, gradually tapering anteriorly and posteriorly; lateral margins of pleomeres 16 expanded and directed ventrally, overlying bases of pereopods; lateral margins of pleomere 7 directed posteriorly; pereomeres 27 each with small setose medioventral tubercle (
<figureCitation id="24DA34D4DA74CC0FFBD7390F50E4B5A9" box="[1072,1163,359,385]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2A</figureCitation>
). Six pairs of pereopods (absent on pereomere 7) (
<figureCitation id="24DA34D4DA74CC0FFE5F39E1567DB58B" box="[440,530,393,419]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2A</figureCitation>
), pereopods subequal in size, each with minute dactylus; propodus, carpus fused into single cylindrical segment distally covered with small spinules, merus small, subcircular, variably distinct from propodus/carpus; ischium, basis distinctly separated (
<figureCitation id="24DA34D4DA74CC0FFD0A39A757E5B5C1" box="[749,906,463,489]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2A, B, D</figureCitation>
).
</paragraph>
<paragraph id="BC5E2851DA74CC0FFF2039995610B65B" blockId="6.[151,1437,150,628]" pageId="6" pageNumber="372">
<emphasis id="8E95F443DA74CC0FFF2039995567B623" box="[199,264,497,523]" italics="true" pageId="6" pageNumber="372">Pleon</emphasis>
with 5 cylindrical segments plus pleotelson (
<figureCitation id="24DA34D4DA74CC0FFCF839995712B623" box="[799,893,497,523]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2A</figureCitation>
). Pleomeres distinctly narrower posteriorly, all separated, with horizontal fold in posterior portion of each segment, when folded, each anterior pleomere overlies base of following segment, pleopods absent. Pleotelson with 2 short, distally rounded, spinulose posterolateral lobes (
<figureCitation id="24DA34D4DA74CC0FFF473A315577B65C" box="[160,280,601,628]" captionStart="FIGURE 2" captionStartId="4.[151,250,1920,1944]" captionTargetBox="[158,1424,189,1892]" captionTargetId="figure-17@4.[151,1436,181,1896]" captionTargetPageId="4" captionText="FIGURE 2. Mature male of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, cephalon and pereomere 1, ventral view; C, dorsolateral view showing fusion of cephalon and pereomere 1; D, pereopod 1; E, pleomere 5 and pleotelson. Scale bars: A = 1 mm; B, C = 0.2 mm; D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4315891" httpUri="https://zenodo.org/record/4315891/files/figure.png" pageId="6" pageNumber="372">Fig. 2A, E</figureCitation>
); anal cone and uropods absent.
</paragraph>
<caption id="E89E78D9DA74CC0FFF703EA45111B2CD" ID-DOI="http://doi.org/10.5281/zenodo.4315893" ID-Zenodo-Dep="4315893" box="[151,1406,1740,1765]" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="6" pageNumber="372" startId="6.[151,250,1740,1764]" targetBox="[209,1380,658,1710]" targetPageId="6">
<paragraph id="BC5E2851DA74CC0FFF703EA45111B2CD" blockId="6.[151,1406,1740,1765]" box="[151,1406,1740,1765]" pageId="6" pageNumber="372">
<emphasis id="8E95F443DA74CC0FFF703EA45562B2CC" bold="true" box="[151,269,1740,1764]" pageId="6" pageNumber="372">FIGURE 3</emphasis>
. Epicaridium larva of
<taxonomicName id="7BE153D2DA74CC0FFE163EA556A7B2CC" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[497,712,1740,1764]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="372" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA74CC0FFE163EA556A7B2CC" box="[497,712,1740,1764]" italics="true" pageId="6" pageNumber="372">Cancrion khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA74CC0FFD293EA45770B2CC" bold="true" box="[718,799,1740,1764]" pageId="6" pageNumber="372">
<taxonomicNameLabel id="95A64938DA74CC0FFD293EA45770B2CC" box="[718,799,1740,1764]" pageId="6" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm.
</paragraph>
</caption>
<paragraph id="BC5E2851DA74CC0FFF203F64578DB3D0" blockId="6.[151,1437,1804,2040]" pageId="6" pageNumber="372">
<emphasis id="8E95F443DA74CC0FFF203F6455F3B30F" bold="true" box="[199,412,1804,1831]" italics="true" pageId="6" pageNumber="372">Epicaridium larva</emphasis>
(
<figureCitation id="24DA34D4DA74CC0FFE493F65559CB30F" box="[430,499,1805,1831]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="6" pageNumber="372">Fig. 3</figureCitation>
) approximately
<quantity id="7B1985B4DA74CC0FFD563F65572DB300" box="[689,834,1805,1832]" metricMagnitude="-1" metricUnit="m" metricValue="2.575" metricValueMax="2.65" metricValueMin="2.5" pageId="6" pageNumber="372" unit="mm" value="257.5" valueMax="265.0" valueMin="250.0">250265 µm</quantity>
long (anterior margin of cephalon to end of telson),
<quantity id="7B1985B4DA74CC0FFF703F585577B362" box="[151,280,1840,1866]" metricMagnitude="-2" metricUnit="m" metricValue="9.9" metricValueMax="10.2" metricValueMin="9.6" pageId="6" pageNumber="372" unit="mm" value="99.0" valueMax="102.0" valueMin="96.0">96102 µm</quantity>
wide (excluding pereopods).
<emphasis id="8E95F443DA74CC0FFD853F5856F3B362" box="[610,668,1840,1866]" italics="true" pageId="6" pageNumber="372">Body</emphasis>
tear-drop shaped; anterior margin of head rounded; black-pigmented posterolateral eyespots.
<emphasis id="8E95F443DA74CC0FFE4F3F3A5674B344" box="[424,539,1874,1900]" italics="true" pageId="6" pageNumber="372">Antennula</emphasis>
of 2 rounded articles, article 1 wider but scarcely longer than terminal article; terminal article with 2 or 3 long setae and several short setae (
<figureCitation id="24DA34D4DA74CC0FFCB53F1D57C3B3A7" box="[850,940,1909,1935]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="6" pageNumber="372">Fig. 3A</figureCitation>
).
<emphasis id="8E95F443DA74CC0FFC273F1D5072B3A7" box="[960,1053,1909,1935]" italics="true" pageId="6" pageNumber="372">Antenna</emphasis>
elongated, nearly as long as body (
<figureCitation id="24DA34D4DA74CC0FFF473FF05494B39A" box="[160,251,1944,1970]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="6" pageNumber="372">Fig. 3A</figureCitation>
); composed of 6 articles, articles 1 and 2 subequal in size; articles 3 and 4 longer, flagellar article 1 approximately 1.5 times as long as terminal article, with 2 distolateral setae; terminal article bearing long medial seta nearly half as long as entire antenna (
<figureCitation id="24DA34D4DA74CC0FFDDE3FB556FCB3DF" box="[569,659,2013,2039]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="6" pageNumber="372">Fig. 3A</figureCitation>
).
<emphasis id="8E95F443DA74CC0FFD403FB55772B3DF" box="[679,797,2013,2039]" italics="true" pageId="6" pageNumber="372">Mandibles</emphasis>
within oral cone.
</paragraph>
<paragraph id="BC5E2851DA75CC0EFF2038FF5004B5DD" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="373">
<emphasis id="8E95F443DA75CC0EFF2038FF5578B498" box="[199,279,151,176]" italics="true" pageId="7" pageNumber="373">Pereon</emphasis>
with 5 anterior pairs of gnathopodal pereopods, subequal in size, each with slender, slightly curved dactylus extending approximately half length of propodus, tip of dactylus hooked; propodus, carpus fused with ventral area of blunt spinules; merus small, subrectangular, ischium and basis cylindrical (
<figureCitation id="24DA34D4DA75CC0EFBA838B750C4B4D1" box="[1103,1195,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="7" pageNumber="373">Fig. 3A</figureCitation>
).
<emphasis id="8E95F443DA75CC0EFB2538B7512AB4D0" box="[1218,1349,222,249]" italics="true" pageId="7" pageNumber="373">Pereopod 6</emphasis>
greatly elongated, dactylus reduced, not adpressed against carpopropodus; propodus, carpus fused, larger than carpopropodus of other pereopods, ventrally with rounded expansion and distoventral finger-like extension; elongate, flattened, distally expanded and distomedially indented process articulated with propodus/carpus dorsal to dactylus, slightly longer than carpopropodus; 2 lateral setae on stalk and 2 long, simple setae at distolateral margins of process; merus triangular, approximately 1/3 length of ischium; ischium cylindrical, approximately subequal in size to basis; basis cylindrical; merus approximately 3 times longer than on other pereopods; ischium approximately 3 times than on other pereopods; basis approximately 2 times longer than on other pereopods (
<figureCitation id="24DA34D4DA75CC0EFBE139B35032B5DD" box="[1030,1117,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="7" pageNumber="373">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="BC5E2851DA75CC0EFF20399751E0B6AD" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="373">
<emphasis id="8E95F443DA75CC0EFF2039975567B631" box="[199,264,511,537]" italics="true" pageId="7" pageNumber="373">Pleon</emphasis>
with 5 pairs of uniramous pleopods (
<figureCitation id="24DA34D4DA75CC0EFD4639975695B631" box="[673,762,511,537]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="7" pageNumber="373">Fig. 3A</figureCitation>
); triangular sympod bearing 2 long setae at inner distal point, cylindrical exopod articulated with outer distal point of sympod bearing 3 long setae.
<emphasis id="8E95F443DA75CC0EFBAE3A4B50C5B615" box="[1097,1194,547,573]" italics="true" pageId="7" pageNumber="373">Uropods</emphasis>
biramous, cylindrical peduncle flared slightly at distal end with 1 stout seta on outer angle, distally with 2 slender rami, exopod ending in 2 thin, short setae and 1 long, robust seta, endopod subequal in length to exopod, with 1 long, robust seta (
<figureCitation id="24DA34D4DA75CC0EFACF3A0351EDB6AD" box="[1320,1410,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1740,1764]" captionTargetBox="[209,1380,658,1710]" captionTargetId="figure-227@6.[201,1387,651,1716]" captionTargetPageId="6" captionText="FIGURE 3. Epicaridium larva of Cancrion khanhensis sp. nov. A, habitus, ventral view; B, pereopod 6. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4315893" httpUri="https://zenodo.org/record/4315893/files/figure.png" pageId="7" pageNumber="373">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="BC5E2851DA75CC0EFF203AE75644B089" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="373">
<emphasis id="8E95F443DA75CC0EFF203AE755CFB681" bold="true" box="[199,416,654,681]" italics="true" pageId="7" pageNumber="373">Cryptoniscus larva</emphasis>
(
<figureCitation id="24DA34D4DA75CC0EFE493AE7566AB681" box="[430,517,655,681]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
) body fusiform, length
<quantity id="7B1985B4DA75CC0EFCEA3AE7570AB681" box="[781,869,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="373" unit="mm" value="700.0">700 µm</quantity>
, maximum width at pereomeres 3 and 4.
<emphasis id="8E95F443DA75CC0EFAC83AE751F3B681" box="[1327,1436,655,681]" italics="true" pageId="7" pageNumber="373">Cephalon</emphasis>
anterior margin round (anterior portion of sole specimen damaged). Body pigmentation lacking.
<emphasis id="8E95F443DA75CC0EFB033ADB5138B6E5" box="[1252,1367,691,717]" italics="true" pageId="7" pageNumber="373">Antennula</emphasis>
missing.
<emphasis id="8E95F443DA75CC0EFF2E3ABF5549B6D9" box="[201,294,727,753]" italics="true" pageId="7" pageNumber="373">Antenna</emphasis>
with four peduncular and 1+ flagellar articles (damaged) (
<figureCitation id="24DA34D4DA75CC0EFC533ABF5061B6D9" box="[948,1038,727,753]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
), distal 2 peduncular articles longer than proximal 2; flagellar article approximately half width of peduncular articles (
<figureCitation id="24DA34D4DA75CC0EFBA73A9350F3B73D" box="[1088,1180,763,789]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
). Oral cone triangular, anteriorly directed (
<figureCitation id="24DA34D4DA75CC0EFE923B7755BFB711" box="[373,464,799,825]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
).
<emphasis id="8E95F443DA75CC0EFE033B7756F6B711" box="[484,665,799,825]" italics="true" pageId="7" pageNumber="373">Pereomeres 17</emphasis>
with entire (not toothed) coxal plates (
<figureCitation id="24DA34D4DA75CC0EFBA93B7750C6B711" box="[1102,1193,799,825]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
).
<emphasis id="8E95F443DA75CC0EFB5A3B77510AB711" box="[1213,1381,799,825]" italics="true" pageId="7" pageNumber="373">Pereopods 15</emphasis>
subequal in shape with long, curved dactylus, propodus expanded proximally with distoventral ridge corresponding to dactylus position bearing small bulges; carpus slender, distally forking; merus triangular, approximately half length of carpus; ischium subrectangular, approximately 2.5 times as long as merus; basis cylindrical, approximately twice as long as ischium (
<figureCitation id="24DA34D4DA75CC0EFE953BC755A5B7E1" box="[370,458,943,969]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4B</figureCitation>
). Pereopods 1 and 2 subequal in size, 35 subequal in size and slightly larger than 1 and 2 (
<figureCitation id="24DA34D4DA75CC0EFF523BBB557FB7C5" box="[181,272,979,1005]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
). Pereopod 6 and 7 missing. Each pleomere with a median ventral acute projection (
<figureCitation id="24DA34D4DA75CC0EFB253BBB5172B7C5" box="[1218,1309,979,1005]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4A</figureCitation>
); 5 pairs of uniramous pleopods; sympod with distomesial short, subquadrate, extension, proximomesial and distomesial corner with few long setae; exopods recurved with few long terminal setae (
<figureCitation id="24DA34D4DA75CC0EFC7E3C73579DB01D" box="[921,1010,1051,1077]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4C</figureCitation>
). Pleotelson subquadrate.
<emphasis id="8E95F443DA75CC0EFAF23C735119B01D" box="[1301,1398,1051,1077]" italics="true" pageId="7" pageNumber="373">Uropods</emphasis>
biramous, composed of wide subquadrate sympod with single long seta at distolateral corner, endopod slightly longer than exopod, pair of long distal setae on endopods and exopods, short seta at distolateral and distomesial margins of endopods and exopods (
<figureCitation id="24DA34D4DA75CC0EFE233CEF5670B089" box="[452,543,1159,1185]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[209,1378,189,1911]" captionTargetId="figure-17@8.[200,1387,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Cryptoniscus larva of Cancrion khanhensis sp. nov. A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="7" pageNumber="373">Fig. 4D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA75CC0EFF203CC251FEB0ED" box="[199,1425,1194,1221]" pageId="7" pageNumber="373" type="distribution">
<paragraph id="BC5E2851DA75CC0EFF203CC251FEB0ED" blockId="7.[151,1437,150,1977]" box="[199,1425,1194,1221]" pageId="7" pageNumber="373">
<emphasis id="8E95F443DA75CC0EFF203CC2550EB0EC" bold="true" box="[199,353,1194,1220]" pageId="7" pageNumber="373">Distribution.</emphasis>
Known only from
<taxonomicName id="7BE153D2DA75CC0EFDD03CC3569EB0ED" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[567,753,1195,1221]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="8E95F443DA75CC0EFDD03CC3569EB0ED" box="[567,753,1195,1221]" italics="true" pageId="7" pageNumber="373">Monomia haanii</emphasis>
</taxonomicName>
collected in Nha Trang bay,
<collectingRegion id="7E25E6B3DA75CC0EFBD43CC3514FB0ED" box="[1075,1312,1195,1221]" country="Vietnam" name="Khanh Hoa" pageId="7" pageNumber="373">Khanh Hoa province</collectingRegion>
,
<collectingCountry id="C4F668C1DA75CC0EFACD3CC351E2B0ED" box="[1322,1421,1195,1221]" name="Vietnam" pageId="7" pageNumber="373">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA75CC00FF203CA75603B25D" lastPageId="9" lastPageNumber="375" pageId="7" pageNumber="373" type="discussion">
<paragraph id="BC5E2851DA75CC0EFF203CA757C9B25D" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="373">
<emphasis id="8E95F443DA75CC0EFF203CA75554B0C1" bold="true" box="[199,315,1231,1257]" pageId="7" pageNumber="373">Remarks.</emphasis>
The new entoniscid isopod appears to belong to the genus
<taxonomicName id="7BE153D2DA75CC0EFC3F3CA7502DB0C1" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[984,1090,1231,1257]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="8E95F443DA75CC0EFC3F3CA7502DB0C1" box="[984,1090,1231,1257]" italics="true" pageId="7" pageNumber="373">Cancrion</emphasis>
</taxonomicName>
with some of whose species it shares the following characters: oostegite 1 of the female of 3 parts; female without ventral ovarian processes; female with two pairs of pleural lamellae; epicaridium larva with elongate pereopod 6 with a highly modified process arising from the distodorsal margin of the propodus. However, it needs to be noted that
<taxonomicName id="7BE153D2DA75CC0EFB653D535083B17D" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[1154,1260,1339,1365]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="8E95F443DA75CC0EFB653D535083B17D" box="[1154,1260,1339,1365]" italics="true" pageId="7" pageNumber="373">Cancrion</emphasis>
</taxonomicName>
is a potentially heterogeneous grouping in that not all the females have the same number of pleural lamellae (
<quantity id="7B1985B4DA75CC0EFB5E3D37508AB151" box="[1209,1253,1375,1401]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="373" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="7BE153D2DA75CC0EFB0B3D3755C9B1B5" authority="Shields &amp; Earley, 1993" authorityName="Shields &amp; Earley" authorityYear="1993" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA75CC0EFB0B3D375090B151" box="[1260,1279,1375,1401]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFAEA3D3751F3B151" box="[1293,1436,1375,1401]" italics="true" pageId="7" pageNumber="373">australiensis</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFF703DEB55C9B1B5" author="Shields, J. D. &amp; Earley, C. G." box="[151,422,1411,1437]" pageId="7" pageNumber="373" pagination="601 - 608" refId="ref14016" refString="Shields, J. D. &amp; Earley, C. G. (1993) Cancrion australiensis new species (Isopoda: Entoniscidae) found in Thalamita sima (Brachyura: Portunidae) from Australia. International Journal for Parasitology, 23, 601 - 608. https: // doi. org / 10.1016 / 0020 - 7519 (93) 90166 - V" type="journal article" year="1993">Shields &amp; Earley, 1993</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7BE153D2DA75CC0EFE543DEB5739B1B5" authority="Pearse &amp; Walker, 1939" authorityName="Pearse &amp; Walker" authorityYear="1939" box="[435,854,1411,1437]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="carolinus">
<emphasis id="8E95F443DA75CC0EFE543DEB55A9B1B5" box="[435,454,1411,1437]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFE303DEB562FB1B5" box="[471,576,1411,1437]" italics="true" pageId="7" pageNumber="373">carolinus</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFDAE3DEB5739B1B5" author="Pearse, A. S. &amp; Walker, H. A." box="[585,854,1411,1437]" pageId="7" pageNumber="373" pagination="19 - 23" refId="ref13600" refString="Pearse, A. S. &amp; Walker, H. A. (1939) Two new parasitic isopods from the eastern coast of North America. Proceedings of the United States National Museum, 87, 19 - 23. https: // doi. org / 10.5479 / si. 00963801.87 - 3067.19" type="journal article" year="1939">Pearse &amp; Walker, 1939</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7BE153D2DA75CC0EFC843DEB50E5B1B5" authority="Shiino, 1942" authorityName="Shiino" authorityYear="1942" box="[867,1162,1411,1437]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA75CC0EFC843DEB5719B1B5" box="[867,886,1411,1437]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFC603DEB5783B1B5" box="[903,1004,1411,1437]" italics="true" pageId="7" pageNumber="373">deltoides</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFC113DEB50E5B1B5" author="Shiino, S. M." box="[1014,1162,1411,1437]" pageId="7" pageNumber="373" pagination="37 - 76" refId="ref14147" refString="Shiino, S. M. (1942) On the parasitic isopods of the family Entoniscidae, especially those found in the vicinity of Seto. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17 (1), 37 - 76." type="journal article" year="1942">Shiino, 1942</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="7BE153D2DA75CC0EFB2D3DEB5521B1E9" authority="Pearse &amp; Walker, 1939" authorityName="Pearse &amp; Walker" authorityYear="1939" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="needleri">
<emphasis id="8E95F443DA75CC0EFB2D3DEB50B2B1B5" box="[1226,1245,1411,1437]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFB093DEB5125B1B5" box="[1262,1354,1411,1437]" italics="true" pageId="7" pageNumber="373">needleri</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFAB43DEB5521B1E9" author="Pearse, A. S. &amp; Walker, H. A." pageId="7" pageNumber="373" pagination="19 - 23" refId="ref13600" refString="Pearse, A. S. &amp; Walker, H. A. (1939) Two new parasitic isopods from the eastern coast of North America. Proceedings of the United States National Museum, 87, 19 - 23. https: // doi. org / 10.5479 / si. 00963801.87 - 3067.19" type="journal article" year="1939">Pearse &amp; Walker, 1939</bibRefCitation>
</taxonomicName>
; 2 or 3 (unclear) in
<taxonomicName id="7BE153D2DA75CC0EFDD53DCF5704B1E9" authority="Muller, 1864" authorityName="Muller" authorityYear="1864" box="[562,875,1447,1473]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="cancrorum">
<emphasis id="8E95F443DA75CC0EFDD53DCF562AB1E9" box="[562,581,1447,1473]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFDB33DCF56A1B1E8" box="[596,718,1447,1472]" italics="true" pageId="7" pageNumber="373">cancrorum</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFD313DCF5704B1E9" author="Muller, F." box="[726,875,1447,1473]" pageId="7" pageNumber="373" refId="ref13568" refString="Muller, F. (1864) Fur Darwin. Wilhelm Engelmann, Leipzig, 91 pp. https: // doi. org / 10.5962 / bhl. title. 168646" type="book" year="1864">Müller, 1864</bibRefCitation>
</taxonomicName>
; more than
<quantity id="7B1985B4DA75CC0EFC153DCF5071B1E9" box="[1010,1054,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="373" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="7BE153D2DA75CC0EFBC13DCF51F8B1E9" authority="Giard &amp; Bonnier, 1887" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[1062,1431,1447,1473]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="miser">
<emphasis id="8E95F443DA75CC0EFBC13DCF5056B1E9" box="[1062,1081,1447,1473]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFBAF3DCF50E9B1E8" box="[1096,1158,1447,1472]" italics="true" pageId="7" pageNumber="373">miser</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFB693DCF51F8B1E9" author="Giard, A. &amp; Bonnier, J." box="[1166,1431,1447,1473]" pageId="7" pageNumber="373" pagination="1 - 272" refId="ref13061" refString="Giard, A. &amp; Bonnier, J. (1887) Contributions a` l'etude des Bopyriens. Traveaux de l'Institut zoologique de Lille et du Laboratoire de Zoologie maritime de Wimereux, 5, 1 - 272, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 64223" type="journal article" year="1887">Giard &amp; Bonnier, 1887</bibRefCitation>
</taxonomicName>
; unknown number in
<taxonomicName id="7BE153D2DA75CC0EFE663DA35767B1CD" authority="Giard &amp; Bonnier, 1887" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[385,776,1483,1509]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="floridus">
<emphasis id="8E95F443DA75CC0EFE663DA355FBB1CD" box="[385,404,1483,1509]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFE453DA35596B1CD" box="[418,505,1483,1509]" italics="true" pageId="7" pageNumber="373">floridus</emphasis>
<bibRefCitation id="D87055A0DA75CC0EFDE63DA35767B1CD" author="Giard, A. &amp; Bonnier, J." box="[513,776,1483,1509]" pageId="7" pageNumber="373" pagination="1 - 272" refId="ref13061" refString="Giard, A. &amp; Bonnier, J. (1887) Contributions a` l'etude des Bopyriens. Traveaux de l'Institut zoologique de Lille et du Laboratoire de Zoologie maritime de Wimereux, 5, 1 - 272, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 64223" type="journal article" year="1887">Giard &amp; Bonnier, 1887</bibRefCitation>
</taxonomicName>
. Additionally, the epicaridium larvae are known from only three other species:
<taxonomicName id="7BE153D2DA75CC0EFE923D87564BB221" authorityName="Shields &amp; Earley" authorityYear="1993" box="[373,548,1519,1545]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA75CC0EFE923D8755E7B221" box="[373,392,1519,1545]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFE723D87564BB221" box="[405,548,1519,1545]" italics="true" pageId="7" pageNumber="373">australiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="7BE153D2DA75CC0EFDD53D8756D8B221" authorityName="Shiino" authorityYear="1942" box="[562,695,1519,1545]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA75CC0EFDD53D87562AB221" box="[562,581,1519,1545]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFDB53D8756D8B221" box="[594,695,1519,1545]" italics="true" pageId="7" pageNumber="373">deltoides</emphasis>
</taxonomicName>
, and
<taxonomicName id="7BE153D2DA75CC0EFD143D875713B221" authorityName="Pearse &amp; Walker" authorityYear="1939" box="[755,892,1519,1545]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="carolinus">
<emphasis id="8E95F443DA75CC0EFD143D875769B221" box="[755,774,1519,1545]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFCF43D875713B221" box="[787,892,1519,1545]" italics="true" pageId="7" pageNumber="373">carolinus</emphasis>
</taxonomicName>
, and only
<taxonomicName id="7BE153D2DA75CC0EFC163D8750CFB221" authorityName="Shields &amp; Earley" authorityYear="1993" box="[1009,1184,1519,1545]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA75CC0EFC163D87506BB221" box="[1009,1028,1519,1545]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFBF63D8750CFB221" box="[1041,1184,1519,1545]" italics="true" pageId="7" pageNumber="373">australiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="7BE153D2DA75CC0EFB323D875134B221" authorityName="Shiino" authorityYear="1942" box="[1237,1371,1519,1545]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA75CC0EFB323D875087B221" box="[1237,1256,1519,1545]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFB113D875134B221" box="[1270,1371,1519,1545]" italics="true" pageId="7" pageNumber="373">deltoides</emphasis>
</taxonomicName>
show the long process arising from the propodus of pereopod 6 as seen in
<taxonomicName id="7BE153D2DA75CC0EFC7F3E7B505AB205" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[920,1077,1555,1581]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA75CC0EFC7F3E7B57C4B205" box="[920,939,1555,1581]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFC5E3E7B505AB205" box="[953,1077,1555,1581]" italics="true" pageId="7" pageNumber="373">khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA75CC0EFBDB3E7B50F9B205" bold="true" box="[1084,1174,1555,1581]" pageId="7" pageNumber="373">
<taxonomicNameLabel id="95A64938DA75CC0EFBDB3E7B50F9B205" box="[1084,1174,1555,1581]" pageId="7" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It is possible that
<taxonomicName id="7BE153D2DA75CC0EFA833E7B54BDB278" authorityName="Giard &amp; Bonnier" authorityYear="1887" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="8E95F443DA75CC0EFA833E7B54BDB278" italics="true" pageId="7" pageNumber="373">Cancrion</emphasis>
</taxonomicName>
is paraphyletic but a complete redescription of
<taxonomicName id="7BE153D2DA75CC0EFD083E5F5721B278" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[751,846,1591,1617]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="miser">
<emphasis id="8E95F443DA75CC0EFD083E5F576DB279" box="[751,770,1591,1617]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFCF73E5F5721B278" box="[784,846,1591,1616]" italics="true" pageId="7" pageNumber="373">miser</emphasis>
</taxonomicName>
, the
<typeStatus id="635A96F3DA75CC0EFC6F3E5F57D7B279" box="[904,952,1591,1617]" pageId="7" pageNumber="373">type</typeStatus>
species, is needed, including details of the epicaridium larval morphology, before any conclusions can be drawn.
</paragraph>
<paragraph id="BC5E2851DA75CC0EFF203E1756F7B391" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="373">
In comparison to other species of
<emphasis id="8E95F443DA75CC0EFDB53E175703B2B1" box="[594,876,1663,1689]" italics="true" pageId="7" pageNumber="373">
<taxonomicName id="7BE153D2DA75CC0EFDB53E1756D1B2B1" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[594,702,1663,1689]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="genus">Cancrion</taxonomicName>
,
<taxonomicName id="7BE153D2DA75CC0EFD2B3E175703B2B1" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[716,876,1663,1689]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">C. khanhensis</taxonomicName>
</emphasis>
<emphasis id="8E95F443DA75CC0EFC913E1757BDB2B1" bold="true" box="[886,978,1663,1689]" pageId="7" pageNumber="373">
<taxonomicNameLabel id="95A64938DA75CC0EFC913E1757BDB2B1" box="[886,978,1663,1689]" pageId="7" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most resembles
<taxonomicName id="7BE153D2DA75CC0EFB723E175128B2B1" authorityName="Shields &amp; Earley" authorityYear="1993" box="[1173,1351,1663,1689]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA75CC0EFB723E175128B2B1" box="[1173,1351,1663,1689]" italics="true" pageId="7" pageNumber="373">C. australiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="7BE153D2DA75CC0EFA643E175493B295" authorityName="Shiino" authorityYear="1942" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA75CC0EFA643E175493B295" italics="true" pageId="7" pageNumber="373">C. deltoides</emphasis>
</taxonomicName>
in all known life history stages based on the characters noted above (cryptoniscus larvae for the latter two species are unknown). However, females, males and epicaridium larvae of
<taxonomicName id="7BE153D2DA75CC0EFC0F3EAF50E9B2C9" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1000,1158,1735,1761]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA75CC0EFC0F3EAF50E9B2C9" box="[1000,1158,1735,1761]" italics="true" pageId="7" pageNumber="373">C. khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA75CC0EFB693EAF5086B2C9" bold="true" box="[1166,1257,1735,1761]" pageId="7" pageNumber="373">
<taxonomicNameLabel id="95A64938DA75CC0EFB693EAF5086B2C9" box="[1166,1257,1735,1761]" pageId="7" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possess several characters that distinguish this species from the other two. The length of both females (
<quantity id="7B1985B4DA75CC0EFB703E825179B32D" box="[1175,1302,1770,1797]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="7" pageNumber="373" unit="mm" value="29.5" valueMax="35.0" valueMin="24.0">2435 mm</quantity>
) and males (
<quantity id="7B1985B4DA75CC0EFF783F675546B301" box="[159,297,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.9" metricValueMin="3.1" pageId="7" pageNumber="373" unit="mm" value="4.0" valueMax="4.9" valueMin="3.1">3.14.9 mm</quantity>
) is considerably larger than for the other two species (
<taxonomicName id="7BE153D2DA75CC0EFC6B3F675053B301" authorityName="Shields &amp; Earley" authorityYear="1993" box="[908,1084,1807,1833]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA75CC0EFC6B3F6757F0B301" box="[908,927,1807,1833]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFC4A3F675053B301" box="[941,1084,1807,1833]" italics="true" pageId="7" pageNumber="373">australiensis</emphasis>
</taxonomicName>
: females
<quantity id="7B1985B4DA75CC0EFB4D3F675120B301" box="[1194,1359,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.8799999999999997" metricValueMax="2.12" metricValueMin="1.64" pageId="7" pageNumber="373" unit="mm" value="18.799999999999997" valueMax="21.2" valueMin="16.4">16.421.2 mm</quantity>
, males 2.12.6;
<taxonomicName id="7BE153D2DA75CC0EFF1D3F5B5510B365" authorityName="Shiino" authorityYear="1942" box="[250,383,1843,1869]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA75CC0EFF1D3F5B5562B365" box="[250,269,1843,1869]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFEFD3F5B5510B365" box="[282,383,1843,1869]" italics="true" pageId="7" pageNumber="373">deltoides</emphasis>
</taxonomicName>
: female
<quantity id="7B1985B4DA75CC0EFE073F5B5657B365" box="[480,568,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="373" unit="mm" value="8.7">8.7 mm</quantity>
, male
<quantity id="7B1985B4DA75CC0EFD673F5A5687B365" box="[640,744,1842,1869]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="7" pageNumber="373" unit="mm" value="1.58">1.58 mm</quantity>
); for females this is probably a function of host size limitation but male size is not strongly correlated to host size due to their being the much smaller sex. The size of epicaridium larvae for
<taxonomicName id="7BE153D2DA75CC0EFEE83F1355C2B3BD" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[271,429,1915,1941]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA75CC0EFEE83F13554DB3BD" box="[271,290,1915,1941]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFED63F1355C2B3BD" box="[305,429,1915,1941]" italics="true" pageId="7" pageNumber="373">khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA75CC0EFE503F13567CB3BD" bold="true" box="[439,531,1915,1941]" pageId="7" pageNumber="373">
<taxonomicNameLabel id="95A64938DA75CC0EFE503F13567CB3BD" box="[439,531,1915,1941]" pageId="7" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<quantity id="7B1985B4DA75CC0EFDC43F1256D8B3BD" box="[547,695,1914,1941]" metricMagnitude="-1" metricUnit="m" metricValue="2.575" metricValueMax="2.65" metricValueMin="2.5" pageId="7" pageNumber="373" unit="mm" value="257.5" valueMax="265.0" valueMin="250.0">250265 µm</quantity>
) is intermediate between those of the other two species (
<taxonomicName id="7BE153D2DA75CC0EFAAE3F135490B391" authorityName="Shields &amp; Earley" authorityYear="1993" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA75CC0EFAAE3F135133B3BD" box="[1353,1372,1915,1941]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFA8B3F135490B391" italics="true" pageId="7" pageNumber="373">australiensis</emphasis>
</taxonomicName>
<quantity id="7B1985B4DA75CC0EFEE13FF755F7B391" box="[262,408,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="2.335" metricValueMax="2.3899999999999997" metricValueMin="2.28" pageId="7" pageNumber="373" unit="mm" value="233.5" valueMax="239.0" valueMin="228.0">228239 µm</quantity>
;
<taxonomicName id="7BE153D2DA75CC0EFE433FF75645B391" authorityName="Shiino" authorityYear="1942" box="[420,554,1951,1977]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="373" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA75CC0EFE433FF755D8B391" box="[420,439,1951,1977]" italics="true" pageId="7" pageNumber="373">C</emphasis>
.
<emphasis id="8E95F443DA75CC0EFE223FF75645B391" box="[453,554,1951,1977]" italics="true" pageId="7" pageNumber="373">deltoides</emphasis>
</taxonomicName>
<quantity id="7B1985B4DA75CC0EFDD63FF656E3B391" box="[561,652,1950,1977]" metricMagnitude="-1" metricUnit="m" metricValue="3.45" pageId="7" pageNumber="373" unit="mm" value="345.0">345 µm</quantity>
).
</paragraph>
<caption id="E89E78D9DA7ACC01FF703FFF5064B3FC" ID-DOI="http://doi.org/10.5281/zenodo.4315895" ID-Zenodo-Dep="4315895" httpUri="https://zenodo.org/record/4315895/files/figure.png" pageId="8" pageNumber="374" startId="8.[151,250,1943,1967]" targetBox="[209,1378,189,1911]" targetPageId="8">
<paragraph id="BC5E2851DA7ACC01FF703FFF5064B3FC" blockId="8.[151,1436,1943,2004]" pageId="8" pageNumber="374">
<emphasis id="8E95F443DA7ACC01FF703FFF5561B387" bold="true" box="[151,270,1943,1967]" pageId="8" pageNumber="374">FIGURE 4</emphasis>
. Cryptoniscus larva of
<taxonomicName id="7BE153D2DA7ACC01FE1A3FF056BAB398" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[509,725,1944,1968]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="374" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7ACC01FE1A3FF056BAB398" box="[509,725,1944,1968]" italics="true" pageId="8" pageNumber="374">Cancrion khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7ACC01FD3C3FF05743B398" bold="true" box="[731,812,1944,1968]" pageId="8" pageNumber="374">
<taxonomicNameLabel id="95A64938DA7ACC01FD3C3FF05743B398" box="[731,812,1944,1968]" pageId="8" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, habitus, ventral view, dashed lines indicate missing parts; B, pereopod 1; C, pleopod 1; D, left uropod. Scale bars: A = 100 µm; B, C, D = 10 µm.
</paragraph>
</caption>
<paragraph id="BC5E2851DA7BCC00FF2038FF556CB5A1" blockId="9.[151,1437,151,1653]" pageId="9" pageNumber="375">
The females of
<taxonomicName id="7BE153D2DA7BCC00FE9138FF567DB499" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[374,530,151,177]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7BCC00FE9138FF55E6B499" box="[374,393,151,177]" italics="true" pageId="9" pageNumber="375">C</emphasis>
.
<emphasis id="8E95F443DA7BCC00FE7138FF567DB499" box="[406,530,151,177]" italics="true" pageId="9" pageNumber="375">khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7BCC00FDFE38FF561CB499" bold="true" box="[537,627,151,177]" pageId="9" pageNumber="375">
<taxonomicNameLabel id="95A64938DA7BCC00FDFE38FF561CB499" box="[537,627,151,177]" pageId="9" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
each possess 4 pairs of dorsolateral ovarian processes, while those of
<taxonomicName id="7BE153D2DA7BCC00FA6438FF5549B4FD" authorityName="Shields &amp; Earley" authorityYear="1993" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA7BCC00FA6438FF5549B4FD" italics="true" pageId="9" pageNumber="375">C. australiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="7BE153D2DA7BCC00FE8738D35588B4FD" authorityName="Shiino" authorityYear="1942" box="[352,487,187,213]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA7BCC00FE8738D35588B4FD" box="[352,487,187,213]" italics="true" pageId="9" pageNumber="375">C. deltoides</emphasis>
</taxonomicName>
bear 3 and 2 pairs, respectively. In addition, the recurrent lamella of
<taxonomicName id="7BE153D2DA7BCC00FB1938D351F3B4FD" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1278,1436,187,213]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7BCC00FB1938D351F3B4FD" box="[1278,1436,187,213]" italics="true" pageId="9" pageNumber="375">C. khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7BCC00FF7038B7549EB4D1" bold="true" box="[151,241,223,249]" pageId="9" pageNumber="375">
<taxonomicNameLabel id="95A64938DA7BCC00FF7038B7549EB4D1" box="[151,241,223,249]" pageId="9" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is developed into an elaborate flap with dorsal flower-shaped folds not seen in the other two species. The males of the new species have horizontal ridges with small median spines on pereomeres 27 and horizontal folds on pleomeres 15 that are lacking in the other two species. The epicaridium larva has a distally expanded and medially indented process on pereopod 6 whereas this process is spoon-shaped in
<taxonomicName id="7BE153D2DA7BCC00FBFF392350A6B54D" authorityName="Shields &amp; Earley" authorityYear="1993" box="[1048,1225,331,357]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="8E95F443DA7BCC00FBFF39235044B54D" box="[1048,1067,331,357]" italics="true" pageId="9" pageNumber="375">C</emphasis>
.
<emphasis id="8E95F443DA7BCC00FBDD392350A6B54D" box="[1082,1225,331,357]" italics="true" pageId="9" pageNumber="375">australiensis</emphasis>
</taxonomicName>
and rounded in
<taxonomicName id="7BE153D2DA7BCC00FA6439235493B5A1" authorityName="Shiino" authorityYear="1942" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="deltoides">
<emphasis id="8E95F443DA7BCC00FA64392351F9B54D" box="[1411,1430,331,357]" italics="true" pageId="9" pageNumber="375">C</emphasis>
.
<emphasis id="8E95F443DA7BCC00FF7039075493B5A1" box="[151,252,367,393]" italics="true" pageId="9" pageNumber="375">deltoides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BC5E2851DA7BCC00FF2039FB550BB631" blockId="9.[151,1437,151,1653]" pageId="9" pageNumber="375">
This is the first description, albeit a partial one, of any cryptoniscus larva of a species of
<taxonomicName id="7BE153D2DA7BCC00FB5E39FB514CB585" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[1209,1315,403,429]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="8E95F443DA7BCC00FB5E39FB514CB585" box="[1209,1315,403,429]" italics="true" pageId="9" pageNumber="375">Cancrion</emphasis>
</taxonomicName>
. Only two cryptoniscus larvae were found on hosts gills, one on each of two crabs. Unfortunately, the first cryptoniscus was damaged during removal and the latter was covered by a dark membrane, possibly caused by host melanization (see
<bibRefCitation id="D87055A0DA7BCC00FF7039975538B631" author="Kuris, A. M. &amp; Poinar, G. O. &amp; Hess, R. T." box="[151,343,511,537]" pageId="9" pageNumber="375" pagination="211 - 232" refId="ref13241" refString="Kuris, A. M., Poinar, G. O. &amp; Hess, R. T. (1980) Post-larval mortality of the endoparasitic castrator Portunion conformis (Epicaridea: Entoniscidae) in the shore crab, Hemigrapsus oregonensis, with a description of the host response. Parasitology, 80, 211 - 232, pls. 1 - 4. https: // doi. org / 10.1017 / S 003118200000069 X" type="journal article" year="1980">
Kuris
<emphasis id="8E95F443DA7BCC00FF3B39975561B631" box="[220,270,511,537]" italics="true" pageId="9" pageNumber="375">et al</emphasis>
. 1980
</bibRefCitation>
).
</paragraph>
<paragraph id="BC5E2851DA7BCC00FF203A4B551DB6E5" blockId="9.[151,1437,151,1653]" pageId="9" pageNumber="375">
<bibRefCitation id="D87055A0DA7BCC00FF203A4B55F7B615" author="Kuris, A. M. &amp; Poinar, G. O. &amp; Hess, R. T." box="[199,408,547,573]" pageId="9" pageNumber="375" pagination="211 - 232" refId="ref13241" refString="Kuris, A. M., Poinar, G. O. &amp; Hess, R. T. (1980) Post-larval mortality of the endoparasitic castrator Portunion conformis (Epicaridea: Entoniscidae) in the shore crab, Hemigrapsus oregonensis, with a description of the host response. Parasitology, 80, 211 - 232, pls. 1 - 4. https: // doi. org / 10.1017 / S 003118200000069 X" type="journal article" year="1980">
Kuris
<emphasis id="8E95F443DA7BCC00FEEA3A4B5550B615" box="[269,319,547,573]" italics="true" pageId="9" pageNumber="375">et al</emphasis>
. (1980)
</bibRefCitation>
divided the growth of female entoniscids into 12 stages based on development of morphological features and reproductive condition. We have only observed female entoniscids corresponding to four of these stages (2, 5, 8, 12) as well as a stage not reported by
<bibRefCitation id="D87055A0DA7BCC00FCCF3A035794B6AD" author="Kuris, A. M. &amp; Poinar, G. O. &amp; Hess, R. T." box="[808,1019,619,645]" pageId="9" pageNumber="375" pagination="211 - 232" refId="ref13241" refString="Kuris, A. M., Poinar, G. O. &amp; Hess, R. T. (1980) Post-larval mortality of the endoparasitic castrator Portunion conformis (Epicaridea: Entoniscidae) in the shore crab, Hemigrapsus oregonensis, with a description of the host response. Parasitology, 80, 211 - 232, pls. 1 - 4. https: // doi. org / 10.1017 / S 003118200000069 X" type="journal article" year="1980">
Kuris
<emphasis id="8E95F443DA7BCC00FC8A3A0357F0B6AD" box="[877,927,619,645]" italics="true" pageId="9" pageNumber="375">et al</emphasis>
. (1980)
</bibRefCitation>
: a cylindrical vermiform stage which appears to be one immediately following molting of the cryptoniscus exuvia as it is the most reduced in structure and simplest shape.
</paragraph>
<paragraph id="BC5E2851DA7BCC00FF203ABE5054B039" blockId="9.[151,1437,151,1653]" pageId="9" pageNumber="375">
<emphasis id="8E95F443DA7BCC00FF203ABE5523B6D8" bold="true" box="[199,332,726,752]" pageId="9" pageNumber="375">Discussion.</emphasis>
<emphasis id="8E95F443DA7BCC00FEB43ABF5675B6D8" box="[339,538,727,752]" italics="true" pageId="9" pageNumber="375">Parasitic position</emphasis>
. As with all other
<taxonomicName id="7BE153D2DA7BCC00FD0D3ABF573BB6D9" authorityName="Giard &amp; Bonnier" authorityYear="1887" box="[746,852,727,753]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="8E95F443DA7BCC00FD0D3ABF573BB6D9" box="[746,852,727,753]" italics="true" pageId="9" pageNumber="375">Cancrion</emphasis>
</taxonomicName>
species,
<taxonomicName id="7BE153D2DA7BCC00FC5E3ABF503AB6D9" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[953,1109,727,753]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7BCC00FC5E3ABF503AB6D9" box="[953,1109,727,753]" italics="true" pageId="9" pageNumber="375">C. khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7BCC00FBBB3ABF50D9B6D9" bold="true" box="[1116,1206,727,753]" pageId="9" pageNumber="375">
<taxonomicNameLabel id="95A64938DA7BCC00FBBB3ABF50D9B6D9" box="[1116,1206,727,753]" pageId="9" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lies in the hemocoel of the host and is enclosed by the host response sheath (
<figureCitation id="24DA34D4DA7BCC00FCEA3A9357E7B73D" box="[781,904,763,789]" captionStart="FIGURE 9" captionStartId="15.[151,250,1231,1255]" captionTargetBox="[151,1435,181,1207]" captionTargetId="figure-25@15.[151,1435,181,1207]" captionTargetPageId="15" captionText="FIGURE 9. A, Uninfested Monomia haani; B, infested M. haani showing more highly vaulted carapace; C, ventral view of M. haanii carapace showing ovigerous female Cancrion khanhensis sp. nov. (arrow); D, double infestation of C. khanhensis sp. nov. (arrows). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4315905" httpUri="https://zenodo.org/record/4315905/files/figure.png" pageId="9" pageNumber="375">Fig. 9C, D</figureCitation>
; see also
<bibRefCitation id="D87055A0DA7BCC00FC133A9350EFB73D" author="Shiino, S. M." box="[1012,1152,763,789]" pageId="9" pageNumber="375" pagination="37 - 76" refId="ref14147" refString="Shiino, S. M. (1942) On the parasitic isopods of the family Entoniscidae, especially those found in the vicinity of Seto. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17 (1), 37 - 76." type="journal article" year="1942">Shiino 1942</bibRefCitation>
;
<bibRefCitation id="D87055A0DA7BCC00FB6B3A9351FFB73D" author="Shields, J. D. &amp; Earley, C. G." box="[1164,1424,763,789]" pageId="9" pageNumber="375" pagination="601 - 608" refId="ref14016" refString="Shields, J. D. &amp; Earley, C. G. (1993) Cancrion australiensis new species (Isopoda: Entoniscidae) found in Thalamita sima (Brachyura: Portunidae) from Australia. International Journal for Parasitology, 23, 601 - 608. https: // doi. org / 10.1016 / 0020 - 7519 (93) 90166 - V" type="journal article" year="1993">Shields &amp; Earley 1993</bibRefCitation>
). The female is curved dorsally with the whole marsupium, including the cephalon and pereon, laying in one side of the hosts hemocoel cavity with the “hood” (distal extension of the oostegites) extending to the anterior border of the host carapace while the abdomen is curved under the hosts stomach, reaching to the other side of the host (
<figureCitation id="24DA34D4DA7BCC00FA923B0F54B2B78D" captionStart="FIGURE 9" captionStartId="15.[151,250,1231,1255]" captionTargetBox="[151,1435,181,1207]" captionTargetId="figure-25@15.[151,1435,181,1207]" captionTargetPageId="15" captionText="FIGURE 9. A, Uninfested Monomia haani; B, infested M. haani showing more highly vaulted carapace; C, ventral view of M. haanii carapace showing ovigerous female Cancrion khanhensis sp. nov. (arrow); D, double infestation of C. khanhensis sp. nov. (arrows). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4315905" httpUri="https://zenodo.org/record/4315905/files/figure.png" pageId="9" pageNumber="375">Fig 9C, D</figureCitation>
). The exit pore, via which the parasite isopod communicates with the external environment, was not observed but was likely entering into the branchial chamber (see
<bibRefCitation id="D87055A0DA7BCC00FCEF3BC75061B7E1" author="McDermott, J. J. &amp; Williams, J. D. &amp; Boyko, C. B." box="[776,1038,943,969]" pageId="9" pageNumber="375" pagination="97 - 114" refId="ref13460" refString="McDermott, J. J., Williams, J. D. &amp; Boyko, C. B. (2019) A new genus and species of parasitic isopod (Bopyroidea: Entoniscidae) infesting pinnotherid crabs (Brachyura: Pinnotheridae) on the Atlantic Coast of the USA, with notes on the life cycle of entoniscids. Journal of Crustacean Biology, 41, 97 - 114. [first published online 16 Dec 2019, with fixed content and layout in compliance with Art. 8.1. 3.2 ICZN] https: // doi. org / 10.1093 / jcbiol / ruz 088" type="journal article" year="2019">
McDermott
<emphasis id="8E95F443DA7BCC00FC753BC757ACB7E1" box="[914,963,943,969]" italics="true" pageId="9" pageNumber="375">et al</emphasis>
. 2019
</bibRefCitation>
). The male is of the
<typeStatus id="635A96F3DA7BCC00FB133BC7514BB7E1" box="[1268,1316,943,969]" pageId="9" pageNumber="375">type</typeStatus>
which has a very strongly curved body (
<bibRefCitation id="D87055A0DA7BCC00FE053BBB5602B7C5" author="Shiino, S. M." box="[482,621,979,1005]" pageId="9" pageNumber="375" pagination="37 - 76" refId="ref14147" refString="Shiino, S. M. (1942) On the parasitic isopods of the family Entoniscidae, especially those found in the vicinity of Seto. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17 (1), 37 - 76." type="journal article" year="1942">Shiino 1942</bibRefCitation>
) and is usually found in the brood-chamber of the female. The marsupia of most females that were not hyperparasitized were filled with epicaridium larvae.
</paragraph>
<paragraph id="BC5E2851DA7BCC00FF203C7350C0B1B5" blockId="9.[151,1437,151,1653]" pageId="9" pageNumber="375">
<emphasis id="8E95F443DA7BCC00FF203C73558EB01C" box="[199,481,1051,1077]" italics="true" pageId="9" pageNumber="375">Prevalence and intensity.</emphasis>
Out of a total of 366
<taxonomicName id="7BE153D2DA7BCC00FD343C7357E9B01D" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[723,902,1051,1077]" class="Malacostraca" family="Portunidae" genus="Monomia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="haani">
<emphasis id="8E95F443DA7BCC00FD343C7357E9B01D" box="[723,902,1051,1077]" italics="true" pageId="9" pageNumber="375">Monomia haani</emphasis>
</taxonomicName>
examined, there were 37 crabs infested with
<taxonomicName id="7BE153D2DA7BCC00FA643C73557CB071" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7BCC00FA643C73557CB071" italics="true" pageId="9" pageNumber="375">C. khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7BCC00FEFC3C575519B071" bold="true" box="[283,374,1087,1113]" pageId="9" pageNumber="375">
<taxonomicNameLabel id="95A64938DA7BCC00FEFC3C575519B071" box="[283,374,1087,1113]" pageId="9" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; a prevalence of 10.1%. The prevalence of isopod infestation in female crab hosts was 13.2% (18/136) and was significantly higher than the 0.83% (19/230) found in male hosts (Chi-square test, χ2 = 127.75, df = 1,
<emphasis id="8E95F443DA7BCC00FF0C3CEF5496B088" box="[235,249,1159,1184]" italics="true" pageId="9" pageNumber="375">p</emphasis>
&lt;0.01). Each host was parasitized by one (
<figureCitation id="24DA34D4DA7BCC00FD0C3CEF572DB089" box="[747,834,1159,1185]" captionStart="FIGURE 9" captionStartId="15.[151,250,1231,1255]" captionTargetBox="[151,1435,181,1207]" captionTargetId="figure-25@15.[151,1435,181,1207]" captionTargetPageId="15" captionText="FIGURE 9. A, Uninfested Monomia haani; B, infested M. haani showing more highly vaulted carapace; C, ventral view of M. haanii carapace showing ovigerous female Cancrion khanhensis sp. nov. (arrow); D, double infestation of C. khanhensis sp. nov. (arrows). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4315905" httpUri="https://zenodo.org/record/4315905/files/figure.png" pageId="9" pageNumber="375">Fig. 9C</figureCitation>
) or two (
<figureCitation id="24DA34D4DA7BCC00FC4B3CEF506AB089" box="[940,1029,1159,1185]" captionStart="FIGURE 9" captionStartId="15.[151,250,1231,1255]" captionTargetBox="[151,1435,181,1207]" captionTargetId="figure-25@15.[151,1435,181,1207]" captionTargetPageId="15" captionText="FIGURE 9. A, Uninfested Monomia haani; B, infested M. haani showing more highly vaulted carapace; C, ventral view of M. haanii carapace showing ovigerous female Cancrion khanhensis sp. nov. (arrow); D, double infestation of C. khanhensis sp. nov. (arrows). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4315905" httpUri="https://zenodo.org/record/4315905/files/figure.png" pageId="9" pageNumber="375">Fig. 9D</figureCitation>
) female entoniscids with double infestations observed in 12 of the hosts. Of these 12 double infestations, there were eight crabs infested by one larger female entoniscid with a marsupium full of epicaridium larvae and one smaller female with eggs; three crabs bore one mature and one immature female entoniscid; and one crab was infested with two juvenile entoniscid females. Females and males of
<taxonomicName id="7BE153D2DA7BCC00FE7D3D7F5656B119" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[410,569,1303,1329]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7BCC00FE7D3D7F5656B119" box="[410,569,1303,1329]" italics="true" pageId="9" pageNumber="375">C. khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7BCC00FDA53D7F56F1B119" bold="true" box="[578,670,1303,1329]" pageId="9" pageNumber="375">
<taxonomicNameLabel id="95A64938DA7BCC00FDA53D7F56F1B119" box="[578,670,1303,1329]" pageId="9" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were not always present together, with the number of males being equal to or often fewer than the number of females in the same host. Specifically, 12 of the 25 single female infestations and two of the 12 double female infestations had no males present, whereas one male accompanying a pair of females occurred in six hosts, and two males present with two females occurred in four hosts.
</paragraph>
<paragraph id="BC5E2851DA7BCC00FF203DCF5603B25D" blockId="9.[151,1437,151,1653]" pageId="9" pageNumber="375">
Infested host crabs can often be recognized by a vertical swelling of the carapace when the female entoniscids are large (
<figureCitation id="24DA34D4DA7BCC00FEE03DA35532B1CD" box="[263,349,1483,1509]" captionStart="FIGURE 9" captionStartId="15.[151,250,1231,1255]" captionTargetBox="[151,1435,181,1207]" captionTargetId="figure-25@15.[151,1435,181,1207]" captionTargetPageId="15" captionText="FIGURE 9. A, Uninfested Monomia haani; B, infested M. haani showing more highly vaulted carapace; C, ventral view of M. haanii carapace showing ovigerous female Cancrion khanhensis sp. nov. (arrow); D, double infestation of C. khanhensis sp. nov. (arrows). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4315905" httpUri="https://zenodo.org/record/4315905/files/figure.png" pageId="9" pageNumber="375">Fig. 9B</figureCitation>
) but modification of the host is not apparent when the isopod parasite is small and immature and the carapace appears the same as that of unifested hosts (
<figureCitation id="24DA34D4DA7BCC00FCF03D87571EB221" box="[791,881,1519,1545]" captionStart="FIGURE 9" captionStartId="15.[151,250,1231,1255]" captionTargetBox="[151,1435,181,1207]" captionTargetId="figure-25@15.[151,1435,181,1207]" captionTargetPageId="15" captionText="FIGURE 9. A, Uninfested Monomia haani; B, infested M. haani showing more highly vaulted carapace; C, ventral view of M. haanii carapace showing ovigerous female Cancrion khanhensis sp. nov. (arrow); D, double infestation of C. khanhensis sp. nov. (arrows). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4315905" httpUri="https://zenodo.org/record/4315905/files/figure.png" pageId="9" pageNumber="375">Fig. 9A</figureCitation>
). The sizes (carapace width) of infested crabs are in the range of
<quantity id="7B1985B4DA7BCC00FEA43E7B55BCB205" box="[323,467,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" metricValueMax="8.2" metricValueMin="4.7" pageId="9" pageNumber="375" unit="mm" value="64.5" valueMax="82.0" valueMin="47.0">47 to 82 mm</quantity>
for females and
<quantity id="7B1985B4DA7BCC00FD763E7A574EB205" box="[657,801,1554,1581]" metricMagnitude="-2" metricUnit="m" metricValue="7.35" metricValueMax="9.2" metricValueMin="5.5" pageId="9" pageNumber="375" unit="mm" value="73.5" valueMax="92.0" valueMin="55.0">55 to 92 mm</quantity>
for males. Castration of hosts infested by
<taxonomicName id="7BE153D2DA7BCC00FB183E7B51F3B205" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1279,1436,1555,1581]" class="Malacostraca" family="Entoniscidae" genus="Cancrion" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="375" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA7BCC00FB183E7B51F3B205" box="[1279,1436,1555,1581]" italics="true" pageId="9" pageNumber="375">C. khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA7BCC00FF703E5F549DB279" bold="true" box="[151,242,1591,1617]" pageId="9" pageNumber="375">
<taxonomicNameLabel id="95A64938DA7BCC00FF703E5F549DB279" box="[151,242,1591,1617]" pageId="9" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was not observed in male or female hosts, as 14 of 18 infested female (1 being ovigerous) and 17 of 19 infested male hosts were sexually mature.
</paragraph>
</subSubSection>
</treatment>
</document>