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<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben &amp; Tattersfield, Peter" docDate="2013" docId="4511E41DD828FFCAFD84FB2CF1CEFD01" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella browni subsp. mafiensis Rowson &amp; Tattersfield 2013, subsp. nov." docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="13" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rowson, Ben</mods:namePart>
<mods:nameIdentifier type="LSID">05D46259-DA45-45A4-9127-5AF361D566A7</mods:nameIdentifier>
<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tattersfield, Peter</mods:namePart>
<mods:nameIdentifier type="LSID">8C55642E-B18A-461B-9BF9-31CB6DB7A070</mods:nameIdentifier>
<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-02-04</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>37</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>46</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.5852/ejt.2013.37</mods:identifier>
<mods:identifier type="GBIF-Dataset">9682b45b-5e22-4965-a293-e203660e0ed3</mods:identifier>
<mods:identifier type="ISSN">2118-9773</mods:identifier>
<mods:identifier type="Zenodo-Dep">3806770</mods:identifier>
<mods:identifier type="ZooBank">ADF6394E-77B5-4309-87FE-4097FDE0A3FD</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3815743" ID-GBIF-Taxon="163971601" ID-Zenodo-Dep="3815743" LSID="urn:lsid:plazi:treatment:4511E41DD828FFCAFD84FB2CF1CEFD01" httpUri="http://treatment.plazi.org/id/4511E41DD828FFCAFD84FB2CF1CEFD01" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection box="[544,1044,1196,1223]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph blockId="12.[544,1044,1196,1258]" box="[544,1044,1196,1223]" pageId="12" pageNumber="13">
<heading box="[544,1044,1196,1223]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName ID-CoL="5DZBT" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[544,905,1196,1223]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="browni" status="subsp. nov." subSpecies="mafiensis">
<emphasis bold="true" box="[544,905,1196,1223]" italics="true" pageId="12" pageNumber="13">Dadagulella browni mafiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[913,1044,1196,1223]" pageId="12" pageNumber="13" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[655,932,1231,1258]" pageId="12" pageNumber="13" type="description">
<paragraph blockId="12.[544,1044,1196,1258]" box="[655,932,1231,1258]" pageId="12" pageNumber="13">
<figureCitation box="[655,745,1232,1258]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="12" pageNumber="13">Figs 21</figureCitation>
,
<figureCitation box="[757,789,1231,1258]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="12" pageNumber="13">54</figureCitation>
,
<figureCitation box="[801,834,1232,1258]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="12" pageNumber="13">84</figureCitation>
;
<tableCitation box="[845,932,1232,1258]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="12" pageNumber="13" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection box="[189,849,1298,1371]" pageId="12" pageNumber="13" type="etymology">
<paragraph blockId="12.[189,324,1298,1324]" box="[189,324,1298,1324]" pageId="12" pageNumber="13">
<heading bold="true" box="[189,324,1298,1324]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis bold="true" box="[189,324,1298,1324]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[189,849,1344,1371]" box="[189,849,1344,1371]" pageId="12" pageNumber="13">From Mafia Island, with the final a elided for euphony.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="materials_examined">
<paragraph blockId="12.[189,492,1410,1436]" box="[189,492,1410,1436]" pageId="12" pageNumber="13">
<heading bold="true" box="[189,492,1410,1436]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis bold="true" box="[189,492,1410,1436]" pageId="12" pageNumber="13">
<typeStatus box="[189,250,1410,1436]" pageId="12" pageNumber="13">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[189,1398,1456,1553]" pageId="12" pageNumber="13">
<collectionCode box="[189,304,1457,1483]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="12" pageNumber="13" type="Museum">
<collectionCodeEnum box="[189,304,1457,1483]" pageId="12" pageNumber="13">NHMUK</collectionCodeEnum>
</collectionCode>
.20110174:
<specimenCount box="[443,507,1457,1483]" pageId="12" pageNumber="13" type="generic">1 ad.</specimenCount>
, Mlula, Mafia
<collectionCode box="[694,706,1457,1483]" country="Romania" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="12" pageNumber="13" type="Herbarium">I</collectionCode>
. (approx.
<geoCoordinate box="[834,915,1457,1483]" degrees="7.87" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="555" value="-7.87">
7.87°
<collectionCode box="[900,915,1457,1483]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="12" pageNumber="13" type="Herbarium">S</collectionCode>
</geoCoordinate>
,
<geoCoordinate box="[932,1032,1456,1483]" degrees="39.76" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="555" value="39.76">
39.76°
<collectionCode box="[1013,1032,1456,1483]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="12" pageNumber="13">E</collectionCode>
</geoCoordinate>
), evergreen coastal thicket on clay loam coral rag, leg. Frontier
<collectingCountry box="[598,707,1492,1518]" name="Tanzania" pageId="12" pageNumber="13">Tanzania</collectingCountry>
,
<date box="[721,907,1492,1518]" pageId="12" pageNumber="13" value="1990-10" valueMax="1990-11-31" valueMin="1990-10-00">
Oct.-
<date box="[778,907,1492,1518]" month="11" monthName="Nov" pageId="12" pageNumber="13" year="1990">
Nov.
<date box="[846,907,1492,1518]" pageId="12" pageNumber="13" value="1990" year="1990">1990</date>
</date>
</date>
, labelled “
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1038,1212,1492,1518]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="radius">
<emphasis box="[1038,1212,1492,1518]" italics="true" pageId="12" pageNumber="13">
<taxonomicName box="[1038,1126,1492,1518]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Gulella</taxonomicName>
radius
</emphasis>
</taxonomicName>
aggreg.” by
<collectionCode box="[1372,1395,1493,1519]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="12" pageNumber="13" type="Herbarium">B</collectionCode>
. Verdcourt.
</paragraph>
<paragraph blockId="12.[189,505,1593,1619]" box="[189,505,1593,1619]" pageId="12" pageNumber="13">
<heading bold="true" box="[189,505,1593,1619]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis bold="true" box="[189,505,1593,1619]" pageId="12" pageNumber="13">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[189,260,1641,1667]" box="[189,260,1641,1667]" pageId="12" pageNumber="13">None.</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph blockId="12.[189,334,1705,1731]" box="[189,334,1705,1731]" pageId="12" pageNumber="13">
<heading bold="true" box="[189,334,1705,1731]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis bold="true" box="[189,334,1705,1731]" pageId="12" pageNumber="13">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[189,1399,1752,2027]" pageId="12" pageNumber="13">
<collectionCode box="[189,257,1753,1779]" pageId="12" pageNumber="13">
<collectionCodeEnum box="[189,257,1753,1779]" pageId="12" pageNumber="13">SHELL</collectionCodeEnum>
</collectionCode>
(
<figureCitation box="[278,371,1752,1779]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="12" pageNumber="13">Figs 21</figureCitation>
,
<figureCitation box="[387,420,1752,1779]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="12" pageNumber="13">54</figureCitation>
). Small (
<quantity box="[531,641,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="13" unit="mm" value="3.3">3.30 mm</quantity>
high x
<quantity box="[739,848,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.80 mm</quantity>
wide), of 6.0 whorls. Ovate-acuminate, spire broadly acuminate (spire angle 65°). Apex conical. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (12 per mm on penultimate whorl). Sutures relatively shallow. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 5-fold, consisting of: one lamella-like parietal tooth and one additional parietal denticle; one slab-like palatal tooth, forming a narrow parieto-palatal sinus; one basal denticle; and a deep-set, folded columellar baffle, but no shallower columellar denticles. Shells and anatomy of juveniles unknown.
</paragraph>
<paragraph blockId="13.[189,421,267,293]" box="[189,421,267,293]" pageId="13" pageNumber="14">
<heading bold="true" box="[189,421,267,293]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis bold="true" box="[189,421,267,293]" pageId="13" pageNumber="14">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[189,716,314,341]" box="[189,716,314,341]" pageId="13" pageNumber="14">
Evergreen coastal thicket at the
<typeStatus box="[564,615,315,341]" pageId="13" pageNumber="14">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="discussion">
<paragraph blockId="13.[189,303,380,406]" box="[189,303,380,406]" pageId="13" pageNumber="14">
<heading bold="true" box="[189,303,380,406]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis bold="true" box="[189,303,380,406]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[189,1399,427,742]" pageId="13" pageNumber="14">
This taxon is in some respects intermediate between the
<typeStatus box="[856,907,428,454]" pageId="13" pageNumber="14">type</typeStatus>
specimens of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[1076,1211,428,454]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[1076,1211,428,454]" italics="true" pageId="13" pageNumber="14">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1219,1349,427,454]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
and
<emphasis box="[189,336,463,489]" italics="true" pageId="13" pageNumber="14">
D. b.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[254,336,463,489]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">browni</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[344,476,463,490]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
, and thus distinct from either, yet difficult to place. It is larger and not as squat as other
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[295,416,499,525]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
<emphasis box="[295,416,499,525]" italics="true" pageId="13" pageNumber="14">D. browni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[427,560,499,526]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[570,605,499,525]" italics="true" pageId="13" pageNumber="14">s.l.</emphasis>
, and lacks the additional basal denticle and shallower columellar dentition. Conversely, it has a parietal denticle not seen in
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[913,1027,535,561]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[913,1027,535,561]" italics="true" pageId="13" pageNumber="14">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1038,1172,535,562]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
s.l, is smaller and squatter than most
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[416,529,571,597]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[416,529,571,597]" italics="true" pageId="13" pageNumber="14">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[538,669,571,598]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[678,712,571,597]" italics="true" pageId="13" pageNumber="14">s.l.</emphasis>
, and has a longer and narrower parieto-palatal sinus than any
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[239,351,607,633]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[239,351,607,633]" italics="true" pageId="13" pageNumber="14">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[359,489,607,634]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[497,532,607,633]" italics="true" pageId="13" pageNumber="14">s.l.</emphasis>
Mafia Island, from which no other
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[954,1101,607,633]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[954,1101,607,633]" italics="true" pageId="13" pageNumber="14">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1108,1216,608,634]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
are yet known, lies at almost the same latitude (
<geoCoordinate box="[575,641,643,669]" degrees="7.8" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="5555" value="-7.8">
7.8°
<collectionCode box="[625,641,643,669]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="13" pageNumber="14" type="Herbarium">S</collectionCode>
</geoCoordinate>
) as the apparent northernmost limit of
<emphasis box="[1109,1259,643,669]" italics="true" pageId="13" pageNumber="14">
D. b.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[1177,1259,643,669]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">browni</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[1268,1399,643,670]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
in mainland
<collectingCountry box="[335,444,679,705]" name="Tanzania" pageId="13" pageNumber="14">Tanzania</collectingCountry>
. Given the morphology of this specimen, its latitude and its isolation as an island population, we treat it as a subspecies of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[670,788,715,741]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
<emphasis box="[670,788,715,741]" italics="true" pageId="13" pageNumber="14">D. browni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[796,925,715,742]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>