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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.164.2201" ID-GBIF-Dataset="18e175c4-93e1-4cba-a7fd-3b05815a9af4" ID-PMC="PMC3272619" ID-Pensoft-Pub="1313-2970-164-1" ID-PubMed="22328849" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-164-1" ModsDocOrigin="ZooKeys 164" ModsDocTitle="Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico" checkinTime="1451249389041" checkinUser="pensoft" docAuthor="Zaldivar-Riveron, Alejandro, Martinez, Juan Jose, Ceccarelli, Fadia Sara &amp; Shaw, Scott R." docDate="2012" docId="AB5E8E67CAA0BCD4BF06D210BFC9CA2A" docLanguage="en" docName="ZooKeys 164: 1-23" docOrigin="ZooKeys 164" docSource="http://dx.doi.org/10.3897/zookeys.164.2201" docTitle="Heerz macrophthalma Martinez, Zaldivar-Riveron, Ceccarelli &amp; Shaw, sp. n." docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="4B553355FFC5FFA9AC7BFFE1FD18FFD2" masterDocTitle="Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico" masterLastPageNumber="23" masterPageNumber="1" pageNumber="8" updateTime="1668153157884" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zaldivar-Riveron, Alejandro</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martinez, Juan Jose</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ceccarelli, Fadia Sara</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaw, Scott R.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>164</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>23</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.164.2201</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.164.2201</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-164-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152033021" LSID="urn:lsid:plazi:treatment:AB5E8E67CAA0BCD4BF06D210BFC9CA2A" httpUri="http://treatment.plazi.org/id/AB5E8E67CAA0BCD4BF06D210BFC9CA2A" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection pageId="7" pageNumber="8" type="nomenclature">
<paragraph pageId="7" pageNumber="8">
<taxonomicName LSID="urn:lsid:zoobank.org:act:2F240FAD-E0C2-43FD-ACBC-B66E432EA6B2" authority="Martinez, Zaldivar-Riveron, Ceccarelli &amp; Shaw" class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz macrophthalma" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="macrophthalma">
Heerz macrophthalma
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
,
<normalizedToken originalValue="Zaldívar-Riverón">Zaldivar-Riveron</normalizedToken>
, Ceccarelli &amp; Shaw
</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
Figs 3
<normalizedToken originalValue="AD">A-D</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="diagnosis">
<paragraph pageId="7" pageNumber="8">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="8">
<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz macrophthalma" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="macrophthalma">Heerz macrophthalma</taxonomicName>
distinguishes from the remaining species of the genus by its considerably larger eyes (Fig. 3B). It can also be distinguished from the Central and South American species,
<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz tooya" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tooya">Heerz tooya</taxonomicName>
and
<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz lukenatcha" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lukenatcha">Heerz lukenatcha</taxonomicName>
, by its entirely hyaline wings (Fig. 3A) [wings partially or totally infuscate in the latter two species (Fig. 2B)] and from
<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz ecmahla" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ecmahla">Heerz ecmahla</taxonomicName>
by the striate sculpture on the second metasomal tergite (Fig. 3D) [coriaceous in
<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz ecmahla" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ecmahla">Heerz ecmahla</taxonomicName>
(Fig. 2C)].
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph pageId="7" pageNumber="8">Description.</paragraph>
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Female. Colour: Body mostly honey yellow, antennae honey yellow, turning darker apically; legs creamish white, with apex of tarsomeres darker; wings hyaline, veins, pterostigma and tegula light brown. Body length: 4.7 mm. Head: vertex, frons and temple striate, face acinose; eyes large, 1.3 times higher than wide (lateral view); malar space about 0.2 times eye height; ocello-ocular distance less than 0.8 times diameter of lateral ocellus and as long as posterior ocellar line; antennae with 31 flagellomeres. Mesosoma: length of mesosoma 1.8 times its maximum height; pronotum smooth, pronotal groove scrobiculate; mesoscutal lobes coriaceous; notauli deep and scrobiculate, obscured in a posterior striate-rugose median area; scutellum smooth, scutellar sulcus deep and scrobiculate, with six longitudinal carinae; mesopleuron largely smooth, striate near subalar furrow; posterior mesopleural sulcus distinct and scrobiculate; subalar groove scrobiculate; precoxal sulcus smooth; metapleuron smooth; propodeal areola distinctly delimited by carinae and with crossing transversal rugae. Legs: fore tibia with a row of 15 spines; hind coxa striate dorsally, smooth ventrally, with a distinct basoventral tooth. Wings: fore wing 2.8 times longer than wide; pterostigma 0.6 times the length of vein R; vein r 1.5 times longer than vein 3RSa; vein 3RSa 0.8 times as long as vein r-m; vein m-cu distinctly antefurcal, vein (Rs+M)b present, 0.2 times length of vein 2RS; hind wing with vein M+CU as long as vein 1M and twice length of vein r-m; vein m-cu slightly curved towards wing apex. Metasoma: first metasomal tergite short, about as long as its apical width, with median dorsal area delimited by two dorsolateral carinae, median area rugulose-coriaceous, lateral areas striate; second metasomal tergite striate medially, with coriaceous sculpture in between striations, with two more conspicuous slightly convergent furrows; third metasomal terigte smooth, turning coriaceous later
<pageBreakToken pageId="8" pageNumber="9" start="start">ally</pageBreakToken>
, with a transverse scrobiculate furrow; remaining metasomal tergites smooth and polished; ovipositor 0.5 times length of metasoma, without distinct nodes.
</paragraph>
<paragraph pageId="8" pageNumber="9">Male. Smaller than female, body length 2.7-2.8 mm.</paragraph>
<paragraph pageId="8" pageNumber="9">Variation. Female: Body length 4.1-4.6 mm; eyes 1.3-1.4 times higher than wide (lateral view); malar space 0.1-0.2 times eye height (lateral view); ocello-ocular distance 0.7-0.8 times diameter of lateral ocellus; antennae with 28-29 flagellomeres; fore wing length 2.8-2.9 times its maximum width; length of pterostigma 2.6-2.8 times its maximum width; hind wing vein M+CU 1.0-1.1 times longer than vein 1M.</paragraph>
<paragraph pageId="8" pageNumber="9">
Holotype. IB-UNAM CNIN. Female. Mexico, Jalisco,
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
Chamela, cerca laboratorio, 19.49, -105.04, 5.v.2011, 99-122 msnm, light trap, selva baja caducifolia, Cham-084,
<normalizedToken originalValue="Zaldívar">Zaldivar</normalizedToken>
, Zaragoza, Ibarra col. DNA voucher no. CNIN795, GenBank accession nos JQ268745, JQ268749 (IB-UNAM CNIN).
</paragraph>
<paragraph lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Paratypes. IB-UNAM CNIN, MACN, UWIM. Three specimens. One female, Mexico, Jalisco,
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
Chamela, camino
<normalizedToken originalValue="Búho">Buho</normalizedToken>
, 19.49/19.49 N, -105.04/-105.04 E, 25.ii.2010, 106 msnm, sweeping net, selva baja caducifolia, A.
<normalizedToken originalValue="Zaldívar">Zaldivar</normalizedToken>
col., DNA voucher no. ASDOR761, GenBank accession no. HQ200977; two males, Mexico, Jalisco,
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
Chamela, cerca laboratorio, 19.49 N, -105.04 E, 20.ii.2011, 99-122 msnm, light trap, selva baja caducifolia, A.
<normalizedToken originalValue="Zaldívar">Zaldivar</normalizedToken>
, col., DNA voucher nos A
<pageBreakToken pageId="9" pageNumber="10" start="start">SDOR</pageBreakToken>
551 (Cham-500), ASDOR555 (Cham-504), GenBank accession nos HQ200979, HQ200967 (IB-UNAM CNIN).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="etymology">
<paragraph pageId="9" pageNumber="10">Etymology.</paragraph>
<paragraph pageId="9" pageNumber="10">The specific epithet derives from the greek words makros and ophthalmos, in reference to the very large compound eyes of this species.</paragraph>
<caption pageId="9" pageNumber="10">
<paragraph pageId="9" pageNumber="10">
Figure 3.
<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz macrophthalma" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macrophthalma">Heerz macrophthalma</taxonomicName>
sp. n. (holotype): A habitus, lateral view B head, frontal view C mesosoma, dorsal view D propodeum and basal half of mesosoma, dorsolateral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>