487 lines
62 KiB
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487 lines
62 KiB
XML
<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben & Tattersfield, Peter" docDate="2013" docId="4511E41DD82EFFCBFE16F960F0C8FB62" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella browni subsp. browni browni (van Bruggen 1969, comb. nov." docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="11" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
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<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rowson, Ben</mods:namePart>
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<mods:affiliation>Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tattersfield, Peter</mods:namePart>
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<mods:nameIdentifier type="LSID">8C55642E-B18A-461B-9BF9-31CB6DB7A070</mods:nameIdentifier>
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<mods:affiliation>Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-02-04</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>37</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.5852/ejt.2013.37</mods:identifier>
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<mods:identifier type="GBIF-Dataset">9682b45b-5e22-4965-a293-e203660e0ed3</mods:identifier>
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<mods:identifier type="ISSN">2118-9773</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3806770</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3815763" ID-GBIF-Taxon="163971596" ID-Zenodo-Dep="3815763" LSID="urn:lsid:plazi:treatment:4511E41DD82EFFCBFE16F960F0C8FB62" httpUri="http://treatment.plazi.org/id/4511E41DD82EFFCBFE16F960F0C8FB62" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<subSubSection box="[434,1154,1760,1788]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph blockId="10.[434,1154,1760,1823]" box="[434,1154,1760,1788]" pageId="10" pageNumber="11">
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<heading box="[434,1154,1760,1788]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
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<taxonomicName ID-CoL="5DZBS" authority="browni (van Bruggen, 1969)" authorityName="browni (van Bruggen" authorityYear="1969" box="[434,1017,1760,1788]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" species="browni" status="comb. nov." subSpecies="browni">
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<emphasis bold="true" box="[434,766,1760,1787]" italics="true" pageId="10" pageNumber="11">Dadagulella browni browni</emphasis>
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(van
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<bibRefCitation author="Bruggen A. C. van" box="[833,1009,1761,1788]" pageId="10" pageNumber="11" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen, 1969</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[1024,1154,1761,1787]" pageId="10" pageNumber="11" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[550,1038,1797,1823]" pageId="10" pageNumber="11" type="description">
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<paragraph blockId="10.[434,1154,1760,1823]" box="[550,1038,1797,1823]" pageId="10" pageNumber="11">
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<figureCitation box="[550,679,1797,1823]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="10" pageNumber="11">Figs 23-26</figureCitation>
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,
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<figureCitation box="[691,762,1797,1823]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="10" pageNumber="11">47-48</figureCitation>
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,
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<figureCitation box="[774,850,1797,1823]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="10" pageNumber="11">71, 74</figureCitation>
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,
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<figureCitation box="[862,894,1797,1823]" captionStart="Figs 77-83" captionStartId="38.[189,241,1789,1815]" captionTargetBox="[201,1381,776,1726]" captionTargetId="figure@38.[189,1398,759,1748]" captionTargetPageId="38" captionText="Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806808" httpUri="https://zenodo.org/record/3806808/files/figure.png" pageId="10" pageNumber="11">82</figureCitation>
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,
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<figureCitation box="[906,939,1797,1823]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="10" pageNumber="11">84</figureCitation>
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;
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||
<tableCitation box="[951,1038,1797,1823]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="10" pageNumber="11" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="reference_group">
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<paragraph blockId="10.[189,822,1863,1890]" box="[189,822,1863,1890]" pageId="10" pageNumber="11">
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<treatmentCitationGroup box="[189,822,1863,1890]" pageId="10" pageNumber="11">
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<treatmentCitation author="Bruggen A. C. van" box="[189,684,1863,1889]" page="71" pageId="10" pageNumber="11" year="1969">
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<taxonomicName ID-CoL="3HLNG" authority="van Bruggen, 1969: 69 - 71" authorityName="van Bruggen" authorityPageNumber="69 - 71" authorityYear="1969" box="[189,684,1863,1889]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="browni">
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<emphasis box="[189,366,1863,1889]" italics="true" pageId="10" pageNumber="11">Gulella browni</emphasis>
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van
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||
<bibRefCitation author="Bruggen A. C. van" box="[423,684,1863,1889]" pageId="10" pageNumber="11" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen, 1969: 69-71</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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, figs 25-26.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph blockId="10.[189,1399,1929,2027]" pageId="10" pageNumber="11">
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||
<treatmentCitationGroup pageId="10" pageNumber="11">
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||
<emphasis box="[189,388,1929,1955]" italics="true" pageId="10" pageNumber="11">
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||
<taxonomicName ID-CoL="3HLNG" authorityName="van Bruggen" authorityYear="1969" box="[189,366,1929,1955]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="browni">Gulella browni</taxonomicName>
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–
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</emphasis>
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||
van
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<treatmentCitation author="Bruggen A. C. van & Appleton C. C." box="[444,843,1929,1955]" page="31" pageId="10" pageNumber="11" year="1977">
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<bibRefCitation author="Bruggen A. C. van & Appleton C. C." box="[444,843,1929,1955]" pageId="10" pageNumber="11" pagination="3 - 44" refId="ref26218" refString="Bruggen A. C. van & Appleton C. C. 1977. Studies on the ecology and systematics of the terrestrial molluscs of the Lake Sibaya area of Zululand, South Africa. Zoologische Verhandelingen 154: 3 - 44, pls. 1 - 4." type="journal article" year="1977">Bruggen & Appleton 1977: 31-32</bibRefCitation>
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||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation author="Aiken D. W." box="[889,1142,1929,1955]" page="321" pageId="10" pageNumber="11" year="1981">
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||
<bibRefCitation author="Aiken D. W." box="[889,1142,1929,1955]" pageId="10" pageNumber="11" pagination="317 - 324" refId="ref25988" refString="Aiken D. W. 1981. Differentiation of the radula of South African species of Gulella into three types. Journal of Conchology 30 (5): 317 - 324." type="journal article" year="1981">Aiken 1981: 321-323</bibRefCitation>
|
||
</treatmentCitation>
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||
, fig. 5. —
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||
<treatmentCitation author="Richardson C. L." page="60" pageId="10" pageNumber="11" year="1988">
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||
<bibRefCitation author="Richardson C. L." pageId="10" pageNumber="11" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson 1988: 60</bibRefCitation>
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||
</treatmentCitation>
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||
. —
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||
<treatmentCitation author="Verdcourt B." box="[348,597,1965,1991]" page="349" pageId="10" pageNumber="11" year="1990">
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||
<bibRefCitation author="Verdcourt B." box="[348,597,1965,1991]" pageId="10" pageNumber="11" pagination="345 - 354" refId="ref28003" refString="Verdcourt B. 1990. Two Ethiopian streptaxids (Gastropoda: Pulmonata: Streptaxidae). Journal of Conchology 33 (6): 345 - 354." type="journal article" year="1990">Verdcourt 1990: 349</bibRefCitation>
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</treatmentCitation>
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||
. — van
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<treatmentCitation author="Bruggen A. C. van & Van Goethem J. L." box="[698,1119,1965,1991]" page="40" pageId="10" pageNumber="11" year="1999">
|
||
<bibRefCitation author="Bruggen A. C. van & Van Goethem J. L." box="[698,1119,1965,1991]" pageId="10" pageNumber="11" pagination="31 - 45" refId="ref26326" refString="Bruggen A. C. van & Van Goethem J. L. 1999. Dr. William Adam's iconography of Central and West African Gulella species (Gastropoda Pulmonata: Streptaxidae). Part 3: Nine new species from the D. R. Congo. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie 69: 31 - 45." type="journal article" year="1999">Bruggen & Van Goethem 1999: 40</bibRefCitation>
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||
</treatmentCitation>
|
||
. — van
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||
<treatmentCitation author="Bruggen A. C. van" page="233" pageId="10" pageNumber="11" year="2000">
|
||
<bibRefCitation author="Bruggen A. C. van" pageId="10" pageNumber="11" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">Bruggen 2000: 233-234</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation author="Herbert D. & Kilburn D." box="[338,686,2001,2027]" page="212" pageId="10" pageNumber="11" year="2004">
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||
<bibRefCitation author="Herbert D. & Kilburn D." box="[338,686,2001,2027]" pageId="10" pageNumber="11" refId="ref26665" refString="Herbert D. & Kilburn D. 2004. Field guide to the land snails and slugs of eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert & Kilburn 2004: 212</bibRefCitation>
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||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation author="Rowson B. & Lange C. N." box="[733,1054,2001,2027]" page="31" pageId="10" pageNumber="11" year="2007">
|
||
<bibRefCitation author="Rowson B. & Lange C. N." box="[733,1054,2001,2027]" pageId="10" pageNumber="11" pagination="21 - 32" refId="ref27291" refString="Rowson B. & Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">Rowson & Lange 2007: 31</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation author="Muratov I. V." box="[1102,1333,2001,2027]" page="277" pageId="10" pageNumber="11" year="2010">
|
||
<bibRefCitation author="Muratov I. V." box="[1102,1333,2001,2027]" pageId="10" pageNumber="11" pagination="255 - 288" refId="ref26968" refString="Muratov I. V. 2010. Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique. African Invertebrates 51 (2): 255 - 288. http: // dx. doi. org / 10.5733 / afin. 051.0203" type="journal article" year="2010">Muratov 2010: 277</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,952,267,329]" box="[189,952,267,294]" pageId="11" pageNumber="12">
|
||
“a closely related species from Zululand”
|
||
<emphasis box="[683,698,268,293]" italics="true" pageId="11" pageNumber="12">–</emphasis>
|
||
<bibRefCitation author="Verdcourt B." box="[704,947,267,294]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref27966" refString="Verdcourt B. 1985. New taxa of Gulella L. Pfr. And Ptychotrema Morch (Mollusca, Streptaxidae) from Eastern Africa. Journal of Conchology 32 (2): 109 - 121." type="journal article" year="1985">Verdcourt 1985: 119</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,952,267,329]" box="[189,829,302,329]" pageId="11" pageNumber="12">
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<treatmentCitationGroup box="[189,829,302,329]" pageId="11" pageNumber="12">
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<taxonomicName ID-CoL="3HLWD" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[189,397,303,329]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis box="[189,277,303,329]" italics="true" pageId="11" pageNumber="12">Gulella</emphasis>
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||
cf.
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||
<emphasis box="[322,397,303,329]" italics="true" pageId="11" pageNumber="12">radius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis box="[404,419,303,328]" italics="true" pageId="11" pageNumber="12">–</emphasis>
|
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<treatmentCitation author="Tattersfield P. & Seddon M. B. & Ngereza C. & Rowson B." box="[426,737,302,329]" page="52" pageId="11" pageNumber="12" year="2006">
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<bibRefCitation author="Tattersfield P. & Seddon M. B. & Ngereza C. & Rowson B." box="[426,737,302,329]" pageId="11" pageNumber="12" pagination="47 - 60" refId="ref27727" refString="Tattersfield P., Seddon M. B., Ngereza C. & Rowson B. 2006. Elevational variation in diversity and composition of land-snail faunas in a Tanzanian forest. African Journal of Ecology 44: 47 - 60. http: // dx. doi. org / 10.1111 / j. 1365 - 2028.2006.00612. x" type="journal article" year="2006">
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||
Tattersfield
|
||
<emphasis box="[564,623,303,329]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||
2006: 52
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||
</bibRefCitation>
|
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</treatmentCitation>
|
||
, 54, 58.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
|
||
<paragraph blockId="11.[189,492,368,394]" box="[189,492,368,394]" pageId="11" pageNumber="12">
|
||
<heading bold="true" box="[189,492,368,394]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis bold="true" box="[189,492,368,394]" pageId="11" pageNumber="12">
|
||
<typeStatus box="[189,250,368,394]" pageId="11" pageNumber="12">Type</typeStatus>
|
||
material examined
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,415,512]" pageId="11" pageNumber="12">
|
||
<materialsCitation ID-GBIF-Occurrence="2610647309" collectingDate="1966-06-12" collectionCode="RMNH" collectorName="D. S. Brown" country="South Africa" latitude="-27.34" location="KwaZulu-Natal" longLatPrecision="742" longitude="32.71" pageId="11" pageNumber="12" specimenCode="RMNH.MOL.54894" specimenCount="1" specimenCount-adult="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
|
||
<collectingCountry box="[189,411,415,441]" name="South Africa" pageId="11" pageNumber="12">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
<typeStatus box="[429,532,415,441]" pageId="11" pageNumber="12">holotype</typeStatus>
|
||
<specimenCode box="[546,792,415,442]" pageId="11" pageNumber="12">RMNH.MOL.54894</specimenCode>
|
||
:
|
||
<specimenCount box="[810,876,415,441]" pageId="11" pageNumber="12" type="adult">1 ad.</specimenCount>
|
||
, Lake Sibayi (approx.
|
||
<geoCoordinate box="[1170,1267,415,441]" degrees="27.34" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-27.34">27.34°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1286,1386,415,441]" degrees="32.71" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="32.71">32.71°E</geoCoordinate>
|
||
),
|
||
<collectingRegion box="[189,376,450,476]" country="South Africa" name="KwaZulu-Natal" pageId="11" pageNumber="12">KwaZulu-Natal</collectingRegion>
|
||
, dune forest on the east shore, leg.
|
||
<collectorName box="[841,990,450,477]" pageId="11" pageNumber="12">D.S. Brown</collectorName>
|
||
,
|
||
<date box="[1008,1172,450,477]" pageId="11" pageNumber="12" value="1966-06-12">
|
||
<collectingDate box="[1008,1172,450,477]" pageId="11" pageNumber="12" value="1966-06-12">12 Jun. 1966</collectingDate>
|
||
</date>
|
||
(examined digital photograph only).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,505,551,577]" box="[189,505,551,577]" pageId="11" pageNumber="12">
|
||
<heading bold="true" box="[189,505,551,577]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis bold="true" box="[189,505,551,577]" pageId="11" pageNumber="12">Other material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,597,1083]" pageId="11" pageNumber="12">
|
||
<collectingCountry box="[189,343,598,624]" name="Tanzania" pageId="11" pageNumber="12">TANZANIA</collectingCountry>
|
||
:
|
||
<collectionCode box="[364,442,598,624]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z.2003.001.00008: 6 ads, 5 juvs, Mt. Mwanihana Forest Reserve (
|
||
<geoCoordinate box="[1311,1393,598,624]" degrees="7.82" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-7.82">7.82°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[189,288,633,660]" degrees="36.83" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="36.83">36.83°E</geoCoordinate>
|
||
), Udzungwa Mts National Park, Kilombero District, lowland forest at approx.
|
||
<quantity box="[1265,1399,633,659]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="11" pageNumber="12" unit="m" value="750.0" valueMax="900.0" valueMin="600.0">600-900 m</quantity>
|
||
alt., leg.
|
||
<collectionCode box="[297,339,669,695]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="11" pageNumber="12">BR</collectionCode>
|
||
,
|
||
<collectionCode box="[354,389,669,695]" pageId="11" pageNumber="12">PT</collectionCode>
|
||
,
|
||
<collectionCode box="[404,466,668,694]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13544" name="Manchester Banksian Society" pageId="11" pageNumber="12" type="Herbarium">MBS</collectionCode>
|
||
&
|
||
<collectionCode box="[510,570,668,694]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12862" name="Clifton College" pageId="11" pageNumber="12" type="Herbarium">CFN</collectionCode>
|
||
,
|
||
<date box="[585,744,668,695]" pageId="11" pageNumber="12" value="2003-01-29">29 Jan. 2003</date>
|
||
.
|
||
<collectionCode box="[759,837,668,694]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z.2003.001.00009: 4 ads, data as previous but
|
||
<quantity box="[189,263,704,730]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="m" value="700.0">700 m</quantity>
|
||
alt.
|
||
<collectionCode box="[313,392,704,730]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z.1997.007.00003: 5 ads, 2 juvs, Mzelezi Forest Reserve (
|
||
<geoCoordinate box="[1075,1158,704,730]" degrees="8.79" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-8.79">8.79°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1170,1269,703,730]" degrees="36.72" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="36.72">36.72°E</geoCoordinate>
|
||
), Mahenge Mts, Ulanga District, forest on limestone, approx.
|
||
<quantity box="[809,887,739,766]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="11" pageNumber="12" unit="m" value="645.0">645 m</quantity>
|
||
alt., leg.
|
||
<collectionCode box="[1009,1044,739,765]" pageId="11" pageNumber="12">PT</collectionCode>
|
||
,
|
||
<collectionCode box="[1060,1122,739,765]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13544" name="Manchester Banksian Society" pageId="11" pageNumber="12" type="Herbarium">MBS</collectionCode>
|
||
&
|
||
<collectionCode box="[1168,1228,739,765]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12862" name="Clifton College" pageId="11" pageNumber="12" type="Herbarium">CFN</collectionCode>
|
||
,
|
||
<date box="[1243,1394,739,765]" pageId="11" pageNumber="12" value="1997-02-06">6 Feb. 1997</date>
|
||
.
|
||
<collectionCode box="[189,267,774,800]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z.1997.007.00004:
|
||
<specimenCount box="[506,571,774,800]" pageId="11" pageNumber="12" type="generic">1 ad.</specimenCount>
|
||
, data as previous.
|
||
<collectionCode box="[810,889,774,801]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z.1997.007.00005: 2 ads, 6 juvs, data as previous.
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,597,1083]" pageId="11" pageNumber="12">
|
||
<collectingCountry box="[189,396,845,871]" name="Mozambique" pageId="11" pageNumber="12">MOZAMBIQUE</collectingCountry>
|
||
:
|
||
<collectionCode box="[407,485,845,871]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z.2012.040.00001: 2 ads, Pomene Bay,
|
||
<collectingRegion box="[960,1205,845,871]" country="Mozambique" name="Inhambane" pageId="11" pageNumber="12">Inhambane Province</collectingRegion>
|
||
, near lighthouse S of old hotel (
|
||
<geoCoordinate box="[359,487,880,906]" degrees="22.9417" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="-22.9417">22.9417°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[497,627,880,907]" degrees="35.5913" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="35.5913">35.5913°E</geoCoordinate>
|
||
), degraded dune forest, in leaf-litter, leg. D. Herbert,
|
||
<date box="[1240,1355,880,906]" pageId="11" pageNumber="12" value="2002-10">Oct. 2002</date>
|
||
(ex
|
||
<collectionCode box="[189,279,915,942]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="12" type="Museum">NMSA</collectionCode>
|
||
.L5797).
|
||
<collectionCode box="[382,472,915,942]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="12" type="Museum">NMSA</collectionCode>
|
||
.L5797: 2 ads, data as previous.
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,597,1083]" pageId="11" pageNumber="12">
|
||
<collectingCountry box="[189,412,951,977]" name="South Africa" pageId="11" pageNumber="12">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
<collectionCode box="[431,510,951,977]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
|
||
.Z. 2012.040.00002: 3 ads, Kosi Bay mouth,
|
||
<collectingRegion box="[1088,1272,950,976]" country="South Africa" name="KwaZulu-Natal" pageId="11" pageNumber="12">KwaZulu-Natal</collectingRegion>
|
||
(
|
||
<geoCoordinate box="[1296,1394,950,976]" degrees="26.89" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-26.89">26.89°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[189,288,986,1012]" degrees="32.88" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="32.88">32.88°E</geoCoordinate>
|
||
), dune forest, in sandy leaf-litter, leg. D. Herbert & L. Davis, “11-41”,
|
||
<date box="[1107,1257,986,1012]" pageId="11" pageNumber="12" value="2011-10-13">13 Oct. 2011</date>
|
||
(ex
|
||
<collectionCode box="[1306,1395,986,1012]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="12" type="Museum">NMSA</collectionCode>
|
||
. W8517).
|
||
<collectionCode box="[307,422,1021,1047]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="12" type="Museum">NHMUK</collectionCode>
|
||
.20110173: 2 ads, Lake Sibaya area of Tongaland (approx.
|
||
<geoCoordinate box="[1170,1267,1021,1048]" degrees="27.35" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-27.35">27.35°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1286,1385,1021,1048]" degrees="32.67" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="32.67">32.67°E</geoCoordinate>
|
||
), Zululand, S. Africa, leg. C.C. Appleton,
|
||
<date box="[662,784,1057,1083]" pageId="11" pageNumber="12" value="1972-12">Dec. 1972</date>
|
||
-
|
||
<date box="[792,918,1057,1083]" pageId="11" pageNumber="12" value="1973-11">Nov. 1973</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
|
||
<paragraph blockId="11.[189,334,1122,1148]" box="[189,334,1122,1148]" pageId="11" pageNumber="12">
|
||
<heading bold="true" box="[189,334,1122,1148]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis bold="true" box="[189,334,1122,1148]" pageId="11" pageNumber="12">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,1168,1654]" pageId="11" pageNumber="12">
|
||
SHELL (
|
||
<figureCitation box="[275,404,1169,1195]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="11" pageNumber="12">Figs 23-26</figureCitation>
|
||
,
|
||
<figureCitation box="[416,488,1169,1195]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="11" pageNumber="12">47-48</figureCitation>
|
||
). Small (
|
||
<quantity box="[592,760,1169,1195]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="11" pageNumber="12" unit="mm" value="2.9" valueMax="3.2" valueMin="2.6">
|
||
2.60
|
||
<emphasis box="[645,655,1170,1195]" italics="true" pageId="11" pageNumber="12">-</emphasis>
|
||
3.20 mm
|
||
</quantity>
|
||
high x
|
||
<quantity box="[848,1014,1169,1195]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="11" pageNumber="12" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">
|
||
1.60
|
||
<emphasis box="[899,909,1170,1195]" italics="true" pageId="11" pageNumber="12">-</emphasis>
|
||
1.90 mm
|
||
</quantity>
|
||
wide), of 5.5
|
||
<emphasis box="[1170,1180,1170,1195]" italics="true" pageId="11" pageNumber="12">-</emphasis>
|
||
6.0 whorls. Ovateacuminate, spire broadly acuminate (spire angle 66
|
||
<emphasis box="[804,814,1205,1230]" italics="true" pageId="11" pageNumber="12">-</emphasis>
|
||
76°). Apex pointed. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (11
|
||
<emphasis box="[857,867,1240,1265]" italics="true" pageId="11" pageNumber="12">-</emphasis>
|
||
15 per mm on penultimate whorl). Sutures relatively shallow. Umbilicus closed or narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth and another much weaker one corresponding to the basal tooth. Dentition 7-fold to 8-fold, consisting of: one lamella-like parietal tooth and (almost always) one additional parietal denticle; one slab-like palatal tooth, forming a narrow parieto-palatal sinus; two basal denticles; a deep-set columellar baffle, sometimes folded, and one or two shallower columellar denticles. Dentition of 8-fold individuals alternatively recognisable as 7-fold, depending on the interpretation of the columellar and parietal denticles (van
|
||
<bibRefCitation author="Bruggen A. C. van" box="[1106,1279,1486,1513]" pageId="11" pageNumber="12" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen 1969</bibRefCitation>
|
||
). Juvenile shells of
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[293,410,1522,1548]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis box="[293,410,1522,1548]" italics="true" pageId="11" pageNumber="12">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[417,546,1522,1548]" pageId="11" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
|
||
were not known to van
|
||
<bibRefCitation author="Bruggen A. C. van" box="[825,1003,1522,1548]" pageId="11" pageNumber="12" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen (1969</bibRefCitation>
|
||
, 2000).
|
||
<bibRefCitation author="Herbert D. & Kilburn D." box="[1097,1398,1522,1548]" pageId="11" pageNumber="12" refId="ref26665" refString="Herbert D. & Kilburn D. 2004. Field guide to the land snails and slugs of eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert & Kilburn (2004)</bibRefCitation>
|
||
describe them simply as dentate, but with dentition different from that of the adult. Juveniles from southern
|
||
<collectingCountry box="[300,405,1592,1618]" name="Tanzania" pageId="11" pageNumber="12">Tanzania</collectingCountry>
|
||
(
|
||
<figureCitation box="[425,558,1592,1618]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="11" pageNumber="12">Figs 47-48</figureCitation>
|
||
) have dentition similar to, but weaker than, juveniles of
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Tattersfield" baseAuthorityYear="2013" box="[1258,1398,1592,1618]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
|
||
<emphasis box="[1258,1398,1592,1618]" italics="true" pageId="11" pageNumber="12">D. r. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[189,319,1627,1654]" pageId="11" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation box="[334,466,1627,1654]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="11" pageNumber="12">Figs 43-45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,1693,1754]" pageId="11" pageNumber="12">CEPHALOPODIUM. Pale cream, with pink-orange tentacle retractors and a few purple-brown speckles in the mantle.</paragraph>
|
||
<paragraph blockId="11.[189,1399,1794,1856]" pageId="11" pageNumber="12">
|
||
SALIVARY GLANDS (
|
||
<figureCitation box="[401,488,1794,1820]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="11" pageNumber="12">Fig. 71</figureCitation>
|
||
). United, soft, not tumid, elongate, bilobed; each duct leaving at the whitened apex of the lobe and evenly thick throughout.
|
||
</paragraph>
|
||
<paragraph blockId="11.[189,1399,1895,2027]" pageId="11" pageNumber="12">
|
||
RADULA. Described and figured by
|
||
<bibRefCitation author="Aiken D. W." box="[596,753,1895,1921]" pageId="11" pageNumber="12" pagination="317 - 324" refId="ref25988" refString="Aiken D. W. 1981. Differentiation of the radula of South African species of Gulella into three types. Journal of Conchology 30 (5): 317 - 324." type="journal article" year="1981">Aiken (1981)</bibRefCitation>
|
||
as having a unicuspid, ‘heart-shaped’ central tooth and nine laterals in each half-row, of which at least the inner six are tricuspid, diminishing gradually in size laterally. The outer cusps of most teeth are smaller than the inner cusp. According to
|
||
<bibRefCitation author="Aiken D. W." box="[1196,1355,1965,1991]" pageId="11" pageNumber="12" pagination="317 - 324" refId="ref25988" refString="Aiken D. W. 1981. Differentiation of the radula of South African species of Gulella into three types. Journal of Conchology 30 (5): 317 - 324." type="journal article" year="1981">Aiken (1981)</bibRefCitation>
|
||
the teeth are flexible when pressed by a cover slip and there are 47 rows.
|
||
</paragraph>
|
||
<paragraph blockId="12.[189,1399,267,436]" pageId="12" pageNumber="13">
|
||
GENITALIA (
|
||
<figureCitation box="[322,413,267,293]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="12" pageNumber="13">Figs 74</figureCitation>
|
||
,
|
||
<figureCitation box="[426,459,267,293]" captionStart="Figs 77-83" captionStartId="38.[189,241,1789,1815]" captionTargetBox="[201,1381,776,1726]" captionTargetId="figure@38.[189,1398,759,1748]" captionTargetPageId="38" captionText="Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806808" httpUri="https://zenodo.org/record/3806808/files/figure.png" pageId="12" pageNumber="13">82</figureCitation>
|
||
). Upper parts of spermoviduct not recovered. Vas deferens appearing thickened prior to insertion on penis but actually with a very short, broad parallel diverticulum. Penial sheath absent. Interior of penis with weak radial pilasters. Apical, muscular part of penis with a single large hook, associated with a spatulate “scoop” with microscopically serrated tip. Elsewhere in penis, a single longitudinal row of short, simple hooks.
|
||
</paragraph>
|
||
<paragraph blockId="12.[189,421,475,501]" box="[189,421,475,501]" pageId="12" pageNumber="13">
|
||
<heading bold="true" box="[189,421,475,501]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis bold="true" box="[189,421,475,501]" pageId="12" pageNumber="13">Range and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="12.[189,1399,522,655]" pageId="12" pageNumber="13">
|
||
Coast of north-eastern
|
||
<collectingCountry box="[453,606,522,548]" name="South Africa" pageId="12" pageNumber="13">South Africa</collectingCountry>
|
||
: dune forest, coastal forest and bush; also central
|
||
<collectingCountry box="[1177,1337,522,548]" name="Mozambique" pageId="12" pageNumber="13">Mozambique</collectingCountry>
|
||
: high forest in Chiluvo [Xiluvo] hills (
|
||
<geoCoordinate box="[580,677,557,583]" degrees="19.24" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="555" value="-19.24">19.24°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[691,789,557,584]" degrees="34.06" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="555" value="34.06">34.06°E</geoCoordinate>
|
||
) at
|
||
<quantity box="[836,907,557,583]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="12" pageNumber="13" unit="ft" value="700.0">700 ft</quantity>
|
||
(approx.
|
||
<quantity box="[1024,1101,557,584]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="12" pageNumber="13" unit="m" value="213.0">213 m</quantity>
|
||
) elevation (van
|
||
<bibRefCitation author="Bruggen A. C. van" pageId="12" pageNumber="13" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen 1969</bibRefCitation>
|
||
; van
|
||
<bibRefCitation author="Bruggen A. C. van & Appleton C. C." box="[312,627,592,619]" pageId="12" pageNumber="13" pagination="3 - 44" refId="ref26218" refString="Bruggen A. C. van & Appleton C. C. 1977. Studies on the ecology and systematics of the terrestrial molluscs of the Lake Sibaya area of Zululand, South Africa. Zoologische Verhandelingen 154: 3 - 44, pls. 1 - 4." type="journal article" year="1977">Bruggen & Appleton 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Herbert D. & Kilburn D." box="[639,928,592,619]" pageId="12" pageNumber="13" refId="ref26665" refString="Herbert D. & Kilburn D. 2004. Field guide to the land snails and slugs of eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert & Kilburn 2004</bibRefCitation>
|
||
). Also in lowland forest, up to
|
||
<quantity box="[1287,1363,593,619]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="m" value="900.0">900 m</quantity>
|
||
, in southern
|
||
<collectingCountry box="[297,406,628,654]" name="Tanzania" pageId="12" pageNumber="13">Tanzania</collectingCountry>
|
||
, as far north as
|
||
<geoCoordinate box="[590,658,628,654]" degrees="7.8" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="5555" value="-7.8">7.8°S</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph blockId="12.[189,303,694,720]" box="[189,303,694,720]" pageId="12" pageNumber="13">
|
||
<heading bold="true" box="[189,303,694,720]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis bold="true" box="[189,303,694,720]" pageId="12" pageNumber="13">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="12.[189,1399,740,1157]" pageId="12" pageNumber="13">
|
||
We agree with van
|
||
<bibRefCitation author="Bruggen A. C. van" box="[423,611,740,767]" pageId="12" pageNumber="13" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen (1969)</bibRefCitation>
|
||
that the complex dentition of
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[977,1097,740,766]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis box="[977,1097,740,766]" italics="true" pageId="12" pageNumber="13">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1107,1238,740,767]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[1247,1282,740,766]" italics="true" pageId="12" pageNumber="13">s.l.</emphasis>
|
||
separates it from
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[282,396,776,802]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
|
||
<emphasis box="[282,396,776,802]" italics="true" pageId="12" pageNumber="13">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[406,538,776,802]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[548,582,776,802]" italics="true" pageId="12" pageNumber="13">s.l.</emphasis>
|
||
, which never seems to have 8-fold dentition, in particular the extra parietal and basal denticles. The shells of
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[677,795,811,837]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis box="[677,795,811,837]" italics="true" pageId="12" pageNumber="13">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[802,931,811,838]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[938,972,811,837]" italics="true" pageId="12" pageNumber="13">s.l.</emphasis>
|
||
are also much smaller, less elongate (i.e. have a lower height/width ratio), and have a much narrower and more horizontal parieto-palatal sinus than is usual for
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[461,574,882,908]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
|
||
<emphasis box="[461,574,882,908]" italics="true" pageId="12" pageNumber="13">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[583,715,882,909]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[723,758,882,908]" italics="true" pageId="12" pageNumber="13">s.l.</emphasis>
|
||
Although some specimens of
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Tattersfield" baseAuthorityYear="2013" box="[1121,1258,882,908]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
|
||
<emphasis box="[1121,1258,882,908]" italics="true" pageId="12" pageNumber="13">D. r. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1267,1399,882,909]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
approach
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[306,426,918,944]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis box="[306,426,918,944]" italics="true" pageId="12" pageNumber="13">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[436,568,918,944]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[577,612,918,944]" italics="true" pageId="12" pageNumber="13">s.l.</emphasis>
|
||
in one or more of these respects (e.g.
|
||
<figureCitation box="[1080,1263,918,944]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="12" pageNumber="13">Figs 10, 14, 17</figureCitation>
|
||
) all can be readily assigned to one or the other and the
|
||
<typeStatus box="[689,740,954,980]" pageId="12" pageNumber="13">type</typeStatus>
|
||
specimens are morphologically quite different (
|
||
<figureCitation box="[1294,1393,953,980]" captionStart="Figs 1-5" captionStartId="6.[189,242,1930,1956]" captionTargetBox="[363,1200,672,1890]" captionTargetId="figure@6.[363,1200,672,1890]" captionTargetPageId="6" captionText="Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view." figureDoi="http://doi.org/10.5281/zenodo.3806774" httpUri="https://zenodo.org/record/3806774/files/figure.png" pageId="12" pageNumber="13">Figs 1-5</figureCitation>
|
||
,
|
||
<figureCitation box="[189,219,989,1015]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
|
||
,
|
||
<figureCitation box="[235,264,988,1015]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="12" pageNumber="13">52</figureCitation>
|
||
vs.
|
||
<figureCitation box="[320,410,989,1016]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="12" pageNumber="13">Fig. 26</figureCitation>
|
||
). Material intermediate between
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[811,932,989,1015]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis box="[811,932,989,1015]" italics="true" pageId="12" pageNumber="13">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[943,1077,989,1015]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1140,1255,989,1015]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
|
||
<emphasis box="[1140,1255,989,1015]" italics="true" pageId="12" pageNumber="13">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1266,1399,989,1015]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
might be expected in the intervening area of southern
|
||
<collectingCountry box="[825,930,1024,1050]" name="Tanzania" pageId="12" pageNumber="13">Tanzania</collectingCountry>
|
||
or northern
|
||
<collectingCountry box="[1078,1237,1024,1050]" name="Mozambique" pageId="12" pageNumber="13">Mozambique</collectingCountry>
|
||
. However, all the relevant southern Tanzanian material here (e.g. from Mafia and Mahenge) is closer to
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[1278,1398,1060,1086]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis box="[1278,1398,1060,1086]" italics="true" pageId="12" pageNumber="13">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[189,319,1095,1122]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[325,360,1095,1121]" italics="true" pageId="12" pageNumber="13">s.l.</emphasis>
|
||
in dentition and other aspects. A recent survey in northern
|
||
<collectingCountry box="[1050,1205,1095,1121]" name="Mozambique" pageId="12" pageNumber="13">Mozambique</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Muratov I. V." box="[1220,1390,1095,1121]" pageId="12" pageNumber="13" pagination="255 - 288" refId="ref26968" refString="Muratov I. V. 2010. Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique. African Invertebrates 51 (2): 255 - 288. http: // dx. doi. org / 10.5733 / afin. 051.0203" type="journal article" year="2010">
|
||
Muratov
|
||
<date box="[1330,1390,1095,1121]" pageId="12" pageNumber="13" value="2010">2010</date>
|
||
</bibRefCitation>
|
||
) encountered only
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Muratov" baseAuthorityYear="2010" box="[399,530,1130,1156]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="delgada" status="comb. nov.">
|
||
<emphasis box="[399,530,1130,1156]" italics="true" pageId="12" pageNumber="13">D. delgada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[537,667,1130,1157]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |