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682 lines
82 KiB
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<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben & Tattersfield, Peter" docDate="2013" docId="4511E41DD826FFC2FDCAFA05F388FB39" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella Rowson & Tattersfield 2013, gen. nov." docType="treatment" docUuid="1C3959C3-50E5-4DFF-A5AC-67BB9F05A042" docUuidSource="ZooBank" docVersion="9" lastPageNumber="6" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="3" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
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<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rowson, Ben</mods:namePart>
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<mods:nameIdentifier type="LSID">05D46259-DA45-45A4-9127-5AF361D566A7</mods:nameIdentifier>
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<mods:affiliation>Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tattersfield, Peter</mods:namePart>
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<mods:nameIdentifier type="LSID">8C55642E-B18A-461B-9BF9-31CB6DB7A070</mods:nameIdentifier>
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<mods:affiliation>Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-02-04</mods:number>
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<mods:detail type="issue">
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<mods:number>37</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.5852/ejt.2013.37</mods:identifier>
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<mods:identifier type="GBIF-Dataset">9682b45b-5e22-4965-a293-e203660e0ed3</mods:identifier>
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<mods:identifier type="ISSN">2118-9773</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3815757" ID-GBIF-Taxon="163971582" ID-Zenodo-Dep="3815757" LSID="urn:lsid:zoobank.org:act:1C3959C3-50E5-4DFF-A5AC-67BB9F05A042" httpUri="http://treatment.plazi.org/id/4511E41DD826FFC2FDCAFA05F388FB39" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection box="[622,966,1413,1440]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph blockId="2.[363,1224,1413,1510]" box="[622,966,1413,1440]" pageId="2" pageNumber="3">
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<heading box="[622,966,1413,1440]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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Genus
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<taxonomicName ID-CoL="7P2R7" ID-ENA="1929309" authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[704,852,1413,1440]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[704,852,1413,1440]" italics="true" pageId="2" pageNumber="3">Dadagulella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[859,966,1413,1439]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="description">
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urn:lsid:zoobank.org:act:
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<uuid box="[652,1224,1448,1475]" pageId="2" pageNumber="3">1C3959C3-50E5-4DFF-A5AC-67BB9F05A042</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="2.[363,1224,1413,1510]" box="[687,901,1484,1510]" pageId="2" pageNumber="3">
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<figureCitation box="[687,802,1484,1510]" captionStart-0="Figs 1-5" captionStart-1="Figs 6-26" captionStart-10="Fig" captionStart-2="Figs 27-35" captionStart-3="Figs 36-40" captionStart-4="Figs 41-42" captionStart-5="Figs 43-50" captionStart-6="Figs 51-64" captionStart-7="Figs 65-68" captionStart-8="Figs 69-76" captionStart-9="Figs 77-83" captionStartId-0="6.[189,242,1930,1956]" captionStartId-1="9.[189,241,1754,1780]" captionStartId-10="40.[189,232,1930,1956]" captionStartId-2="16.[189,241,1824,1850]" captionStartId-3="24.[189,241,1860,1886]" captionStartId-4="30.[189,241,1930,1956]" captionStartId-5="34.[189,241,1895,1921]" captionStartId-6="35.[189,241,1719,1745]" captionStartId-7="36.[189,241,1895,1921]" captionStartId-8="37.[189,241,1789,1815]" captionStartId-9="38.[189,241,1789,1815]" captionTargetBox-0="[363,1200,672,1890]" captionTargetBox-1="[295,1292,265,1714]" captionTargetBox-10="[193,1393,858,1884]" captionTargetBox-2="[189,1399,348,1784]" captionTargetBox-3="[189,1399,1268,1819]" captionTargetBox-4="[314,1273,931,1890]" captionTargetBox-5="[189,1399,682,1855]" captionTargetBox-6="[203,1389,740,1651]" captionTargetBox-7="[189,1399,903,1855]" captionTargetBox-8="[202,1390,364,1721]" captionTargetBox-9="[201,1381,776,1726]" captionTargetId-0="figure@6.[363,1200,672,1890]" captionTargetId-1="figure@9.[295,1292,265,1714]" captionTargetId-10="figure@40.[189,1399,855,1890]" captionTargetId-2="figure@16.[189,1399,348,1784]" captionTargetId-3="figure@24.[189,1399,1268,1819]" captionTargetId-4="figure@30.[314,1273,931,1890]" captionTargetId-5="figure@34.[189,1399,682,1855]" captionTargetId-6="figure@35.[189,1398,725,1678]" captionTargetId-7="figure@36.[189,1399,903,1855]" captionTargetId-8="figure@37.[189,1398,356,1749]" captionTargetId-9="figure@38.[189,1398,759,1748]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionTargetPageId-10="40" captionTargetPageId-2="16" captionTargetPageId-3="24" captionTargetPageId-4="30" captionTargetPageId-5="34" captionTargetPageId-6="35" captionTargetPageId-7="36" captionTargetPageId-8="37" captionTargetPageId-9="38" captionText-0="Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view." captionText-1="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." captionText-10="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." captionText-2="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." captionText-3="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." captionText-4="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." captionText-5="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." captionText-6="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." captionText-7="Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm." captionText-8="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." captionText-9="Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens." figureDoi-0="http://doi.org/10.5281/zenodo.3806774" figureDoi-1="http://doi.org/10.5281/zenodo.3806778" figureDoi-10="http://doi.org/10.5281/zenodo.3806812" figureDoi-2="http://doi.org/10.5281/zenodo.3806780" figureDoi-3="http://doi.org/10.5281/zenodo.3806784" figureDoi-4="http://doi.org/10.5281/zenodo.3806788" figureDoi-5="http://doi.org/10.5281/zenodo.3806792" figureDoi-6="http://doi.org/10.5281/zenodo.3806796" figureDoi-7="http://doi.org/10.5281/zenodo.3806800" figureDoi-8="http://doi.org/10.5281/zenodo.3806804" figureDoi-9="http://doi.org/10.5281/zenodo.3806808" httpUri-0="https://zenodo.org/record/3806774/files/figure.png" httpUri-1="https://zenodo.org/record/3806778/files/figure.png" httpUri-10="https://zenodo.org/record/3806812/files/figure.png" httpUri-2="https://zenodo.org/record/3806780/files/figure.png" httpUri-3="https://zenodo.org/record/3806784/files/figure.png" httpUri-4="https://zenodo.org/record/3806788/files/figure.png" httpUri-5="https://zenodo.org/record/3806792/files/figure.png" httpUri-6="https://zenodo.org/record/3806796/files/figure.png" httpUri-7="https://zenodo.org/record/3806800/files/figure.png" httpUri-8="https://zenodo.org/record/3806804/files/figure.png" httpUri-9="https://zenodo.org/record/3806808/files/figure.png" pageId="2" pageNumber="3">Figs 1-84</figureCitation>
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;
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<tableCitation box="[814,901,1484,1510]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="2" pageNumber="3" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="type_taxon">
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<paragraph blockId="2.[189,344,1550,1576]" box="[189,344,1550,1576]" pageId="2" pageNumber="3">
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<heading bold="true" box="[189,344,1550,1576]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis bold="true" box="[189,344,1550,1576]" pageId="2" pageNumber="3">
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<typeStatus box="[189,250,1550,1576]" pageId="2" pageNumber="3">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[189,523,1597,1624]" box="[189,523,1597,1624]" pageId="2" pageNumber="3">
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<taxonomicName authority="Preston, 1910" authorityName="Preston" authorityYear="1910" box="[189,518,1597,1624]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="radius">
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<emphasis box="[189,346,1597,1623]" italics="true" pageId="2" pageNumber="3">Ennea radius</emphasis>
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<bibRefCitation author="Preston H. B." box="[354,518,1597,1624]" pageId="2" pageNumber="3" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston, 1910</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="discussion">
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<paragraph blockId="2.[189,393,1663,1689]" box="[189,393,1663,1689]" pageId="2" pageNumber="3">
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<heading bold="true" box="[189,393,1663,1689]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis bold="true" box="[189,393,1663,1689]" pageId="2" pageNumber="3">Included species</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[189,1399,1710,1808]" pageId="2" pageNumber="3">
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Sixteen. In addition,
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<taxonomicName authority="(Preston, 1910)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[433,730,1710,1736]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
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<emphasis box="[433,543,1710,1736]" italics="true" pageId="2" pageNumber="3">D. radius</emphasis>
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(
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<bibRefCitation author="Preston H. B." box="[558,722,1710,1736]" pageId="2" pageNumber="3" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston, 1910</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[737,866,1710,1736]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
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and
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<taxonomicName authority="(Morelet, 1877)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[922,1263,1710,1737]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
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<emphasis box="[922,1078,1710,1736]" italics="true" pageId="2" pageNumber="3">D. minuscula</emphasis>
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(
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<bibRefCitation author="Morelet A." box="[1093,1254,1710,1737]" pageId="2" pageNumber="3" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet,1877</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[1270,1399,1710,1736]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
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are treated as having two subspecies and
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<taxonomicName authority="(van Bruggen, 1969)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[688,1065,1745,1772]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
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<emphasis box="[688,808,1745,1771]" italics="true" pageId="2" pageNumber="3">D. browni</emphasis>
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(van
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<bibRefCitation author="Bruggen A. C. van" box="[879,1057,1745,1772]" pageId="2" pageNumber="3" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen, 1969</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1075,1207,1745,1772]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
|
|
as having three (see below).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="diagnosis">
|
|
<paragraph blockId="2.[189,310,1847,1873]" box="[189,310,1847,1873]" pageId="2" pageNumber="3">
|
|
<heading bold="true" box="[189,310,1847,1873]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
|
<emphasis bold="true" box="[189,310,1847,1873]" pageId="2" pageNumber="3">Diagnosis</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph blockId="2.[189,1399,1894,2028]" lastBlockId="4.[189,1399,267,437]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
|
|
Shell small (
|
|
<quantity box="[341,481,1894,1921]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="5.5" metricValueMin="2.3" pageId="2" pageNumber="3" unit="mm" value="3.9" valueMax="5.5" valueMin="2.3">
|
|
2.3
|
|
<emphasis box="[379,389,1895,1920]" italics="true" pageId="2" pageNumber="3">-</emphasis>
|
|
5.5 mm
|
|
</quantity>
|
|
high), ovate to ovate-acuminate with an acuminate (sometimes coeloconoid or cyrtoconoid) spire; strongly and often sinuously radially ribbed, never smooth. Peristome reflected. Aperture dentate, with at least one parietal and one palatal tooth. Juvenile aperture (known from 7-8 species) dentate (except possibly in
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Lange" baseAuthorityYear="2007" box="[684,825,2002,2027]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="nictitans" status="comb. nov.">
|
|
<emphasis box="[684,825,2002,2027]" italics="true" pageId="2" pageNumber="3">D. nictitans</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[837,974,2001,2027]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
|
|
), aperture not downturned. Radula (known from three species) bearing multicuspid teeth, the outer cusps much smaller than the inner. Other anatomy (known from six species): salivary gland bilobed or Y-shaped. Vas deferens thickened or with diverticulum prior to insertion on penis, penis lacking sheath, with a muscular apex in which is embedded a spatulate scoop with a microscopically serrated tip, usually associated with one or two large apical hooks, and a few (<40) smaller hooks lower down. For dimensions, see
|
|
<tableCitation box="[1111,1199,410,436]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="4" pageNumber="5" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption ID-Table-UUID="99C70583D827FFC4FF1FFDA6F602FD6F" httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="3" pageNumber="4" targetBox="[202,1925,710,1733]" targetIsTable="true" targetPageId="3">
|
|
<paragraph blockId="3.[187,1941,550,648]" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[187,288,550,578]" pageId="3" pageNumber="4">Table 1.</emphasis>
|
|
Measurements and dentition scores for adult shells of
|
|
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[944,1091,550,578]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus" status="gen. nov.">
|
|
<emphasis box="[944,1091,550,578]" italics="true" pageId="3" pageNumber="4">Dadagulella</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1100,1208,550,578]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="3" pageNumber="4">
|
|
<table box="[202,1925,710,1733]" gridcols="12" gridrows="22" pageId="3" pageNumber="4">
|
|
<tr box="[202,1925,710,779]" gridrow="0" pageId="3" pageNumber="4">
|
|
<th box="[202,416,710,779]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[299,369,732,758]" pageId="3" pageNumber="4">Taxon</emphasis>
|
|
</th>
|
|
<th box="[495,531,710,779]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[506,521,732,758]" pageId="3" pageNumber="4">n</emphasis>
|
|
</th>
|
|
<th box="[565,686,710,779]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[585,666,756,779]" pageId="3" pageNumber="4">H (mm)</emphasis>
|
|
</th>
|
|
<th box="[722,1018,710,779]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[793,961,710,736]" pageId="3" pageNumber="4">Size and shape</emphasis>
|
|
<emphasis bold="true" box="[740,965,756,779]" pageId="3" pageNumber="4">W (mm) Wh</emphasis>
|
|
</th>
|
|
<th box="[1047,1193,710,779]" gridcol="5" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[1047,1193,756,779]" pageId="3" pageNumber="4">Spire angle (°)</emphasis>
|
|
</th>
|
|
<th box="[1224,1335,710,779]" gridcol="6" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[1224,1335,710,736]" pageId="3" pageNumber="4">Sculpture</emphasis>
|
|
<emphasis bold="true" box="[1227,1332,756,779]" pageId="3" pageNumber="4">Ribs / mm</emphasis>
|
|
</th>
|
|
<th box="[1366,1447,710,779]" gridcol="7" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[1366,1447,756,779]" pageId="3" pageNumber="4">Parietal</emphasis>
|
|
</th>
|
|
<th box="[1478,1826,710,779]" colspan="3" colspanRight="2" gridcol="8" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[1553,1739,710,736]" pageId="3" pageNumber="4">Number of teeth</emphasis>
|
|
<emphasis bold="true" box="[1478,1826,756,779]" pageId="3" pageNumber="4">Palatal Basal Columellar</emphasis>
|
|
</th>
|
|
<th box="[1858,1925,710,779]" gridcol="11" gridrow="0" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[1865,1917,756,779]" pageId="3" pageNumber="4">Total</emphasis>
|
|
</th>
|
|
</tr>
|
|
<tr box="[202,1925,802,825]" gridrow="1" pageId="3" pageNumber="4">
|
|
<th box="[202,416,802,825]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">
|
|
<taxonomicName authority="radius" authorityName="radius (Preston" authorityYear="1910" baseAuthorityName="radius (Preston" baseAuthorityYear="1910" box="[202,334,802,825]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="variety" species="radius" variety="radius">
|
|
<emphasis bold="true" box="[202,334,802,825]" italics="true" pageId="3" pageNumber="4">radius radius</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,802,825]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">44</td>
|
|
<td box="[565,686,802,825]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">2.8-4.6 (3.6)</td>
|
|
<td box="[722,842,802,825]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">1.6-2.4 (1.9)</td>
|
|
<td box="[875,1018,802,825]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">6.0-7.0 (6.5)</td>
|
|
<td box="[1047,1193,802,825]" gridcol="5" gridrow="1" pageId="3" pageNumber="4">55-70 (62)</td>
|
|
<td box="[1224,1335,802,1686]" gridcol="6" gridrow="1" pageId="3" pageNumber="4" rowspan="20" rowspanBelow="19">7-16 (12) 5-10 (7) 11-15 (13) 12 12 7 8-14 (10) 13 10-13 (11) 8 12-15 (13) 13-15 (14) 5-7 (6) 12-13 16 27 10-13 15-23 (18) 12 9</td>
|
|
<td box="[1366,1447,802,825]" gridcol="7" gridrow="1" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,802,825]" gridcol="8" gridrow="1" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,802,825]" gridcol="9" gridrow="1" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,802,825]" gridcol="10" gridrow="1" pageId="3" pageNumber="4">2-4 (2)</td>
|
|
<td box="[1858,1925,802,825]" gridcol="11" gridrow="1" pageId="3" pageNumber="4">5-7 (5)</td>
|
|
</tr>
|
|
<tr box="[202,1925,847,870]" gridrow="2" pageId="3" pageNumber="4">
|
|
<th box="[202,416,847,870]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
|
|
<taxonomicName box="[202,274,847,870]" pageId="3" pageNumber="4" rank="subSpecies" subSpecies="calva">
|
|
<emphasis bold="true" box="[202,274,847,870]" italics="true" pageId="3" pageNumber="4">r. calva</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,847,870]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">7</td>
|
|
<td box="[565,686,847,870]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">4.0-4.6 (4.4)</td>
|
|
<td box="[722,842,847,870]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">2.1-2.5 (2.3)</td>
|
|
<td box="[875,1018,847,870]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">6.5-8.0 (7.0)</td>
|
|
<td box="[1047,1193,847,870]" gridcol="5" gridrow="2" pageId="3" pageNumber="4">49-62 (57)</td>
|
|
<td box="[1366,1447,847,870]" gridcol="7" gridrow="2" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,847,870]" gridcol="8" gridrow="2" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,847,870]" gridcol="9" gridrow="2" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,847,870]" gridcol="10" gridrow="2" pageId="3" pageNumber="4">1-2 (2)</td>
|
|
<td box="[1858,1925,847,870]" gridcol="11" gridrow="2" pageId="3" pageNumber="4">4-5 (5)</td>
|
|
</tr>
|
|
<tr box="[202,1925,892,916]" gridrow="3" pageId="3" pageNumber="4">
|
|
<th box="[202,416,892,916]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[202,347,892,915]" italics="true" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[202,271,892,915]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="browni">browni</taxonomicName>
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[278,347,892,915]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="browni">browni</taxonomicName>
|
|
</emphasis>
|
|
</th>
|
|
<td box="[495,531,892,916]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">17</td>
|
|
<td box="[565,686,892,916]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">2.6-3.3 (2.9)</td>
|
|
<td box="[722,842,892,916]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">1.6-1.9 (1.8)</td>
|
|
<td box="[875,1018,892,916]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">5.5-6.0 (5.5)</td>
|
|
<td box="[1047,1193,892,916]" gridcol="5" gridrow="3" pageId="3" pageNumber="4">66-76 (70)</td>
|
|
<td box="[1366,1447,892,916]" gridcol="7" gridrow="3" pageId="3" pageNumber="4">1-2 (2)</td>
|
|
<td box="[1478,1547,892,916]" gridcol="8" gridrow="3" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,892,916]" gridcol="9" gridrow="3" pageId="3" pageNumber="4">1-2 (2)</td>
|
|
<td box="[1735,1826,892,916]" gridcol="10" gridrow="3" pageId="3" pageNumber="4">2-3 (3)</td>
|
|
<td box="[1858,1925,892,916]" gridcol="11" gridrow="3" pageId="3" pageNumber="4">6-8 (8)</td>
|
|
</tr>
|
|
<tr box="[202,1925,938,961]" gridrow="4" pageId="3" pageNumber="4">
|
|
<th box="[202,416,938,961]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[202,319,938,961]" italics="true" pageId="3" pageNumber="4">b. mafiensis</emphasis>
|
|
</th>
|
|
<td box="[495,531,938,961]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">1</td>
|
|
<td box="[565,686,938,961]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">3.3</td>
|
|
<td box="[722,842,938,961]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">1.8</td>
|
|
<td box="[875,1018,938,961]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">6.0</td>
|
|
<td box="[1047,1193,938,961]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">65</td>
|
|
<td box="[1366,1447,938,961]" gridcol="7" gridrow="4" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1478,1547,938,961]" gridcol="8" gridrow="4" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,938,961]" gridcol="9" gridrow="4" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,938,961]" gridcol="10" gridrow="4" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1858,1925,938,961]" gridcol="11" gridrow="4" pageId="3" pageNumber="4">5</td>
|
|
</tr>
|
|
<tr box="[202,1925,983,1006]" gridrow="5" pageId="3" pageNumber="4">
|
|
<th box="[202,416,983,1006]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[202,359,983,1006]" italics="true" pageId="3" pageNumber="4">b. semulikiensis</emphasis>
|
|
</th>
|
|
<td box="[495,531,983,1006]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">2</td>
|
|
<td box="[565,686,983,1006]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">3.2-3.3</td>
|
|
<td box="[722,842,983,1006]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">1.9</td>
|
|
<td box="[875,1018,983,1006]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">6.0-6.5</td>
|
|
<td box="[1047,1193,983,1006]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">67</td>
|
|
<td box="[1366,1447,983,1006]" gridcol="7" gridrow="5" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1478,1547,983,1006]" gridcol="8" gridrow="5" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,983,1006]" gridcol="9" gridrow="5" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1735,1826,983,1006]" gridcol="10" gridrow="5" pageId="3" pageNumber="4">3</td>
|
|
<td box="[1858,1925,983,1006]" gridcol="11" gridrow="5" pageId="3" pageNumber="4">8</td>
|
|
</tr>
|
|
<tr box="[202,1925,1029,1052]" gridrow="6" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1029,1052]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Muratov" baseAuthorityYear="2010" box="[202,279,1029,1052]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="delgada">
|
|
<emphasis bold="true" box="[202,279,1029,1052]" italics="true" pageId="3" pageNumber="4">delgada</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1029,1052]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">2</td>
|
|
<td box="[565,686,1029,1052]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">3.8-4.0</td>
|
|
<td box="[722,842,1029,1052]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">1.8</td>
|
|
<td box="[875,1018,1029,1052]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">6.5-7.5</td>
|
|
<td box="[1047,1193,1029,1052]" gridcol="5" gridrow="6" pageId="3" pageNumber="4">43-49</td>
|
|
<td box="[1366,1447,1029,1052]" gridcol="7" gridrow="6" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1029,1052]" gridcol="8" gridrow="6" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1029,1052]" gridcol="9" gridrow="6" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1029,1052]" gridcol="10" gridrow="6" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1858,1925,1029,1052]" gridcol="11" gridrow="6" pageId="3" pageNumber="4">5</td>
|
|
</tr>
|
|
<tr box="[202,1925,1074,1097]" gridrow="7" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1074,1097]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[202,298,1074,1097]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="ecclesiola">
|
|
<emphasis bold="true" box="[202,298,1074,1097]" italics="true" pageId="3" pageNumber="4">ecclesiola</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1074,1097]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">15</td>
|
|
<td box="[565,686,1074,1097]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">3.2-3.7 (3.4)</td>
|
|
<td box="[722,842,1074,1097]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">1.8-1.9 (1.8)</td>
|
|
<td box="[875,1018,1074,1097]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">5.5-7.0 (6.5)</td>
|
|
<td box="[1047,1193,1074,1097]" gridcol="5" gridrow="7" pageId="3" pageNumber="4">53-65 (60)</td>
|
|
<td box="[1366,1447,1074,1097]" gridcol="7" gridrow="7" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1074,1097]" gridcol="8" gridrow="7" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1074,1097]" gridcol="9" gridrow="7" pageId="3" pageNumber="4">1 (hidden)</td>
|
|
<td box="[1735,1826,1074,1097]" gridcol="10" gridrow="7" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1858,1925,1074,1097]" gridcol="11" gridrow="7" pageId="3" pageNumber="4">4</td>
|
|
</tr>
|
|
<tr box="[202,1925,1119,1142]" gridrow="8" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1119,1142]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[202,298,1119,1142]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="cresswelli">
|
|
<emphasis bold="true" box="[202,298,1119,1142]" italics="true" pageId="3" pageNumber="4">cresswelli</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1119,1142]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">1</td>
|
|
<td box="[565,686,1119,1142]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">3.7</td>
|
|
<td box="[722,842,1119,1142]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">1.8</td>
|
|
<td box="[875,1018,1119,1142]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">7.0</td>
|
|
<td box="[1047,1193,1119,1142]" gridcol="5" gridrow="8" pageId="3" pageNumber="4">52</td>
|
|
<td box="[1366,1447,1119,1142]" gridcol="7" gridrow="8" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1119,1142]" gridcol="8" gridrow="8" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1119,1142]" gridcol="9" gridrow="8" pageId="3" pageNumber="4">0</td>
|
|
<td box="[1735,1826,1119,1142]" gridcol="10" gridrow="8" pageId="3" pageNumber="4">3</td>
|
|
<td box="[1858,1925,1119,1142]" gridcol="11" gridrow="8" pageId="3" pageNumber="4">5</td>
|
|
</tr>
|
|
<tr box="[202,1925,1165,1188]" gridrow="9" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1165,1188]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[202,291,1165,1188]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="minareta">
|
|
<emphasis bold="true" box="[202,291,1165,1188]" italics="true" pageId="3" pageNumber="4">minareta</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1165,1188]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">18</td>
|
|
<td box="[565,686,1165,1188]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">3.4-4.0 (3.6)</td>
|
|
<td box="[722,842,1165,1188]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">1.8-2.0 (1.9)</td>
|
|
<td box="[875,1018,1165,1188]" gridcol="4" gridrow="9" pageId="3" pageNumber="4">6.0-7.0 (6.5)</td>
|
|
<td box="[1047,1193,1165,1188]" gridcol="5" gridrow="9" pageId="3" pageNumber="4">48-63 (55)</td>
|
|
<td box="[1366,1447,1165,1188]" gridcol="7" gridrow="9" pageId="3" pageNumber="4">1-2 (1)</td>
|
|
<td box="[1478,1547,1165,1188]" gridcol="8" gridrow="9" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1165,1188]" gridcol="9" gridrow="9" pageId="3" pageNumber="4">0-1 (0)</td>
|
|
<td box="[1735,1826,1165,1188]" gridcol="10" gridrow="9" pageId="3" pageNumber="4">2-3 (2)</td>
|
|
<td box="[1858,1925,1165,1188]" gridcol="11" gridrow="9" pageId="3" pageNumber="4">5-7 (5)</td>
|
|
</tr>
|
|
<tr box="[202,1925,1210,1233]" gridrow="10" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1210,1233]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Verdcourt" authorityYear="1962" box="[202,297,1210,1233]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="cuspidata">
|
|
<emphasis bold="true" box="[202,297,1210,1233]" italics="true" pageId="3" pageNumber="4">cuspidata</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1210,1233]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">3</td>
|
|
<td box="[565,686,1210,1233]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">4.3-4.8 (4.6)</td>
|
|
<td box="[722,842,1210,1233]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">2.4-2.5 (2.5)</td>
|
|
<td box="[875,1018,1210,1233]" gridcol="4" gridrow="10" pageId="3" pageNumber="4">7.75</td>
|
|
<td box="[1047,1193,1210,1233]" gridcol="5" gridrow="10" pageId="3" pageNumber="4">58</td>
|
|
<td box="[1366,1447,1210,1233]" gridcol="7" gridrow="10" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1210,1233]" gridcol="8" gridrow="10" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1581,1680,1210,1233]" gridcol="9" gridrow="10" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1210,1233]" gridcol="10" gridrow="10" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1858,1925,1210,1233]" gridcol="11" gridrow="10" pageId="3" pageNumber="4">6</td>
|
|
</tr>
|
|
<tr box="[202,1925,1255,1278]" gridrow="11" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1255,1278]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[202,305,1255,1278]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pembensis">
|
|
<emphasis bold="true" box="[202,305,1255,1278]" italics="true" pageId="3" pageNumber="4">pembensis</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1255,1278]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">11</td>
|
|
<td box="[565,686,1255,1278]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">4.8-5.5 (5.1)</td>
|
|
<td box="[722,842,1255,1278]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">2.0-2.5 (2.4)</td>
|
|
<td box="[875,1018,1255,1278]" gridcol="4" gridrow="11" pageId="3" pageNumber="4">6.5-7.5 (7.0)</td>
|
|
<td box="[1047,1193,1255,1278]" gridcol="5" gridrow="11" pageId="3" pageNumber="4">51- 65 (57)</td>
|
|
<td box="[1366,1447,1255,1278]" gridcol="7" gridrow="11" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1255,1278]" gridcol="8" gridrow="11" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1255,1278]" gridcol="9" gridrow="11" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1255,1278]" gridcol="10" gridrow="11" pageId="3" pageNumber="4">3</td>
|
|
<td box="[1858,1925,1255,1278]" gridcol="11" gridrow="11" pageId="3" pageNumber="4">6</td>
|
|
</tr>
|
|
<tr box="[202,1925,1301,1324]" gridrow="12" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1301,1324]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Lange" baseAuthorityYear="2007" box="[202,287,1301,1324]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="nictitans">
|
|
<emphasis bold="true" box="[202,287,1301,1324]" italics="true" pageId="3" pageNumber="4">nictitans</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1301,1324]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">9</td>
|
|
<td box="[565,686,1301,1324]" gridcol="2" gridrow="12" pageId="3" pageNumber="4">3.6-3.9 (3.8)</td>
|
|
<td box="[722,842,1301,1324]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">2.0-2.2 (2.1)</td>
|
|
<td box="[875,1018,1301,1324]" gridcol="4" gridrow="12" pageId="3" pageNumber="4">5.5</td>
|
|
<td box="[1047,1193,1301,1324]" gridcol="5" gridrow="12" pageId="3" pageNumber="4">64-69 (67)</td>
|
|
<td box="[1366,1447,1301,1324]" gridcol="7" gridrow="12" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1301,1324]" gridcol="8" gridrow="12" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1301,1324]" gridcol="9" gridrow="12" pageId="3" pageNumber="4">1 (hidden)</td>
|
|
<td box="[1735,1826,1301,1324]" gridcol="10" gridrow="12" pageId="3" pageNumber="4">2-3 (3)</td>
|
|
<td box="[1858,1925,1301,1324]" gridcol="11" gridrow="12" pageId="3" pageNumber="4">5-6 (6)</td>
|
|
</tr>
|
|
<tr box="[202,1925,1346,1369]" gridrow="13" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1346,1369]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[202,331,1346,1369]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="frontierarum">
|
|
<emphasis bold="true" box="[202,331,1346,1369]" italics="true" pageId="3" pageNumber="4">frontierarum</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1346,1369]" gridcol="1" gridrow="13" pageId="3" pageNumber="4">5</td>
|
|
<td box="[565,686,1346,1369]" gridcol="2" gridrow="13" pageId="3" pageNumber="4">3.1-3.4 (3.3)</td>
|
|
<td box="[722,842,1346,1369]" gridcol="3" gridrow="13" pageId="3" pageNumber="4">1.8-2.0 (1.9)</td>
|
|
<td box="[875,1018,1346,1369]" gridcol="4" gridrow="13" pageId="3" pageNumber="4">4.5-5.0 (4.75)</td>
|
|
<td box="[1047,1193,1346,1369]" gridcol="5" gridrow="13" pageId="3" pageNumber="4">68-77 (73)</td>
|
|
<td box="[1366,1447,1346,1369]" gridcol="7" gridrow="13" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1346,1369]" gridcol="8" gridrow="13" pageId="3" pageNumber="4">3</td>
|
|
<td box="[1581,1680,1346,1369]" gridcol="9" gridrow="13" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1346,1369]" gridcol="10" gridrow="13" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1858,1925,1346,1369]" gridcol="11" gridrow="13" pageId="3" pageNumber="4">6</td>
|
|
</tr>
|
|
<tr box="[202,1925,1391,1415]" gridrow="14" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1391,1415]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[202,250,1391,1414]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="delta">
|
|
<emphasis bold="true" box="[202,250,1391,1414]" italics="true" pageId="3" pageNumber="4">delta</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1391,1415]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">2</td>
|
|
<td box="[565,686,1391,1415]" gridcol="2" gridrow="14" pageId="3" pageNumber="4">4.2-4.3</td>
|
|
<td box="[722,842,1391,1415]" gridcol="3" gridrow="14" pageId="3" pageNumber="4">2.1-2.4</td>
|
|
<td box="[875,1018,1391,1415]" gridcol="4" gridrow="14" pageId="3" pageNumber="4">6.5</td>
|
|
<td box="[1047,1193,1391,1415]" gridcol="5" gridrow="14" pageId="3" pageNumber="4">56-66</td>
|
|
<td box="[1366,1447,1391,1415]" gridcol="7" gridrow="14" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1391,1415]" gridcol="8" gridrow="14" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1391,1415]" gridcol="9" gridrow="14" pageId="3" pageNumber="4">0</td>
|
|
<td box="[1735,1826,1391,1415]" gridcol="10" gridrow="14" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1858,1925,1391,1415]" gridcol="11" gridrow="14" pageId="3" pageNumber="4">3</td>
|
|
</tr>
|
|
<tr box="[202,1925,1437,1460]" gridrow="15" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1437,1460]" gridcol="0" gridrow="15" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="van Bruggen & Van Goethem" authorityYear="1999" box="[202,263,1437,1460]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="selene">
|
|
<emphasis bold="true" box="[202,263,1437,1460]" italics="true" pageId="3" pageNumber="4">selene</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1437,1460]" gridcol="1" gridrow="15" pageId="3" pageNumber="4">2</td>
|
|
<td box="[565,686,1437,1460]" gridcol="2" gridrow="15" pageId="3" pageNumber="4">3.6-3.7</td>
|
|
<td box="[722,842,1437,1460]" gridcol="3" gridrow="15" pageId="3" pageNumber="4">2.1-2.2</td>
|
|
<td box="[875,1018,1437,1460]" gridcol="4" gridrow="15" pageId="3" pageNumber="4">6.0-6.25</td>
|
|
<td box="[1047,1193,1437,1460]" gridcol="5" gridrow="15" pageId="3" pageNumber="4">70</td>
|
|
<td box="[1366,1447,1437,1460]" gridcol="7" gridrow="15" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1437,1460]" gridcol="8" gridrow="15" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1437,1460]" gridcol="9" gridrow="15" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1437,1460]" gridcol="10" gridrow="15" pageId="3" pageNumber="4">2</td>
|
|
<td box="[1858,1925,1437,1460]" gridcol="11" gridrow="15" pageId="3" pageNumber="4">5</td>
|
|
</tr>
|
|
<tr box="[202,1925,1482,1505]" gridrow="16" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1482,1505]" gridcol="0" gridrow="16" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="van Bruggen" authorityYear="2000" box="[202,313,1482,1505]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="meredithae">
|
|
<emphasis bold="true" box="[202,313,1482,1505]" italics="true" pageId="3" pageNumber="4">meredithae</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1482,1505]" gridcol="1" gridrow="16" pageId="3" pageNumber="4">23</td>
|
|
<td box="[565,686,1482,1505]" gridcol="2" gridrow="16" pageId="3" pageNumber="4">2.3-3.1 (2.6)</td>
|
|
<td box="[722,842,1482,1505]" gridcol="3" gridrow="16" pageId="3" pageNumber="4">1.4-1.6 (1.5)</td>
|
|
<td box="[875,1018,1482,1505]" gridcol="4" gridrow="16" pageId="3" pageNumber="4">5.25-5.75 (5.5)</td>
|
|
<td box="[1047,1193,1482,1505]" gridcol="5" gridrow="16" pageId="3" pageNumber="4">58-77 (71)</td>
|
|
<td box="[1366,1447,1482,1505]" gridcol="7" gridrow="16" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1482,1505]" gridcol="8" gridrow="16" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1482,1505]" gridcol="9" gridrow="16" pageId="3" pageNumber="4">0</td>
|
|
<td box="[1735,1826,1482,1505]" gridcol="10" gridrow="16" pageId="3" pageNumber="4">1-2 (2)</td>
|
|
<td box="[1858,1925,1482,1505]" gridcol="11" gridrow="16" pageId="3" pageNumber="4">3-4 (3)</td>
|
|
</tr>
|
|
<tr box="[202,1925,1527,1551]" gridrow="17" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1527,1551]" gridcol="0" gridrow="17" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[202,416,1527,1550]" italics="true" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[202,306,1527,1550]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[312,416,1527,1550]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
|
|
</emphasis>
|
|
</th>
|
|
<td box="[495,531,1527,1551]" gridcol="1" gridrow="17" pageId="3" pageNumber="4">2</td>
|
|
<td box="[565,686,1527,1551]" gridcol="2" gridrow="17" pageId="3" pageNumber="4">3.0-4.1</td>
|
|
<td box="[722,842,1527,1551]" gridcol="3" gridrow="17" pageId="3" pageNumber="4">1.9-2.0</td>
|
|
<td box="[875,1018,1527,1551]" gridcol="4" gridrow="17" pageId="3" pageNumber="4">6.5-7.0</td>
|
|
<td box="[1047,1193,1527,1551]" gridcol="5" gridrow="17" pageId="3" pageNumber="4">48- 63</td>
|
|
<td box="[1366,1447,1527,1551]" gridcol="7" gridrow="17" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1527,1551]" gridcol="8" gridrow="17" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1527,1551]" gridcol="9" gridrow="17" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1527,1551]" gridcol="10" gridrow="17" pageId="3" pageNumber="4">1-2</td>
|
|
<td box="[1858,1925,1527,1551]" gridcol="11" gridrow="17" pageId="3" pageNumber="4">4-5</td>
|
|
</tr>
|
|
<tr box="[202,1925,1573,1596]" gridrow="18" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1573,1596]" gridcol="0" gridrow="18" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[202,415,1573,1596]" italics="true" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[202,306,1573,1596]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
|
|
mahorana
|
|
</emphasis>
|
|
</th>
|
|
<td box="[495,531,1573,1596]" gridcol="1" gridrow="18" pageId="3" pageNumber="4">7</td>
|
|
<td box="[565,686,1573,1596]" gridcol="2" gridrow="18" pageId="3" pageNumber="4">4.3-4.6 (4.4)</td>
|
|
<td box="[722,842,1573,1596]" gridcol="3" gridrow="18" pageId="3" pageNumber="4">1.9-2.2 (2.0)</td>
|
|
<td box="[875,1018,1573,1596]" gridcol="4" gridrow="18" pageId="3" pageNumber="4">6.5-7.5 (7.0)</td>
|
|
<td box="[1047,1193,1573,1596]" gridcol="5" gridrow="18" pageId="3" pageNumber="4">50-57 (53)</td>
|
|
<td box="[1366,1447,1573,1596]" gridcol="7" gridrow="18" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1573,1596]" gridcol="8" gridrow="18" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1573,1596]" gridcol="9" gridrow="18" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1735,1826,1573,1596]" gridcol="10" gridrow="18" pageId="3" pageNumber="4">3</td>
|
|
<td box="[1858,1925,1573,1596]" gridcol="11" gridrow="18" pageId="3" pageNumber="4">6</td>
|
|
</tr>
|
|
<tr box="[202,1925,1618,1641]" gridrow="19" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1618,1641]" gridcol="0" gridrow="19" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[202,310,1618,1641]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="rondoensis">
|
|
<emphasis bold="true" box="[202,310,1618,1641]" italics="true" pageId="3" pageNumber="4">rondoensis</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1618,1641]" gridcol="1" gridrow="19" pageId="3" pageNumber="4">1</td>
|
|
<td box="[565,686,1618,1641]" gridcol="2" gridrow="19" pageId="3" pageNumber="4">4.1</td>
|
|
<td box="[722,842,1618,1641]" gridcol="3" gridrow="19" pageId="3" pageNumber="4">2.2</td>
|
|
<td box="[875,1018,1618,1641]" gridcol="4" gridrow="19" pageId="3" pageNumber="4">6.25</td>
|
|
<td box="[1047,1193,1618,1641]" gridcol="5" gridrow="19" pageId="3" pageNumber="4">53</td>
|
|
<td box="[1366,1447,1618,1641]" gridcol="7" gridrow="19" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1618,1641]" gridcol="8" gridrow="19" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1618,1641]" gridcol="9" gridrow="19" pageId="3" pageNumber="4">0</td>
|
|
<td box="[1735,1826,1618,1641]" gridcol="10" gridrow="19" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1858,1925,1618,1641]" gridcol="11" gridrow="19" pageId="3" pageNumber="4">3</td>
|
|
</tr>
|
|
<tr box="[202,1925,1663,1686]" gridrow="20" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1663,1686]" gridcol="0" gridrow="20" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[202,291,1663,1686]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="conoidea">
|
|
<emphasis bold="true" box="[202,291,1663,1686]" italics="true" pageId="3" pageNumber="4">conoidea</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[495,531,1663,1686]" gridcol="1" gridrow="20" pageId="3" pageNumber="4">1</td>
|
|
<td box="[565,686,1663,1686]" gridcol="2" gridrow="20" pageId="3" pageNumber="4">5.5</td>
|
|
<td box="[722,842,1663,1686]" gridcol="3" gridrow="20" pageId="3" pageNumber="4">2.8</td>
|
|
<td box="[875,1018,1663,1686]" gridcol="4" gridrow="20" pageId="3" pageNumber="4">8</td>
|
|
<td box="[1047,1193,1663,1686]" gridcol="5" gridrow="20" pageId="3" pageNumber="4">48</td>
|
|
<td box="[1366,1447,1663,1686]" gridcol="7" gridrow="20" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1478,1547,1663,1686]" gridcol="8" gridrow="20" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1581,1680,1663,1686]" gridcol="9" gridrow="20" pageId="3" pageNumber="4">0</td>
|
|
<td box="[1735,1826,1663,1686]" gridcol="10" gridrow="20" pageId="3" pageNumber="4">1</td>
|
|
<td box="[1858,1925,1663,1686]" gridcol="11" gridrow="20" pageId="3" pageNumber="4">3</td>
|
|
</tr>
|
|
<tr box="[202,1925,1709,1733]" gridrow="21" pageId="3" pageNumber="4">
|
|
<th box="[202,416,1709,1733]" gridcol="0" gridrow="21" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[260,408,1709,1732]" pageId="3" pageNumber="4">Overall values</emphasis>
|
|
</th>
|
|
<td box="[495,531,1709,1733]" gridcol="1" gridrow="21" pageId="3" pageNumber="4">173</td>
|
|
<td box="[565,686,1709,1733]" gridcol="2" gridrow="21" pageId="3" pageNumber="4">2.3-5.5 (3.7)</td>
|
|
<td box="[722,842,1709,1733]" gridcol="3" gridrow="21" pageId="3" pageNumber="4">1.4-2.8 (2.0)</td>
|
|
<td box="[875,1018,1709,1733]" gridcol="4" gridrow="21" pageId="3" pageNumber="4">4.5-8 (6.5)</td>
|
|
<td box="[1047,1193,1709,1733]" gridcol="5" gridrow="21" pageId="3" pageNumber="4">43-77 (60)</td>
|
|
<td box="[1224,1335,1709,1733]" gridcol="6" gridrow="21" pageId="3" pageNumber="4">5-27 (12)</td>
|
|
<td box="[1366,1447,1709,1733]" gridcol="7" gridrow="21" pageId="3" pageNumber="4">1-2 (1)</td>
|
|
<td box="[1478,1547,1709,1733]" gridcol="8" gridrow="21" pageId="3" pageNumber="4">1-3 (1)</td>
|
|
<td box="[1581,1680,1709,1733]" gridcol="9" gridrow="21" pageId="3" pageNumber="4">0-2 (1)</td>
|
|
<td box="[1735,1826,1709,1733]" gridcol="10" gridrow="21" pageId="3" pageNumber="4">1-4 (3)</td>
|
|
<td box="[1858,1925,1709,1733]" gridcol="11" gridrow="21" pageId="3" pageNumber="4">3-8 (5)</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<subSubSection pageId="4" pageNumber="5" type="etymology">
|
|
<paragraph blockId="4.[189,324,476,502]" box="[189,324,476,502]" pageId="4" pageNumber="5">
|
|
<heading bold="true" box="[189,324,476,502]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
|
<emphasis bold="true" box="[189,324,476,502]" pageId="4" pageNumber="5">Etymology</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph blockId="4.[189,1399,523,585]" pageId="4" pageNumber="5">
|
|
Prefix from Swahili noun “
|
|
<emphasis box="[511,570,523,549]" italics="true" pageId="4" pageNumber="5">dada</emphasis>
|
|
” meaning sister, with reference to the apparent relationships and East African centre of diversity of the group.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
|
<paragraph blockId="4.[189,334,624,650]" box="[189,334,624,650]" pageId="4" pageNumber="5">
|
|
<heading bold="true" box="[189,334,624,650]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
|
<emphasis bold="true" box="[189,334,624,650]" pageId="4" pageNumber="5">Description</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph blockId="4.[189,1399,671,1448]" pageId="4" pageNumber="5">
|
|
SHELL (
|
|
<figureCitation box="[273,388,672,699]" captionStart-0="Figs 1-5" captionStart-1="Figs 6-26" captionStart-2="Figs 27-35" captionStart-3="Figs 36-40" captionStart-4="Figs 41-42" captionStart-5="Figs 43-50" captionStart-6="Figs 51-64" captionStartId-0="6.[189,242,1930,1956]" captionStartId-1="9.[189,241,1754,1780]" captionStartId-2="16.[189,241,1824,1850]" captionStartId-3="24.[189,241,1860,1886]" captionStartId-4="30.[189,241,1930,1956]" captionStartId-5="34.[189,241,1895,1921]" captionStartId-6="35.[189,241,1719,1745]" captionTargetBox-0="[363,1200,672,1890]" captionTargetBox-1="[295,1292,265,1714]" captionTargetBox-2="[189,1399,348,1784]" captionTargetBox-3="[189,1399,1268,1819]" captionTargetBox-4="[314,1273,931,1890]" captionTargetBox-5="[189,1399,682,1855]" captionTargetBox-6="[203,1389,740,1651]" captionTargetId-0="figure@6.[363,1200,672,1890]" captionTargetId-1="figure@9.[295,1292,265,1714]" captionTargetId-2="figure@16.[189,1399,348,1784]" captionTargetId-3="figure@24.[189,1399,1268,1819]" captionTargetId-4="figure@30.[314,1273,931,1890]" captionTargetId-5="figure@34.[189,1399,682,1855]" captionTargetId-6="figure@35.[189,1398,725,1678]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionTargetPageId-2="16" captionTargetPageId-3="24" captionTargetPageId-4="30" captionTargetPageId-5="34" captionTargetPageId-6="35" captionText-0="Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view." captionText-1="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." captionText-2="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." captionText-3="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." captionText-4="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." captionText-5="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." captionText-6="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi-0="http://doi.org/10.5281/zenodo.3806774" figureDoi-1="http://doi.org/10.5281/zenodo.3806778" figureDoi-2="http://doi.org/10.5281/zenodo.3806780" figureDoi-3="http://doi.org/10.5281/zenodo.3806784" figureDoi-4="http://doi.org/10.5281/zenodo.3806788" figureDoi-5="http://doi.org/10.5281/zenodo.3806792" figureDoi-6="http://doi.org/10.5281/zenodo.3806796" httpUri-0="https://zenodo.org/record/3806774/files/figure.png" httpUri-1="https://zenodo.org/record/3806778/files/figure.png" httpUri-2="https://zenodo.org/record/3806780/files/figure.png" httpUri-3="https://zenodo.org/record/3806784/files/figure.png" httpUri-4="https://zenodo.org/record/3806788/files/figure.png" httpUri-5="https://zenodo.org/record/3806792/files/figure.png" httpUri-6="https://zenodo.org/record/3806796/files/figure.png" pageId="4" pageNumber="5">Figs 1-64</figureCitation>
|
|
). Small to medium-sized relative to species of
|
|
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1856" box="[935,1023,672,698]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
|
<emphasis box="[935,1023,672,698]" italics="true" pageId="4" pageNumber="5">Gulella</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<quantity box="[1038,1176,671,698]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="5.5" metricValueMin="2.3" pageId="4" pageNumber="5" unit="mm" value="3.9" valueMax="5.5" valueMin="2.3">
|
|
2.3
|
|
<emphasis box="[1076,1086,673,698]" italics="true" pageId="4" pageNumber="5">-</emphasis>
|
|
5.5 mm
|
|
</quantity>
|
|
high x
|
|
<quantity box="[1263,1399,672,698]" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.8" metricValueMin="1.4" pageId="4" pageNumber="5" unit="mm" value="2.0999999999999996" valueMax="2.8" valueMin="1.4">
|
|
1.4
|
|
<emphasis box="[1299,1309,673,698]" italics="true" pageId="4" pageNumber="5">-</emphasis>
|
|
2.8 mm
|
|
</quantity>
|
|
wide), of 4.5
|
|
<emphasis box="[342,352,708,733]" italics="true" pageId="4" pageNumber="5">-</emphasis>
|
|
8 whorls. Ovate to ovate-acuminate (maximum width being approximately at middle of shell, usually at penultimate whorl) or subconical (maximum width being in bottom third of the shell, at body whorl). Spire characteristically narrowly to broadly acuminate, occasionally coeloconoid or cyrtoconoid (spire angle 43-77°), angle varying a little within most species. Apex (i.e. top of spire) pointed in most species, but rounded in others. Sutures usually deep, shells never completely smoothsided. Umbilicus closed or narrowly open, this sometimes varying within a species. Embryonic whorls usually smoothly granulate, with fine radial striae in two species, or irregularly punctate in another. The last part of the embryonic shell is sculptured with very fine radial striae in most (possibly all) species but these are worn away in all but the freshest individuals. Later whorls never smooth, with radial ribs (5
|
|
<emphasis box="[1359,1369,994,1019]" italics="true" pageId="4" pageNumber="5">-</emphasis>
|
|
27 per mm on penultimate whorl) running from suture to suture, often strong and/or sinuous, lamella-like in some species. Peristome complete, or incomplete parietally, always reflected to some extent. Outer palatal surface of aperture with a depression, sometimes furrow-like, corresponding to the palatal tooth, and sometimes another corresponding to the basal tooth. Dentition 3-fold (rarely, and debatably, 2-fold) to 8-fold. Dentition consisting of at least one parietal tooth and one palatal tooth, the latter often slablike and/or bifid, sometimes forming a parieto-palatal sinus. Usually also with one deep-set columellar baffle, folded or sub-bifid or sub-trifid in some species. Often with additional parietal, palatal, basal and/or columellar teeth or denticles. Dentition is variable in some species but the form, as well as the number, of the teeth is often consistent enough to distinguish species. Juvenile shells (known from 7-8 species) always with 3-fold to 4-fold apertural dentition (
|
|
<figureCitation box="[856,985,1350,1377]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="4" pageNumber="5">Figs 43-50</figureCitation>
|
|
) except in one species,
|
|
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Lange" baseAuthorityYear="2007" box="[1262,1398,1351,1376]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="nictitans" status="comb. nov.">
|
|
<emphasis box="[1262,1398,1351,1376]" italics="true" pageId="4" pageNumber="5">D. nictitans</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[189,322,1385,1412]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
|
|
, where the only known juvenile has no teeth. In several species, some or all of the teeth in juvenile shells appear to be resorbed at intervals.
|
|
</paragraph>
|
|
<paragraph blockId="4.[189,1398,1487,1549]" pageId="4" pageNumber="5">CEPHALOPODIUM. Pale cream or yellow, usually with apricot-coloured to orange tentacle retractors and sometimes with brown speckles on the mantle.</paragraph>
|
|
<paragraph blockId="4.[189,1398,1589,1651]" pageId="4" pageNumber="5">PALLIAL COMPLEX. Sigmurethrous, with long zone of contact between long, oblong kidney and hindgut, pulmonary cavity not strongly vascularised.</paragraph>
|
|
<paragraph blockId="4.[189,1398,1690,1753]" pageId="4" pageNumber="5">
|
|
SALIVARY GLANDS (
|
|
<figureCitation box="[400,532,1690,1717]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="4" pageNumber="5">Figs 69-72</figureCitation>
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). United, soft, not tumid, elongate, bilobed to Y-shaped; each duct leaving at the apex of the lobe and evenly thick throughout.
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</paragraph>
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<paragraph blockId="4.[189,1399,1792,1854]" pageId="4" pageNumber="5">BUCCAL MASS. Very small and elongate, radula correspondingly small, tightly enrolled and difficult to locate or prepare.</paragraph>
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<paragraph blockId="4.[189,1399,1893,2027]" pageId="4" pageNumber="5">
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RADULA (
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<figureCitation box="[302,437,1893,1920]" captionStart="Figs 65-68" captionStartId="36.[189,241,1895,1921]" captionTargetBox="[189,1399,903,1855]" captionTargetId="figure@36.[189,1399,903,1855]" captionTargetPageId="36" captionText="Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.3806800" httpUri="https://zenodo.org/record/3806800/files/figure.png" pageId="4" pageNumber="5">Figs 65-68</figureCitation>
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). (known from three species) With a unicuspid central tooth and 9
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<emphasis box="[1263,1273,1894,1919]" italics="true" pageId="4" pageNumber="5">-</emphasis>
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15 laterals in each half-row, diminishing gradually in size laterally. Most or all laterals distinctively bicuspid or multicuspid, with outer cusps smaller or much smaller than inner cusps. Teeth are somewhat delicate (or flexible;
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<bibRefCitation author="Aiken D. W." box="[291,430,2001,2027]" pageId="4" pageNumber="5" pagination="317 - 324" refId="ref25988" refString="Aiken D. W. 1981. Differentiation of the radula of South African species of Gulella into three types. Journal of Conchology 30 (5): 317 - 324." type="journal article" year="1981">Aiken 1981</bibRefCitation>
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) and are short and flake-like at the ventral end of the radular ribbon.
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</paragraph>
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<paragraph blockId="5.[189,1399,267,870]" pageId="5" pageNumber="6">
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GENITALIA. (
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<figureCitation box="[326,454,267,293]" captionStart-0="Figs 69-76" captionStart-1="Figs 77-83" captionStartId-0="37.[189,241,1789,1815]" captionStartId-1="38.[189,241,1789,1815]" captionTargetBox-0="[202,1390,364,1721]" captionTargetBox-1="[201,1381,776,1726]" captionTargetId-0="figure@37.[189,1398,356,1749]" captionTargetId-1="figure@38.[189,1398,759,1748]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." captionText-1="Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens." figureDoi-0="http://doi.org/10.5281/zenodo.3806804" figureDoi-1="http://doi.org/10.5281/zenodo.3806808" httpUri-0="https://zenodo.org/record/3806804/files/figure.png" httpUri-1="https://zenodo.org/record/3806808/files/figure.png" pageId="5" pageNumber="6">Figs 73-83</figureCitation>
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) (known from six species) Hermaphroditic duct diverticulum (talon) short, large, sausage-shaped, not convoluted. Bursa copulatrix attending albumen gland. Acini of prostate indistinct to distinct. Vagina attenuate. Oviduct often containing a single large egg. Vas deferens either thickened prior to insertion on penis, or appearing as such but actually with a diverticulum lying parallel to it. Vas deferens entering penis subapically. Penial retractor muscle branching off columellar muscle, attaching partly to vas deferens, thus sometimes bifid or nearly so. Penis elongate, tubular. Penial sheath absent, but lower part sometimes surrounded by a thin sheath-like layer contiguous with walls of lower penis. Interior of penis with weak radial pilasters and small rhombic pads, sometimes with a longitudinal pilaster or short rounded lobe. Apical part of penis with a spatulate or broad “scoop”. One end with a microscopically serrated tip, the other end deeply embedded in muscular apex of penis. Scoop usually associated with one or two large, broad hooks, lying just behind or underneath it. Scoop and hook(s) could conceivably act together as a grip or pincer. Elsewhere in penis, a longitudinal row or group of few (<40) short, simple hooks mounted on rhombic pads. “Spermatophore” (see Discussion) detected in two species: comma-shaped, lying with reservoir in vagina with longitudinally ribbed tail extending into mid penis. End of tail probably originally weakly attached to wall of penis [as in one specimen of
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<taxonomicName authority="radius (Preston, 1910)" authorityName="radius (Preston" authorityYear="1910" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="variety" species="radius" status="comb. nov." variety="radius">
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<emphasis italics="true" pageId="5" pageNumber="6">D. radius radius</emphasis>
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(
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<bibRefCitation author="Preston H. B." box="[279,444,808,834]" pageId="5" pageNumber="6" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston, 1910</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[460,593,807,834]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
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] but subsequently detached. Partially digested tail fragments present in bursa of one specimen of
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<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[521,679,843,869]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="pembensis" status="sp. nov.">
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<emphasis box="[521,679,843,869]" italics="true" pageId="5" pageNumber="6">D. pembensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[687,777,844,870]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="distribution">
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<paragraph blockId="5.[189,436,909,936]" box="[189,436,909,936]" pageId="5" pageNumber="6">
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<heading bold="true" box="[189,436,909,936]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis bold="true" box="[189,436,909,936]" pageId="5" pageNumber="6">Known distribution</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="5.[189,1398,957,1019]" pageId="5" pageNumber="6">
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Eastern
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<collectingCountry box="[292,372,958,984]" name="Kenya" pageId="5" pageNumber="6">Kenya</collectingCountry>
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, eastern
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<collectingCountry box="[489,598,957,983]" name="Tanzania" pageId="5" pageNumber="6">Tanzania</collectingCountry>
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, western
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<collectingCountry box="[723,817,957,983]" name="Uganda" pageId="5" pageNumber="6">Uganda</collectingCountry>
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, eastern
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<collectingCountry box="[935,1072,957,984]" name="Democratic Republic of the Congo" pageId="5" pageNumber="6">DR Congo</collectingCountry>
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,
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<collectingCountry box="[1091,1184,957,983]" name="Malawi" pageId="5" pageNumber="6">Malawi</collectingCountry>
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, eastern
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<collectingCountry box="[1302,1395,957,983]" name="Zambia" pageId="5" pageNumber="6">Zambia</collectingCountry>
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,
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<collectingCountry box="[189,347,993,1019]" name="Mozambique" pageId="5" pageNumber="6">Mozambique</collectingCountry>
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, north-eastern
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<collectingCountry box="[521,673,993,1019]" name="South Africa" pageId="5" pageNumber="6">South Africa</collectingCountry>
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,
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<collectingCountry box="[684,792,993,1019]" name="Comoros" pageId="5" pageNumber="6">Comoros</collectingCountry>
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archipelago (
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|
<figureCitation box="[952,1040,993,1019]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="5" pageNumber="6">Fig. 84</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection box="[189,308,1059,1133]" pageId="5" pageNumber="6" type="description">
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<paragraph blockId="5.[189,284,1059,1085]" box="[189,284,1059,1085]" pageId="5" pageNumber="6">
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|
<heading bold="true" box="[189,284,1059,1085]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis bold="true" box="[189,284,1059,1085]" pageId="5" pageNumber="6">Gender</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="5.[189,308,1107,1133]" box="[189,308,1107,1133]" pageId="5" pageNumber="6">Feminine.</paragraph>
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|
</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="discussion">
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<paragraph blockId="5.[189,511,1173,1199]" box="[189,511,1173,1199]" pageId="5" pageNumber="6">
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|
<heading bold="true" box="[189,511,1173,1199]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis bold="true" box="[189,511,1173,1199]" pageId="5" pageNumber="6">Comparison and remarks</emphasis>
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|
</heading>
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</paragraph>
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<paragraph blockId="5.[189,475,1220,1246]" box="[189,475,1220,1246]" pageId="5" pageNumber="6">See General Discussion.</paragraph>
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</subSubSection>
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</treatment>
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</document> |