213 lines
25 KiB
XML
213 lines
25 KiB
XML
<document id="67100CDE9B17733B27D42AB3ED498F5B" ID-DOI="10.11646/zootaxa.5446.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11084604" ID-ZooBank="A2CC8E7D-0766-4EC7-80F6-412A269E465D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714390467925" checkinUser="plazi" docAuthor="Kamiński, Marcin Jan" docDate="2024" docId="03F36C399A18D46DD9FBFACE6745F843" docLanguage="en" docName="zootaxa.5446.1.4.pdf" docOrigin="Zootaxa 5446 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5446.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Toktokkus zofiae Kamiński 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="85" masterDocId="FFCA14419A1FD465D96CFFB26308FFB5" masterDocTitle="New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel" masterLastPageNumber="87" masterPageNumber="77" pageNumber="84" updateTime="1714390994664" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="85F483F45A0753BF098DEB4A2EF80986" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="896427299006F0E7C6E496036F529510">
|
||
<mods:title id="3D842B7A3B4F0DDE8B6A67FBE045A98B">New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="CF39CBE81E46F6826BD723E5EACE92B2" type="personal">
|
||
<mods:role id="ABA131D1959F4767A2317551DDEEB9B8">
|
||
<mods:roleTerm id="ABF2C3EAAEC6A609D65393E32B592AFA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F7F632805456EC2E069DF6345A5CC93D">Kamiński, Marcin Jan</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="569EFBEE95B3E7AF85F64C90BC90E1CF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="5969D082E5F3F6414D7A2BC1FB501F3D" type="host">
|
||
<mods:titleInfo id="EC9A3142ADCC72EA9160C891C81A2A6B">
|
||
<mods:title id="5B6BE140B323212C2203E4CF5540A444">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C4CC456CA869802EA31D96E3BF9C845E">
|
||
<mods:date id="52549B20ACFE365B0C90E5463D295322">2024</mods:date>
|
||
<mods:detail id="406B01CDA72C9443395B4773B7AC287E" type="pubDate">
|
||
<mods:number id="EBDFB085277A83AF8295FE2775DCD222">2024-04-29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A662544CF47216458579DCEF98AE06B6" type="volume">
|
||
<mods:number id="08AE806575114D253CFD8F6D7DB315B3">5446</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D6414E0BEF75A14888669CDD8AAB15B3" type="issue">
|
||
<mods:number id="4BEEC554F89A4AB111D56CA081C32D65">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="DFC76D15811B429948D556FB41193581" unit="page">
|
||
<mods:start id="0E056C1DA25A8333CE475D37BB809C5B">77</mods:start>
|
||
<mods:end id="CBFEFA17AE8215AE40497CDE0B9C2605">87</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="9F567ADB0D441593CDF0B1BABDD4DE0F">
|
||
<mods:url id="24E8831C1E3EEEAE5BEFA2D6D0B2525A">http://dx.doi.org/10.11646/zootaxa.5446.1.4</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="08213198B171C1044C85A6A976A9F88A">journal article</mods:classification>
|
||
<mods:identifier id="08C7BC4FA54204A82E4F4319575D7215" type="DOI">10.11646/zootaxa.5446.1.4</mods:identifier>
|
||
<mods:identifier id="B26ACD69E0CA3D61AED18E23E9E93CF4" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="D5F14FBA308ECC682ED5DC2C84B9416D" type="Zenodo-Dep">11084604</mods:identifier>
|
||
<mods:identifier id="AB00971C82E3B4BF3FFEF95CEAF39791" type="ZooBank">A2CC8E7D-0766-4EC7-80F6-412A269E465D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F36C399A18D46DD9FBFACE6745F843" ID-DOI="http://doi.org/10.5281/zenodo.11084617" ID-Zenodo-Dep="11084617" LSID="urn:lsid:plazi:treatment:03F36C399A18D46DD9FBFACE6745F843" httpUri="http://treatment.plazi.org/id/03F36C399A18D46DD9FBFACE6745F843" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84">
|
||
<subSubSection id="C3408EA49A18D462D9FBFACE62C0FA22" box="[151,456,1404,1431]" pageId="7" pageNumber="84" type="nomenclature">
|
||
<paragraph id="8BE5DD2F9A18D462D9FBFACE62C0FA22" blockId="7.[151,456,1404,1466]" box="[151,456,1404,1431]" pageId="7" pageNumber="84">
|
||
<heading id="D0AD6A439A18D462D9FBFACE62C0FA22" bold="true" box="[151,456,1404,1431]" fontSize="11" level="1" pageId="7" pageNumber="84" reason="1">
|
||
<emphasis id="B92E013D9A18D462D9FBFACE62C0FA22" bold="true" box="[151,456,1404,1431]" pageId="7" pageNumber="84">
|
||
<taxonomicName id="4C5AA6AC9A18D462D9FBFACE626BFA23" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[151,355,1404,1431]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="zofiae" status="sp. nov.">
|
||
<emphasis id="B92E013D9A18D462D9FBFACE626BFA23" bold="true" box="[151,355,1404,1431]" italics="true" pageId="7" pageNumber="84">Toktokkus zofiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DBC469A18D462D806FACF62C0FA22" box="[362,456,1405,1431]" pageId="7" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A18D462D9FBFA126215FA0F" box="[151,285,1440,1466]" pageId="7" pageNumber="84" type="description">
|
||
<paragraph id="8BE5DD2F9A18D462D9FBFA126215FA0F" blockId="7.[151,456,1404,1466]" box="[151,285,1440,1466]" pageId="7" pageNumber="84">
|
||
(
|
||
<figureCitation id="1361C1AA9A18D462D9F3FA12621DFA0F" box="[159,277,1440,1466]" captionStart="FIGURE 6" captionStartId="8.[151,250,1199,1223]" captionTargetBox="[156,1435,183,1168]" captionTargetId="figure-22@8.[151,1436,181,1175]" captionTargetPageId="8" captionText="FIGURE 6. Morphology of Toktokkus zofiae sp. nov. (holotype) (A–F) and T. herero (G, H). Lateral (A), dorsal (B), and ventral (C) views of the body; pronotal disc (D), prosternal process (E, H); label (F); and elytral disc (G)." figureDoi="http://doi.org/10.5281/zenodo.11084625" httpUri="https://zenodo.org/record/11084625/files/figure.png" pageId="7" pageNumber="84">Fig. 6A–F</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A18D462D9FBFA5B63CEF992" pageId="7" pageNumber="84" type="materials_examined">
|
||
<paragraph id="8BE5DD2F9A18D462D9FBFA5B63CEF992" blockId="7.[151,1437,1512,2043]" pageId="7" pageNumber="84">
|
||
<emphasis id="B92E013D9A18D462D9FBFA5B6249F9B7" bold="true" box="[151,321,1512,1539]" pageId="7" pageNumber="84">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B32D7729A18D462D83EFA5B63CEF992" collectionCode="NHMB" collectorName="V. - L. Kai. Exp. Gemsbok Pan." pageId="7" pageNumber="84" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B92E013D9A18D462D83EFA5B62B7F9B6" bold="true" box="[338,447,1513,1539]" pageId="7" pageNumber="84">
|
||
<typeStatus id="54E1638D9A18D462D83EFA5B62B7F9B6" box="[338,447,1513,1539]" pageId="7" pageNumber="84" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED4B45EA9A18D462D8BEFA5B6124F9B6" box="[466,556,1513,1539]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="84">NHMB</collectionCode>
|
||
), female: “
|
||
<collectorName id="26AFB8F99A18D462DBC7FA5B6707F9B6" box="[683,1039,1512,1539]" pageId="7" pageNumber="84">V.-L. Kai. Exp. Gemsbok Pan.</collectorName>
|
||
23/4–5/5/30.”, “
|
||
<taxonomicName id="4C5AA6AC9A18D462DDA2FA5A665EF9B7" authorityName="Kirby" authorityYear="1819" box="[1230,1366,1512,1538]" class="Insecta" family="Tenebrionidae" genus="Psammodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="genus">Psammodes</taxonomicName>
|
||
peezi K.”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A18D462D9ABF982618DF94A" pageId="7" pageNumber="84" type="diagnosis">
|
||
<paragraph id="8BE5DD2F9A18D462D9ABF982618DF94A" blockId="7.[151,1437,1512,2043]" pageId="7" pageNumber="84">
|
||
<emphasis id="B92E013D9A18D462D9ABF982624AF9FF" bold="true" box="[199,322,1584,1610]" pageId="7" pageNumber="84">Diagnosis.</emphasis>
|
||
The species is most similar to
|
||
<taxonomicName id="4C5AA6AC9A18D462DBE3F983609CF9FE" authority="Gearner, 2021" authorityName="Gearner" authorityYear="2021" box="[655,916,1584,1611]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="B92E013D9A18D462DBE3F98361FAF9FF" box="[655,754,1584,1610]" italics="true" pageId="7" pageNumber="84">T. herero</emphasis>
|
||
<bibRefCitation id="EFCBA0DE9A18D462DB9BF983609CF9FE" author="Gearner, O. M. & Kaminski, M. J. & Kanda, K. & Swichtenberg, K. & Smith, A. D." box="[759,916,1585,1611]" pageId="7" pageNumber="84" pagination="1 - 10" refId="ref5773" refString="Gearner, O. M., Kaminski, M. J., Kanda, K., Swichtenberg, K. & Smith, A. D. (2021) Discovery of New Genera Challenges the Subtribal Classification of Tok-Tok Beetles (Coleoptera: Tenebrionidae: Sepidiini). Insect Systematics and Diversity, 5, 1 - 10. https: // doi. org / 10.1093 / isd / ixab 006" type="journal article" year="2021">Gearner, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
by having prominent punctures on the pronotal disc and a golden setation of the elytral disc (
|
||
<bibRefCitation id="EFCBA0DE9A18D462DBFFF9E6608CF9DA" author="Kaminski, M. J. & Gearner, O. M. & Kanda, K & Swichtenberg, K. & Purchart, L. & Smith, A. D." box="[659,900,1620,1647]" pageId="7" pageNumber="84" pagination="883 - 901" refId="ref6154" refString="Kaminski, M. J., Gearner, O. M., Kanda, K .., Swichtenberg, K., Purchart, L. & Smith, A. D. (2021) First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina), and examination of the status of the Psammodes vialis species-group. Zoological Journal of the Linnean Society, 191, 883 - 901. https: // doi. org / 10.1093 / zoolinnean / zlaa 052" type="journal article" year="2021">
|
||
Kaminski
|
||
<emphasis id="B92E013D9A18D462DA65F9E76049F9DB" box="[777,833,1620,1646]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
). Both species can be distinguished by different body size (
|
||
<taxonomicName id="4C5AA6AC9A18D462D879F9CB627CF926" authorityName="Kamiński" authorityYear="2024" box="[277,372,1657,1683]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="zofiae">
|
||
<emphasis id="B92E013D9A18D462D879F9CB627CF926" box="[277,372,1657,1683]" italics="true" pageId="7" pageNumber="84">T. zofiae</emphasis>
|
||
</taxonomicName>
|
||
: 18.5 mm,
|
||
<taxonomicName id="4C5AA6AC9A18D462D894F9CB6156F927" authorityName="Gearner" authorityYear="2021" box="[504,606,1656,1682]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="B92E013D9A18D462D894F9CB6156F927" box="[504,606,1656,1682]" italics="true" pageId="7" pageNumber="84">T. herero</emphasis>
|
||
</taxonomicName>
|
||
: 23.0–25.0 mm), basal margination of pronotal disc (
|
||
<taxonomicName id="4C5AA6AC9A18D462DDD3F9CB6617F926" authorityName="Kamiński" authorityYear="2024" box="[1215,1311,1657,1683]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="zofiae">
|
||
<emphasis id="B92E013D9A18D462DDD3F9CB6617F926" box="[1215,1311,1657,1683]" italics="true" pageId="7" pageNumber="84">T. zofiae</emphasis>
|
||
</taxonomicName>
|
||
: complete,
|
||
<taxonomicName id="4C5AA6AC9A18D462D9FBF92F63F4F903" authorityName="Gearner" authorityYear="2021" box="[151,252,1692,1718]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="B92E013D9A18D462D9FBF92F63F4F903" box="[151,252,1692,1718]" italics="true" pageId="7" pageNumber="84">T. herero</emphasis>
|
||
</taxonomicName>
|
||
: interrupted in the middle) (
|
||
<figureCitation id="1361C1AA9A18D462DB55F92E619BF903" box="[569,659,1692,1718]" captionStart="FIGURE 6" captionStartId="8.[151,250,1199,1223]" captionTargetBox="[156,1435,183,1168]" captionTargetId="figure-22@8.[151,1436,181,1175]" captionTargetPageId="8" captionText="FIGURE 6. Morphology of Toktokkus zofiae sp. nov. (holotype) (A–F) and T. herero (G, H). Lateral (A), dorsal (B), and ventral (C) views of the body; pronotal disc (D), prosternal process (E, H); label (F); and elytral disc (G)." figureDoi="http://doi.org/10.5281/zenodo.11084625" httpUri="https://zenodo.org/record/11084625/files/figure.png" pageId="7" pageNumber="84">Fig. 6D</figureCitation>
|
||
), structure of prosternal process (
|
||
<taxonomicName id="4C5AA6AC9A18D462DD67F92F6761F902" authorityName="Kamiński" authorityYear="2024" box="[1035,1129,1693,1719]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="zofiae">
|
||
<emphasis id="B92E013D9A18D462DD67F92F6761F902" box="[1035,1129,1693,1719]" italics="true" pageId="7" pageNumber="84">T. zofiae</emphasis>
|
||
</taxonomicName>
|
||
: rounded in lateral view,
|
||
<taxonomicName id="4C5AA6AC9A18D462DCE4F92F63E8F96F" authorityName="Gearner" authorityYear="2021" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="B92E013D9A18D462DCE4F92F63E8F96F" italics="true" pageId="7" pageNumber="84">T. herero</emphasis>
|
||
</taxonomicName>
|
||
: angular in lateral view) (
|
||
<figureCitation id="1361C1AA9A18D462DB69F9726177F96E" box="[517,639,1728,1755]" captionStart="FIGURE 6" captionStartId="8.[151,250,1199,1223]" captionTargetBox="[156,1435,183,1168]" captionTargetId="figure-22@8.[151,1436,181,1175]" captionTargetPageId="8" captionText="FIGURE 6. Morphology of Toktokkus zofiae sp. nov. (holotype) (A–F) and T. herero (G, H). Lateral (A), dorsal (B), and ventral (C) views of the body; pronotal disc (D), prosternal process (E, H); label (F); and elytral disc (G)." figureDoi="http://doi.org/10.5281/zenodo.11084625" httpUri="https://zenodo.org/record/11084625/files/figure.png" pageId="7" pageNumber="84">Fig. 6H, E</figureCitation>
|
||
), and tuberculation of elytral disc (
|
||
<taxonomicName id="4C5AA6AC9A18D462DD60F9736763F96E" authorityName="Kamiński" authorityYear="2024" box="[1036,1131,1729,1755]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="zofiae">
|
||
<emphasis id="B92E013D9A18D462DD60F9736763F96E" box="[1036,1131,1729,1755]" italics="true" pageId="7" pageNumber="84">T. zofiae</emphasis>
|
||
</taxonomicName>
|
||
: microtubercles present,
|
||
<taxonomicName id="4C5AA6AC9A18D462DCE4F97363E8F94B" authorityName="Gearner" authorityYear="2021" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="B92E013D9A18D462DCE4F97363E8F94B" italics="true" pageId="7" pageNumber="84">T. herero</emphasis>
|
||
</taxonomicName>
|
||
: microtubercles absent) (
|
||
<figureCitation id="1361C1AA9A18D462D890F9566171F94A" box="[508,633,1764,1791]" captionStart="FIGURE 6" captionStartId="8.[151,250,1199,1223]" captionTargetBox="[156,1435,183,1168]" captionTargetId="figure-22@8.[151,1436,181,1175]" captionTargetPageId="8" captionText="FIGURE 6. Morphology of Toktokkus zofiae sp. nov. (holotype) (A–F) and T. herero (G, H). Lateral (A), dorsal (B), and ventral (C) views of the body; pronotal disc (D), prosternal process (E, H); label (F); and elytral disc (G)." figureDoi="http://doi.org/10.5281/zenodo.11084625" httpUri="https://zenodo.org/record/11084625/files/figure.png" pageId="7" pageNumber="84">Fig. 6A, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A18D46DD9ABF8BA62ECF83F" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84" type="description">
|
||
<paragraph id="8BE5DD2F9A18D462D9ABF8BA663DF896" blockId="7.[151,1437,1512,2043]" box="[199,1333,1800,1827]" pageId="7" pageNumber="84">
|
||
<emphasis id="B92E013D9A18D462D9ABF8BA6141F897" bold="true" box="[199,585,1800,1827]" pageId="7" pageNumber="84">
|
||
Description (based on
|
||
<typeStatus id="54E1638D9A18D462D8BEF8BA6134F897" box="[466,572,1800,1826]" pageId="7" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Length 18.5 mm, width of pronotum 7.0 mm, and elytra 11.0 mm.
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A18D462D9ABF89E6021F84F" blockId="7.[151,1437,1512,2043]" pageId="7" pageNumber="84">
|
||
<emphasis id="B92E013D9A18D462D9ABF89E620CF8F3" box="[199,260,1836,1862]" italics="true" pageId="7" pageNumber="84">Head</emphasis>
|
||
: Hypognathous. Frons finely and irregularly punctate; frontoclypeal suture coarse, with deep groove in middle; apical clypeal margin broadly shallowly emarginate; clypeus projected toward front of body. Eye comma-shaped, with reduced ventral part, strongly emarginate around epistomal base; with deep groove on temporal side. Mentum and submentum missing due to dermestid damage. Antennae broken off at level of 4th segments; slender, moderately covered in recumbent acuminate goldish setae; antennomere 2 short, equal to 0.2 of antennomere 3 length; antennomere 4 about half of antennomere 3 length.
|
||
</paragraph>
|
||
<caption id="DF258DA79A17D46DD9FBFB1D677DFB5E" ID-DOI="http://doi.org/10.5281/zenodo.11084625" ID-Zenodo-Dep="11084625" httpUri="https://zenodo.org/record/11084625/files/figure.png" pageId="8" pageNumber="85" startId="8.[151,250,1199,1223]" targetBox="[156,1435,183,1168]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BE5DD2F9A17D46DD9FBFB1D677DFB5E" blockId="8.[151,1436,1199,1259]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9FBFB1D621AFB72" bold="true" box="[151,274,1199,1223]" pageId="8" pageNumber="85">FIGURE 6.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C5AA6AC9A17D46DD8DAFB1D6155FB72" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[438,605,1199,1223]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="zofiae" status="sp. nov.">
|
||
<emphasis id="B92E013D9A17D46DD8DAFB1D6155FB72" box="[438,605,1199,1223]" italics="true" pageId="8" pageNumber="85">Toktokkus zofiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A17D46DDB0EFB1D61B9FB72" bold="true" box="[610,689,1199,1223]" pageId="8" pageNumber="85">
|
||
<taxonomicNameLabel id="A21DBC469A17D46DDB0EFB1D61B9FB72" box="[610,689,1199,1223]" pageId="8" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype) (A–F) and
|
||
<taxonomicName id="4C5AA6AC9A17D46DDAFFFB0260E5FB72" authorityName="Gearner" authorityYear="2021" box="[915,1005,1199,1223]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="B92E013D9A17D46DDAFFFB0260E5FB72" box="[915,1005,1199,1223]" italics="true" pageId="8" pageNumber="85">T. herero</emphasis>
|
||
</taxonomicName>
|
||
(G, H). Lateral (A), dorsal (B), and ventral (C) views of the body; pronotal disc (D), prosternal process (E, H); label (F); and elytral disc (G).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABFA826794FA02" blockId="8.[151,1437,1328,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABFA826231FAFF" box="[199,313,1328,1354]" italics="true" pageId="8" pageNumber="85">Prothorax</emphasis>
|
||
: Pronotal lateral margin rounded, well visible. Pronotum widest above middle. Disc dull, finely punctate (4–5 diameters apart); anterior and basal margins complete, anterior apices slightly produced. Hypomeron convex, without submarginal groove, impunctate. Prosternal process rounded in lateral view, longitudinally depressed in middle (ventral view). Anterior margin of prosternum straight, with gold setae.
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABFA726628F93B" blockId="8.[151,1437,1328,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABFA726245FA6F" box="[199,333,1472,1498]" italics="true" pageId="8" pageNumber="85">Pterothorax</emphasis>
|
||
: Scutellum densely covered with microtubercles. Elytra widest in half, slightly rounded, covered in scattered gold setae; covered with dense tubercles (1–3 diameters apart) and microtubercles (2–4 diameters apart). Laterally situated tubercles very pronounced. Elytral slope relatively steep, elytral apex flattened. Epipleura impunctate, not tuberculated, clearly distinguishable from neighbouring portion of elytra, widely enfolding fifth ventrite. Mesoventrite with deep median groove and elevated sides. Metaventrite impunctate, densely setose. Lateral regions of metaventrite (between coxae) extremely short. Metaepisternal suture abbreviated posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABF92B6676F963" blockId="8.[151,1437,1328,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABF92B63F4F907" box="[199,252,1689,1714]" italics="true" pageId="8" pageNumber="85">Legs</emphasis>
|
||
: Covered with dense gold setae. Procoxa exposed basally. Apex of protibia with denticle on outer margin, lateral carina terminating in basal third. Spurs on meso- and metatibiae of equal length. Tarsi narrowed laterally.
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABF9526658F8AB" blockId="8.[151,1437,1328,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABF9526239F94F" box="[199,305,1760,1786]" italics="true" pageId="8" pageNumber="85">Abdomen</emphasis>
|
||
: Ventrites 1–3 medially covered with goldish setae, moderately punctate (2–3 diameters apart) and weakly rugulose; ventrites 4–5 finely punctate (1–3 diameters apart); ventrite 5 without submarginal sulcus.
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABF89A62ECF83F" blockId="8.[151,1437,1328,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABF89A6249F8F7" box="[199,321,1832,1858]" italics="true" pageId="8" pageNumber="85">Terminalia</emphasis>
|
||
: Due to scarcity of materials, terminalia were not dissected. Nevertheless, the tip of the ovipositor was visible on the analyzed specimen. The morphology of coxites is consistent with the bauplan reported for the genus (
|
||
<bibRefCitation id="EFCBA0DE9A17D46DD98BF8C262D0F83F" author="Kaminski, M. J. & Gearner, O. M. & Kanda, K & Swichtenberg, K. & Purchart, L. & Smith, A. D." box="[231,472,1904,1930]" pageId="8" pageNumber="85" pagination="883 - 901" refId="ref6154" refString="Kaminski, M. J., Gearner, O. M., Kanda, K .., Swichtenberg, K., Purchart, L. & Smith, A. D. (2021) First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina), and examination of the status of the Psammodes vialis species-group. Zoological Journal of the Linnean Society, 191, 883 - 901. https: // doi. org / 10.1093 / zoolinnean / zlaa 052" type="journal article" year="2021">
|
||
Kamiński
|
||
<emphasis id="B92E013D9A17D46DD831F8C3629DF83F" box="[349,405,1904,1930]" italics="true" pageId="8" pageNumber="85">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A17D46DD9ABF8266252F867" pageId="8" pageNumber="85" type="etymology">
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABF8266252F867" blockId="8.[151,1437,1328,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABF8266246F81B" bold="true" box="[199,334,1940,1966]" pageId="8" pageNumber="85">Etymology.</emphasis>
|
||
This species is named in honor of my daughter, Zofia Irena Kamińska, born on
|
||
<date id="FFE4FBEF9A17D46DDDA7F8266694F81B" box="[1227,1436,1940,1966]" pageId="8" pageNumber="85" value="2018-11-03">November 3, 2018</date>
|
||
(Flagstaff,
|
||
<collectingCountry id="F34D9DBF9A17D46DD87EF80A6247F867" box="[274,335,1976,2002]" name="United States of America" pageId="8" pageNumber="85">USA</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A17D46DD9ABF86E6745F843" box="[199,1101,2012,2039]" pageId="8" pageNumber="85" type="distribution">
|
||
<paragraph id="8BE5DD2F9A17D46DD9ABF86E6745F843" blockId="8.[151,1437,1328,2039]" box="[199,1101,2012,2039]" pageId="8" pageNumber="85">
|
||
<emphasis id="B92E013D9A17D46DD9ABF86E6269F843" bold="true" box="[199,353,2012,2038]" pageId="8" pageNumber="85">Distribution.</emphasis>
|
||
<collectingCountry id="F34D9DBF9A17D46DD80BF86E62FFF843" box="[359,503,2012,2038]" name="South Africa" pageId="8" pageNumber="85">South Africa</collectingCountry>
|
||
—only known from the
|
||
<typeStatus id="54E1638D9A17D46DDA68F86F603CF842" box="[772,820,2013,2039]" pageId="8" pageNumber="85">type</typeStatus>
|
||
locality (Gemsbok Pan).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |