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<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben &amp; Tattersfield, Peter" docDate="2013" docId="4511E41DD837FFD4FE46FE8BF0D2FA35" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella cuspidata Rowson &amp; Tattersfield 2013, comb. nov." docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="20" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rowson, Ben</mods:namePart>
<mods:nameIdentifier type="LSID">05D46259-DA45-45A4-9127-5AF361D566A7</mods:nameIdentifier>
<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tattersfield, Peter</mods:namePart>
<mods:nameIdentifier type="LSID">8C55642E-B18A-461B-9BF9-31CB6DB7A070</mods:nameIdentifier>
<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-02-04</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>37</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>46</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.5852/ejt.2013.37</mods:identifier>
<mods:identifier type="GBIF-Dataset">9682b45b-5e22-4965-a293-e203660e0ed3</mods:identifier>
<mods:identifier type="ISSN">2118-9773</mods:identifier>
<mods:identifier type="Zenodo-Dep">3806770</mods:identifier>
<mods:identifier type="ZooBank">ADF6394E-77B5-4309-87FE-4097FDE0A3FD</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3815775" ID-GBIF-Taxon="163971590" ID-Zenodo-Dep="3815775" LSID="urn:lsid:plazi:treatment:4511E41DD837FFD4FE46FE8BF0D2FA35" httpUri="http://treatment.plazi.org/id/4511E41DD837FFD4FE46FE8BF0D2FA35" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection box="[482,1106,267,294]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph blockId="19.[482,1106,267,330]" box="[482,1106,267,294]" pageId="19" pageNumber="20">
<heading box="[482,1106,267,294]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<taxonomicName ID-CoL="33XQD" authority="(Verdcourt, 1962)" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1962" box="[482,969,267,294]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="cuspidata" status="comb. nov.">
<emphasis bold="true" box="[482,753,267,294]" italics="true" pageId="19" pageNumber="20">Dadagulella cuspidata</emphasis>
(
<bibRefCitation author="Verdcourt B." box="[768,961,267,294]" pageId="19" pageNumber="20" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">
Verdcourt,
<date box="[901,961,267,294]" pageId="19" pageNumber="20" value="1962">1962</date>
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel box="[976,1106,267,294]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[655,932,303,330]" pageId="19" pageNumber="20" type="description">
<paragraph blockId="19.[482,1106,267,330]" box="[655,932,303,330]" pageId="19" pageNumber="20">
<figureCitation box="[655,745,303,329]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson &amp; Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="19" pageNumber="20">Figs 31</figureCitation>
,
<figureCitation box="[757,789,303,330]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="19" pageNumber="20">59</figureCitation>
,
<figureCitation box="[801,834,303,329]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="19" pageNumber="20">84</figureCitation>
;
<tableCitation box="[845,932,303,329]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="19" pageNumber="20" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="reference_group">
<paragraph blockId="19.[189,857,369,395]" box="[189,857,369,395]" pageId="19" pageNumber="20">
<treatmentCitationGroup box="[189,857,369,395]" pageId="19" pageNumber="20">
<treatmentCitation author="Verdcourt B." box="[189,626,369,395]" page="3" pageId="19" pageNumber="20" year="1962">
<taxonomicName ID-CoL="6KTKP" authority="Verdcourt, 1962: 3" authorityName="Verdcourt" authorityPageNumber="3" authorityYear="1962" box="[189,626,369,395]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="cuspidata">
<emphasis box="[189,398,369,395]" italics="true" pageId="19" pageNumber="20">Gulella cuspidata</emphasis>
<bibRefCitation author="Verdcourt B." box="[406,626,369,395]" pageId="19" pageNumber="20" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">
Verdcourt,
<date box="[536,599,369,395]" pageId="19" pageNumber="20" value="1962">1962</date>
: 3
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 27-28, pl. 3, fig. 2.
</treatmentCitationGroup>
</paragraph>
<treatmentCitationGroup pageId="19" pageNumber="20">
<paragraph blockId="19.[189,1399,435,569]" box="[189,810,435,462]" pageId="19" pageNumber="20">
“
<emphasis box="[202,386,435,461]" italics="true" pageId="19" pageNumber="20">
<taxonomicName ID-CoL="7P4LY" authorityName="L. Pfeiffer" authorityYear="1856" box="[202,290,435,461]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">Gulella</taxonomicName>
sp. nov.
</emphasis>
”
<emphasis box="[406,421,436,461]" italics="true" pageId="19" pageNumber="20">–</emphasis>
Verdcourt
<date box="[551,614,435,462]" pageId="19" pageNumber="20" value="1958">1958</date>
: 94, 100, fig. 10.
</paragraph>
<paragraph blockId="19.[189,1399,435,569]" pageId="19" pageNumber="20">
<emphasis box="[189,431,471,497]" italics="true" pageId="19" pageNumber="20">
<taxonomicName authorityName="Verdcourt" authorityYear="1962" box="[189,403,471,497]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="cuspidata">Gulella cuspidata</taxonomicName>
–
</emphasis>
van
<treatmentCitation author="Bruggen A. C. van" box="[498,724,471,497]" page="71" pageId="19" pageNumber="20" year="1969">
<bibRefCitation author="Bruggen A. C. van" box="[498,724,471,497]" pageId="19" pageNumber="20" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">
Bruggen
<date box="[612,675,471,497]" pageId="19" pageNumber="20" value="1969">1969</date>
: 71
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Verdcourt B." box="[781,1037,471,497]" page="232" pageId="19" pageNumber="20" year="1983">
<bibRefCitation author="Verdcourt B." box="[781,1037,471,497]" pageId="19" pageNumber="20" pagination="200 - 239" refId="ref27933" refString="Verdcourt B. 1983. A list of the non-marine Mollusca of East Africa (Kenya, Uganda, Tanzania, excluding Lake Malawi). Achatina 11: 200 - 239." type="journal article" year="1983">
Verdcourt
<date box="[910,973,471,497]" pageId="19" pageNumber="20" value="1983">1983</date>
: 232
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Richardson C. L." box="[1094,1353,471,497]" page="72" pageId="19" pageNumber="20" year="1988">
<bibRefCitation author="Richardson C. L." box="[1094,1353,471,497]" pageId="19" pageNumber="20" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">
Richardson
<date box="[1241,1304,471,497]" pageId="19" pageNumber="20" value="1988">1988</date>
: 72
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Verdcourt B." box="[189,434,506,532]" page="136" pageId="19" pageNumber="20" year="1996">
<bibRefCitation author="Verdcourt B." box="[189,434,506,532]" pageId="19" pageNumber="20" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">
Verdcourt
<date box="[313,376,506,532]" pageId="19" pageNumber="20" value="1996">1996</date>
: 136
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Tattersfield P." box="[484,746,506,532]" page="37" pageId="19" pageNumber="20" year="1998">
<bibRefCitation author="Tattersfield P." box="[484,746,506,532]" pageId="19" pageNumber="20" pagination="31 - 41" refId="ref27614" refString="Tattersfield P. 1998 b. Three new species and a new subgenus of Gulella (Gastropoda: Streptaxidae) from Tanzania. Journal of Conchology 36 (2): 31 - 41." type="journal article" year="1998">
Tattersfield
<date box="[623,685,506,532]" pageId="19" pageNumber="20" value="1998">1998</date>
b: 37
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Tattersfield P. &amp; Seddon M. B. &amp; Meena C. &amp; Kayumbo N. &amp; Kasigwa P." box="[794,1123,506,533]" page="135" pageId="19" pageNumber="20" year="1998">
<bibRefCitation author="Tattersfield P. &amp; Seddon M. B. &amp; Meena C. &amp; Kayumbo N. &amp; Kasigwa P." box="[794,1123,506,533]" pageId="19" pageNumber="20" pagination="119 - 138" refId="ref27649" refString="Tattersfield P., Seddon M. B., Meena C., Kayumbo N. &amp; Kasigwa P. 1998. Ecology and conservation of the land-snails of the Eastern Arc Mountains. Journal of East African Natural History 87 (1): 119 - 138. http: // dx. doi. org / 10.2982 / 0012 - 8317 (1998) 87 [119: EACOTL] 2.0. CO; 2" type="journal article" year="1998">
Tattersfield
<emphasis box="[934,993,506,532]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
<date box="[1001,1064,506,532]" pageId="19" pageNumber="20" value="1998">1998</date>
: 135
</bibRefCitation>
</treatmentCitation>
. — van
<treatmentCitation author="Bruggen A. C. van" page="233" pageId="19" pageNumber="20" year="2000">
<bibRefCitation author="Bruggen A. C. van" pageId="19" pageNumber="20" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">
Bruggen
<date box="[1332,1395,506,532]" pageId="19" pageNumber="20" value="2000">2000</date>
: 233
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Verdcourt B." box="[284,514,542,568]" page="46" pageId="19" pageNumber="20" year="2006">
<bibRefCitation author="Verdcourt B." box="[284,514,542,568]" pageId="19" pageNumber="20" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">
Verdcourt
<date box="[407,470,542,568]" pageId="19" pageNumber="20" value="2006">2006</date>
: 46
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Rowson B. &amp; Lange C. N." box="[562,882,542,569]" page="31" pageId="19" pageNumber="20" year="2007">
<bibRefCitation author="Rowson B. &amp; Lange C. N." box="[562,882,542,569]" pageId="19" pageNumber="20" pagination="21 - 32" refId="ref27291" refString="Rowson B. &amp; Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">
Rowson &amp; Lange
<date box="[776,839,542,568]" pageId="19" pageNumber="20" value="2007">2007</date>
: 31
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Muratov I. V." box="[931,1161,542,569]" page="277" pageId="19" pageNumber="20" year="2010">
<bibRefCitation author="Muratov I. V." box="[931,1161,542,569]" pageId="19" pageNumber="20" pagination="255 - 288" refId="ref26968" refString="Muratov I. V. 2010. Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique. African Invertebrates 51 (2): 255 - 288. http: // dx. doi. org / 10.5733 / afin. 051.0203" type="journal article" year="2010">
Muratov
<date box="[1039,1102,542,568]" pageId="19" pageNumber="20" value="2010">2010</date>
: 277
</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="materials_examined">
<paragraph blockId="19.[189,492,608,634]" box="[189,492,608,634]" pageId="19" pageNumber="20">
<heading bold="true" box="[189,492,608,634]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,492,608,634]" pageId="19" pageNumber="20">
<typeStatus box="[189,250,608,634]" pageId="19" pageNumber="20">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,1399,655,718]" pageId="19" pageNumber="20">
<materialsCitation ID-GBIF-Occurrence="2610647307" collectingDate="1956-12" collectionCode="MRAC" country="Tanzania" latitude="-4.7" location="Shume" longLatPrecision="783" longitude="38.2" municipality="Worlds View" pageId="19" pageNumber="20" specimenCode="MRAC.792352" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
<collectingCountry box="[189,343,655,681]" name="Tanzania" pageId="19" pageNumber="20">TANZANIA</collectingCountry>
:
<typeStatus box="[359,462,655,681]" pageId="19" pageNumber="20">holotype</typeStatus>
<specimenCode box="[475,661,655,682]" pageId="19" pageNumber="20">
<collectionCode45 box="[475,566,655,682]" pageId="19" pageNumber="20">MRAC</collectionCode45>
.792352
</specimenCode>
:
<specimenCount box="[678,743,655,681]" pageId="19" pageNumber="20" type="adult">1 ad.</specimenCount>
,
<collectingMunicipality box="[761,921,655,681]" pageId="19" pageNumber="20">Worlds View</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:4511E41DD837FFD4FE46FE8BF0D2FA35:C86703D0D837FFD4FC0EFD0FF1ADFD4E" box="[938,1022,655,681]" country="Tanzania" latitude="-4.7" longLatPrecision="783" longitude="38.2" municipality="Worlds View" name="Shume" pageId="19" pageNumber="20">Shume</location>
,
<location LSID="urn:lsid:plazi:treatment:4511E41DD837FFD4FE46FE8BF0D2FA35:C86703D0D837FFD4FBB4FD10F75AFD4D" box="[1040,1289,655,682]" country="Tanzania" latitude="-4.7" longLatPrecision="783" longitude="38.2" municipality="Worlds View" name="West Usambara Mts" pageId="19" pageNumber="20">West Usambara Mts</location>
(
<geoCoordinate box="[1311,1393,655,681]" degrees="4.70" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="-4.7">4.70°S</geoCoordinate>
,
<geoCoordinate box="[189,289,691,717]" degrees="38.20" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="38.2">38.20°E</geoCoordinate>
), in rather dry evergreen forest, leg. B. &amp; L. Verdcourt,
<date box="[943,1066,691,718]" pageId="19" pageNumber="20" value="1956-12">
<collectingDate box="[943,1066,691,718]" pageId="19" pageNumber="20" value="1956-12">Dec. 1956</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="19.[189,505,757,783]" box="[189,505,757,783]" pageId="19" pageNumber="20">
<heading bold="true" box="[189,505,757,783]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,505,757,783]" pageId="19" pageNumber="20">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,260,804,830]" box="[189,260,804,830]" pageId="19" pageNumber="20">None.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="description">
<paragraph blockId="19.[189,334,870,896]" box="[189,334,870,896]" pageId="19" pageNumber="20">
<heading bold="true" box="[189,334,870,896]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,334,870,896]" pageId="19" pageNumber="20">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,1399,917,1229]" pageId="19" pageNumber="20">
<collectionCode box="[189,257,917,943]" pageId="19" pageNumber="20">
<collectionCode45 box="[189,257,917,943]" pageId="19" pageNumber="20">SHELL</collectionCode45>
</collectionCode>
(
<figureCitation box="[273,360,917,943]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson &amp; Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="19" pageNumber="20">Figs 31</figureCitation>
,
<figureCitation box="[370,403,917,944]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="19" pageNumber="20">59</figureCitation>
). Large (
<quantity box="[503,668,917,943]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="19" pageNumber="20" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">
4.30
<emphasis box="[556,566,918,943]" italics="true" pageId="19" pageNumber="20">-</emphasis>
4.80 mm
</quantity>
high x 1.80 x
<quantity box="[826,928,917,944]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="2.5">2.50 mm</quantity>
wide), of 7.75 whorls. Ovate-acuminate, spire coeloconoid (spire angle 58°). Apex sharply pointed. Embryonic whorls appear smooth but with fine regular radial striae. Later whorls with relatively few, coarse ribs (about 8 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a furrow-like depression corresponding to the upper palatal tooth. Dentition 6-fold (alternatively recognisable as 5-fold), consisting of: one lamella-like parietal tooth; two palatal teeth, the upper much larger and forming a parieto-palatal sinus; one basal, in-running tooth; and a strong, squarish columellar tooth, running in to form a columellar baffle which is connected to it. Juvenile shells and anatomy unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="multiple">
<paragraph blockId="19.[189,421,1268,1295]" box="[189,421,1268,1295]" pageId="19" pageNumber="20">
<heading bold="true" box="[189,421,1268,1295]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,421,1268,1295]" pageId="19" pageNumber="20">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,1375,1315,1342]" box="[189,1375,1315,1342]" pageId="19" pageNumber="20">
Elevation not stated but probably between
<quantity box="[691,848,1315,1342]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="19" pageNumber="20" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">
<date box="[691,818,1315,1342]" pageId="19" pageNumber="20" value="1500" valueMax="2000">1500-2000</date>
m
</quantity>
at the
<typeStatus box="[927,978,1316,1342]" pageId="19" pageNumber="20">type</typeStatus>
locality in northeastern
<collectingCountry box="[1262,1371,1315,1341]" name="Tanzania" pageId="19" pageNumber="20">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="discussion">
<paragraph blockId="19.[189,303,1382,1408]" box="[189,303,1382,1408]" pageId="19" pageNumber="20">
<heading bold="true" box="[189,303,1382,1408]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,303,1382,1408]" pageId="19" pageNumber="20">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,1399,1428,1490]" pageId="19" pageNumber="20">
This species differs from
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[487,627,1429,1454]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="minareta" status="sp. nov.">
<emphasis box="[487,627,1429,1454]" italics="true" pageId="19" pageNumber="20">D. minareta</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[634,724,1429,1455]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
in having two palatal teeth, in having a basal tooth, in the shape of the parietal tooth, and in size.
</paragraph>
</subSubSection>
</treatment>
</document>
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