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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.912.47719" ID-GBIF-Dataset="26396469-500e-42b8-acba-f6a22f652923" ID-PMC="PMC7040049" ID-Pensoft-Pub="1313-2970-912-65" ID-Pensoft-UUID="D1FDF644344C5B5B83E6024904E40A05" ID-PubMed="32123500" ID-ZooBank="4F361E38B2684B70944DD6B6FCC66542" ModsDocID="1313-2970-912-65" checkinTime="1581958640118" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2020" docId="177CACE4373A5708864BF61B0A5A9451" docLanguage="en" docName="ZooKeys 912: 65-124" docOrigin="ZooKeys 912" docSource="http://dx.doi.org/10.3897/zookeys.912.47719" docTitle="Theopea mouhoti Baly 1864" docType="treatment" docVersion="4" id="D1FDF644344C5B5B83E6024904E40A05" lastPageNumber="65" masterDocId="D1FDF644344C5B5B83E6024904E40A05" masterDocTitle="Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species" masterLastPageNumber="124" masterPageNumber="65" pageNumber="65" updateTime="1668168482312" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bezdek, Jan</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>912</mods:number>
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<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>124</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.912.47719</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.912.47719</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-912-65</mods:identifier>
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<treatment ID-GBIF-Taxon="162147242" LSID="urn:lsid:plazi:treatment:177CACE4373A5708864BF61B0A5A9451" httpUri="http://treatment.plazi.org/id/177CACE4373A5708864BF61B0A5A9451" lastPageNumber="65" pageId="0" pageNumber="65">
<subSubSection pageId="0" pageNumber="65" type="nomenclature">
<paragraph pageId="0" pageNumber="65">
<taxonomicName LSID="177CACE4-373A-5708-864B-F61B0A5A9451" authority="Baly, 1864" authorityName="Baly" authorityYear="1864" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea mouhoti" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mouhoti">Theopea mouhoti Baly, 1864</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">Figs 1G-I</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="reference_group">
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Baly" authorityYear="1864" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea mouhoti" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mouhoti">Theopea mouhoti</taxonomicName>
Baly, 1864: 238 (Thailand); Wilcox, 1973: 631 (catalogue); Kimoto, 1989: 200 (Laos); Staines &amp; Staines, 1999: 522 (catalogue).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="types">
<paragraph pageId="0" pageNumber="65">Types.</paragraph>
<paragraph pageId="0" pageNumber="65">Holotype ♂ (NHMUK, by monotypy), labeled: &quot;Type [p, w, circle label with red border] // Theopea / Mouhoti / Baly / Siam [h, g] // Baly Coll. [p, w]&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="materials_examined">
<paragraph pageId="0" pageNumber="65">Other specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">CAMBODIA</emphasis>
. 1♂ (NHMUK), Chautd;
<emphasis bold="true" pageId="0" pageNumber="65">LAOS</emphasis>
. Attapu: 5♂♂, 2♀♀ (NHMUK), Bolaven Plateau, 15 km SE of Ban Huangkong, Nong Lom (Lake) env.,
<geoCoordinate degrees="15" direction="north" minutes="02" orientation="latitude" precision="925" value="15.033334">15°02'N</geoCoordinate>
<geoCoordinate degrees="106" direction="east" minutes="35" orientation="longitude" precision="925" value="106.583336">106°35'E</geoCoordinate>
, 800 m, 18-30.IV.1999, leg. E. Jendek and O.
<normalizedToken originalValue="Šauša">Sausa</normalizedToken>
; Boli Kham Xai: 1♂, 3♀♀ (RBCN), Ban Nok env.,
<geoCoordinate degrees="18" direction="north" minutes="08.7" orientation="latitude" precision="92" value="18.145">18°08.7'N</geoCoordinate>
<geoCoordinate degrees="104" direction="east" minutes="28.1" orientation="longitude" precision="92" value="104.46833">104°28.1'E</geoCoordinate>
, Route no 8, 220 m, 9-14.V.1998, leg. E. Jendek, and O.
<normalizedToken originalValue="Šauša">Sausa</normalizedToken>
; Champasak: 4♀♀ (NHMB), Ban Nong Panouan env., 15°02'N 106°31-34'E, 770-800 m, leg. M. Geiser and D. Hauck; Khammouane: 2♂♂, 8♀♀ (NHMB), Ban Khoun Ngeun,
<geoCoordinate degrees="18" direction="north" minutes="07" orientation="latitude" precision="925" value="18.116667">18°07'N</geoCoordinate>
<geoCoordinate degrees="104" direction="east" minutes="29" orientation="longitude" precision="925" value="104.48333">104°29'E</geoCoordinate>
, 200 m, 24-29.IV.2001, leg.
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
; Vientiane: 1♂ (ZSM), III.-VI.1963 (identified by
<bibRefCitation author="Kimoto, S" journalOrPublisher="Esakia" pageId="0" pageNumber="65" pagination="1 - 241" refId="B10" refString="Kimoto, S, 1989. Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos, and Vietnam. IV. Gallerucinaea. Esakia 27: 1 - 241" title="Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos, and Vietnam. IV. Gallerucinaea." volume="27" year="1989">Kimoto (1989)</bibRefCitation>
;
<emphasis bold="true" pageId="0" pageNumber="65">THAILAND</emphasis>
. 4♂♂ (NHMUK); Loei: 1♀ (RBCN), Phu Rua N.P.,
<geoCoordinate degrees="17" direction="north" minutes="30" orientation="latitude" precision="925" value="17.5">17°30'N</geoCoordinate>
<geoCoordinate degrees="101" direction="east" minutes="21" orientation="longitude" precision="925" value="101.35">101°21'E</geoCoordinate>
, 6-9.IV.1999, leg. M.
<normalizedToken originalValue="Říha">Řiha</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="redescription">
<paragraph pageId="0" pageNumber="65">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="65">
Length 6.5-8.0 mm, width 2.4-3.1 mm. Body color (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">1G-I</figureCitation>
) reddish brown; meso- and metathoracic ventrites, abdomen, and legs dark brown or blackish brown; antenna black but antennomere XI reddish brown. Antennae filiform in males, but antennomere VI-VIII moderately swollen (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4A</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.7: 0.9: 1.0: 1.0: 1.0: 1.0: 0.9: 0.8: 1.1, length to width ratios of antennomeres I-XI 3.3: 1.5: 2.8: 3.1: 3.1: 2.8: 2.6: 2.6: 3.0: 3.0: 4.5; filiform in females (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.7: 0.9: 0.9: 0.9: 0.9: 0.9: 0.8: 0.8: 1.0, length to width ratios of antennomeres I-XI 3.2: 1.5: 2.8: 3.7: 3.6: 3.6: 3.6: 3.5: 3.8: 3.9: 4.6. Elytra elongate, parallel-sided, 1.8-2.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one distinct longitudinal ridge between two longitudinal rows of punctures. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4C-E</figureCitation>
) slender, 8.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; sides widest at middle, gradually narrowed towards basal 1/4, moderately and apically narrowed, apex with shallow notch; tectum well sclerotized, basally broadened, broader than aedeagus, with hollow area behind base of tectum; moderately curved in lateral view; ventral surface with deep notch from near apex, apically extending into basal opening, more approximate in apical 1/5; triangular sclerites small; internal sac with one median, elongate sclerite, 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, apically tapering from basal 1/3, apex acute, connected by short, broad sclerite at base; with one pair of elongate, longitudinal rows of stout setae, and one pair of short, longitudinal rows of stout setae dorsally and basally. Gonocoxae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4H</figureCitation>
) elongate, widest at apical 1/9, both gonocoxae combined from basal 1/7 to apical 1/7; apices narrowly rounded, each gonocoxa with eight setae along lateral margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4G</figureCitation>
) elongate and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4I</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
<caption doi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 4.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Baly" authorityYear="1864" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea mouhoti" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea mouhoti</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
Antenna, male
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Antenna, female
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Same, lateral view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Aedeagus, variation, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
Abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Gonocoxae
<emphasis bold="true" pageId="0" pageNumber="65">I</emphasis>
Spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="variation">
<paragraph pageId="0" pageNumber="65">Variations.</paragraph>
<paragraph pageId="0" pageNumber="65">
Males of Laos have apically tapering and truncate apex of aedeagus and its ventral surface have median notch not extending into basal opening (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Baly" authorityYear="1864" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea mouhoti" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea mouhoti</emphasis>
</taxonomicName>
Baly,
<taxonomicName lsidName="T. bicolor" pageId="0" pageNumber="65" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicolor</emphasis>
</taxonomicName>
Kimoto, and
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. are characterized by their reddish brown elytra.
<taxonomicName authorityName="Baly" authorityYear="1864" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea mouhoti" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea mouhoti</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">1G-I</figureCitation>
) can be easily separately from
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicolor</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">1A-F</figureCitation>
) by the reddish brown head, prothorax, and scutellum, and distinct and convex ridges on the elytra. Further, males of
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
have median elongate internal aedeagal sclerites without transverse rows of hair-like setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4C, D</figureCitation>
). This differs from those of
<taxonomicName lsidName="T. bicolor" pageId="0" pageNumber="65" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov., which possess a median elongate internal aedeagal sclerite with hair-like setae in transverse rows (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2C, D</figureCitation>
;
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic characters of Theopea bicoloroides sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure3" httpUri="https://binary.pensoft.net/fig/380658" pageId="0" pageNumber="65">3C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="distribution">
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="65">Cambodia, Laos, Thailand.</paragraph>
</subSubSection>
</treatment>
</document>