228 lines
30 KiB
XML
228 lines
30 KiB
XML
<document id="A93CAEE9DEC58BFCF7EF63B068EC8542" ID-DOI="10.5281/zenodo.13319951" ID-ISSN="0253-116X" ID-Zenodo-Dep="13319951" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722341515648" checkinUser="felipe" docAuthor="Assing, V." docDate="2006" docId="DE7E8782C331761D3003FCE1FC4A77BF" docLanguage="en" docName="LinzerbiolBeitr.2006.38.2.1129-1137.pdf" docOrigin="Linzer biologische Beiträge 38 (2)" docStyle="DocumentStyle:6260912E6386EE56293601EB6FBA9D25.4:LinzerbiolBeitr.2006-.journal_article.type2" docStyleId="6260912E6386EE56293601EB6FBA9D25" docStyleName="LinzerbiolBeitr.2006-.journal_article.type2" docStyleVersion="4" docTitle="Atheta tenebrarum Assing, 2006, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1137" masterDocId="2247FFFAC33776153157FFDAFF89740F" masterDocTitle="Three new species of Staphylinidae from Spain, with a new synonymy (Insecta: Coleoptera)" masterLastPageNumber="1137" masterPageNumber="1129" pageNumber="1129" updateTime="1723635347682" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="EAB3C69865F06ADE04A213D9BBAC90E0">
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<mods:title id="154CF265C2754E2B5EA42CB51CE270E5">Three new species of Staphylinidae from Spain, with a new synonymy (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="E19BA41ECD715A8D9CB7408F52D28CAD" type="personal">
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<mods:role id="D478361AA39D00877C40CAC9DB391771">
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<mods:roleTerm id="2490528308E82BCF2652F2523F59640E">Author</mods:roleTerm>
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<mods:namePart id="8B0D9F4FB27D7415B4F8B559499A6CC2">Assing, V.</mods:namePart>
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<mods:typeOfResource id="610EED84A7DAAE22F5C7B13EDDAB069F">text</mods:typeOfResource>
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<mods:titleInfo id="24DFB4A0C83D12558D1AF0715C1E8155">
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<mods:title id="94E10CD32EE8BD400856774E24E1DB8A">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="20E13EEC14836584162A627A101BD234">
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<mods:date id="B72C22CC78E28DACFAEF1B45317E9020">2006</mods:date>
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<mods:detail id="E21988A89216057AE45C7FC2AF0167B8" type="pubDate">
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<mods:number id="3063DB0947AA05E5BF2F93FE749674C0">2006-12-29</mods:number>
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</mods:detail>
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<mods:detail id="3E21D39C45442F780AE7C80ABFBA7CD5" type="volume">
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<mods:number id="6A8CC5E99F4FE1751F4B35103E224938">38</mods:number>
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</mods:detail>
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<mods:detail id="EFE39336EFA50A5A54AF87D7DA02C2A6" type="issue">
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<mods:number id="A408B33BE35544ABE53EEB6E55CBAE3F">2</mods:number>
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</mods:detail>
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<mods:extent id="21AFDD6A6DF3CD272AB72AF7886208BA" unit="page">
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<mods:start id="7B3BC6E03A27A1BF115E6DBB6215A371">1129</mods:start>
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<mods:end id="E1A3195B7A52E301361F64FAE1E3D501">1137</mods:end>
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<mods:classification id="DAF63FB4EF8063CE2B0B7F31C9D2C57A">journal article</mods:classification>
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<mods:identifier id="BD60E4C6D8112D7BCAC2B8BB30701C6C" type="DOI">10.5281/zenodo.13319951</mods:identifier>
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<mods:identifier id="24BD09BB7ABDA0CA5870EE0804BA9D55" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="0220692E2DDFC651675F27E138F38D1D" type="Zenodo-Dep">13319951</mods:identifier>
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</mods:mods>
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<treatment id="DE7E8782C331761D3003FCE1FC4A77BF" LSID="urn:lsid:plazi:treatment:DE7E8782C331761D3003FCE1FC4A77BF" httpUri="http://treatment.plazi.org/id/DE7E8782C331761D3003FCE1FC4A77BF" lastPageId="8" pageId="6">
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<subSubSection id="1ECD651FC33176133003FCE1FCCF775E" box="[340,838,827,849]" pageId="6" type="nomenclature">
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<paragraph id="56683694C33176133003FCE1FCCF775E" blockId="6.[340,838,827,849]" box="[340,838,827,849]" pageId="6">
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<heading id="0D2081F8C33176133003FCE1FCCF775E" box="[340,838,827,849]" fontSize="8" level="2" pageId="6" reason="2">
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<emphasis id="64A3EA86C33176133003FCE1FCCF775E" bold="true" box="[340,838,827,849]" pageId="6">
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<taxonomicName id="91D74D17C33176133003FCE1FD90775E" ID-CoL="688QK" authorityName="Assing" authorityYear="2006" box="[340,537,827,849]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tenebrarum" status="sp. nov.">
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<emphasis id="64A3EA86C33176133003FCE1FD90775E" bold="true" box="[340,537,827,849]" italics="true" pageId="6">Atheta tenebrarum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7F9057FDC33176133377FCE1FDE5775E" box="[544,620,827,849]" pageId="6" rank="species">nov.sp.</taxonomicNameLabel>
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(
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<figureCitation id="CEEC2A11C3317613332EFCE1FD65775E" box="[633,748,827,849]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Figs 22-33</figureCitation>
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;
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<figureCitation id="CEEC2A11C331761333AFFCE1FCB7775E" box="[760,830,827,849]" captionStart="Map 1" captionStartId="3.[340,385,1351,1371]" captionTargetBox="[340,1247,707,1340]" captionTargetId="figure-370@3.[340,1247,707,1340]" captionTargetPageId="3" captionText="Map 1: Distributions of Phloeocharis baenai nov.sp. (open circle), Geostiba jaenica nov.sp. (filled circles), and Atheta tenebrarum nov.sp. (square) in southern Spain." figureDoi="http://doi.org/10.5281/zenodo.13319957" httpUri="https://zenodo.org/record/13319957/files/figure.png" pageId="6">Map 1</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1ECD651FC331761D3003FCBBFC4A77BF" lastPageId="8" pageId="6" type="materials_examined">
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<paragraph id="56683694C33176133003FCBBFE1F77AD" blockId="6.[340,1247,864,930]" pageId="6">
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<materialsCitation id="E6BF3CC9C33176133003FCBBFBD8777A" box="[340,1105,864,885]" collectingDate="2006-03-18" collectorName="La Iruela" location="Cueva Secreta del Sagreo" pageId="6" specimenCount="1" typeStatus="holotype">
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<typeStatus id="896C8836C33176133003FCBBFE20777A" box="[340,425,865,885]" pageId="6" type="holotype">Holotype</typeStatus>
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3: E - Jaén,
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<collectorName id="FB225342C3317613337DFCBBFD09777A" box="[554,640,865,885]" pageId="6">La Iruela</collectorName>
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,
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<location id="5308604FC331761333DCFCBBFCF2777A" LSID="urn:lsid:plazi:treatment:DE7E8782C331761D3003FCE1FC4A77BF:5308604FC331761333DCFCBBFCF2777A" box="[651,891,865,885]" name="Cueva Secreta del Sagreo" pageId="6">Cueva Secreta del Sagreo</location>
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,
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<date id="22691054C331761332D1FCBBFC67777A" box="[902,1006,865,885]" pageId="6" value="2006-03-18">
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<collectingDate id="322DE9BCC331761332D1FCBBFC67777A" box="[902,1006,865,885]" pageId="6" value="2006-03-18">18.III.2006</collectingDate>
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</date>
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, leg. GEV
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</materialsCitation>
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/
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<typeStatus id="896C8836C33176133531FCBBFB4C777A" box="[1126,1221,865,885]" pageId="6" type="holotype">Holotypus</typeStatus>
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3
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<taxonomicName id="91D74D17C33176133003FCA2FD887783" authorityName="Assing" authorityYear="2006" box="[340,513,888,908]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tenebrarum" status="sp. nov.">
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<emphasis id="64A3EA86C33176133003FCA2FD887783" box="[340,513,888,908]" italics="true" pageId="6">Atheta tenebrarum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7F9057FDC3317613335FFCA2FDBA7783" box="[520,563,888,908]" pageId="6" rank="species">sp.n.</taxonomicNameLabel>
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det.
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<collectionCode id="30C6AE51C33176133335FCA2FDFC7783" box="[610,629,888,908]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
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. Assing 2006 (cAss).
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<typeStatus id="896C8836C33176133214FCA2FC167783" box="[835,927,888,908]" pageId="6" type="paratype">Paratypes</typeStatus>
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: 233,
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<specimenCount id="40D1FD1DC331761332B0FCA2FB8C779E" box="[999,1029,888,913]" count="1" pageId="6" type="female">1♀</specimenCount>
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: same data as
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<typeStatus id="896C8836C331761335C7FCA2FB567783" box="[1168,1247,888,908]" pageId="6" type="holotype">holotype</typeStatus>
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(cAss).
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</paragraph>
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<paragraph id="56683694C33176133003FC68FBD57010" blockId="6.[340,1248,945,1055]" pageId="6">
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D e s c r i p t i o n: Species of moderately large size,
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<quantity id="912F9B71C331761332C6FC6BFB8277C8" box="[913,1035,945,967]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="6" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8-4.2 mm</quantity>
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; facies as in
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<figureCitation id="CEEC2A11C331761335C7FC6BFB5577C8" box="[1168,1244,945,967]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 22</figureCitation>
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. Coloration: head dark brown; pronotum reddish brown; elytra yellowish to yellowish brown; abdomen reddish brown, with the preapical segments somewhat infuscate; legs yellowish; antennae dark brown, with the basal 3-4 antennomeres yellowish.
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</paragraph>
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<paragraph id="56683694C33176133003FBF4FB3270D9" blockId="6.[340,1248,1070,1238]" pageId="6">
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Head slightly wider than long; eyes of normal size (
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<figureCitation id="CEEC2A11C3317613322FFBF4FC4F704B" box="[888,966,1070,1092]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 24</figureCitation>
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), approximately as long as postocular region in dorsal view, or slightly longer; postgenae finely carinate ventrally; integument with distinct microreticulation; puncturation rather dense and fine, barely visible in the microsculpture (
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<figureCitation id="CEEC2A11C331761333D2FB5CFD587093" box="[645,721,1158,1180]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 23</figureCitation>
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). Antennae slender; antennomere III slightly longer than II, IV weakly oblong, V-IX approximately as long as wide, X weakly transverse, and XI approximately as long as the combined length of IX-X or slightly longer (
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<figureCitation id="CEEC2A11C33176133532FB1AFB3970D9" box="[1125,1200,1216,1238]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 25</figureCitation>
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).
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</paragraph>
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<paragraph id="56683694C33176133003FB3CFDD07181" blockId="6.[340,1248,1254,1422]" pageId="6">
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Pronotum approximately 1.3 times as wide as head and 1.35 times as wide as long; maximal width a short distance anterior to middle; posterior angles obtuse, weakly marked; puncturation and microreticulation similar to those of head (
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<figureCitation id="CEEC2A11C33176133562FAFAFB0C7139" box="[1077,1157,1312,1334]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 23</figureCitation>
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); pubescence directed cephalad in anterior 1/4 of midline, caudad in posterior 3/4, and diagonally latero-caudad in lateral area; marginal setae short, slightly shorter than maximal width of antennomere III.
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</paragraph>
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<paragraph id="56683694C33176133003FA47FBDE71E1" blockId="6.[340,1248,1437,1518]" pageId="6">
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Elytra about 1.25 times as wide and at suture almost as long as pronotum; microsculpture and puncturation similar to those of head and pronotum; pubescence short and depressed (
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<figureCitation id="CEEC2A11C3317613300BFA02FE2E71E1" box="[348,423,1496,1518]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 23</figureCitation>
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). Hind wings fully developed. Tibiae without long prominent setae.
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</paragraph>
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<paragraph id="56683694C33176133003FA27FC327241" blockId="6.[340,1248,1533,1614]" pageId="6">
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Abdomen slightly narrower than elytra, widest at segments IV/
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<collectionCode id="30C6AE51C331761332BAFA27FC76721C" box="[1005,1023,1533,1555]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" type="Museum">V</collectionCode>
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(
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<figureCitation id="CEEC2A11C33176133547FA27FBE9721C" box="[1040,1120,1533,1555]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 22</figureCitation>
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); tergal surfaces with fine transverse microscupture everywhere; puncturation relatively sparse and very fine, barely noticeable; pubescence dark and depressed.
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</paragraph>
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<paragraph id="56683694C33176133003F987FE1672A2" blockId="6.[340,1248,1629,1709]" pageId="6">
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3: posterior margin of tergite VIII weakly concave and weakly serrate (
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<figureCitation id="CEEC2A11C3317613356BF987FB00727C" box="[1084,1161,1629,1651]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 26</figureCitation>
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), that of sternite VIII convex (
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<figureCitation id="CEEC2A11C33176133366F9A0FDF6729F" box="[561,639,1658,1680]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 27</figureCitation>
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); median lobe of aedeagus of highly distinctive shape (
|
|
<figureCitation id="CEEC2A11C331761335E1F9A0FE1A72A2" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 28-30</figureCitation>
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).
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</paragraph>
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<paragraph id="56683694C33176133003F919FE2A7302" blockId="6.[340,1248,1725,1805]" pageId="6">
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<specimenCount id="40D1FD1DC33176133003F919FEE072D7" box="[340,361,1731,1752]" count="1" pageId="6" type="female">♀</specimenCount>
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: posterior margin of tergite VIII in the middle weakly concave (
|
|
<figureCitation id="CEEC2A11C33176133551F967FBDB72DC" box="[1030,1106,1725,1747]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 31</figureCitation>
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), that of sternite VIII broadly convex and with long thin marginal setae (
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<figureCitation id="CEEC2A11C33176133291F900FB9C72FF" box="[966,1045,1754,1776]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 32</figureCitation>
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); spermatheca as in
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<figureCitation id="CEEC2A11C33176133003F92DFE167302" box="[340,415,1783,1805]" captionStart="Figs 22-33" captionStartId="7.[340,380,1613,1633]" captionTargetBox="[340,1240,425,1579]" captionTargetId="figure-86@7.[340,1246,653,1129]" captionTargetPageId="7" captionText="Figs 22-33: Atheta tenebrarum nov.sp.: (22) habitus; (23) forebody; (24) head in lateral view; (25) antenna; (26) 3 tergite VIII; (27) posterior margin of 3 sternite VIII; (28-30) median lobe of aedeagus in lateral and in ventral view; (31) ♀ tergite VIII; (32) ♀ sternite VIII; (33) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="6">Fig. 33</figureCitation>
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.
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</paragraph>
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<paragraph id="56683694C33076123255FEBFFCBB7575" blockId="7.[770,818,357,378]" box="[770,818,357,378]" pageId="7">1136</paragraph>
|
|
<caption id="02A8661CC33076123003F997FC2672AA" ID-DOI="http://doi.org/10.5281/zenodo.13319961" ID-Zenodo-Dep="13319961" httpUri="https://zenodo.org/record/13319961/files/figure.png" pageId="7" startId="7.[340,380,1613,1633]" targetBox="[340,1240,425,1579]" targetPageId="7" targetType="figure">
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<paragraph id="56683694C33076123003F997FC2672AA" blockId="7.[340,1249,1613,1701]" pageId="7">
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<emphasis id="64A3EA86C33076123003F997FE31726E" bold="true" box="[340,440,1613,1633]" pageId="7">Figs 22-33</emphasis>
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|
:
|
|
<taxonomicName id="91D74D17C33076123092F997FDF8726E" authorityName="Assing" authorityYear="2006" box="[453,625,1613,1633]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tenebrarum" status="sp. nov.">
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|
<emphasis id="64A3EA86C33076123092F997FDF8726E" box="[453,625,1613,1633]" italics="true" pageId="7">Atheta tenebrarum</emphasis>
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|
</taxonomicName>
|
|
<taxonomicNameLabel id="7F9057FDC33076123320F997FD34726E" box="[631,701,1613,1633]" pageId="7" rank="species">nov.sp.</taxonomicNameLabel>
|
|
: (
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|
<emphasis id="64A3EA86C3307612339AF997FD6D726E" bold="true" box="[717,740,1613,1633]" pageId="7">22</emphasis>
|
|
) habitus; (
|
|
<emphasis id="64A3EA86C3307612321FF997FCD6726E" bold="true" box="[840,863,1613,1633]" pageId="7">23</emphasis>
|
|
) forebody; (
|
|
<emphasis id="64A3EA86C33076123284F997FC63726E" bold="true" box="[979,1002,1613,1633]" pageId="7">24</emphasis>
|
|
) head in lateral view; (
|
|
<emphasis id="64A3EA86C33076123596F997FB51726E" bold="true" box="[1217,1240,1613,1633]" pageId="7">25</emphasis>
|
|
) antenna; (
|
|
<emphasis id="64A3EA86C330761230E5F9BEFE407277" bold="true" box="[434,457,1636,1656]" pageId="7">26</emphasis>
|
|
) 3 tergite VIII; (
|
|
<emphasis id="64A3EA86C3307612332EF9BEFD197277" bold="true" box="[633,656,1636,1656]" pageId="7">27</emphasis>
|
|
) posterior margin of 3 sternite VIII; (
|
|
<emphasis id="64A3EA86C3307612355EF9BEFBB77277" bold="true" box="[1033,1086,1636,1656]" pageId="7">28-30</emphasis>
|
|
) median lobe of aedeagus in lateral and in ventral view; (
|
|
<emphasis id="64A3EA86C3307612339FF9A0FD567281" bold="true" box="[712,735,1658,1678]" pageId="7">31</emphasis>
|
|
) ♀ tergite VIII; (
|
|
<emphasis id="64A3EA86C330761232D5F9A0FC107281" bold="true" box="[898,921,1658,1678]" pageId="7">32</emphasis>
|
|
) ♀ sternite VIII; (
|
|
<emphasis id="64A3EA86C33076123513F9A0FBD27281" bold="true" box="[1092,1115,1658,1678]" pageId="7">33</emphasis>
|
|
) spermatheca. Scales: 22-23: 1.0 mm; 24-27, 31-32: 0.5 mm; 28-30, 33: 0.2 mm.
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</paragraph>
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</caption>
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<paragraph id="56683694C33F761D3255FEBFFCBB7575" blockId="8.[770,818,357,378]" box="[770,818,357,378]" pageId="8">1137</paragraph>
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|
<paragraph id="56683694C33F761D3003FE75FD7475EF" blockId="8.[340,1247,430,480]" pageId="8">E t y m o l o g y: The name (Lat.: noun, genitive: of the darkness) alludes to the fact that this species was discovered in a cave.</paragraph>
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|
<paragraph id="56683694C33F761D3003FE34FDFF7713" blockId="8.[340,1248,494,796]" pageId="8">
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|
C o m p a r a t i v e n o t e s:
|
|
<taxonomicName id="91D74D17C33F761D33E5FE34FCFB760B" authorityName="Assing" authorityYear="2006" box="[690,882,494,516]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="tenebrarum">
|
|
<emphasis id="64A3EA86C33F761D33E5FE34FCFB760B" box="[690,882,494,516]" italics="true" pageId="8">Atheta tenebrarum</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from all its congeners especially by the highly distinctive morphology of the median lobe of the aedeagus and also by the shape of the spermatheca. Based on external morphology (pronotal pubescence pattern, chaetotaxy) and on the primary and secondary sexual characters, the new species is closely related to
|
|
<taxonomicName id="91D74D17C33F761D33E3FD85FC14767B" authority="(KRAATZ)" baseAuthorityName="KRAATZ" box="[692,925,606,628]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="trinotata">
|
|
<emphasis id="64A3EA86C33F761D33E3FD85FCA5767B" box="[692,812,607,628]" italics="true" pageId="8">A. trinotata</emphasis>
|
|
(KRAATZ)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="91D74D17C33F761D32FBFD85FBE1767B" authority="SHARP" authorityName="SHARP" box="[940,1128,606,628]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
|
|
<emphasis id="64A3EA86C33F761D32FBFD85FB92767B" box="[940,1051,606,628]" italics="true" pageId="8">A. hybrida</emphasis>
|
|
SHARP
|
|
</taxonomicName>
|
|
, and allied species, its closest relative being
|
|
<taxonomicName id="91D74D17C33F761D33E6FDA1FC97769F" box="[689,798,634,656]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
|
|
<emphasis id="64A3EA86C33F761D33E6FDA1FC97769F" box="[689,798,634,656]" italics="true" pageId="8">A. hybrida</emphasis>
|
|
</taxonomicName>
|
|
, as is suggested particularly by the similar internal structures and shape of the median lobe of the aedeagus. From both
|
|
<taxonomicName id="91D74D17C33F761D353EFD4DFB6976A3" box="[1129,1248,663,684]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="trinotata">
|
|
<emphasis id="64A3EA86C33F761D353EFD4DFB6976A3" box="[1129,1248,663,684]" italics="true" pageId="8">A. trinotata</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="91D74D17C33F761D30D7FD69FE6576C7" box="[384,492,690,712]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
|
|
<emphasis id="64A3EA86C33F761D30D7FD69FE6576C7" box="[384,492,690,712]" italics="true" pageId="8">A. hybrida</emphasis>
|
|
</taxonomicName>
|
|
, it is separated also by the longer antennae, the relatively larger eyes, the paler and broader pronotum, the uniformly yellowish elytra, the different coloration of the abdomen (in
|
|
<taxonomicName id="91D74D17C33F761D3359FD31FD01770F" box="[526,648,747,768]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="trinotata">
|
|
<emphasis id="64A3EA86C33F761D3359FD31FD01770F" box="[526,648,747,768]" italics="true" pageId="8">A. trinotata</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="91D74D17C33F761D3394FD31FCBA770F" box="[707,819,746,768]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hybrida">
|
|
<emphasis id="64A3EA86C33F761D3394FD31FCBA770F" box="[707,819,746,768]" italics="true" pageId="8">A. hybrida</emphasis>
|
|
</taxonomicName>
|
|
almost uniformly blackish), and by the secondary sexual characters.
|
|
</paragraph>
|
|
<paragraph id="56683694C33F761D3003FCF1FC4A77BF" blockId="8.[340,1248,810,944]" pageId="8">
|
|
D i s t r i b u t i o n a n d b i o n o m i c s: The species is currently known only from one cave near La Iruela in the Sierra de Cazorla, eastern Jaén province (altitude: slightly less than
|
|
<quantity id="912F9B71C33F761D3352FCB8FDDC7777" box="[517,597,866,888]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="8" unit="m" value="1200.0">1200 m</quantity>
|
|
). The
|
|
<taxonomicName id="91D74D17C33F761D33C2FCB8FCA97777" box="[661,800,866,888]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
|
|
from this cave were collected using various methods, so that the particular circumstances, under which the
|
|
<specimenCount id="40D1FD1DC33F761D328FFCA4FBBF779B" box="[984,1078,894,916]" count="4" pageId="8" type="generic" typeStatus="type">four type</specimenCount>
|
|
specimens of
|
|
<taxonomicName id="91D74D17C33F761D359EFCA5FE4377BF" authorityName="Assing" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="tenebrarum">
|
|
<emphasis id="64A3EA86C33F761D359EFCA5FE4377BF" italics="true" pageId="8">A. tenebrarum</emphasis>
|
|
</taxonomicName>
|
|
were found, are unknown (
|
|
<collectionCode id="30C6AE51C33F761D33B6FC40FCA377BF" box="[737,810,922,944]" pageId="8">BAENA</collectionCode>
|
|
, pers. comm.).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |