442 lines
61 KiB
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442 lines
61 KiB
XML
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<mods:title id="E37B25CB49565C98C851F181848AC1D4">Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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<mods:namePart id="4DFD9681D5A11FF1321EFCAABB8ADE3D">Rowson, Ben</mods:namePart>
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<mods:affiliation id="8FD9BDCF4C29139D0094E9C3A3B15C28">Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
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<mods:affiliation id="21974335D54F5E689C80ED403FF2A61E">Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
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<mods:title id="923A3E2C01E68B238986501606BAD664">European Journal of Taxonomy</mods:title>
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<mods:date id="D83DC72D70E1D8F53438A36BE2A87186">2013</mods:date>
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<subSubSection id="85A20680D82CFFCFFE08F9C2F6DBF9BA" box="[428,1160,1602,1629]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="CD07550BD82CFFCFFE08F9C2F6DBF9BA" blockId="8.[428,1160,1602,1666]" box="[428,1160,1602,1629]" pageId="8" pageNumber="9">
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<heading id="964FE267D82CFFCFFE08F9C2F6DBF9BA" box="[428,1160,1602,1629]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<taxonomicName id="0AB82E88D82CFFCFFE08F9C2F1F9F9BA" ID-CoL="33XQP" ID-ENA="940762" authority="(Connolly, 1922)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[428,938,1602,1629]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="radius" status="comb. et stat. nov.">
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<emphasis id="FFCC8919D82CFFCFFE08F9C2F08FF9BA" bold="true" box="[428,732,1602,1629]" italics="true" pageId="8" pageNumber="9">Dadagulella radius calva</emphasis>
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(
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<bibRefCitation id="A92928FAD82CFFCFFD4FF9C2F1F1F9BA" author="Connolly M." box="[747,930,1602,1629]" pageId="8" pageNumber="9" pagination="485 - 517" refId="ref26391" refString="Connolly M. 1922. Notes on African Non-marine Mollusca, with Descriptions of Many New Species. Annals and Magazine of Natural History Series 9 10 (59): 485 - 517, pl. XIV. http: // dx. doi. org / 10.1080 / 00222932208632797" type="journal article" year="1922">Connolly, 1922</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="E4FF3462D82CFFCFFC16F9C2F6DBF9BA" box="[946,1160,1602,1629]" pageId="8" pageNumber="9">comb. et stat. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="85A20680D82CFFCFFD2EF9E7F1FAF965" box="[650,937,1639,1666]" pageId="8" pageNumber="9" type="description">
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<paragraph id="CD07550BD82CFFCFFD2EF9E7F1FAF965" blockId="8.[428,1160,1602,1666]" box="[650,937,1639,1666]" pageId="8" pageNumber="9">
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<figureCitation id="5583498ED82CFFCFFD2EF9E7F0BEF966" box="[650,749,1639,1665]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="8" pageNumber="9">Figs 6-8</figureCitation>
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,
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<figureCitation id="5583498ED82CFFCFFD5EF9E7F149F965" box="[762,794,1639,1666]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="8" pageNumber="9">51</figureCitation>
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,
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<figureCitation id="5583498ED82CFFCFFC82F9E8F114F965" box="[806,839,1640,1666]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="8" pageNumber="9">84</figureCitation>
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;
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||
<tableCitation id="803A60B0D82CFFCFFCF6F9E7F1FAF965" box="[850,937,1639,1666]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="8" pageNumber="9" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
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</paragraph>
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<subSubSection id="85A20680D82CFFCFFF19F92BF605F8CA" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="CD07550BD82CFFCFFF19F92BF14AF921" blockId="8.[189,793,1707,1734]" box="[189,793,1707,1734]" pageId="8" pageNumber="9">
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<treatmentCitation id="4C19731AD82CFFCFFF19F92BF005F921" author="Connolly M." box="[189,598,1707,1734]" page="495" pageId="8" pageNumber="9" year="1922">
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<taxonomicName id="0AB82E88D82CFFCFFF19F92BF005F921" ID-CoL="f070a937-dd91-4e03-82c7-8e36a0a07b9b" authority="Connolly, 1922: 495" authorityName="Connolly" authorityPageNumber="495" authorityYear="1922" box="[189,598,1707,1734]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="calva">
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<emphasis id="FFCC8919D82CFFCFFF19F92BF30FF922" box="[189,348,1707,1733]" italics="true" pageId="8" pageNumber="9">Gulella calva</emphasis>
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<bibRefCitation id="A92928FAD82CFFCFFEC7F92BF005F921" author="Connolly M." box="[355,598,1707,1734]" pageId="8" pageNumber="9" pagination="485 - 517" refId="ref26391" refString="Connolly M. 1922. Notes on African Non-marine Mollusca, with Descriptions of Many New Species. Annals and Magazine of Natural History Series 9 10 (59): 485 - 517, pl. XIV. http: // dx. doi. org / 10.1080 / 00222932208632797" type="journal article" year="1922">Connolly, 1922: 495</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl. XIV, fig. 35.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="CD07550BD82CFFCFFF19F96EF605F8CA" blockId="8.[189,1398,1774,1838]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="EDA87225D82CFFCFFF19F96EF605F8CA" pageId="8" pageNumber="9">
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<emphasis id="FFCC8919D82CFFCFFF19F96EF339F8EF" box="[189,362,1774,1800]" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="0AB82E88D82CFFCFFF19F96EF304F8EF" ID-CoL="f070a937-dd91-4e03-82c7-8e36a0a07b9b" authorityName="Connolly" authorityYear="1922" box="[189,343,1774,1800]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="calva">Gulella calva</taxonomicName>
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–
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</emphasis>
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<treatmentCitation id="4C19731AD82CFFCFFED4F96EF01EF8EF" author="Verdcourt B." box="[368,589,1774,1800]" page="17" pageId="8" pageNumber="9" year="1962">
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<bibRefCitation id="A92928FAD82CFFCFFED4F96EF01EF8EF" author="Verdcourt B." box="[368,589,1774,1800]" pageId="8" pageNumber="9" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">Verdcourt 1962: 17</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="4C19731AD82CFFCFFDFCF96EF09DF8EF" author="Verdcourt B." box="[600,718,1774,1800]" page="234" pageId="8" pageNumber="9" year="1983">
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<bibRefCitation id="A92928FAD82CFFCFFDFCF96EF09DF8EF" author="Verdcourt B." box="[600,718,1774,1800]" pageId="8" pageNumber="9" pagination="200 - 239" refId="ref27933" refString="Verdcourt B. 1983. A list of the non-marine Mollusca of East Africa (Kenya, Uganda, Tanzania, excluding Lake Malawi). Achatina 11: 200 - 239." type="journal article" year="1983">1983: 234</bibRefCitation>
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</treatmentCitation>
|
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. —
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<treatmentCitation id="4C19731AD82CFFCFFD5EF96EF1BAF8EF" author="Richardson C. L." box="[762,1001,1774,1800]" page="62" pageId="8" pageNumber="9" year="1988">
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<bibRefCitation id="A92928FAD82CFFCFFD5EF96EF1BAF8EF" author="Richardson C. L." box="[762,1001,1774,1800]" pageId="8" pageNumber="9" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson 1988: 62</bibRefCitation>
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</treatmentCitation>
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||
. —
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<treatmentCitation id="4C19731AD82CFFCFFBB1F96EF752F8EE" author="Verdcourt B." box="[1045,1281,1774,1801]" page="215" pageId="8" pageNumber="9" year="2000">
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<bibRefCitation id="A92928FAD82CFFCFFBB1F96EF752F8EE" author="Verdcourt B." box="[1045,1281,1774,1801]" pageId="8" pageNumber="9" pagination="219 - 224" refId="ref28100" refString="Verdcourt B. 2000. Molluscs. In: Burgess N. D. & Clarke G. P. (eds) Coastal Forests of Eastern Africa: 219 - 224. IUCN, Gland, Switzerland." type="book chapter" year="2000">Verdcourt 2000: 215</bibRefCitation>
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</treatmentCitation>
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;
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||
<treatmentCitation id="4C19731AD82CFFCFFAAFF96EF721F8EF" author="Verdcourt B." box="[1291,1394,1774,1800]" page="49" pageId="8" pageNumber="9" year="2006">
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<bibRefCitation id="A92928FAD82CFFCFFAAFF96EF721F8EF" author="Verdcourt B." box="[1291,1394,1774,1800]" pageId="8" pageNumber="9" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">2006: 49</bibRefCitation>
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</treatmentCitation>
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.? “
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||
<taxonomicName id="0AB82E88D82CFFCFFF7AF893F3AFF8CA" ID-CoL="3HLWD" authority="(Preston)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[222,508,1811,1837]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="radius">
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<emphasis id="FFCC8919D82CFFCFFF7AF893F3DAF8CA" box="[222,393,1811,1837]" italics="true" pageId="8" pageNumber="9">Gulella radius</emphasis>
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(Preston)
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</taxonomicName>
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var. (K, Mrima Hill Forest)”
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<emphasis id="FFCC8919D82CFFCFFCF3F894F135F8CA" box="[855,870,1812,1837]" italics="true" pageId="8" pageNumber="9">–</emphasis>
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<treatmentCitation id="4C19731AD82CFFCFFCC8F893F601F8CA" author="Verdcourt B." box="[876,1106,1811,1837]" page="22" pageId="8" pageNumber="9" year="1962">
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<bibRefCitation id="A92928FAD82CFFCFFCC8F893F601F8CA" author="Verdcourt B." box="[876,1106,1811,1837]" pageId="8" pageNumber="9" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">Verdcourt 1962: 22</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
|
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</paragraph>
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</subSubSection>
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|
||
<paragraph id="CD07550BD82CFFCFFF19F8D6F3BFF897" blockId="8.[189,492,1878,1904]" box="[189,492,1878,1904]" pageId="8" pageNumber="9">
|
||
<heading id="964FE267D82CFFCFFF19F8D6F3BFF897" bold="true" box="[189,492,1878,1904]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="FFCC8919D82CFFCFFF19F8D6F3BFF897" bold="true" box="[189,492,1878,1904]" pageId="8" pageNumber="9">Type material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="CD07550BD82CFFCFFF19F807F758F80C" blockId="8.[189,1399,1927,2027]" pageId="8" pageNumber="9">
|
||
<collectingCountry id="B5AF159BD82CFFCFFF19F807F376F846" box="[189,293,1927,1953]" name="Kenya" pageId="8" pageNumber="9">KENYA</collectingCountry>
|
||
:
|
||
<typeStatus id="1203EBA9D82CFFCFFE91F807F3F2F846" box="[309,417,1927,1953]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
|
||
(here designated)
|
||
<collectionCode id="ABA9CDCED82CFFCFFD2CF807F0A8F846" box="[648,763,1927,1953]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK</collectionCode>
|
||
.1937.12.30.486:
|
||
<specimenCount id="DBBE9E82D82CFFCFFC6EF807F659F846" box="[970,1034,1927,1953]" pageId="8" pageNumber="9" type="generic">1 ad.</specimenCount>
|
||
, Taru Desert (i.e. a semi-arid area of southeastern Kenya now partly in the Tsavo East National Park, approx.
|
||
<geoCoordinate id="A88C33CCD82CFFCFFBD0F82CF696F821" box="[1140,1221,1964,1990]" degrees="3.40" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="-3.4">3.40°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A88C33CCD82CFFCFFB76F82CF765F821" box="[1234,1334,1964,1990]" degrees="39.00" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="39.0">39.00°E</geoCoordinate>
|
||
), leg. Percival, labelled “type”, and apparently the shell figured in
|
||
<bibRefCitation id="A92928FAD82CFFCFFC27F851F613F80C" author="Connolly M." box="[899,1088,2001,2027]" pageId="8" pageNumber="9" pagination="485 - 517" refId="ref26391" refString="Connolly M. 1922. Notes on African Non-marine Mollusca, with Descriptions of Many New Species. Annals and Magazine of Natural History Series 9 10 (59): 485 - 517, pl. XIV. http: // dx. doi. org / 10.1080 / 00222932208632797" type="journal article" year="1922">Connolly (1922</bibRefCitation>
|
||
: pl. XIV, fig. 35).
|
||
</paragraph>
|
||
<caption id="99C70583D82DFFCEFF19F95AF3BDF80B" ID-DOI="http://doi.org/10.5281/zenodo.3806778" ID-Zenodo-Dep="3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,241,1754,1780]" targetBox="[295,1292,265,1714]" targetPageId="9">
|
||
<paragraph id="CD07550BD82DFFCEFF19F95AF3BDF80B" blockId="9.[189,1399,1754,2028]" pageId="9" pageNumber="10">
|
||
<emphasis id="FFCC8919D82DFFCEFF19F95AF367F913" bold="true" box="[189,308,1754,1780]" pageId="9" pageNumber="10">Figs 6-26.</emphasis>
|
||
Adult shells of
|
||
<taxonomicName id="0AB82E88D82DFFCEFE40F95AF12DF913" authority="(Preston, 1910)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[484,894,1754,1780]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
|
||
<emphasis id="FFCC8919D82DFFCEFE40F95AF094F913" box="[484,711,1754,1780]" italics="true" pageId="9" pageNumber="10">Dadagulella radius</emphasis>
|
||
(
|
||
<bibRefCitation id="A92928FAD82DFFCEFD70F95AF125F913" author="Preston H. B." box="[724,886,1754,1780]" pageId="9" pageNumber="10" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston, 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82DFFCEFC27F95AF656F912" box="[899,1029,1754,1781]" pageId="9" pageNumber="10" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="0AB82E88D82DFFCEFBABF95BF725F912" authority="(van Bruggen, 1969)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[1039,1398,1754,1781]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis id="FFCC8919D82DFFCEFBABF95BF6D1F913" box="[1039,1154,1754,1780]" italics="true" pageId="9" pageNumber="10">D. browni</emphasis>
|
||
(van
|
||
<bibRefCitation id="A92928FAD82DFFCEFB65F95BF73DF912" author="Bruggen A. C. van" box="[1217,1390,1754,1781]" pageId="9" pageNumber="10" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen, 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82DFFCEFF19F97DF36FF8FF" box="[189,316,1789,1816]" pageId="9" pageNumber="10" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and their respective subspecies.
|
||
<emphasis id="FFCC8919D82DFFCEFD15F97DF0BCF8F0" bold="true" box="[689,751,1789,1815]" pageId="9" pageNumber="10">6-20.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFD57F97EF133F8F0" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[755,864,1789,1815]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
|
||
<emphasis id="FFCC8919D82DFFCEFD57F97EF133F8F0" box="[755,864,1789,1815]" italics="true" pageId="9" pageNumber="10">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82DFFCEFCC1F97DF1B7F8FF" box="[869,996,1789,1816]" pageId="9" pageNumber="10" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis id="FFCC8919D82DFFCEFC4DF97DF65FF8F0" box="[1001,1036,1789,1815]" italics="true" pageId="9" pageNumber="10">s.l.</emphasis>
|
||
<emphasis id="FFCC8919D82DFFCEFBB4F97DF66CF8F0" bold="true" box="[1040,1087,1789,1815]" pageId="9" pageNumber="10">6-8.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFBE0F97EF6E3F8F0" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1092,1200,1789,1815]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82DFFCEFBE0F97EF6E3F8F0" box="[1092,1200,1789,1815]" italics="true" pageId="9" pageNumber="10">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="0AB82E88D82DFFCEFB11F97DF355F8DC" authority="(Connolly, 1922)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Connolly" baseAuthorityYear="1922" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="calva">
|
||
<emphasis id="FFCC8919D82DFFCEFB11F97DF6A6F8F0" box="[1205,1269,1789,1815]" italics="true" pageId="9" pageNumber="10">calva</emphasis>
|
||
(
|
||
<bibRefCitation id="A92928FAD82DFFCEFAA6F97DF2AEF8DC" author="Connolly M." pageId="9" pageNumber="10" pagination="485 - 517" refId="ref26391" refString="Connolly M. 1922. Notes on African Non-marine Mollusca, with Descriptions of Many New Species. Annals and Magazine of Natural History Series 9 10 (59): 485 - 517, pl. XIV. http: // dx. doi. org / 10.1080 / 00222932208632797" type="journal article" year="1922">Connolly, 1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="FFCC8919D82DFFCEFEB4F8A1F375F8DC" bold="true" box="[272,294,1825,1851]" pageId="9" pageNumber="10">6.</emphasis>
|
||
Lectotype, Taru Desert.
|
||
<emphasis id="FFCC8919D82DFFCEFDE7F8A1F009F8DC" bold="true" box="[579,602,1825,1851]" pageId="9" pageNumber="10">7.</emphasis>
|
||
Taru Desert near Malindi.
|
||
<emphasis id="FFCC8919D82DFFCEFC34F8A1F1F5F8DC" bold="true" box="[912,934,1825,1851]" pageId="9" pageNumber="10">8.</emphasis>
|
||
Near Mombasa.
|
||
<emphasis id="FFCC8919D82DFFCEFBC8F8A1F6F9F8DC" bold="true" box="[1132,1194,1825,1851]" pageId="9" pageNumber="10">9-20.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFB14F8A2F73DF8DC" authority="radius" authorityName="radius (Preston" authorityYear="1910" baseAuthorityName="radius (Preston" baseAuthorityYear="1910" box="[1200,1390,1825,1851]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="variety" species="radius" variety="radius">
|
||
<emphasis id="FFCC8919D82DFFCEFB14F8A2F73DF8DC" box="[1200,1390,1825,1851]" italics="true" pageId="9" pageNumber="10">D. radius radius</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="FFCC8919D82DFFCEFF19F8C4F28DF8B9" bold="true" box="[189,222,1860,1886]" pageId="9" pageNumber="10">11.</emphasis>
|
||
Lectotype, Shimba Hills.
|
||
<emphasis id="FFCC8919D82DFFCEFD82F8C4F018F8B9" bold="true" box="[550,587,1860,1886]" pageId="9" pageNumber="10">12.</emphasis>
|
||
Gazi.
|
||
<emphasis id="FFCC8919D82DFFCEFD05F8C4F095F8B9" bold="true" box="[673,710,1860,1886]" pageId="9" pageNumber="10">13.</emphasis>
|
||
also Gazi.
|
||
<emphasis id="FFCC8919D82DFFCEFCF3F8C4F12FF8B9" bold="true" box="[855,892,1860,1886]" pageId="9" pageNumber="10">14.</emphasis>
|
||
Diani Beach.
|
||
<emphasis id="FFCC8919D82DFFCEFB95F8C4F605F8B9" bold="true" box="[1073,1110,1860,1886]" pageId="9" pageNumber="10">15.</emphasis>
|
||
Amboni.
|
||
<emphasis id="FFCC8919D82DFFCEFB77F8C4F6ABF8B9" bold="true" box="[1235,1272,1860,1886]" pageId="9" pageNumber="10">16.</emphasis>
|
||
Kimboza.
|
||
<emphasis id="FFCC8919D82DFFCEFF19F8E7F2B1F866" bold="true" box="[189,226,1895,1921]" pageId="9" pageNumber="10">17.</emphasis>
|
||
Mkungwe.
|
||
<emphasis id="FFCC8919D82DFFCEFED0F8E7F3CAF866" bold="true" box="[372,409,1895,1921]" pageId="9" pageNumber="10">18.</emphasis>
|
||
Pugu.
|
||
<emphasis id="FFCC8919D82DFFCEFE55F8E7F045F866" bold="true" box="[497,534,1895,1921]" pageId="9" pageNumber="10">19.</emphasis>
|
||
Mbudya I.
|
||
<emphasis id="FFCC8919D82DFFCEFD03F8E7F09FF866" bold="true" box="[679,716,1895,1921]" pageId="9" pageNumber="10">20.</emphasis>
|
||
Jozani.
|
||
<emphasis id="FFCC8919D82DFFCEFC97F8E7F12CF866" bold="true" box="[819,895,1895,1921]" pageId="9" pageNumber="10">21-26.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFC2DF8E8F652F866" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[905,1025,1895,1921]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis id="FFCC8919D82DFFCEFC2DF8E8F652F866" box="[905,1025,1895,1921]" italics="true" pageId="9" pageNumber="10">D. browni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82DFFCEFBA8F8E7F6C3F865" box="[1036,1168,1895,1922]" pageId="9" pageNumber="10" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis id="FFCC8919D82DFFCEFB3EF8E7F6EEF866" box="[1178,1213,1895,1921]" italics="true" pageId="9" pageNumber="10">s.l.</emphasis>
|
||
:
|
||
<emphasis id="FFCC8919D82DFFCEFB6BF8E7F6A7F866" bold="true" box="[1231,1268,1895,1921]" pageId="9" pageNumber="10">21.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFB5AF8E8F379F843" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="subSpecies" species="browni" subSpecies="mafiensis">
|
||
<emphasis id="FFCC8919D82DFFCEFB5AF8E8F379F843" italics="true" pageId="9" pageNumber="10">D. browni mafiensis</emphasis>
|
||
</taxonomicName>
|
||
subsp. nov., holotype, Mlula.
|
||
<emphasis id="FFCC8919D82DFFCEFD2FF80AF0FCF843" bold="true" box="[651,687,1930,1956]" pageId="9" pageNumber="10">22.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFD13F80BF19CF843" box="[695,975,1930,1956]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="subSpecies" species="browni" subSpecies="semulikiensis">
|
||
<emphasis id="FFCC8919D82DFFCEFD13F80BF19CF843" box="[695,975,1930,1956]" italics="true" pageId="9" pageNumber="10">D. browni semulikiensis</emphasis>
|
||
</taxonomicName>
|
||
subsp. nov., holotype, Semuliki NP.
|
||
<emphasis id="FFCC8919D82DFFCEFF19F82DF35AF820" bold="true" box="[189,265,1965,1991]" pageId="9" pageNumber="10">23-26.</emphasis>
|
||
<taxonomicName id="0AB82E88D82DFFCEFEB7F82EF3BBF820" authority="browni" authorityName="browni (van Bruggen" authorityYear="1969" baseAuthorityName="browni (van Bruggen" baseAuthorityYear="1969" box="[275,488,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="subSpecies" species="browni" subSpecies="browni">
|
||
<emphasis id="FFCC8919D82DFFCEFEB7F82EF3BBF820" box="[275,488,1965,1991]" italics="true" pageId="9" pageNumber="10">D. browni browni</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="FFCC8919D82DFFCEFE5EF82DF04CF820" bold="true" box="[506,543,1965,1991]" pageId="9" pageNumber="10">23.</emphasis>
|
||
Mwanihana.
|
||
<emphasis id="FFCC8919D82DFFCEFD61F82DF0B9F820" bold="true" box="[709,746,1965,1991]" pageId="9" pageNumber="10">24.</emphasis>
|
||
Mzelezi.
|
||
<emphasis id="FFCC8919D82DFFCEFCC7F82DF1DBF820" bold="true" box="[867,904,1965,1991]" pageId="9" pageNumber="10">25.</emphasis>
|
||
Pomene Bay.
|
||
<emphasis id="FFCC8919D82DFFCEFB9DF82DF60CF820" bold="true" box="[1081,1119,1965,1991]" pageId="9" pageNumber="10">26.</emphasis>
|
||
Holotype, Lake Sibayi (after van
|
||
<bibRefCitation id="A92928FAD82DFFCEFE92F851F3B2F80B" author="Bruggen A. C. van" box="[310,481,2001,2028]" pageId="9" pageNumber="10" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen 1969</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CD07550BD82EFFCDFF19FE8BF3AAFEC2" blockId="10.[189,505,267,293]" box="[189,505,267,293]" pageId="10" pageNumber="11">
|
||
<heading id="964FE267D82EFFCDFF19FE8BF3AAFEC2" bold="true" box="[189,505,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="FFCC8919D82EFFCDFF19FE8BF3AAFEC2" bold="true" box="[189,505,267,293]" pageId="10" pageNumber="11">Other material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="CD07550BD82EFFCDFF19FEBAF724FE27" blockId="10.[189,1399,314,449]" pageId="10" pageNumber="11">
|
||
<collectingCountry id="B5AF159BD82EFFCDFF19FEBAF377FEB3" box="[189,292,314,340]" name="Kenya" pageId="10" pageNumber="11">KENYA</collectingCountry>
|
||
:
|
||
<collectionCode id="ABA9CDCED82EFFCDFE94FEBAF32FFEB2" box="[304,380,314,341]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="11">NMW</collectionCode>
|
||
.1955.158.25052: 5 ads, “ca. lat. 3°5’, long. 39°27’”, standing as “
|
||
<taxonomicName id="0AB82E88D82EFFCDFB2EFEBAF6AFFEB3" authority="Co." authorityName="Connolly" authorityYear="1922" box="[1162,1276,314,340]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFB2EFEBAF69AFEB3" box="[1162,1225,314,340]" italics="true" pageId="10" pageNumber="11">calva</emphasis>
|
||
Co.
|
||
</taxonomicName>
|
||
” and later labelled “
|
||
<taxonomicName id="0AB82E88D82EFFCDFE8FFEDEF3CFFE9F" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[299,412,350,376]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
cf.
|
||
<emphasis id="FFCC8919D82EFFCDFEF6FEDEF3CFFE9F" box="[338,412,350,376]" italics="true" pageId="10" pageNumber="11">radius</emphasis>
|
||
</taxonomicName>
|
||
” by B. Verdcourt. Assuming that the latitude refers to a point south rather than north of the equator, the coordinates correspond to a point in the Taru Desert, ±
|
||
<quantity id="0A40F8EED82EFFCDFBE2FE02F6C1FE7B" box="[1094,1170,386,413]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="11" unit="km" value="75.0">75 km</quantity>
|
||
inland of Malindi.
|
||
<collectionCode id="ABA9CDCED82EFFCDFF19FE26F34AFE27" box="[189,281,422,448]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="11" type="Museum">RMNH</collectionCode>
|
||
.MOL.288087:
|
||
<specimenCount id="DBBE9E82D82EFFCDFE6DFE26F050FE27" box="[457,515,422,448]" pageId="10" pageNumber="11" type="generic">1 ad.</specimenCount>
|
||
, near
|
||
<collectingRegion id="0F7C9BE9D82EFFCDFDE3FE26F0E9FE27" box="[583,698,422,448]" country="Kenya" name="Mombasa" pageId="10" pageNumber="11">Mombasa</collectingRegion>
|
||
(approx.
|
||
<geoCoordinate id="A88C33CCD82EFFCDFC8CFE26F125FE27" box="[808,886,422,448]" degrees="4.04" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="555" value="-4.04">4.04°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A88C33CCD82EFFCDFC2FFE26F1BDFE26" box="[907,1006,422,449]" degrees="39.66" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="555" value="39.66">39.66°E</geoCoordinate>
|
||
), amongst river debris,
|
||
<date id="B90673CBD82EFFCDFB58FE26F720FE27" box="[1276,1395,422,448]" pageId="10" pageNumber="11" value="1987-09">Sep. 1987</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="85A20680D82EFFCDFF19FE68F19FFCB7" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="CD07550BD82EFFCDFF19FE68F31DFDE5" blockId="10.[189,334,488,514]" box="[189,334,488,514]" pageId="10" pageNumber="11">
|
||
<heading id="964FE267D82EFFCDFF19FE68F31DFDE5" bold="true" box="[189,334,488,514]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="FFCC8919D82EFFCDFF19FE68F31DFDE5" bold="true" box="[189,334,488,514]" pageId="10" pageNumber="11">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="CD07550BD82EFFCDFF19FD97F19FFCB7" blockId="10.[189,1399,535,849]" pageId="10" pageNumber="11">
|
||
SHELL (
|
||
<figureCitation id="5583498ED82EFFCDFEB3FD97F32CFDD6" box="[279,383,535,561]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="10" pageNumber="11">Figs 6-8</figureCitation>
|
||
,
|
||
<figureCitation id="5583498ED82EFFCDFE34FD97F3E2FDD5" box="[400,433,535,562]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="10" pageNumber="11">51</figureCitation>
|
||
). Large (4.00
|
||
<emphasis id="FFCC8919D82EFFCDFDF3FD98F032FDD6" box="[599,609,536,561]" italics="true" pageId="10" pageNumber="11">-</emphasis>
|
||
<quantity id="0A40F8EED82EFFCDFDC5FD97F09DFDD5" box="[609,718,535,562]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="10" pageNumber="11" unit="mm" value="4.6">4.60 mm</quantity>
|
||
high x
|
||
<quantity id="0A40F8EED82EFFCDFC90FD97F18CFDD5" box="[820,991,535,562]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="10" pageNumber="11" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">
|
||
2.10
|
||
<emphasis id="FFCC8919D82EFFCDFCCCFD98F121FDD6" box="[872,882,536,561]" italics="true" pageId="10" pageNumber="11">-</emphasis>
|
||
2.50 mm
|
||
</quantity>
|
||
wide), of 6.5
|
||
<emphasis id="FFCC8919D82EFFCDFB2DFD98F6C0FDD6" box="[1161,1171,536,561]" italics="true" pageId="10" pageNumber="11">-</emphasis>
|
||
8.0 whorls. Ovateacuminate, but more columnar than other
|
||
<taxonomicName id="0AB82E88D82EFFCDFD14FDBBF110FDB2" authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[688,835,571,597]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="FFCC8919D82EFFCDFD14FDBBF110FDB2" box="[688,835,571,597]" italics="true" pageId="10" pageNumber="11">Dadagulella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFCE8FDBBF1E8FDB2" box="[844,955,571,597]" pageId="10" pageNumber="11" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, spire narrowly to broadly acuminate (spire angle 49
|
||
<emphasis id="FFCC8919D82EFFCDFE24FDE0F3D9FD9E" box="[384,394,608,633]" italics="true" pageId="10" pageNumber="11">-</emphasis>
|
||
62°). Apex pointed. Embryonic whorls smoothly granulate. Later whorls with characteristically coarse, few and widely spaced ribs (5
|
||
<emphasis id="FFCC8919D82EFFCDFCC6FD04F13FFD7A" box="[866,876,644,669]" italics="true" pageId="10" pageNumber="11">-</emphasis>
|
||
10 per mm on penultimate whorl). Sutures deep. Umbilicus closed or nearly so. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition characteristically simple, 4-fold to 5-fold, consisting of: one lamella-like parietal tooth; one slab-like palatal tooth without parieto-palatal sinus; one basal denticle; and one deep-set columellar baffle, always visible, sometimes with a very slight, shallower columellar tooth. Juvenile shells and anatomy unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="85A20680D82EFFCDFF19FCF8F6D5FBB6" pageId="10" pageNumber="11" type="multiple">
|
||
<paragraph id="CD07550BD82EFFCDFF19FCF8F3F6FC75" blockId="10.[189,421,888,914]" box="[189,421,888,914]" pageId="10" pageNumber="11">
|
||
<heading id="964FE267D82EFFCDFF19FCF8F3F6FC75" bold="true" box="[189,421,888,914]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="FFCC8919D82EFFCDFF19FCF8F3F6FC75" bold="true" box="[189,421,888,914]" pageId="10" pageNumber="11">Range and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="CD07550BD82EFFCDFF19FC27F6D5FBB6" blockId="10.[189,1399,935,1106]" pageId="10" pageNumber="11">
|
||
Lowlands of extreme southeastern
|
||
<collectingCountry id="B5AF159BD82EFFCDFDF5FC28F0F2FC25" box="[593,673,936,962]" name="Kenya" pageId="10" pageNumber="11">Kenya</collectingCountry>
|
||
, including Taru Desert. This subspecies has also been recorded from
|
||
<collectingCountry id="B5AF159BD82EFFCDFEA6FC4BF30CFC02" box="[258,351,971,997]" name="Kenya" pageId="10" pageNumber="11">Malindi</collectingCountry>
|
||
(
|
||
<bibRefCitation id="A92928FAD82EFFCDFED6FC4BF032FC02" author="Verdcourt B." box="[370,609,971,997]" pageId="10" pageNumber="11" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">Verdcourt 1962: 17</bibRefCitation>
|
||
) and Mrima Hill (
|
||
<geoCoordinate id="A88C33CCD82EFFCDFCEEFC4BF1CFFC02" box="[842,924,971,997]" degrees="4.48" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="555" value="-4.48">4.48°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A88C33CCD82EFFCDFC09FC4BF65CFC01" box="[941,1039,971,998]" degrees="39.25" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="555" value="39.25">39.25°E</geoCoordinate>
|
||
) (
|
||
<bibRefCitation id="A92928FAD82EFFCDFB8EFC4BF6BFFC01" author="Verdcourt B." box="[1066,1260,971,998]" pageId="10" pageNumber="11" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">Verdcourt 2006</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="A92928FAD82EFFCDFAA7FC4BF358FBEE" author="Verdcourt B." pageId="10" pageNumber="11" pagination="219 - 224" refId="ref28100" refString="Verdcourt B. 2000. Molluscs. In: Burgess N. D. & Clarke G. P. (eds) Coastal Forests of Eastern Africa: 219 - 224. IUCN, Gland, Switzerland." type="book chapter" year="2000">Verdcourt (2000)</bibRefCitation>
|
||
suggested the habitat was “bushland to forest”, in contrast to “woodland/forest” for
|
||
<taxonomicName id="0AB82E88D82EFFCDFB54FC70F725FBEE" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Tattersfield" baseAuthorityYear="2013" box="[1264,1398,1007,1033]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFB54FC70F725FBEE" box="[1264,1398,1007,1033]" italics="true" pageId="10" pageNumber="11">D. r. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFF19FB93F36CFBC9" box="[189,319,1043,1070]" pageId="10" pageNumber="11" rank="species">comb. nov.</taxonomicNameLabel>
|
||
Shells of both
|
||
<taxonomicName id="0AB82E88D82EFFCDFE54FB94F025FBCA" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Tattersfield" baseAuthorityYear="2013" box="[496,630,1043,1069]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFE54FB94F025FBCA" box="[496,630,1043,1069]" italics="true" pageId="10" pageNumber="11">D. r. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFDD9FB93F0ACFBC9" box="[637,767,1043,1070]" pageId="10" pageNumber="11" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="0AB82E88D82EFFCDFC9CFB94F1E0FBCA" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Connolly" baseAuthorityYear="1922" box="[824,947,1043,1069]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" status="comb. et stat. nov." subSpecies="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFC9CFB94F1E0FBCA" box="[824,947,1043,1069]" italics="true" pageId="10" pageNumber="11">D. r. calva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFC1EFB93F6DCFBC9" box="[954,1167,1043,1070]" pageId="10" pageNumber="11">comb. et stat. nov.</taxonomicNameLabel>
|
||
have been found in river debris near
|
||
<collectingCountry id="B5AF159BD82EFFCDFE21FBB7F3AFFBB6" box="[389,508,1079,1105]" name="Kenya" pageId="10" pageNumber="11">Mombasa</collectingCountry>
|
||
, but may have been washed in from different localities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="85A20680D82EFFCDFF19FBF9F151F95E" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="CD07550BD82EFFCDFF19FBF9F37CFB74" blockId="10.[189,303,1145,1171]" box="[189,303,1145,1171]" pageId="10" pageNumber="11">
|
||
<heading id="964FE267D82EFFCDFF19FBF9F37CFB74" bold="true" box="[189,303,1145,1171]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="FFCC8919D82EFFCDFF19FBF9F37CFB74" bold="true" box="[189,303,1145,1171]" pageId="10" pageNumber="11">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="CD07550BD82EFFCDFF19FB28F151F95E" blockId="10.[189,1399,1192,1721]" pageId="10" pageNumber="11">
|
||
The
|
||
<typeStatus id="1203EBA9D82EFFCDFF57FB28F30CFB25" box="[243,351,1192,1218]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
|
||
and similar material of this subspecies (
|
||
<figureCitation id="5583498ED82EFFCDFC9EFB28F1F3FB25" box="[826,928,1192,1218]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="10" pageNumber="11">Figs 6-8</figureCitation>
|
||
,
|
||
<figureCitation id="5583498ED82EFFCDFC09FB28F198FB24" box="[941,971,1192,1219]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="10" pageNumber="11">51</figureCitation>
|
||
) differ from
|
||
<taxonomicName id="0AB82E88D82EFFCDFBC2FB29F6BEFB25" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Tattersfield" baseAuthorityYear="2013" box="[1126,1261,1192,1218]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFBC2FB29F6BEFB25" box="[1126,1261,1192,1218]" italics="true" pageId="10" pageNumber="11">D. r. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFB51FB28F724FB24" box="[1269,1399,1192,1219]" pageId="10" pageNumber="11" rank="species">comb. nov.</taxonomicNameLabel>
|
||
in the more widely spaced ribs, more columnar shape and simpler dentition, although some specimens seem to show intermediate characters. Verdcourt evidently had difficulty separating them. Although
|
||
<bibRefCitation id="A92928FAD82EFFCDFF19FA94F3E7FAC9" author="Verdcourt B." box="[189,436,1300,1326]" pageId="10" pageNumber="11" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">Verdcourt (1962: 17)</bibRefCitation>
|
||
suggested
|
||
<taxonomicName id="0AB82E88D82EFFCDFD9DFA94F0CEFAC9" authorityName="Connolly" authorityYear="1922" box="[569,669,1300,1326]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFD9DFA94F0CEFAC9" box="[569,669,1300,1326]" italics="true" pageId="10" pageNumber="11">G. calva</emphasis>
|
||
</taxonomicName>
|
||
was “scarcely more than a variety of
|
||
<taxonomicName id="0AB82E88D82EFFCDFBFAFA94F69DFAC9" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1118,1230,1300,1326]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFBFAFA94F69DFAC9" box="[1118,1230,1300,1326]" italics="true" pageId="10" pageNumber="11">G. radius</emphasis>
|
||
</taxonomicName>
|
||
”, it keyed out in a separate part of his key, with
|
||
<taxonomicName id="0AB82E88D82EFFCDFDF1FAB8F0F3FAB5" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[597,672,1336,1362]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFDF1FAB8F0F3FAB5" box="[597,672,1336,1362]" italics="true" pageId="10" pageNumber="11">radius</emphasis>
|
||
</taxonomicName>
|
||
appearing in two other places (1962: 13, 22). The second of these (p. 22) was listed as “
|
||
<taxonomicName id="0AB82E88D82EFFCDFDA4FADCF14CFA91" authority="(Preston)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[512,799,1372,1398]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFDA4FADCF0F8FA91" box="[512,683,1372,1398]" italics="true" pageId="10" pageNumber="11">Gulella radius</emphasis>
|
||
(Preston)
|
||
</taxonomicName>
|
||
var. (K [
|
||
<collectingCountry id="B5AF159BD82EFFCDFC2DFADBF18DFA91" box="[905,990,1371,1398]" name="Kenya" pageId="10" pageNumber="11">Kenya</collectingCountry>
|
||
], Mrima Hill Forest)”. Verdcourt’s measurements, description, and range of apertural tooth formulas allow attribution of this taxon to
|
||
<taxonomicName id="0AB82E88D82EFFCDFAE7FA00F2AEFA5A" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Connolly" baseAuthorityYear="1922" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" status="comb. et stat. nov" subSpecies="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFAE7FA00F2AEFA5A" italics="true" pageId="10" pageNumber="11">D. r. calva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFEA0FA23F387FA59" box="[260,468,1443,1470]" pageId="10" pageNumber="11">comb. et stat. nov</taxonomicNameLabel>
|
||
.. In his later checklists,
|
||
<bibRefCitation id="A92928FAD82EFFCDFD49FA23F1B8FA59" author="Verdcourt B." box="[749,1003,1443,1470]" pageId="10" pageNumber="11" pagination="200 - 239" refId="ref27933" refString="Verdcourt B. 1983. A list of the non-marine Mollusca of East Africa (Kenya, Uganda, Tanzania, excluding Lake Malawi). Achatina 11: 200 - 239." type="journal article" year="1983">Verdcourt (1983: 234</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A92928FAD82EFFCDFC53FA23F6D5FA59" author="Verdcourt B." box="[1015,1158,1443,1470]" pageId="10" pageNumber="11" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">2006: 48-49</bibRefCitation>
|
||
) maintained the two as separate species, and gave Mrima Hill as a locality for
|
||
<taxonomicName id="0AB82E88D82EFFCDFCC0FA47F1F7FA06" authorityName="Connolly" authorityYear="1922" box="[868,932,1479,1505]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFCC0FA47F1F7FA06" box="[868,932,1479,1505]" italics="true" pageId="10" pageNumber="11">calva</emphasis>
|
||
</taxonomicName>
|
||
but not
|
||
<taxonomicName id="0AB82E88D82EFFCDFBA1FA47F603FA06" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1029,1104,1479,1505]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFBA1FA47F603FA06" box="[1029,1104,1479,1505]" italics="true" pageId="10" pageNumber="11">radius</emphasis>
|
||
</taxonomicName>
|
||
. He also kept them apart in his list of coastal molluscs (2000). Thus it appears likely that he later decided that (1962: 22) “
|
||
<taxonomicName id="0AB82E88D82EFFCDFA8FFA6BF725F9E2" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1323,1398,1515,1541]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFA8FFA6BF725F9E2" box="[1323,1398,1515,1541]" italics="true" pageId="10" pageNumber="11">radius</emphasis>
|
||
</taxonomicName>
|
||
var.” belonged to what we treat as
|
||
<taxonomicName id="0AB82E88D82EFFCDFDF6F990F09EF9CE" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Connolly" baseAuthorityYear="1922" box="[594,717,1551,1577]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" status="comb. et stat. nov." subSpecies="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFDF6F990F09EF9CE" box="[594,717,1551,1577]" italics="true" pageId="10" pageNumber="11">D. r. calva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFD77F98FF1FAF9CD" box="[723,937,1551,1578]" pageId="10" pageNumber="11">comb. et stat. nov.</taxonomicNameLabel>
|
||
and not to what we treat as
|
||
<taxonomicName id="0AB82E88D82EFFCDFB55F990F725F9CE" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Tattersfield" baseAuthorityYear="2013" box="[1265,1398,1551,1577]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFB55F990F725F9CE" box="[1265,1398,1551,1577]" italics="true" pageId="10" pageNumber="11">D. r. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4FF3462D82EFFCDFF19F9B3F312F9A9" box="[189,321,1587,1614]" pageId="10" pageNumber="11" rank="species">comb. nov.</taxonomicNameLabel>
|
||
It also appears that he decided not to separate the two taxa. We too found this a difficult decision to take, owing to the morphological, habitat and distributional differences, which may of course be interrelated. To acknowledge this difficulty, and the potential difficulties in assigning future material to either, we rank
|
||
<taxonomicName id="0AB82E88D82EFFCDFE34F91FF383F95E" authorityName="Connolly" authorityYear="1922" box="[400,464,1695,1721]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="calva">
|
||
<emphasis id="FFCC8919D82EFFCDFE34F91FF383F95E" box="[400,464,1695,1721]" italics="true" pageId="10" pageNumber="11">calva</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName id="0AB82E88D82EFFCDFD14F91FF0A8F95E" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[688,763,1695,1721]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis id="FFCC8919D82EFFCDFD14F91FF0A8F95E" box="[688,763,1695,1721]" italics="true" pageId="10" pageNumber="11">radius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |