treatments-xml/data/C6/39/87/C63987A74C6CE567E7AF4701FF73FD50.xml
2024-06-21 12:51:07 +02:00

880 lines
114 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A245796644249F799481ADD0EFF9D8B7" ID-CLB-Dataset="31704" ID-DOI="10.11646/zootaxa.4341.1.5" ID-GBIF-Dataset="c74b6c1b-3efc-4630-9825-adf109a8fa37" ID-ISSN="1175-5326" ID-Zenodo-Dep="1038746" ID-ZooBank="481167BF-B57C-4BE4-BAAD-4396BF8F9336" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509343700906" checkinUser="plazi" docAuthor="Gill, Anthony C., Tea, Yi-Kai &amp; Senou, Hiroshi" docDate="2017" docId="C63987A74C6CE567E7AF4701FF73FD50" docLanguage="en" docName="zootaxa.4341.1.5.pdf" docOrigin="Zootaxa 4341 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudanthias tequila Gill &amp; Tea &amp; Senou 2017, new species" docType="treatment" docVersion="8" lastPageNumber="75" masterDocId="3A00FFDF4C6DE56FE738431DFFB0FFA0" masterDocTitle="Pseudanthias tequila, a new species of anthiadine serranid from the Ogasawara and Mariana Islands" masterLastPageNumber="76" masterPageNumber="67" pageNumber="68" updateTime="1702000979995" updateUser="plazi">
<mods:mods id="D6853EAD82F592BDA57827EC74081D13" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7845ABCB65B980D5703D485629081F2A">
<mods:title id="1ABEBC1F1C8878B644F6B0216BFD8D31">Pseudanthias tequila, a new species of anthiadine serranid from the Ogasawara and Mariana Islands</mods:title>
</mods:titleInfo>
<mods:name id="F6924BD77229DC3272731CA8A60FD0A1" type="personal">
<mods:role id="B2CC1F1586EB8D9DA106AF1590E6181C">
<mods:roleTerm id="079FC18F8831827B3D1723A32B2A8089">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F125F1526661FA3E9152E483718F4A9E">Gill, Anthony C.</mods:namePart>
</mods:name>
<mods:name id="F5982BA73A56B6589175CC645417CC3C" type="personal">
<mods:role id="37B91F38628C3718A0301CFFEA709725">
<mods:roleTerm id="AB673B045F9B2F25CBA2A62D9DA0210E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C79C4DAD1ED2158315555A61D866131B">Tea, Yi-Kai</mods:namePart>
</mods:name>
<mods:name id="7E482D3FB0E0FC825BC79595D2F0492D" type="personal">
<mods:role id="17CBAB7C61FF99F564EB2BF839E773A2">
<mods:roleTerm id="CCFAAF30C7FDBB935A31EDF9C3E0406F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A0002743A2163871BFEEAF7643AFEAAD">Senou, Hiroshi</mods:namePart>
</mods:name>
<mods:typeOfResource id="017AB071558CCEDB9344082C693D1201">text</mods:typeOfResource>
<mods:relatedItem id="449E32DE69574415C2E5F3AB2DD3C582" type="host">
<mods:titleInfo id="C25A70F8A26F89BCA83C2F74DBF9BFF8">
<mods:title id="D0AC7D59497CA2D07FEFB5FEBE58EDA0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="60A851A1B1FD80A321BCFDC5C45375CB">
<mods:date id="2AE94C6752211638611DB3ABD62FAD08">2017</mods:date>
<mods:detail id="CF6930CE99EE6ECB913E804125FC86AD" type="pubDate">
<mods:number id="32F15E583F6EAA88A02A07C885B356EA">2017-10-30</mods:number>
</mods:detail>
<mods:detail id="B1A95BFCE4FA62B11E4ABCFDAE1FF8CA" type="volume">
<mods:number id="73757D9BD5E81CBD507E4F076D90B5B5">4341</mods:number>
</mods:detail>
<mods:detail id="4C205F66152322231C8A730BE86D9E6C" type="issue">
<mods:number id="168D2EBD207D48E0388C8A4871255F8C">1</mods:number>
</mods:detail>
<mods:extent id="67693CA89ABEF6F787B12DC40013FE05" unit="page">
<mods:start id="C91721EE85C3F167F77BF8DFFF545223">67</mods:start>
<mods:end id="6FCED165D52A5563822C045AF1A78A12">76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7A2C510DE96870C5343163AC76CD76F5">journal article</mods:classification>
<mods:identifier id="5FA776F7F23C0D0033B79EAC530CE9F3" type="CLB-Dataset">31704</mods:identifier>
<mods:identifier id="23ABF3324EDD261AD2A591F06AC7F0EB" type="DOI">10.11646/zootaxa.4341.1.5</mods:identifier>
<mods:identifier id="414E2418C160F566A4B9586116465498" type="GBIF-Dataset">c74b6c1b-3efc-4630-9825-adf109a8fa37</mods:identifier>
<mods:identifier id="08D9AEC5B22AAE076C87E0E8CD305460" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="55A9999BCF6D4777737C9B3FEC433E1B" type="Zenodo-Dep">1038746</mods:identifier>
<mods:identifier id="1810B30D0C0238A38A47372C43502A87" type="ZooBank">481167BF-B57C-4BE4-BAAD-4396BF8F9336</mods:identifier>
</mods:mods>
<treatment id="C63987A74C6CE567E7AF4701FF73FD50" ID-DOI="http://doi.org/10.5281/zenodo.6041597" ID-GBIF-Taxon="135477071" ID-Zenodo-Dep="6041597" LSID="urn:lsid:plazi:treatment:C63987A74C6CE567E7AF4701FF73FD50" httpUri="http://treatment.plazi.org/id/C63987A74C6CE567E7AF4701FF73FD50" lastPageId="8" lastPageNumber="75" pageId="1" pageNumber="68">
<subSubSection id="068A653A4C6CE56EE7AF4701FECCFB3F" pageId="1" pageNumber="68" type="nomenclature">
<paragraph id="4E2F36B14C6CE56EE7AF4701FD9BFB96" blockId="1.[151,660,1052,1183]" box="[151,555,1052,1078]" pageId="1" pageNumber="68">
<heading id="156781DD4C6CE56EE7AF4701FD9BFB96" bold="true" box="[151,555,1052,1078]" fontSize="11" level="1" pageId="1" pageNumber="68" reason="1">
<emphasis id="7CE4EAA34C6CE56EE7AF4701FD9BFB96" bold="true" box="[151,555,1052,1078]" pageId="1" pageNumber="68">
<taxonomicName id="89904D324C6CE56EE7AF4701FE24FB96" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[151,404,1052,1078]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C6CE56EE7AF4701FE24FB96" bold="true" box="[151,404,1052,1078]" italics="true" pageId="1" pageNumber="68">Pseudanthias tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C6CE56EE6A34701FD9BFB96" box="[411,555,1052,1078]" pageId="1" pageNumber="68" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4E2F36B14C6CE56EE7AF4722FECCFB3F" blockId="1.[151,660,1052,1183]" pageId="1" pageNumber="68">
New standard Japanese name: Bonin-hanadai English common name: Cave anthias
<figureCitation id="D6AB2A344C6CE56EE7AF479BFEAEFB3F" box="[151,286,1158,1183]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,755,777]" captionStartId-1="3.[151,250,1370,1392]" captionStartId-2="4.[151,250,970,992]" captionStartId-3="5.[151,250,1242,1264]" captionStartId-4="6.[151,250,1147,1169]" captionStartId-5="7.[151,250,1966,1988]" captionStartId-6="8.[151,250,1650,1672]" captionTargetBox-0="[340,1247,195,733]" captionTargetBox-1="[340,1247,857,1348]" captionTargetBox-2="[151,1436,193,948]" captionTargetBox-3="[151,1436,193,1220]" captionTargetBox-4="[151,1436,301,1126]" captionTargetBox-5="[151,1436,981,1945]" captionTargetBox-6="[177,1393,807,1627]" captionTargetId-0="figure@3.[340,1247,195,733]" captionTargetId-1="figure@3.[340,1247,857,1348]" captionTargetId-2="figure@4.[151,1436,193,948]" captionTargetId-3="figure@5.[151,1436,193,1220]" captionTargetId-4="figure@6.[151,1436,301,1126]" captionTargetId-5="figure@7.[151,1436,981,1945]" captionTargetId-6="figure@8.[167,1403,807,1637]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURE 1. Pseudanthias tequila n. sp., KPM-NI 3759, 59.3 mm SL male holotype, Ototo-jima Island, Chichi-jima Islands, Ogasawara Islands, Japan. Photo (KPM-NR 51086 B) by H. Senou." captionText-1="FIGURE 2. Pseudanthias tequila n. sp., KPM-NI 3758, 31.5 mm SL female paratype, Ototo-jima Island, Chichi-jima Islands, Ogasawara Islands, Japan. Photo (KPM-NR 51087 B) by H. Senou." captionText-2="FIGURE 3. Pseudanthias tequila n. sp., KPM-NI 3759, 59.3 mm SL, holotype, x-radiograph. Radiograph (KPM-NR 51086 C) by H. Senou." captionText-3="FIGURE 4. Males of selected Pseudanthias species: A) P. tequila n. sp., Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan (KPM-NR 179391 A, photo by O. Morishita); B) P. tequila n. sp., Tinian, Mariana Islands (photo by N. Tsuji); C) P. pulcherrimus, aquarium specimen (not retained) from the Maldives (photo by Y. K. Tea); D) P. randalli, Kwajalein, Marshall Islands (photo by J. Johnson); E) P. flavicauda, Fiji (photo by J. E. Randall); F) P. sp., Kirimati, Kiribati (photo by D. J. Linehan)." captionText-4="FIGURE 5. Pseudanthias tequila n. sp., female, Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan. Photo by S. Kobayashi." captionText-5="FIGURE 6. Group of Pseudanthias tequila n. sp., Tinian, Mariana Islands, Micronesia. Photo by N. Tsuji." captionText-6="FIGURE 7. Distribution records for Pseudanthias tequila." httpUri-0="https://zenodo.org/record/1038748/files/figure.png" httpUri-1="https://zenodo.org/record/1038750/files/figure.png" httpUri-2="https://zenodo.org/record/1038752/files/figure.png" httpUri-3="https://zenodo.org/record/1038754/files/figure.png" httpUri-4="https://zenodo.org/record/1038756/files/figure.png" httpUri-5="https://zenodo.org/record/1038758/files/figure.png" httpUri-6="https://zenodo.org/record/1038760/files/figure.png" pageId="1" pageNumber="68">Figures 17</figureCitation>
,
<tableCitation id="0312030A4C6CE56EE612479BFECCFB3F" box="[298,380,1158,1183]" captionStart="TABLE 1" captionStartId="2.[151,239,544,567]" captionText="TABLE 1. Morphometric values for Pseudanthias tequila expressed as percentage SL." pageId="1" pageNumber="68">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6CE56EE7AF47D3FCA2FA82" pageId="1" pageNumber="68" type="reference_group">
<paragraph id="4E2F36B14C6CE56EE7AF47D3FDEFFAA3" blockId="1.[151,1388,1230,1436]" pageId="1" pageNumber="68">
<treatmentCitationGroup id="6E80119F4C6CE56EE7AF47D3FDEFFAA3" pageId="1" pageNumber="68">
<taxonomicName id="89904D324C6CE56EE7AF47D3FEC7FB44" ID-CoL="4NKPH" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[151,375,1230,1252]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C6CE56EE7AF47D3FEC7FB44" box="[151,375,1230,1252]" italics="true" pageId="1" pageNumber="68">Pseudanthias randalli</emphasis>
</taxonomicName>
(
<emphasis id="7CE4EAA34C6CE56EE6BE47CDFE1CFB45" box="[390,428,1232,1253]" italics="true" pageId="1" pageNumber="68">non</emphasis>
<bibRefCitation id="2A014B404C6CE56EE68A47D3FD18FB44" author="Lubbock" box="[434,680,1230,1252]" pageId="1" pageNumber="68" refString="Lubbock, R. &amp; Allen, G. R. (1978) A distinctive new Anthias (Teleostei: Serranidae) from the western Pacific. Records of the Western Australian Museum, 6, 259 - 268." type="journal article" year="1978">Lubbock &amp; Allen, 1978</bibRefCitation>
);
<treatmentCitation id="CF3110A04C6CE56EE58247D2FC5DFB44" author="Myers" box="[698,1005,1230,1252]" page="316" pageId="1" pageNumber="68" year="1980">
<bibRefCitation id="2A014B404C6CE56EE58247D2FC0DFB44" author="Myers" box="[698,957,1230,1252]" pageId="1" pageNumber="68" refString="Myers, R. F. &amp; Shepard, J. W. (1980) New records of fishes from Guam, with notes on the ichthyofauna of the southern Marianas. Micronesica, 16, 304 - 347." type="journal article" year="1980">Myers &amp; Shephard, 1980</bibRefCitation>
: 316
</treatmentCitation>
(Blue Hole, Orote Peninsula, Guam).
<taxonomicName id="89904D324C6CE56EE7AF47F0FEAFFAA3" ID-CoL="6YRF" authorityName="Bleeker" authorityYear="1871" box="[151,287,1261,1283]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6CE56EE7AF47F0FEAFFAA3" box="[151,287,1261,1283]" italics="true" pageId="1" pageNumber="68">Pseudanthias</emphasis>
</taxonomicName>
sp.;
<treatmentCitation id="CF3110A04C6CE56EE66847F3FDB3FAA3" author="Myers" box="[336,515,1261,1283]" page="140" pageId="1" pageNumber="68" year="1988">
<bibRefCitation id="2A014B404C6CE56EE66847F3FE63FAA3" author="Myers" box="[336,467,1261,1283]" pageId="1" pageNumber="68" refString="Myers, R. F. (1988) An annotated checklist of the fishes of the Mariana Islands. Micronesica, 21, 115 - 180." type="journal article" year="1988">Myers, 1988</bibRefCitation>
: 140
</treatmentCitation>
(Guam).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E2F36B14C6CE56EE7AF4611FCA2FA82" blockId="1.[151,1388,1230,1436]" box="[151,786,1292,1314]" pageId="1" pageNumber="68">
<treatmentCitationGroup id="6E80119F4C6CE56EE7AF4611FCA2FA82" box="[151,786,1292,1314]" pageId="1" pageNumber="68">
<taxonomicName id="89904D324C6CE56EE7AF4611FEAFFA82" ID-CoL="6YRF" authorityName="Bleeker" authorityYear="1871" box="[151,287,1292,1314]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6CE56EE7AF4611FEAFFA82" box="[151,287,1292,1314]" italics="true" pageId="1" pageNumber="68">Pseudanthias</emphasis>
</taxonomicName>
sp. 3;
<treatmentCitation id="CF3110A04C6CE56EE65B4611FD98FA82" author="Michael" box="[355,552,1292,1314]" page="576" pageId="1" pageNumber="68" year="1998">
<bibRefCitation id="2A014B404C6CE56EE65B4611FE48FA82" author="Michael" box="[355,504,1292,1314]" pageId="1" pageNumber="68" refString="Michael, S. W. (1998) Reef Fishes: A Guide to Their Identification, Behaviour, and Captive Care. Vol. 1. Microcosm Ltd, Shelburne, 624 pp." type="book" year="1998">Michael, 1998</bibRefCitation>
: 576
</treatmentCitation>
(colour photo; Guam).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6CE56EE7AF4637FCEDFAE0" box="[151,861,1322,1344]" pageId="1" pageNumber="68" type="description">
<paragraph id="4E2F36B14C6CE56EE7AF4637FCEDFAE0" blockId="1.[151,1388,1230,1436]" box="[151,861,1322,1344]" pageId="1" pageNumber="68">
<treatmentCitationGroup id="6E80119F4C6CE56EE7AF4637FCEDFAE0" box="[151,861,1322,1344]" pageId="1" pageNumber="68">
<taxonomicName id="89904D324C6CE56EE7AF4637FEAFFAE0" ID-CoL="6YRF" authorityName="Bleeker" authorityYear="1871" box="[151,287,1322,1344]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6CE56EE7AF4637FEAFFAE0" box="[151,287,1322,1344]" italics="true" pageId="1" pageNumber="68">Pseudanthias</emphasis>
</taxonomicName>
sp. B;
<treatmentCitation id="CF3110A04C6CE56EE65F4636FDA6FAE0" author="Myers" box="[359,534,1322,1344]" page="109" pageId="1" pageNumber="68" year="1999">
<bibRefCitation id="2A014B404C6CE56EE65F4636FE55FAE0" author="Myers" box="[359,485,1322,1344]" pageId="1" pageNumber="68" refString="Myers, R. F. (1999) Micronesian Reef Fishes. A Practical Guide to the Identification of the Coral Reef Fishes of the Tropical Central and Western Pacific. 3 rd Edition. Coral Graphics, Barrigada, 330 pp., 192 pls." type="book" year="1999">Myers 1999</bibRefCitation>
: 109
</treatmentCitation>
, pl. 35 G (colour photo; Guam).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6CE56EE7AF4654FBB4FA3C" pageId="1" pageNumber="68" type="reference_group">
<paragraph id="4E2F36B14C6CE56EE7AF4654FBB4FA3C" blockId="1.[151,1388,1230,1436]" pageId="1" pageNumber="68">
<treatmentCitationGroup id="6E80119F4C6CE56EE7AF4654FBB4FA3C" pageId="1" pageNumber="68">
<taxonomicName id="89904D324C6CE56EE7AF4654FE20FAFF" ID-CoL="4NKNC" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[151,400,1353,1375]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C6CE56EE7AF4654FE20FAFF" box="[151,400,1353,1375]" italics="true" pageId="1" pageNumber="68">Pseudanthias flavicauda</emphasis>
</taxonomicName>
[non
<bibRefCitation id="2A014B404C6CE56EE6F44654FD1CFAFF" author="Randall" box="[460,684,1353,1375]" pageId="1" pageNumber="68" refString="Randall, J. E. &amp; Pyle, R. L. (2001) Four new serranid fishes of the anthiine genus Pseudanthias from the South Pacific. Raffles Bulletin of Zoology, 49, 19 - 34." type="journal article" year="2001">Randall &amp; Pyle, 2001</bibRefCitation>
];
<treatmentCitation id="CF3110A04C6CE56EE5844657FC4EFAFF" author="Myers" box="[700,1022,1353,1375]" page="616" pageId="1" pageNumber="68" year="2003">
<bibRefCitation id="2A014B404C6CE56EE5844657FC7EFAFF" author="Myers" box="[700,974,1353,1375]" pageId="1" pageNumber="68" refString="Myers, R. F. &amp; Donaldson, T. J. (2003) The fishes of the Mariana Islands. Micronesica, 35 - 36, 594 - 648." type="book chapter" year="2003">Myers &amp; Donaldson, 2003</bibRefCitation>
: 616
</treatmentCitation>
(list, Marianas Islands).
<taxonomicName id="89904D324C6CE56EE7AF4675FEAFFADE" ID-CoL="6YRF" authorityName="Bleeker" authorityYear="1871" box="[151,287,1384,1406]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6CE56EE7AF4675FEAFFADE" box="[151,287,1384,1406]" italics="true" pageId="1" pageNumber="68">Pseudanthias</emphasis>
</taxonomicName>
cf
<taxonomicName id="89904D324C6CE56EE6074675FE20FADE" ID-CoL="4NKPH" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[319,400,1384,1406]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C6CE56EE6074675FE20FADE" box="[319,400,1384,1406]" italics="true" pageId="1" pageNumber="68">randalli</emphasis>
</taxonomicName>
;
<treatmentCitation id="CF3110A04C6CE56EE6A64675FDF6FADE" author="Kuiter" box="[414,582,1384,1406]" page="61" pageId="1" pageNumber="68" year="2004">
<bibRefCitation id="2A014B404C6CE56EE6A64675FD92FADE" author="Kuiter" box="[414,546,1384,1406]" pageId="1" pageNumber="68" refString="Kuiter, R. H. (2004) Basslets, Hamlets and Their Relatives. A Comprehensive Guide to Selected Serranidae and Plesiopidae. TMC Publishing, Chorleywood, 216 pp." type="book" year="2004">Kuiter, 2004</bibRefCitation>
: 61
</treatmentCitation>
, unnumbered colour figs A and B (colour photos; Palau, Saipan and Guam).
<taxonomicName id="89904D324C6CE56EE7AF469BFEAFFA3C" ID-CoL="6YRF" authorityName="Bleeker" authorityYear="1871" box="[151,287,1414,1436]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6CE56EE7AF469BFEAFFA3C" box="[151,287,1414,1436]" italics="true" pageId="1" pageNumber="68">Pseudanthias</emphasis>
</taxonomicName>
cf
<taxonomicName id="89904D324C6CE56EE607469BFE1AFA3C" ID-CoL="4NKNC" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[319,426,1414,1436]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C6CE56EE607469BFE1AFA3C" box="[319,426,1414,1436]" italics="true" pageId="1" pageNumber="68">flavicauda</emphasis>
</taxonomicName>
;
<treatmentCitation id="CF3110A04C6CE56EE680469BFD55FA3C" author="Kuiter" box="[440,741,1414,1436]" page="298" pageId="1" pageNumber="68" year="2006">
<bibRefCitation id="2A014B404C6CE56EE680469BFD06FA3C" author="Kuiter" box="[440,694,1414,1436]" pageId="1" pageNumber="68" refString="Kuiter, R. H. &amp; Debelius, H. (2006) World Atlas of Marine Fishes. IKAN-Unterwasserarchiv, Frankfurt, 720 pp." type="book" year="2006">Kuiter &amp; Debelius, 2006</bibRefCitation>
: 298
</treatmentCitation>
(colour photo; Micronesia).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6CE56EE7AF46D4FC45F98A" pageId="1" pageNumber="68" type="materials_examined">
<paragraph id="4E2F36B14C6CE56EE7AF46D4FD0CF9A6" blockId="1.[151,1437,1481,2010]" pageId="1" pageNumber="68">
<materialsCitation id="FEF83CEC4C6CE56EE7AF46D4FD0CF9A6" ID-GBIF-Occurrence="1676226877" collectingDate="1997-03-16" collectionCode="KPM-NI" collectorName="O. Morishita" country="Japan" county="Ototo-jima Island" elevation="40" location="Ogasawara Islands" municipality="Chichi-jima Islands" pageId="1" pageNumber="68" specimenCode="KPM-NI 3759, 59.3" specimenCount="1" typeStatus="holotype">
<emphasis id="7CE4EAA34C6CE56EE7AF46D4FEBBFA42" bold="true" box="[151,267,1481,1506]" pageId="1" pageNumber="68">
<typeStatus id="912B88134C6CE56EE7AF46D4FEB6FA42" box="[151,262,1481,1506]" pageId="1" pageNumber="68" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="1E369ECA4C6CE56EE62B46D7FE0AFA42" box="[275,442,1481,1506]" collectionCode="KPM-NI" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35223" name="Kanagawa Prefectural Museum of Natural History" pageId="1" pageNumber="68">KPM-NI 3759</specimenCode>
,
<quantity id="89689B544C6CE56EE6FF46D4FD9DFA41" box="[455,557,1481,1506]" metricMagnitude="-2" metricUnit="m" metricValue="5.93" pageId="1" pageNumber="68" unit="mm" value="59.3">
<specimenCode id="1E369ECA4C6CE56EE6FF46D4FE48FA42" box="[455,504,1481,1506]" collectionCode="KPM-NI" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35223" name="Kanagawa Prefectural Museum of Natural History" pageId="1" pageNumber="68">59.3</specimenCode>
mm
</quantity>
SL, male,
<collectingCountry id="368776214C6CE56EE59446D7FD5FFA41" box="[684,751,1482,1505]" name="Japan" pageId="1" pageNumber="68">Japan</collectingCountry>
,
<location id="4B4F606A4C6CE56EE5C346D7FC62FA42" LSID="urn:lsid:plazi:treatment:C63987A74C6CE567E7AF4701FF73FD50:4B4F606A4C6CE56EE5C346D7FC62FA42" box="[763,978,1481,1506]" country="Japan" county="Ototo-jima Island" municipality="Chichi-jima Islands" name="Ogasawara Islands" pageId="1" pageNumber="68">Ogasawara Islands</location>
,
<collectingMunicipality id="AE4BACCB4C6CE56EE4E746D4FB70FA42" box="[991,1216,1481,1506]" pageId="1" pageNumber="68">Chichi-jima Islands</collectingMunicipality>
,
<collectingCounty id="A74E4E3D4C6CE56EE3F546D4FA28FA42" box="[1229,1432,1481,1506]" pageId="1" pageNumber="68">Ototo-jima Island</collectingCounty>
,
<quantity id="89689B544C6CE56EE7AF46F3FF62F9A5" box="[151,210,1518,1542]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="68" unit="m" value="40.0">
<elevation id="C5BDD1824C6CE56EE7AF46F3FF62F9A5" box="[151,210,1518,1542]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="68" unit="m" value="40.0">40 m</elevation>
</quantity>
, collected by
<collectorName id="E36553674C6CE56EE65546F3FE4FF9A6" box="[365,511,1517,1542]" pageId="1" pageNumber="68">O. Morishita</collectorName>
,
<date id="3A2E10714C6CE56EE53246F0FD08F9A6" box="[522,696,1517,1542]" pageId="1" pageNumber="68" value="1997-03-16">
<collectingDate id="2A6AE9994C6CE56EE53246F0FD08F9A6" box="[522,696,1517,1542]" pageId="1" pageNumber="68" value="1997-03-16">16 March 1997</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="4E2F36B14C6CE56EE7FF450CFC45F98A" blockId="1.[151,1437,1481,2010]" box="[199,1013,1553,1578]" pageId="1" pageNumber="68">
<materialsCitation id="FEF83CEC4C6CE56EE7FF450CFC45F98A" ID-GBIF-Occurrence="1676226876" box="[199,1013,1553,1578]" collectingDate="1997-03-16" collectionCode="KPM-NI" collectorName="O. Morishita" country="Japan" county="Ototo-jima Island" elevation="40" location="Ogasawara Islands" municipality="Chichi-jima Islands" pageId="1" pageNumber="68" specimenCode="KPM-NI 3758, 31.5" specimenCount="1" typeStatus="paratype">
<emphasis id="7CE4EAA34C6CE56EE7FF450CFE8BF98A" bold="true" box="[199,315,1553,1578]" pageId="1" pageNumber="68">
<typeStatus id="912B88134C6CE56EE7FF450CFE86F98A" box="[199,310,1553,1578]" pageId="1" pageNumber="68" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="1E369ECA4C6CE56EE67A450FFE57F98A" box="[322,487,1553,1578]" collectionCode="KPM-NI" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35223" name="Kanagawa Prefectural Museum of Natural History" pageId="1" pageNumber="68">KPM-NI 3758</specimenCode>
,
<quantity id="89689B544C6CE56EE6CA450CFDE6F989" box="[498,598,1553,1578]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="1" pageNumber="68" unit="mm" value="31.5">
<specimenCode id="1E369ECA4C6CE56EE6CA450CFD93F98A" box="[498,547,1553,1578]" collectionCode="KPM-NI" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35223" name="Kanagawa Prefectural Museum of Natural History" pageId="1" pageNumber="68">31.5</specimenCode>
mm
</quantity>
SL, female, collected with holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6CE56EE7FF4528FE1FF8A2" pageId="1" pageNumber="68" type="diagnosis">
<paragraph id="4E2F36B14C6CE56EE7FF4528FE1FF8A2" blockId="1.[151,1437,1481,2010]" pageId="1" pageNumber="68">
<emphasis id="7CE4EAA34C6CE56EE7FF4528FEF1F9EE" bold="true" box="[199,321,1589,1614]" pageId="1" pageNumber="68">Diagnosis.</emphasis>
The following combination of characters distinguishes
<taxonomicName id="89904D324C6CE56EE4E74528FB61F9EE" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[991,1233,1589,1614]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="68" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C6CE56EE4E74528FB61F9EE" box="[991,1233,1589,1614]" italics="true" pageId="1" pageNumber="68">Pseudanthias tequila</emphasis>
</taxonomicName>
from congeners: dorsal rays X,16; anal rays III,7; pectoral rays 18; third dorsal spine prolonged in males; third segmented anal-fin ray longest; lateral-line scales 4647; no auxiliary scales on body; interopercle and subopercle with distinct serrations in adults; males in life with upper body beneath anterior part of dorsal fin purple to pink with orange-red stripe, dorsal fin greenish yellow anteriorly, posteriorly red with blue basal area, anterior part of anal fin red, and pelvic fins bright yellow.
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6CE568E7FF4410FBB5FF75" lastPageId="7" lastPageNumber="74" pageId="1" pageNumber="68" type="description">
<paragraph id="4E2F36B14C6CE56DE7FF4410FBBFFF74" blockId="1.[151,1437,1481,2010]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
<emphasis id="7CE4EAA34C6CE56EE7FF4410FEE9F886" bold="true" box="[199,345,1805,1830]" pageId="1" pageNumber="68">Description.</emphasis>
Dorsal rays X,16, all segmented rays branched; each dorsal fin spine with fleshy pennant extending from tip, this best developed on longest (third) spine; anal rays III,7, all segmented rays branched; pectoral rays 18/18, all rays branched except upper 2/2 and lower 1/1 (2/2); pelvic rays 1,5; principal caudal rays 9 + 8; branched caudal-fin rays 7 + 6; upper procurrent caudal-fin rays 8 (9); lower procurrent caudal rays 8 (9); lateral-line scales too damaged (and regrown) on left side of
<typeStatus id="912B88134C6CE56EE4524480FC7FF816" box="[874,975,1949,1974]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
, 46 on right side (49/48); scales above lateral-line to origin of dorsal fin 5/5; scales above lateral-line to base of fifth dorsal spine 4/4 (3/4); scales below lateral line to origin of anal fin 18/19 (20/19); circumpeduncular scales not determinable in
<typeStatus id="912B88134C6FE56DE3F2438AFA9CFF10" box="[1226,1324,151,176]" pageId="2" pageNumber="69" type="holotype">holotype</typeStatus>
(24); gill rakers 10 + 23 (8 + 21); branchiostegal rays 7; pseudobranch filaments 16 (10).
</paragraph>
<paragraph id="4E2F36B14C6FE56DE7FF43C2FED9FE70" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="69">
Vertebrae 10 + 16; supraneurals 3 (posterior two closely applied in
<typeStatus id="912B88134C6FE56DE4FF43FDFB9AFF58" box="[967,1066,224,248]" pageId="2" pageNumber="69" type="paratype">paratype</typeStatus>
, though still separate); predorsal formula 0/0+0/2/1+1; main shaft (proximal component) of first dorsal pterygiophore roughly perpendicular to long axis of body; no trisegmental pterygiophores associated with dorsal and anal fins; ribs present on vertebrae 3 through 10; epineurals present on vertebrae 1 through 12; paired parapophyses present on first caudal vertebra (see
<bibRefCitation id="2A014B404C6FE56DE7AF4272FE8DFE28" author="Baldwin" box="[151,317,367,392]" pageId="2" pageNumber="69" refString="Baldwin, C. C. (1990) Morphology of the larvae of American Anthiinae (Teleostei: Serranidae), with comments on relationships within the subfamily. Copeia, 1990, 913 - 955. https: // doi. org / 10.2307 / 1446477" type="journal article" year="1990">Baldwin 1990</bibRefCitation>
: fig. 21B); parhypural and hypurals autogenous; well-developed hypurapophysis on parhypural; epurals 3; single uroneural (posterior uroneural absent); ventral tip of cleithrum with well-developed posteroventral process (
<figureCitation id="D6AB2A344C6FE56DE7C142AAFEEBFE70" box="[249,347,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,970,992]" captionTargetBox="[151,1436,193,948]" captionTargetId="figure@4.[151,1436,193,948]" captionTargetPageId="4" captionText="FIGURE 3. Pseudanthias tequila n. sp., KPM-NI 3759, 59.3 mm SL, holotype, x-radiograph. Radiograph (KPM-NR 51086 C) by H. Senou." httpUri="https://zenodo.org/record/1038752/files/figure.png" pageId="2" pageNumber="69">Figure 3</figureCitation>
).
</paragraph>
<paragraph id="4E2F36B14C6FE56DE7FF42C1FD5DFE54" blockId="2.[151,1436,151,501]" box="[199,749,476,501]" pageId="2" pageNumber="69">
Morphometric values are summarised in
<tableCitation id="0312030A4C6FE56DE5AA42C1FD56FE54" box="[658,742,476,501]" captionStart="TABLE 1" captionStartId="2.[151,239,544,567]" captionText="TABLE 1. Morphometric values for Pseudanthias tequila expressed as percentage SL." pageId="2" pageNumber="69">Table 1</tableCitation>
.
</paragraph>
<caption id="1AEF66394C6FE56DE7AF413DFB86FD97" box="[151,1078,543,567]" pageId="2" pageNumber="69">
<paragraph id="4E2F36B14C6FE56DE7AF413DFB86FD97" blockId="2.[151,1078,543,567]" box="[151,1078,543,567]" pageId="2" pageNumber="69">
<emphasis id="7CE4EAA34C6FE56DE7AF413DFEB3FD97" bold="true" box="[151,259,544,567]" pageId="2" pageNumber="69">TABLE 1</emphasis>
. Morphometric values for
<taxonomicName id="89904D324C6FE56DE51A4102FCB2FD97" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[546,770,543,567]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="69" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C6FE56DE51A4102FCB2FD97" box="[546,770,543,567]" italics="true" pageId="2" pageNumber="69">Pseudanthias tequila</emphasis>
</taxonomicName>
expressed as percentage SL.
</paragraph>
</caption>
<paragraph id="4E2F36B14C6FE56CE40B414EFCDDF94E" blockId="2.[819,1287,594,647]" lastBlockId="3.[151,1437,1461,2026]" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
KPM-NI 3759 KPM-NI 3758
<typeStatus id="912B88134C6FE56DE40B416CFC21FD27" box="[819,913,625,647]" pageId="2" pageNumber="69" type="holotype">Holotype</typeStatus>
<typeStatus id="912B88134C6FE56DE34C416FFB7DFD27" box="[1140,1229,626,647]" pageId="2" pageNumber="69" type="paratype">Paratype</typeStatus>
Mouth large, slightly oblique, posterior margin of maxilla reaching to vertical through posterior edge of pupil; mouth terminal, lower jaw projecting slightly; premaxilla with an enlarged recurved canine anterolaterally, a band of small conical teeth about five rows wide at symphysis reducing to two rows on sides of jaw, with the outer row teeth larger and slightly curved, and the three or four teeth nearest symphysis enlarged and caniniform, lying almost flat against roof of mouth; dentary with an enlarged recurved canine at symphysis, followed by a band of small conical teeth about three or four rows wide; sides of lower jaw with single row of slightly curved, conical teeth, with one or two teeth on middle of jaw greatly enlarged and caniniform; vomer with a triangular patch of small conical teeth, three rows wide in midline; palatine with a narrow band of small conical teeth, two rows wide at widest point; ectopterygoid, mesopterygoid and tongue edentate.
</paragraph>
<paragraph id="4E2F36B14C6FE56DE7A74187FB10F828" pageId="2" pageNumber="69">
<table id="3C90C4114C6F1A90E7A74187FAB7F828" box="[159,1287,666,1928]" gridcols="3" gridrows="11" pageId="2" pageNumber="69">
<tr id="F0A034F34C6F1A90E7A74187FAB7FD10" box="[159,1287,666,688]" gridrow="0" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A74187FD82FD10" box="[159,562,666,688]" gridcol="0" gridrow="0" pageId="2" pageNumber="69">SL (mm)</th>
<th id="B3715D8F4C6F1A90E40B4187FC33FD10" box="[819,899,666,688]" gridcol="1" gridrow="0" pageId="2" pageNumber="69">59.3</th>
<th id="B3715D8F4C6F1A90E34C4187FAB7FD10" box="[1140,1287,666,688]" gridcol="2" gridrow="0" pageId="2" pageNumber="69">31.5</th>
</tr>
<tr id="F0A034F34C6F1A90E7A741D9FAB7FC8C" box="[159,1287,708,812]" gridrow="1" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A741D9FD82FC8C" box="[159,562,708,812]" gridcol="0" gridrow="1" pageId="2" pageNumber="69">Greatest body depth Body depth at anal-fin origin Body width</th>
<td id="B3715D8F4C6F1A90E40B41D9FC33FC8C" box="[819,899,708,812]" gridcol="1" gridrow="1" pageId="2" pageNumber="69">35.2 32.5 16.7</td>
<td id="B3715D8F4C6F1A90E34C41D9FAB7FC8C" box="[1140,1287,708,812]" gridcol="2" gridrow="1" pageId="2" pageNumber="69">33.7 29.5 16.5</td>
</tr>
<tr id="F0A034F34C6F1A90E7A7405DFAB7FC08" box="[159,1287,832,936]" gridrow="2" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A7405DFD82FC08" box="[159,562,832,936]" gridcol="0" gridrow="2" pageId="2" pageNumber="69">Head length Snout length Orbit diameter</th>
<td id="B3715D8F4C6F1A90E40B405DFC33FC08" box="[819,899,832,936]" gridcol="1" gridrow="2" pageId="2" pageNumber="69">30.9 7.4 9.3</td>
<td id="B3715D8F4C6F1A90E34C405DFAB7FC08" box="[1140,1287,832,936]" gridcol="2" gridrow="2" pageId="2" pageNumber="69">30.8 6.3 11.1</td>
</tr>
<tr id="F0A034F34C6F1A90E7A740A1FAB7FB84" box="[159,1287,956,1060]" gridrow="3" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A740A1FD82FB84" box="[159,562,956,1060]" gridcol="0" gridrow="3" pageId="2" pageNumber="69">Bony interorbital width Upper jaw length Maxilla width</th>
<td id="B3715D8F4C6F1A90E40B40A1FC33FB84" box="[819,899,956,1060]" gridcol="1" gridrow="3" pageId="2" pageNumber="69">7.6 15.9 5.7</td>
<td id="B3715D8F4C6F1A90E34C40A1FAB7FB84" box="[1140,1287,956,1060]" gridcol="2" gridrow="3" pageId="2" pageNumber="69">7.6 14.0 5.7</td>
</tr>
<tr id="F0A034F34C6F1A90E7A74725FAB7FB00" box="[159,1287,1080,1184]" gridrow="4" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A74725FD82FB00" box="[159,562,1080,1184]" gridcol="0" gridrow="4" pageId="2" pageNumber="69">Caudal peduncle length Caudal peduncle depth Predorsal length</th>
<td id="B3715D8F4C6F1A90E40B4725FC33FB00" box="[819,899,1080,1184]" gridcol="1" gridrow="4" pageId="2" pageNumber="69">21.9 13.3 27.5</td>
<td id="B3715D8F4C6F1A90E34C4725FAB7FB00" box="[1140,1287,1080,1184]" gridcol="2" gridrow="4" pageId="2" pageNumber="69">24.1 11.7 29.5</td>
</tr>
<tr id="F0A034F34C6F1A90E7A747A9FAB7FABC" box="[159,1287,1204,1308]" gridrow="5" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A747A9FD82FABC" box="[159,562,1204,1308]" gridcol="0" gridrow="5" pageId="2" pageNumber="69">Preanal length Prepelvic length Dorsal fin base length</th>
<td id="B3715D8F4C6F1A90E40B47A9FC33FABC" box="[819,899,1204,1308]" gridcol="1" gridrow="5" pageId="2" pageNumber="69">62.6 34.1 65.8</td>
<td id="B3715D8F4C6F1A90E34C47A9FAB7FABC" box="[1140,1287,1204,1308]" gridcol="2" gridrow="5" pageId="2" pageNumber="69">60.3 32.4 63.2</td>
</tr>
<tr id="F0A034F34C6F1A90E7A7462DFAB7FA38" box="[159,1287,1328,1432]" gridrow="6" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A7462DFD82FA38" box="[159,562,1328,1432]" gridcol="0" gridrow="6" pageId="2" pageNumber="69">First dorsal spine Longest dorsal spine (number) First segmented dorsal ray</th>
<td id="B3715D8F4C6F1A90E40B462DFC33FA38" box="[819,899,1328,1432]" gridcol="1" gridrow="6" pageId="2" pageNumber="69">7.1 22.3 (3) 15.3</td>
<td id="B3715D8F4C6F1A90E34C462DFAB7FA38" box="[1140,1287,1328,1432]" gridcol="2" gridrow="6" pageId="2" pageNumber="69">6.3 18.1 (3) 15.5</td>
</tr>
<tr id="F0A034F34C6F1A90E7A746B1FAB7F9B4" box="[159,1287,1452,1556]" gridrow="7" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A746B1FD82F9B4" box="[159,562,1452,1556]" gridcol="0" gridrow="7" pageId="2" pageNumber="69">Longest segmented dorsal ray (number) Anal fin base length First anal spine</th>
<td id="B3715D8F4C6F1A90E40B46B1FC33F9B4" box="[819,899,1452,1556]" gridcol="1" gridrow="7" pageId="2" pageNumber="69">17.0 (7) 19.4 7.4</td>
<td id="B3715D8F4C6F1A90E34C46B1FAB7F9B4" box="[1140,1287,1452,1556]" gridcol="2" gridrow="7" pageId="2" pageNumber="69">16.5 (5) 17.5 8.9</td>
</tr>
<tr id="F0A034F34C6F1A90E7A74535FAB7F930" box="[159,1287,1576,1680]" gridrow="8" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A74535FD82F930" box="[159,562,1576,1680]" gridcol="0" gridrow="8" pageId="2" pageNumber="69">Second anal spine Third anal spine First segmented anal ray</th>
<td id="B3715D8F4C6F1A90E40B4535FC33F930" box="[819,899,1576,1680]" gridcol="1" gridrow="8" pageId="2" pageNumber="69">18.4 16.4 20.9</td>
<td id="B3715D8F4C6F1A90E34C4535FAB7F930" box="[1140,1287,1576,1680]" gridcol="2" gridrow="8" pageId="2" pageNumber="69">20.6 16.5 21.3</td>
</tr>
<tr id="F0A034F34C6F1A90E7A745B9FAB7F8AC" box="[159,1287,1700,1804]" gridrow="9" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A745B9FD82F8AC" box="[159,562,1700,1804]" gridcol="0" gridrow="9" pageId="2" pageNumber="69">Longest segmented anal ray (number) Caudal fin length Caudal concavity</th>
<td id="B3715D8F4C6F1A90E40B45B9FC33F8AC" box="[819,899,1700,1804]" gridcol="1" gridrow="9" pageId="2" pageNumber="69">24.1 (3) 37.9 20.2</td>
<td id="B3715D8F4C6F1A90E34C45B9FAB7F8AC" box="[1140,1287,1700,1804]" gridcol="2" gridrow="9" pageId="2" pageNumber="69">23.8 (3) 42.5 22.5</td>
</tr>
<tr id="F0A034F34C6F1A90E7A7443DFAB7F828" box="[159,1287,1824,1928]" gridrow="10" pageId="2" pageNumber="69">
<th id="B3715D8F4C6F1A90E7A7443DFD82F828" box="[159,562,1824,1928]" gridcol="0" gridrow="10" pageId="2" pageNumber="69">Pectoral fin length Pelvic fin spine Pelvic fin length</th>
<td id="B3715D8F4C6F1A90E40B443DFC33F828" box="[819,899,1824,1928]" gridcol="1" gridrow="10" pageId="2" pageNumber="69">28.0 18.9 28.0</td>
<td id="B3715D8F4C6F1A90E34C443DFAB7F828" box="[1140,1287,1824,1928]" gridcol="2" gridrow="10" pageId="2" pageNumber="69">27.3 19.7 28.6</td>
</tr>
</table>
</paragraph>
<caption id="1AEF66394C6EE56CE7AF41EEFC8EFC87" httpUri="https://zenodo.org/record/1038748/files/figure.png" pageId="3" pageNumber="70" targetBox="[340,1247,195,733]" targetPageId="3">
<paragraph id="4E2F36B14C6EE56CE7AF41EEFC8EFC87" blockId="3.[151,1436,755,807]" pageId="3" pageNumber="70">
<emphasis id="7CE4EAA34C6EE56CE7AF41EEFEA4FCA8" bold="true" box="[151,276,755,777]" pageId="3" pageNumber="70">FIGURE 1.</emphasis>
<taxonomicName id="89904D324C6EE56CE62441EEFE42FCA9" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[284,498,755,777]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="70" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C6EE56CE62441EEFE42FCA9" box="[284,498,755,777]" italics="true" pageId="3" pageNumber="70">Pseudanthias tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C6EE56CE6C241E9FD84FCA9" box="[506,564,756,777]" pageId="3" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
, KPM-NI 3759, 59.3 mm SL male holotype, Ototo-jima Island, Chichi-jima Islands, Ogasawara Islands, Japan. Photo (KPM-NR 51086B) by H. Senou.
</paragraph>
</caption>
<caption id="1AEF66394C6EE56CE7AF4647FC8EFA2E" httpUri="https://zenodo.org/record/1038750/files/figure.png" pageId="3" pageNumber="70" targetBox="[340,1247,857,1348]" targetPageId="3">
<paragraph id="4E2F36B14C6EE56CE7AF4647FC8EFA2E" blockId="3.[151,1436,1370,1422]" pageId="3" pageNumber="70">
<emphasis id="7CE4EAA34C6EE56CE7AF4647FEA3FACF" bold="true" box="[151,275,1370,1392]" pageId="3" pageNumber="70">FIGURE 2.</emphasis>
<taxonomicName id="89904D324C6EE56CE6214647FE5EFAD0" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[281,494,1370,1392]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="70" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C6EE56CE6214647FE5EFAD0" box="[281,494,1370,1392]" italics="true" pageId="3" pageNumber="70">Pseudanthias tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C6EE56CE6CD4646FD9EFAD0" box="[501,558,1371,1392]" pageId="3" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
, KPM-NI 3758, 31.5 mm SL female paratype, Ototo-jima Island, Chichi-jima Islands, Ogasawara Islands, Japan. Photo (KPM-NR 51087B) by H. Senou.
</paragraph>
</caption>
<paragraph id="4E2F36B14C6EE56CE7FF45E4FB21F84A" blockId="3.[151,1437,1461,2026]" pageId="3" pageNumber="70">
Opercle with 3 flat spines, middle spine longest, upper spine mostly concealed by scales; preopercle with 20 (15) fine serrations on vertical portion, 2 (1) enlarged serrations at angle, and 4 (0) weak serrations on ventral portion; interopercle with 5 (2) relatively large serrations; subopercle with 6 (2) relatively large serrations; posttemporal with 4 (3) fine serrations; supracleithrum with 2 fine serrations. Anterior nostril positioned at middle of snout, tubular with small flap on posterior rim, flap almost reaching posterior nostril when laid back; posterior nostril at mid-upper, anterior border of orbit, covered by thin, narrow membrane anteriorly. No papillae on posterior rim of orbit. Snout of
<typeStatus id="912B88134C6EE56CE6CF44CCFDE9F84A" box="[503,601,2001,2026]" pageId="3" pageNumber="70" type="holotype">holotype</typeStatus>
slightly hypertrophied with papillae at symphysis.
</paragraph>
<caption id="1AEF66394C69E56BE7AF40D7FEACFC5E" httpUri="https://zenodo.org/record/1038752/files/figure.png" pageId="4" pageNumber="71" targetBox="[151,1436,193,948]" targetPageId="4">
<paragraph id="4E2F36B14C69E56BE7AF40D7FEACFC5E" blockId="4.[151,1436,970,1022]" pageId="4" pageNumber="71">
<emphasis id="7CE4EAA34C69E56BE7AF40D7FEA2FC7F" bold="true" box="[151,274,970,992]" pageId="4" pageNumber="71">FIGURE 3.</emphasis>
<taxonomicName id="89904D324C69E56BE62040D7FE5DFC40" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[280,493,970,992]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="71" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C69E56BE62040D7FE5DFC40" box="[280,493,970,992]" italics="true" pageId="4" pageNumber="71">Pseudanthias tequila</emphasis>
</taxonomicName>
<emphasis id="7CE4EAA34C69E56BE6CC40D7FD9CFC7F" bold="true" box="[500,556,970,991]" pageId="4" pageNumber="71">
<taxonomicNameLabel id="67D757D84C69E56BE6CC40D7FD9CFC7F" box="[500,556,970,991]" pageId="4" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, KPM-NI 3759, 59.3 mm SL, holotype, x-radiograph. Radiograph (KPM-NR 51086C) by H. Senou.
</paragraph>
</caption>
<paragraph id="4E2F36B14C69E56BE7FF4736FCFCFB58" blockId="4.[151,1437,1067,1992]" pageId="4" pageNumber="71">
Scales ctenoid with peripheral cteni (
<bibRefCitation id="2A014B404C69E56BE54D4736FCA5FBE3" author="Roberts" box="[629,789,1067,1092]" pageId="4" pageNumber="71" refString="Roberts, C. D. (1993) Comparative morphology of spined scales and their phylogenetic significance in the Teleostei. Bulletin of Marine Science, 52, 60 - 113." type="journal article" year="1993">Roberts 1993</bibRefCitation>
); lateral line broadly arched over pectoral fin following body contour to caudal-fin base; head scaled, including dentary, maxilla, snout, and infraorbitals, no scales on lips; no auxiliary scales on body, a few auxiliary scales on cheeks and operculum; low scaly sheath on soft dorsal and anal fins, with indistinct small scales present between segmented rays; caudal fin with scaly basal sheath, with small scales extending over almost all of fin, except for fin tips and posterior part of membranes of middle rays; pectoral fins with basal, wedge-shaped sheath of small scales.
</paragraph>
<paragraph id="4E2F36B14C69E56BE7FF461EFB57F908" blockId="4.[151,1437,1067,1992]" pageId="4" pageNumber="71">
<emphasis id="7CE4EAA34C69E56BE7FF461EFE79FABC" box="[199,457,1283,1308]" italics="true" pageId="4" pageNumber="71">Colour of males in life</emphasis>
(based on colour photos of
<typeStatus id="912B88134C69E56BE431461EFCDBFABC" box="[777,875,1283,1308]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
when freshly dead, and of live individuals in the Ogasawara and
<collectingCountry id="368776214C69E56BE672463AFDB2FAE0" box="[330,514,1319,1344]" name="Northern Mariana Islands" pageId="4" pageNumber="71">Mariana Islands</collectingCountry>
;
<figureCitation id="D6AB2A344C69E56BE536463AFDC7FA9F" box="[526,631,1319,1344]" captionStart="FIGURE 1" captionStartId="3.[151,250,755,777]" captionTargetBox="[340,1247,195,733]" captionTargetId="figure@3.[340,1247,195,733]" captionTargetPageId="3" captionText="FIGURE 1. Pseudanthias tequila n. sp., KPM-NI 3759, 59.3 mm SL male holotype, Ototo-jima Island, Chichi-jima Islands, Ogasawara Islands, Japan. Photo (KPM-NR 51086 B) by H. Senou." httpUri="https://zenodo.org/record/1038748/files/figure.png" pageId="4" pageNumber="71">Figures 1</figureCitation>
,
<figureCitation id="D6AB2A344C69E56BE5BD463AFD52FA9F" box="[645,738,1319,1343]" captionStart="FIGURE 4" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1436,193,1220]" captionTargetId="figure@5.[151,1436,193,1220]" captionTargetPageId="5" captionText="FIGURE 4. Males of selected Pseudanthias species: A) P. tequila n. sp., Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan (KPM-NR 179391 A, photo by O. Morishita); B) P. tequila n. sp., Tinian, Mariana Islands (photo by N. Tsuji); C) P. pulcherrimus, aquarium specimen (not retained) from the Maldives (photo by Y. K. Tea); D) P. randalli, Kwajalein, Marshall Islands (photo by J. Johnson); E) P. flavicauda, Fiji (photo by J. E. Randall); F) P. sp., Kirimati, Kiribati (photo by D. J. Linehan)." httpUri="https://zenodo.org/record/1038754/files/figure.png" pageId="4" pageNumber="71">4A &amp; B</figureCitation>
): head and body purple to pink; bright purple stripe extending from anterior edge of eye around ventral part of orbit to lower part of pectoral-fin base; snout, interorbital area and area above stripe orange to orange-red; indistinct purple stripe sometimes present from behind middle of eye to upper part of pectoral fin base; iris orange to purple, bright yellow around pupil; upper part of body with broad orange-red stripe extending from behind head to caudal peduncle; similar orange-red stripe extending from pectoral-fin base along midside to caudal peduncle; scales on nape and body often dark basally; dorsal fin greenish yellow in front of eighth to tenth spine, bright red posteriorly, with bright blue base and distal margin; anal fin bright red anteriorly, with bright blue wedge extending from near base of first segmented ray to posterior part of fin, distal margin of red region narrowly edged with bright blue; caudal fin variable (probably mood induced), either bright yellow, greenish yellow with purple base, or solid purple, the upper and lower edges of fin bright blue; pelvic fins bright yellow, with spine dusky red to bright orange-red; pectoral fins orangish hyaline.
</paragraph>
<paragraph id="4E2F36B14C69E56BE7FF45AEFAF4F868" blockId="4.[151,1437,1067,1992]" pageId="4" pageNumber="71">
<emphasis id="7CE4EAA34C69E56BE7FF45AEFE69F96C" box="[199,473,1715,1740]" italics="true" pageId="4" pageNumber="71">Colour of females in life</emphasis>
(based on colour photos of
<typeStatus id="912B88134C69E56BE42C45A9FCC4F96C" box="[788,884,1716,1740]" pageId="4" pageNumber="71" type="paratype">paratype</typeStatus>
when freshly dead, and of live individuals in the Ogasawara and
<collectingCountry id="368776214C69E56BE67145CAFDB0F950" box="[329,512,1751,1776]" name="Northern Mariana Islands" pageId="4" pageNumber="71">Mariana Islands</collectingCountry>
;
<figureCitation id="D6AB2A344C69E56BE53345CAFDC5F94F" box="[523,629,1751,1776]" captionStart="FIGURE 2" captionStartId="3.[151,250,1370,1392]" captionTargetBox="[340,1247,857,1348]" captionTargetId="figure@3.[340,1247,857,1348]" captionTargetPageId="3" captionText="FIGURE 2. Pseudanthias tequila n. sp., KPM-NI 3758, 31.5 mm SL female paratype, Ototo-jima Island, Chichi-jima Islands, Ogasawara Islands, Japan. Photo (KPM-NR 51087 B) by H. Senou." httpUri="https://zenodo.org/record/1038750/files/figure.png" pageId="4" pageNumber="71">Figures 2</figureCitation>
,
<figureCitation id="D6AB2A344C69E56BE5B945CAFD3FF950" box="[641,655,1751,1776]" captionStart="FIGURE 5" captionStartId="6.[151,250,1147,1169]" captionTargetBox="[151,1436,301,1126]" captionTargetId="figure@6.[151,1436,301,1126]" captionTargetPageId="6" captionText="FIGURE 5. Pseudanthias tequila n. sp., female, Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan. Photo by S. Kobayashi." httpUri="https://zenodo.org/record/1038756/files/figure.png" pageId="4" pageNumber="71">5</figureCitation>
&amp;
<figureCitation id="D6AB2A344C69E56BE58A45CAFD73F950" box="[690,707,1751,1776]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[151,1436,981,1945]" captionTargetId="figure@7.[151,1436,981,1945]" captionTargetPageId="7" captionText="FIGURE 6. Group of Pseudanthias tequila n. sp., Tinian, Mariana Islands, Micronesia. Photo by N. Tsuji." httpUri="https://zenodo.org/record/1038758/files/figure.png" pageId="4" pageNumber="71">6</figureCitation>
): head and body purple to orange-pink; snout, lips and upper part of head yellow-grey to bright yellow; iris yellow with submarginal purple ring around pupil; scales of nape and body purplish grey or brown to bright red basally; caudal peduncle yellow; dorsal fin greenish to yellowish hyaline anteriorly, reddish to greenish hyaline posteriorly, with bright purple-blue distal margin; anal fin red to yellow anteriorly, becoming reddish hyaline posteriorly, with bright purple-blue leading edge of fin; caudal fin bright yellow, yellowish hyaline distally on central rays, with upper and lower edges of fin narrowly bright purple-blue; pelvic fins yellowish hyaline, bright purple-blue on leading edge; pectoral fins pinkish to orangish hyaline.
</paragraph>
<caption id="1AEF66394C68E56AE7AF47C7FF49FACA" httpUri="https://zenodo.org/record/1038754/files/figure.png" pageId="5" pageNumber="72" targetBox="[151,1436,193,1220]" targetPageId="5">
<paragraph id="4E2F36B14C68E56AE7AF47C7FF49FACA" blockId="5.[151,1437,1242,1386]" pageId="5" pageNumber="72">
<emphasis id="7CE4EAA34C68E56AE7AF47C7FEA3FB4F" bold="true" box="[151,275,1242,1264]" pageId="5" pageNumber="72">FIGURE 4.</emphasis>
Males of selected
<taxonomicName id="89904D324C68E56AE6EB47C7FDEBFB50" authorityName="Bleeker" authorityYear="1871" box="[467,603,1242,1264]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C68E56AE6EB47C7FDEBFB50" box="[467,603,1242,1264]" italics="true" pageId="5" pageNumber="72">Pseudanthias</emphasis>
</taxonomicName>
species: A)
<taxonomicName id="89904D324C68E56AE5E347C6FC8AFB50" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[731,826,1242,1264]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C68E56AE5E347C6FC8AFB50" box="[731,826,1242,1264]" italics="true" pageId="5" pageNumber="72">P. tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C68E56AE47A47C6FCCAFB50" box="[834,890,1243,1264]" pageId="5" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
, Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan (KPM-NR 179391A, photo by O. Morishita); B)
<taxonomicName id="89904D324C68E56AE5E547E7FC8FFAAE" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[733,831,1272,1295]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C68E56AE5E547E7FC8FFAAE" box="[733,831,1272,1295]" italics="true" pageId="5" pageNumber="72">P. tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C68E56AE47047E4FC33FAAE" box="[840,899,1273,1294]" pageId="5" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
, Tinian, Mariana Islands (photo by N. Tsuji); C)
<taxonomicName id="89904D324C68E56AE2B147E7FEAEFA8D" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C68E56AE2B147E7FEAEFA8D" italics="true" pageId="5" pageNumber="72">P. pulcherrimus</emphasis>
</taxonomicName>
, aquarium specimen (not retained) from the Maldives (photo by Y.K. Tea); D)
<taxonomicName id="89904D324C68E56AE36A4605FB0DFA8D" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[1106,1213,1303,1325]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C68E56AE36A4605FB0DFA8D" box="[1106,1213,1303,1325]" italics="true" pageId="5" pageNumber="72">P. randalli</emphasis>
</taxonomicName>
, Kwajalein, Marshall Islands (photo by J. Johnson); E)
<taxonomicName id="89904D324C68E56AE53C462AFD3DFAEB" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[516,653,1333,1356]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C68E56AE53C462AFD3DFAEB" box="[516,653,1333,1356]" italics="true" pageId="5" pageNumber="72">P. flavicauda</emphasis>
</taxonomicName>
, Fiji (photo by J.E. Randall); F)
<emphasis id="7CE4EAA34C68E56AE4CF462AFBBBFAEC" box="[1015,1035,1335,1356]" italics="true" pageId="5" pageNumber="72">P.</emphasis>
sp., Kirimati, Kiribati (photo by D.J. Linehan).
</paragraph>
</caption>
<paragraph id="4E2F36B14C68E56AE7FF468CFC5FFA6E" blockId="5.[151,1437,1425,2026]" pageId="5" pageNumber="72">
<emphasis id="7CE4EAA34C68E56AE7FF468CFE72FA0A" box="[199,450,1425,1450]" italics="true" pageId="5" pageNumber="72">Colour in preservative</emphasis>
: both sexes generally pale tan, lighter ventrally; basal markings on nape and body scales remain, becoming greyish brown; other markings on body and fins obsolete.
</paragraph>
<paragraph id="4E2F36B14C68E56AE7FF46C4FD05F8B2" blockId="5.[151,1437,1425,2026]" pageId="5" pageNumber="72">
<emphasis id="7CE4EAA34C68E56AE7FF46C4FE45FA52" bold="true" box="[199,501,1497,1522]" pageId="5" pageNumber="72">Habitat and distribution.</emphasis>
<taxonomicName id="89904D324C68E56AE6C446C4FD59FA52" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[508,745,1497,1522]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C68E56AE6C446C4FD59FA52" box="[508,745,1497,1522]" italics="true" pageId="5" pageNumber="72">Pseudanthias tequila</emphasis>
</taxonomicName>
is described on the basis of two specimens from Ototo-jima, Chichi-jima Islands, Ogasawara Islands,
<collectingCountry id="368776214C68E56AE55246E2FD1DF9B6" box="[618,685,1535,1558]" name="Japan" pageId="5" pageNumber="72">Japan</collectingCountry>
. We also record it from
<collectingCountry id="368776214C68E56AE4F046E3FBBFF9B6" box="[968,1039,1534,1558]" name="Guam" pageId="5" pageNumber="72">Guam</collectingCountry>
,
<collectingRegion id="8C54F8534C68E56AE32446E0FBD8F9B6" box="[1052,1128,1533,1558]" country="Northern Mariana Islands" name="Saipan" pageId="5" pageNumber="72">Saipan</collectingRegion>
and
<collectingRegion id="8C54F8534C68E56AE39B46E0FB5BF9B6" box="[1187,1259,1533,1558]" country="Northern Mariana Islands" name="Tinian" pageId="5" pageNumber="72">Tinian</collectingRegion>
in the
<collectingCountry id="368776214C68E56AE20746E0FF56F99A" name="Northern Mariana Islands" pageId="5" pageNumber="72">Mariana Islands</collectingCountry>
on the basis of photographs (
<figureCitation id="D6AB2A344C68E56AE505453CFD11F99A" box="[573,673,1569,1594]" captionStart="FIGURE 7" captionStartId="8.[151,250,1650,1672]" captionTargetBox="[177,1393,807,1627]" captionTargetId="figure@8.[167,1403,807,1637]" captionTargetPageId="8" captionText="FIGURE 7. Distribution records for Pseudanthias tequila." httpUri="https://zenodo.org/record/1038760/files/figure.png" pageId="5" pageNumber="72">Figure 7</figureCitation>
). Kuiters (2004) photograph of an individual of “
<taxonomicName id="89904D324C68E56AE3DE453CFACDF99A" authorityName="Bleeker" authorityYear="1871" box="[1254,1405,1569,1594]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C68E56AE3DE453CFACDF99A" box="[1254,1405,1569,1594]" italics="true" pageId="5" pageNumber="72">Pseudanthias</emphasis>
</taxonomicName>
cf
<taxonomicName id="89904D324C68E56AE7AF4558FF41F9FE" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[151,241,1605,1630]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C68E56AE7AF4558FF41F9FE" box="[151,241,1605,1630]" italics="true" pageId="5" pageNumber="72">randalli</emphasis>
</taxonomicName>
”, stated to be from
<collectingCountry id="368776214C68E56AE6F74558FDA0F9FE" box="[463,528,1605,1630]" name="Palau" pageId="5" pageNumber="72">Palau</collectingCountry>
, is referable to
<taxonomicName id="89904D324C68E56AE584455AFC95F9FE" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[700,805,1605,1630]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C68E56AE584455AFC95F9FE" box="[700,805,1605,1630]" italics="true" pageId="5" pageNumber="72">P. tequila</emphasis>
</taxonomicName>
. However, R.F. Myers (pers. comm.) advised us that the photograph, by Hiroyuki Kimura, is actually from
<collectingRegion id="8C54F8534C68E56AE5D24574FC8AF922" box="[746,826,1641,1666]" country="Northern Mariana Islands" name="Saipan" pageId="5" pageNumber="72">Saipan</collectingRegion>
. Other photographs from
<collectingCountry id="368776214C68E56AE3524574FB18F922" box="[1130,1192,1641,1666]" name="Palau" pageId="5" pageNumber="72">Palau</collectingCountry>
are referrable to the closely related
<taxonomicName id="89904D324C68E56AE6784592FD6DF906" authority="(Lubbock &amp; Allen, 1978)" authorityName="Lubbock &amp; Allen" authorityYear="1978" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[320,733,1677,1702]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C68E56AE6784592FE05F906" box="[320,437,1677,1702]" italics="true" pageId="5" pageNumber="72">P. randalli</emphasis>
(
<bibRefCitation id="2A014B404C68E56AE6FC4590FD64F906" author="Lubbock" box="[452,724,1677,1702]" pageId="5" pageNumber="72" refString="Lubbock, R. &amp; Allen, G. R. (1978) A distinctive new Anthias (Teleostei: Serranidae) from the western Pacific. Records of the Western Australian Museum, 6, 259 - 268." type="journal article" year="1978">Lubbock &amp; Allen, 1978</bibRefCitation>
)
</taxonomicName>
. The original description of
<taxonomicName id="89904D324C68E56AE3214592FB3EF906" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[1049,1166,1677,1702]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C68E56AE3214592FB3EF906" box="[1049,1166,1677,1702]" italics="true" pageId="5" pageNumber="72">P. randalli</emphasis>
</taxonomicName>
also included
<typeStatus id="912B88134C68E56AE2094592FA2CF907" box="[1329,1436,1679,1703]" pageId="5" pageNumber="72" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="368776214C68E56AE7EF45ACFEA9F96A" box="[215,281,1713,1738]" name="Palau" pageId="5" pageNumber="72">Palau</collectingCountry>
.
<taxonomicName id="89904D324C68E56AE61D45ACFDA3F96A" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[293,531,1713,1738]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C68E56AE61D45ACFDA3F96A" box="[293,531,1713,1738]" italics="true" pageId="5" pageNumber="72">Pseudanthias tequila</emphasis>
</taxonomicName>
appears to be mostly found in caves on reef slopes in
<quantity id="89689B544C68E56AE3B645ACFB46F969" box="[1166,1270,1713,1738]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="72" unit="m" value="50.0" valueMax="60.0" valueMin="40.0">4060 m</quantity>
, earning it the popular name of “cave anthias”. In
<collectingRegion id="8C54F8534C68E56AE51045C8FDC4F94E" box="[552,628,1749,1774]" country="Northern Mariana Islands" name="Saipan" pageId="5" pageNumber="72">Saipan</collectingRegion>
and
<collectingRegion id="8C54F8534C68E56AE59445C8FD47F94E" box="[684,759,1749,1774]" country="Northern Mariana Islands" name="Tinian" pageId="5" pageNumber="72">Tinian</collectingRegion>
, however, it occurs in the open on reef slopes dominated by
<taxonomicName id="89904D324C68E56AE7AF45E4FEB5F8B2" authorityName="J.V.Lamouroux" authorityYear="1812" box="[151,261,1785,1810]" class="Ulvophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="5" pageNumber="72" phylum="Chlorophyta" rank="genus">
<emphasis id="7CE4EAA34C68E56AE7AF45E4FEB5F8B2" box="[151,261,1785,1810]" italics="true" pageId="5" pageNumber="72">Halimeda</emphasis>
</taxonomicName>
and
<taxonomicName id="89904D324C68E56AE60445E4FE16F8B2" authorityName="J.V. Lamouroux" authorityYear="1809" box="[316,422,1785,1810]" class="Ulvophyceae" family="Caulerpaceae" genus="Caulerpa" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="5" pageNumber="72" phylum="Chlorophyta" rank="genus">
<emphasis id="7CE4EAA34C68E56AE60445E4FE16F8B2" box="[316,422,1785,1810]" italics="true" pageId="5" pageNumber="72">Caulerpa</emphasis>
</taxonomicName>
algae (
<figureCitation id="D6AB2A344C68E56AE6CE45E4FDC3F8B2" box="[502,627,1785,1810]" captionStart="FIGURE 4" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1436,193,1220]" captionTargetId="figure@5.[151,1436,193,1220]" captionTargetPageId="5" captionText="FIGURE 4. Males of selected Pseudanthias species: A) P. tequila n. sp., Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan (KPM-NR 179391 A, photo by O. Morishita); B) P. tequila n. sp., Tinian, Mariana Islands (photo by N. Tsuji); C) P. pulcherrimus, aquarium specimen (not retained) from the Maldives (photo by Y. K. Tea); D) P. randalli, Kwajalein, Marshall Islands (photo by J. Johnson); E) P. flavicauda, Fiji (photo by J. E. Randall); F) P. sp., Kirimati, Kiribati (photo by D. J. Linehan)." httpUri="https://zenodo.org/record/1038754/files/figure.png" pageId="5" pageNumber="72">Figures 4B</figureCitation>
&amp;
<figureCitation id="D6AB2A344C68E56AE5AF45E4FD17F8B2" box="[663,679,1785,1810]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[151,1436,981,1945]" captionTargetId="figure@7.[151,1436,981,1945]" captionTargetPageId="7" captionText="FIGURE 6. Group of Pseudanthias tequila n. sp., Tinian, Mariana Islands, Micronesia. Photo by N. Tsuji." httpUri="https://zenodo.org/record/1038758/files/figure.png" pageId="5" pageNumber="72">6</figureCitation>
).
</paragraph>
<paragraph id="4E2F36B14C68E56AE7FF4400FA2DF84A" blockId="5.[151,1437,1425,2026]" pageId="5" pageNumber="72">
<emphasis id="7CE4EAA34C68E56AE7FF4400FEDDF896" bold="true" box="[199,365,1821,1846]" pageId="5" pageNumber="72">Comparisons.</emphasis>
<taxonomicName id="89904D324C68E56AE64F4400FDD6F896" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[375,614,1821,1846]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C68E56AE64F4400FDD6F896" box="[375,614,1821,1846]" italics="true" pageId="5" pageNumber="72">Pseudanthias tequila</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="89904D324C68E56AE4BC4402FA9FF896" authority="(Lubbock &amp; Allen, 1978)" authorityName="Lubbock &amp; Allen" authorityYear="1978" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[900,1327,1821,1846]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C68E56AE4BC4402FC4DF896" box="[900,1021,1821,1846]" italics="true" pageId="5" pageNumber="72">P. randalli</emphasis>
(
<bibRefCitation id="2A014B404C68E56AE3364400FA96F896" author="Lubbock" box="[1038,1318,1821,1846]" pageId="5" pageNumber="72" refString="Lubbock, R. &amp; Allen, G. R. (1978) A distinctive new Anthias (Teleostei: Serranidae) from the western Pacific. Records of the Western Australian Museum, 6, 259 - 268." type="journal article" year="1978">Lubbock &amp; Allen, 1978</bibRefCitation>
)
</taxonomicName>
from the West Pacific,
<taxonomicName id="89904D324C68E56AE60D445EFC80F8FA" authority="Heemstra &amp; Randall (1986)" authorityName="Heemstra &amp; Randall" authorityYear="1986" box="[309,816,1857,1882]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C68E56AE60D445EFE58F8FA" box="[309,488,1857,1882]" italics="true" pageId="5" pageNumber="72">P. pulcherrimus</emphasis>
<bibRefCitation id="2A014B404C68E56AE6CA445FFC80F8FA" author="Heemstra" box="[498,816,1857,1882]" pageId="5" pageNumber="72" refString="Heemstra, P. C. &amp; Randall, J. E. (1986) Family No. 166: Serranidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smiths' Sea Fishes. Macmillan South Africa, Johannesburg, pp. 509 - 537." type="book chapter" year="1986">Heemstra &amp; Randall (1986)</bibRefCitation>
</taxonomicName>
from the Indian Ocean,
<taxonomicName id="89904D324C68E56AE375445EFF51F8DE" authority="Randall &amp; Pyle (2001)" authorityName="Randall &amp; Pyle" authorityYear="2001" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C68E56AE375445EFB51F8FA" box="[1101,1249,1857,1882]" italics="true" pageId="5" pageNumber="72">P. flavicauda</emphasis>
<bibRefCitation id="2A014B404C68E56AE3D3445CFF51F8DE" author="Randall" pageId="5" pageNumber="72" refString="Randall, J. E. &amp; Pyle, R. L. (2001) Four new serranid fishes of the anthiine genus Pseudanthias from the South Pacific. Raffles Bulletin of Zoology, 49, 19 - 34." type="journal article" year="2001">Randall &amp; Pyle (2001)</bibRefCitation>
</taxonomicName>
from the south-west Pacific, and
<taxonomicName id="89904D324C68E56AE565447AFB1DF8DE" authority="Williams, Delrieu-Trottin &amp; Planes (2013)" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[605,1197,1893,1918]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="oumati">
<emphasis id="7CE4EAA34C68E56AE565447AFD77F8DE" box="[605,711,1895,1918]" italics="true" pageId="5" pageNumber="72">P. oumati</emphasis>
<bibRefCitation id="2A014B404C68E56AE5F64478FB1DF8DE" box="[718,1197,1893,1918]" pageId="5" pageNumber="72" refString="Williams, J. T., Delrieu-Trottin, E. &amp; Planes, S. (2013) Two new fish species of the subfamily Anthiinae (Perciformes, Serranidae) from the Marquesas. Zootaxa, 3647 (1), 167 - 180." type="journal article">Williams, Delrieu-Trottin &amp; Planes (2013)</bibRefCitation>
</taxonomicName>
from the Marquesas. The five species share the following combination of characters: dorsal rays usually X,16; anal rays usually III,7; third dorsal spine prolonged at least in males; third segmented anal-fin ray longest; lateral-line scales 4050; no auxiliary scales on body; and interopercle and subopercle with distinct serrations in adults. Males of
<taxonomicName id="89904D324C68E56AE23644CEFAC9F84A" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[1294,1401,2001,2026]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C68E56AE23644CEFAC9F84A" box="[1294,1401,2001,2026]" italics="true" pageId="5" pageNumber="72">P. tequila</emphasis>
</taxonomicName>
,
<emphasis id="7CE4EAA34C68E56AE2BF44CEFA2DF84A" box="[1415,1437,2003,2026]" italics="true" pageId="5" pageNumber="72">P.</emphasis>
</paragraph>
<paragraph id="4E2F36B14C6BE569E7AF438AFE43FF58" blockId="6.[151,1436,151,248]" pageId="6" pageNumber="73">
<emphasis id="7CE4EAA34C6BE569E7AF438AFE23FF10" box="[151,403,151,176]" italics="true" pageId="6" pageNumber="73">
<taxonomicName id="89904D324C6BE569E7AF438AFEA0FF10" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[151,272,151,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="flavicauda">flavicauda</taxonomicName>
,
<taxonomicName id="89904D324C6BE569E6244384FE23FF10" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[284,403,151,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="randalli">P. randalli</taxonomicName>
</emphasis>
and
<taxonomicName id="89904D324C6BE569E6F44384FDCDFF10" box="[460,637,151,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C6BE569E6F44384FDCDFF10" box="[460,637,151,176]" italics="true" pageId="6" pageNumber="73">P. pulcherrimus</emphasis>
</taxonomicName>
are also unusual in exhibiting weak to pronounced hypertrophy of the upper lip (see Remarks below). The condition of the upper lip is unknown in
<taxonomicName id="89904D324C6BE569E32C43A0FBCFFF74" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[1044,1151,189,212]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="oumati">
<emphasis id="7CE4EAA34C6BE569E32C43A0FBCFFF74" box="[1044,1151,189,212]" italics="true" pageId="6" pageNumber="73">P. oumati</emphasis>
</taxonomicName>
, as the species is known only from the female
<typeStatus id="912B88134C6BE569E6B143C2FE5FFF58" box="[393,495,223,248]" pageId="6" pageNumber="73" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="1AEF66394C6BE569E7AF4766FEBAFB10" httpUri="https://zenodo.org/record/1038756/files/figure.png" pageId="6" pageNumber="73" targetBox="[151,1436,301,1126]" targetPageId="6">
<paragraph id="4E2F36B14C6BE569E7AF4766FEBAFB10" blockId="6.[151,1435,1147,1200]" pageId="6" pageNumber="73">
<emphasis id="7CE4EAA34C6BE569E7AF4766FEA6FB31" bold="true" box="[151,278,1147,1169]" pageId="6" pageNumber="73">FIGURE 5.</emphasis>
<taxonomicName id="89904D324C6BE569E6194766FE4AFB31" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[289,506,1147,1169]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C6BE569E6194766FE4AFB31" box="[289,506,1147,1169]" italics="true" pageId="6" pageNumber="73">Pseudanthias tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C6BE569E53C4761FDF0FB31" box="[516,576,1148,1169]" pageId="6" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
, female, Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan. Photo by S. Kobayashi.
</paragraph>
</caption>
<paragraph id="4E2F36B14C6BE569E7FF47CAFF40F994" blockId="6.[151,1437,1239,2020]" pageId="6" pageNumber="73">
<taxonomicName id="89904D324C6BE569E7FF47CAFE03FB50" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[199,435,1239,1264]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C6BE569E7FF47CAFE03FB50" box="[199,435,1239,1264]" italics="true" pageId="6" pageNumber="73">Pseudanthias tequila</emphasis>
</taxonomicName>
is distinguished from at least
<taxonomicName id="89904D324C6BE569E43F47C5FCCEFB50" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[775,894,1239,1264]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C6BE569E43F47C5FCCEFB50" box="[775,894,1239,1264]" italics="true" pageId="6" pageNumber="73">P. randalli</emphasis>
</taxonomicName>
,
<taxonomicName id="89904D324C6BE569E4B447C5FB8EFB50" box="[908,1086,1239,1264]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C6BE569E4B447C5FB8EFB50" box="[908,1086,1239,1264]" italics="true" pageId="6" pageNumber="73">P. pulcherrimus</emphasis>
</taxonomicName>
and
<taxonomicName id="89904D324C6BE569E34E47C5FAB8FB50" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[1142,1288,1239,1264]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C6BE569E34E47C5FAB8FB50" box="[1142,1288,1239,1264]" italics="true" pageId="6" pageNumber="73">P. flavicauda</emphasis>
</taxonomicName>
by male live colouration. The live colouration of the single known female specimen of
<taxonomicName id="89904D324C6BE569E4EB47E0FB8CFAB4" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[979,1084,1277,1300]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="oumati">
<emphasis id="7CE4EAA34C6BE569E4EB47E0FB8CFAB4" box="[979,1084,1277,1300]" italics="true" pageId="6" pageNumber="73">P. oumati</emphasis>
</taxonomicName>
is distinctive in being more-orless uniformly yellow.
<bibRefCitation id="2A014B404C6BE569E69C4602FD16FA98" box="[420,678,1311,1336]" pageId="6" pageNumber="73" refString="Williams, J. T., Delrieu-Trottin, E. &amp; Planes, S. (2013) Two new fish species of the subfamily Anthiinae (Perciformes, Serranidae) from the Marquesas. Zootaxa, 3647 (1), 167 - 180." type="journal article">
Williams
<emphasis id="7CE4EAA34C6BE569E52C463DFDE0FA98" box="[532,592,1311,1336]" italics="true" pageId="6" pageNumber="73">et al.</emphasis>
(2013)
</bibRefCitation>
noted that the
<typeStatus id="912B88134C6BE569E4654602FC0FFA98" box="[861,959,1311,1336]" pageId="6" pageNumber="73" type="holotype">holotype</typeStatus>
was collected from a group of similarly coloured individuals. Given that
<taxonomicName id="89904D324C6BE569E53F465EFD2EFAFC" authorityName="Bleeker" authorityYear="1871" box="[519,670,1347,1372]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6BE569E53F465EFD2EFAFC" box="[519,670,1347,1372]" italics="true" pageId="6" pageNumber="73">Pseudanthias</emphasis>
</taxonomicName>
species are protogynous hermaphrodites that typically live in shoals consisting of female and juvenile individuals guarded by a smaller number of males, it is likely that the shoal observed by Williams
<emphasis id="7CE4EAA34C6BE569E6A94690FE79FA04" box="[401,457,1419,1444]" italics="true" pageId="6" pageNumber="73">et al.</emphasis>
included at least one male, and that males of the species are also predominantly yellow. Males of the remaining species are shown in
<figureCitation id="D6AB2A344C6BE569E5A746B2FCB2FA68" box="[671,770,1455,1480]" captionStart="FIGURE 4" captionStartId="5.[151,250,1242,1264]" captionTargetBox="[151,1436,193,1220]" captionTargetId="figure@5.[151,1436,193,1220]" captionTargetPageId="5" captionText="FIGURE 4. Males of selected Pseudanthias species: A) P. tequila n. sp., Ototo-jima, Chichi-jima Islands, Ogasawara Islands, Japan (KPM-NR 179391 A, photo by O. Morishita); B) P. tequila n. sp., Tinian, Mariana Islands (photo by N. Tsuji); C) P. pulcherrimus, aquarium specimen (not retained) from the Maldives (photo by Y. K. Tea); D) P. randalli, Kwajalein, Marshall Islands (photo by J. Johnson); E) P. flavicauda, Fiji (photo by J. E. Randall); F) P. sp., Kirimati, Kiribati (photo by D. J. Linehan)." httpUri="https://zenodo.org/record/1038754/files/figure.png" pageId="6" pageNumber="73">Figure 4</figureCitation>
. We also include an additional, undescribed species in the figure. That species, hereafter termed
<taxonomicName id="89904D324C6BE569E57546CEFD54FA4C" authorityName="Bleeker" authorityYear="1871" box="[589,740,1491,1516]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6BE569E57546CEFD54FA4C" box="[589,740,1491,1516]" italics="true" pageId="6" pageNumber="73">Pseudanthias</emphasis>
</taxonomicName>
sp., is known on the basis of aquarium specimens collected from Kirimati in the Line Islands. Male colouration characters distinguishing the five species are summarised in
<tableCitation id="0312030A4C6BE569E7AF4506FF5DF994" box="[151,237,1563,1588]" captionStart="TABLE 2" captionStartId="7.[151,239,332,355]" captionText="TABLE 2. Summary of male live colouration characters for selected Pseudanthias species." pageId="6" pageNumber="73">Table 2</tableCitation>
.
</paragraph>
<paragraph id="4E2F36B14C6BE569E7FF4522FE8CF890" blockId="6.[151,1437,1239,2020]" pageId="6" pageNumber="73">
Aside from live colouration,
<taxonomicName id="89904D324C6BE569E536455DFDC9F9F8" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[526,633,1599,1624]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C6BE569E536455DFDC9F9F8" box="[526,633,1599,1624]" italics="true" pageId="6" pageNumber="73">P. tequila</emphasis>
</taxonomicName>
differs: from
<taxonomicName id="89904D324C6BE569E42F455DFC19F9F8" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[791,937,1599,1624]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C6BE569E42F455DFC19F9F8" box="[791,937,1599,1624]" italics="true" pageId="6" pageNumber="73">P. flavicauda</emphasis>
</taxonomicName>
in being deeper bodied (33.735.2 vs 30.1 32.5 % SL), with a longer second anal-fin spine (18.420.6 vs 16.817.6 % SL), and fewer circumpeduncular scales (24 vs 2627); from
<taxonomicName id="89904D324C6BE569E6FF4595FD80F93F" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[455,560,1672,1695]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="oumati">
<emphasis id="7CE4EAA34C6BE569E6FF4595FD80F93F" box="[455,560,1672,1695]" italics="true" pageId="6" pageNumber="73">P. oumati</emphasis>
</taxonomicName>
in being shallower bodied (33.735.2 vs 40 % SL), with fewer lower limb gill rakers (2123 vs 28) and fewer circumpeduncular scales (24 vs 27); from
<taxonomicName id="89904D324C6BE569E4E345B0FBE1F964" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[987,1105,1707,1732]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C6BE569E4E345B0FBE1F964" box="[987,1105,1707,1732]" italics="true" pageId="6" pageNumber="73">P. randalli</emphasis>
</taxonomicName>
in having a shorter predorsal length (27.529.5 vs 30.232.4 % SL), a longer second anal-fin spine (18.420.6 vs 15.5 17.0 % SL) and more pectoral-fin rays (18 vs 16-18, usually 17); and from
<taxonomicName id="89904D324C6BE569E43D45E8FC09F8AC" box="[773,953,1779,1804]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C6BE569E43D45E8FC09F8AC" box="[773,953,1779,1804]" italics="true" pageId="6" pageNumber="73">P. pulcherrimus</emphasis>
</taxonomicName>
in having more lateral-line scales (4649 versus 4045).
</paragraph>
<paragraph id="4E2F36B14C6BE568E7FF4421FBB5FF75" blockId="6.[151,1437,1239,2020]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
In describing
<taxonomicName id="89904D324C6BE569E6534420FE0EF8F4" authorityName="Allen &amp; Erdmann" authorityYear="2012" box="[363,446,1853,1876]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="mica">
<emphasis id="7CE4EAA34C6BE569E6534420FE0EF8F4" box="[363,446,1853,1876]" italics="true" pageId="6" pageNumber="73">P. mica</emphasis>
</taxonomicName>
from Lembatta Island (
<collectingRegion id="8C54F8534C6BE569E5E64421FC69F8F4" box="[734,985,1851,1876]" country="Indonesia" name="Nusa Tenggara Timur" pageId="6" pageNumber="73">Lesser Sunda Islands</collectingRegion>
,
<collectingCountry id="368776214C6BE569E4D14426FBEAF8F4" box="[1001,1114,1851,1876]" name="Indonesia" pageId="6" pageNumber="73">Indonesia</collectingCountry>
),
<bibRefCitation id="2A014B404C6BE569E34A4426FA2CF8F4" author="Allen" box="[1138,1436,1851,1876]" pageId="6" pageNumber="73" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef Fishes of the East Indies. Volumes I - III. Tropical Reef Research, Perth, 1292 pp." type="book" year="2012">Allen &amp; Erdmann (2012)</bibRefCitation>
suggested a close relationship to
<taxonomicName id="89904D324C6BE569E520447DFD20F8D8" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[536,656,1887,1912]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C6BE569E520447DFD20F8D8" box="[536,656,1887,1912]" italics="true" pageId="6" pageNumber="73">P. randalli</emphasis>
</taxonomicName>
and
<taxonomicName id="89904D324C6BE569E5F5447DFC31F8D8" box="[717,897,1887,1912]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C6BE569E5F5447DFC31F8D8" box="[717,897,1887,1912]" italics="true" pageId="6" pageNumber="73">P. pulcherrimus</emphasis>
</taxonomicName>
.
<bibRefCitation id="2A014B404C6BE569E4AA4442FB22F8D8" box="[914,1170,1887,1912]" pageId="6" pageNumber="73" refString="Williams, J. T., Delrieu-Trottin, E. &amp; Planes, S. (2013) Two new fish species of the subfamily Anthiinae (Perciformes, Serranidae) from the Marquesas. Zootaxa, 3647 (1), 167 - 180." type="journal article">
Williams
<emphasis id="7CE4EAA34C6BE569E33A447DFB8DF8D8" box="[1026,1085,1887,1912]" italics="true" pageId="6" pageNumber="73">et al.</emphasis>
(2013)
</bibRefCitation>
also suggested a close relationship between the three species and
<taxonomicName id="89904D324C6BE569E5B74498FD4DF83C" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[655,765,1925,1948]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="oumati">
<emphasis id="7CE4EAA34C6BE569E5B74498FD4DF83C" box="[655,765,1925,1948]" italics="true" pageId="6" pageNumber="73">P. oumati</emphasis>
</taxonomicName>
. However, aside from several minor differences (such as number of gill rakers), Allen &amp; Erdmanns description of
<taxonomicName id="89904D324C6BE569E41144B5FCCBF81F" authorityName="Allen &amp; Erdmann" authorityYear="2012" box="[809,891,1960,1983]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="mica">
<emphasis id="7CE4EAA34C6BE569E41144B5FCCBF81F" box="[809,891,1960,1983]" italics="true" pageId="6" pageNumber="73">P. mica</emphasis>
</taxonomicName>
agrees well with Katayama &amp; Masudas (1982) original description of
<emphasis id="7CE4EAA34C6BE569E6A544D6FDC9F844" box="[413,633,1995,2020]" italics="true" pageId="6" pageNumber="73">
Anthias
<taxonomicName id="89904D324C6BE569E6C444D6FDC9F844" baseAuthorityName="Katayama &amp; Masuda" baseAuthorityYear="1982" box="[508,633,1995,2020]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="leucozonus">leucozonus</taxonomicName>
</emphasis>
(now
<taxonomicName id="89904D324C6BE569E5FD44D6FC52F844" baseAuthorityName="Katayama &amp; Masuda" baseAuthorityYear="1982" box="[709,994,1995,2020]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="leucozonus">
<emphasis id="7CE4EAA34C6BE569E5FD44D6FC52F844" box="[709,994,1995,2020]" italics="true" pageId="6" pageNumber="73">Pseudanthias leucozonus</emphasis>
</taxonomicName>
) from southern
<collectingCountry id="368776214C6BE569E3A644D1FB51F843" box="[1182,1249,1996,2019]" name="Japan" pageId="6" pageNumber="73">Japan</collectingCountry>
, and it is highly probable that the two species are closely related. Both nominal species depart from
<emphasis id="7CE4EAA34C6AE568E3024384FA2CFF10" box="[1082,1436,151,176]" italics="true" pageId="7" pageNumber="74">
<taxonomicName id="89904D324C6AE568E3024384FB15FF10" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[1082,1189,151,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="tequila">P. tequila</taxonomicName>
,
<taxonomicName id="89904D324C6AE568E3884384FAACFF10" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[1200,1308,153,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="oumati">P. oumati</taxonomicName>
,
<taxonomicName id="89904D324C6AE568E21E4384FA2CFF10" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[1318,1436,151,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="randalli">P. randalli</taxonomicName>
</emphasis>
and
<taxonomicName id="89904D324C6AE568E7FE43A0FEC7FF75" box="[198,375,188,213]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C6AE568E7FE43A0FEC7FF75" box="[198,375,188,213]" italics="true" pageId="7" pageNumber="74">P. pulcherrimus</emphasis>
</taxonomicName>
in having fewer lateral-lines scales (3638 versus 4050).
</paragraph>
</subSubSection>
<subSubSection id="068A653A4C6AE568E7FF43C2FC79FEBD" pageId="7" pageNumber="74" type="etymology">
<paragraph id="4E2F36B14C6AE568E7FF43C2FC79FEBD" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="74">
<emphasis id="7CE4EAA34C6AE568E7FF43C2FEFEFF58" bold="true" box="[199,334,223,248]" pageId="7" pageNumber="74">Etymology.</emphasis>
The specific epithet refers to the alcoholic beverage
<taxonomicName id="89904D324C6AE568E48F43C2FBB3FF58" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[951,1027,223,248]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="tequila">tequila</taxonomicName>
sunrise, alluding to the vibrant life colours of the males of the species. To be treated as a noun in apposition.
</paragraph>
</subSubSection>
<caption id="1AEF66394C6AE568E7AF4251FBD5FEC3" box="[151,1125,331,355]" pageId="7" pageNumber="74">
<paragraph id="4E2F36B14C6AE568E7AF4251FBD5FEC3" blockId="7.[151,1125,331,355]" box="[151,1125,331,355]" pageId="7" pageNumber="74">
<emphasis id="7CE4EAA34C6AE568E7AF4251FEB9FEC3" bold="true" box="[151,265,332,355]" pageId="7" pageNumber="74">TABLE 2.</emphasis>
Summary of male live colouration characters for selected
<taxonomicName id="89904D324C6AE568E4424256FBB9FEC3" authorityName="Bleeker" authorityYear="1871" box="[890,1033,331,355]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C6AE568E4424256FBB9FEC3" box="[890,1033,331,355]" italics="true" pageId="7" pageNumber="74">Pseudanthias</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="4E2F36B14C6AE568E674429DFAD2FC8F" pageId="7" pageNumber="74">
<table id="3C90C4114C6A1A90E7A74263FA23FC2B" box="[159,1427,382,907]" gridcols="6" gridrows="5" pageId="7" pageNumber="74">
<tr id="F0A034F34C6A1A90E7A74263FA23FE35" box="[159,1427,382,405]" gridrow="0" pageId="7" pageNumber="74" rowspan-0="1">
<th id="B3715D8F4C6A1A90E6744263FDB3FE35" box="[332,515,382,405]" gridcol="1" gridrow="0" pageId="7" pageNumber="74">
<taxonomicName id="89904D324C6AE568E674429DFE60FE34" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[332,464,382,405]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C6AE568E674429DFE60FE34" box="[332,464,382,405]" italics="true" pageId="7" pageNumber="74">P. flavicauda</emphasis>
</taxonomicName>
</th>
<th id="B3715D8F4C6A1A90E5274263FD49FE35" box="[543,761,382,405]" gridcol="2" gridrow="0" pageId="7" pageNumber="74">
<taxonomicName id="89904D324C6AE568E527429DFD0EFE34" box="[543,702,382,405]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C6AE568E527429DFD0EFE34" box="[543,702,382,405]" italics="true" pageId="7" pageNumber="74">P. pulcherrimus</emphasis>
</taxonomicName>
</th>
<th id="B3715D8F4C6A1A90E41D4263FC56FE35" box="[805,998,382,405]" gridcol="3" gridrow="0" pageId="7" pageNumber="74">
<taxonomicName id="89904D324C6AE568E41D429DFC20FE34" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[805,912,382,405]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C6AE568E41D429DFC20FE34" box="[805,912,382,405]" italics="true" pageId="7" pageNumber="74">P. randalli</emphasis>
</taxonomicName>
</th>
<th id="B3715D8F4C6A1A90E4C34263FB71FE35" box="[1019,1217,382,405]" gridcol="4" gridrow="0" pageId="7" pageNumber="74">
<taxonomicName id="89904D324C6AE568E4C3429DFBEAFE34" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[1019,1114,382,405]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C6AE568E4C3429DFBEAFE34" box="[1019,1114,382,405]" italics="true" pageId="7" pageNumber="74">P. tequila</emphasis>
</taxonomicName>
</th>
<th id="B3715D8F4C6A1A90E3EE4263FA23FE35" box="[1238,1427,382,405]" gridcol="5" gridrow="0" pageId="7" pageNumber="74">
<emphasis id="7CE4EAA34C6AE568E3EE429DFB5AFE35" box="[1238,1258,384,405]" italics="true" pageId="7" pageNumber="74">P.</emphasis>
sp.
</th>
</tr>
<tr id="F0A034F34C6A1A90E7A742B5FA23FDBA" box="[159,1427,424,538]" gridrow="1" pageId="7" pageNumber="74">
<th id="B3715D8F4C6A1A90E7A742B5FE89FDBA" box="[159,313,424,538]" gridcol="0" gridrow="1" pageId="7" pageNumber="74">Upper body beneath anterior part of dorsal fin</th>
<td id="B3715D8F4C6A1A90E67442B5FDB3FDBA" box="[332,515,424,538]" gridcol="1" gridrow="1" pageId="7" pageNumber="74">Mostly yellow</td>
<td id="B3715D8F4C6A1A90E52742B5FD49FDBA" box="[543,761,424,538]" gridcol="2" gridrow="1" pageId="7" pageNumber="74">Purple to pink with orange-red stripe</td>
<td id="B3715D8F4C6A1A90E41D42B5FC56FDBA" box="[805,998,424,538]" gridcol="3" gridrow="1" pageId="7" pageNumber="74">Purple to pink with orange-red stripe</td>
<td id="B3715D8F4C6A1A90E4C342B5FB71FDBA" box="[1019,1217,424,538]" gridcol="4" gridrow="1" pageId="7" pageNumber="74">Purple to pink with orange-red stripe</td>
<td id="B3715D8F4C6A1A90E3EE42B5FA23FDBA" box="[1238,1427,424,538]" gridcol="5" gridrow="1" pageId="7" pageNumber="74">Purple to pink with orange-red stripe</td>
</tr>
<tr id="F0A034F34C6A1A90E7A74130FA23FD1E" box="[159,1427,557,702]" gridrow="2" pageId="7" pageNumber="74">
<th id="B3715D8F4C6A1A90E7A74130FE89FD1E" box="[159,313,557,702]" gridcol="0" gridrow="2" pageId="7" pageNumber="74">Dorsal fin</th>
<td id="B3715D8F4C6A1A90E6744130FDB3FD1E" box="[332,515,557,702]" gridcol="1" gridrow="2" pageId="7" pageNumber="74">Yellow, with purple/blue basal area posteriorly</td>
<td id="B3715D8F4C6A1A90E5274130FD49FD1E" box="[543,761,557,702]" gridcol="2" gridrow="2" pageId="7" pageNumber="74">Greenish yellow with blue basal area posteriorly</td>
<td id="B3715D8F4C6A1A90E41D4130FC56FD1E" box="[805,998,557,702]" gridcol="3" gridrow="2" pageId="7" pageNumber="74">Orange-red anteriorly, posteriorly red with blue basal area</td>
<td id="B3715D8F4C6A1A90E4C34130FB71FD1E" box="[1019,1217,557,702]" gridcol="4" gridrow="2" pageId="7" pageNumber="74">Greenish yellow anteriorly, posteriorly red with blue basal area</td>
<td id="B3715D8F4C6A1A90E3EE4130FA23FD1E" box="[1238,1427,557,702]" gridcol="5" gridrow="2" pageId="7" pageNumber="74">Yellow, with purple/blue basal area posteriorly</td>
</tr>
<tr id="F0A034F34C6A1A90E7A741CCFA23FCA6" box="[159,1427,721,774]" gridrow="3" pageId="7" pageNumber="74">
<th id="B3715D8F4C6A1A90E7A741CCFE89FCA6" box="[159,313,721,774]" gridcol="0" gridrow="3" pageId="7" pageNumber="74">Anterior part of anal fin</th>
<td id="B3715D8F4C6A1A90E67441CCFDB3FCA6" box="[332,515,721,774]" gridcol="1" gridrow="3" pageId="7" pageNumber="74">Yellow</td>
<td id="B3715D8F4C6A1A90E52741CCFD49FCA6" box="[543,761,721,774]" gridcol="2" gridrow="3" pageId="7" pageNumber="74">Yellow to orange-red</td>
<td id="B3715D8F4C6A1A90E41D41CCFC56FCA6" box="[805,998,721,774]" gridcol="3" gridrow="3" pageId="7" pageNumber="74">Red</td>
<td id="B3715D8F4C6A1A90E4C341CCFB71FCA6" box="[1019,1217,721,774]" gridcol="4" gridrow="3" pageId="7" pageNumber="74">Red</td>
<td id="B3715D8F4C6A1A90E3EE41CCFA23FCA6" box="[1238,1427,721,774]" gridcol="5" gridrow="3" pageId="7" pageNumber="74">Yellow</td>
</tr>
<tr id="F0A034F34C6A1A90E7A74004FA23FC2B" box="[159,1427,793,907]" gridrow="4" pageId="7" pageNumber="74">
<th id="B3715D8F4C6A1A90E7A74004FE89FC2B" box="[159,313,793,907]" gridcol="0" gridrow="4" pageId="7" pageNumber="74">Pelvic fins</th>
<td id="B3715D8F4C6A1A90E6744004FDB3FC2B" box="[332,515,793,907]" gridcol="1" gridrow="4" pageId="7" pageNumber="74">Yellowish hyaline with purple leading edge</td>
<td id="B3715D8F4C6A1A90E5274004FD49FC2B" box="[543,761,793,907]" gridcol="2" gridrow="4" pageId="7" pageNumber="74">Yellow to orange anteriorly, bright red posteriorly, with pale blue basal wedge</td>
<td id="B3715D8F4C6A1A90E41D4004FC56FC2B" box="[805,998,793,907]" gridcol="3" gridrow="4" pageId="7" pageNumber="74">Bright red, with pale blue basal wedge</td>
<td id="B3715D8F4C6A1A90E4C34004FB71FC2B" box="[1019,1217,793,907]" gridcol="4" gridrow="4" pageId="7" pageNumber="74">Bright yellow</td>
<td id="B3715D8F4C6A1A90E3EE4004FA23FC2B" box="[1238,1427,793,907]" gridcol="5" gridrow="4" pageId="7" pageNumber="74">Bright yellow</td>
</tr>
</table>
</paragraph>
<caption id="1AEF66394C6AE568E7AF44B3FB7EF864" box="[151,1230,1966,1988]" httpUri="https://zenodo.org/record/1038758/files/figure.png" pageId="7" pageNumber="74" targetBox="[151,1436,981,1945]" targetPageId="7">
<paragraph id="4E2F36B14C6AE568E7AF44B3FB7EF864" blockId="7.[151,1230,1966,1988]" box="[151,1230,1966,1988]" pageId="7" pageNumber="74">
<emphasis id="7CE4EAA34C6AE568E7AF44B3FEA2F863" bold="true" box="[151,274,1966,1988]" pageId="7" pageNumber="74">FIGURE 6.</emphasis>
Group of
<taxonomicName id="89904D324C6AE568E64344B3FDE0F864" authority="Gill &amp; Tea &amp; Senou, 2017" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[379,592,1966,1988]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="tequila" status="sp. nov.">
<emphasis id="7CE4EAA34C6AE568E64344B3FDE0F864" box="[379,592,1966,1988]" italics="true" pageId="7" pageNumber="74">Pseudanthias tequila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67D757D84C6AE568E56F44B2FD3FF864" box="[599,655,1967,1988]" pageId="7" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
, Tinian, Mariana Islands, Micronesia. Photo by N. Tsuji.
</paragraph>
</caption>
<subSubSection id="068A653A4C65E567E7FF438AFF73FD50" pageId="8" pageNumber="75" type="discussion">
<paragraph id="4E2F36B14C65E567E7FF438AFF73FD50" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="75">
<emphasis id="7CE4EAA34C65E567E7FF438AFE8BFF10" bold="true" box="[199,315,151,176]" pageId="8" pageNumber="75">Remarks.</emphasis>
Placement of the new species in
<taxonomicName id="89904D324C65E567E5F3438AFCD2FF10" authorityName="Bleeker" authorityYear="1871" box="[715,866,151,176]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E5F3438AFCD2FF10" box="[715,866,151,176]" italics="true" pageId="8" pageNumber="75">Pseudanthias</emphasis>
</taxonomicName>
should be regarded as provisional. The genus is poorly diagnosed and preliminary analysis of morphological characters by the first author suggests some species currently included in the genus may be more closely related to other genera, such as
<taxonomicName id="89904D324C65E567E35743C2FA27FF58" authority="Smith (1954)" authorityName="Smith" authorityYear="1954" box="[1135,1431,223,248]" class="Actinopterygii" family="Serranidae" genus="Nemanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E35743C2FB45FF58" box="[1135,1269,223,248]" italics="true" pageId="8" pageNumber="75">Nemanthias</emphasis>
<bibRefCitation id="2A014B404C65E567E3C643C2FA27FF58" author="Smith" box="[1278,1431,223,248]" pageId="8" pageNumber="75" refString="Smith, J. L. B. (1954) Two interesting new anthiid fishes from East Africa. Annals and Magazine of Natural History, Series 12, 7, 1 - 6. https: // doi. org / 10.1080 / 00222935408651683" type="journal article" year="1954">Smith (1954)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="89904D324C65E567E7AF4218FE43FEBD" authority="Smith &amp; Pope (1906)" authorityName="Smith &amp; Pope" authorityYear="1906" box="[151,499,260,285]" class="Actinopterygii" family="Serranidae" genus="Tosana" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E7AF4218FF57FEBC" box="[151,231,261,284]" italics="true" pageId="8" pageNumber="75">Tosana</emphasis>
<bibRefCitation id="2A014B404C65E567E7CA4219FE43FEBD" author="Smith" box="[242,499,260,285]" pageId="8" pageNumber="75" refString="Smith, H. M. &amp; Pope, T. E. B. (1906) List of fishes collected in Japan in 1903, with descriptions of new genera and species. Proceedings of the United States National Museum, 31 (No. 1489), 459 - 499. https: // doi. org / 10.5479 / si. 00963801.31 - 1489.459" type="journal article" year="1906">Smith &amp; Pope (1906)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="89904D324C65E567E53B4219FCE9FEBD" authority="Kamohara (1953)" authorityName="Kamohara" authorityYear="1953" box="[515,857,260,285]" class="Actinopterygii" family="Serranidae" genus="Tosanoides" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E53B4219FDCDFEBD" box="[515,637,260,285]" italics="true" pageId="8" pageNumber="75">Tosanoides</emphasis>
<bibRefCitation id="2A014B404C65E567E5B44219FCE9FEBD" author="Kamohara" box="[652,857,260,285]" pageId="8" pageNumber="75" refString="Kamohara, T. (1953) Marine fishes newly found in Prov. Tosa, Japan, with descriptions of a new genus and species. Research Reports of Kochi University, 2 (11), 1 - 10, 2 pls." type="journal article" year="1953">Kamohara (1953)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="89904D324C65E567E4524219FB11FEBD" authority="Herre (1936)" authorityName="Herre" authorityYear="1936" box="[874,1185,260,285]" class="Actinopterygii" family="Serranidae" genus="Luzonichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E4524219FC4EFEBD" box="[874,1022,260,285]" italics="true" pageId="8" pageNumber="75">Luzonichthys</emphasis>
<bibRefCitation id="2A014B404C65E567E3314218FB11FEBD" author="Herre" box="[1033,1185,260,285]" pageId="8" pageNumber="75" refString="Herre, A. W. C. T. (1936) Notes on fishes in the Zoological Museum of Stanford University, V. New or rare Philippine fishes from the Herre 1933 Philippine expedition. Philippine Journal of Science, 59, 357 - 373, pls. 1 - 2." type="journal article" year="1936">Herre (1936)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="89904D324C65E567E38A4219FF51FEE0" authority="Allen (1984)" authorityName="Allen" authorityYear="1984" class="Actinopterygii" family="Serranidae" genus="Rabaulichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E38A4219FAE2FEBD" box="[1202,1362,260,285]" italics="true" pageId="8" pageNumber="75">Rabaulichthys</emphasis>
<bibRefCitation id="2A014B404C65E567E2664219FF51FEE0" author="Allen" pageId="8" pageNumber="75" refString="Allen, G. R. (1984) A new genus and species of anthiid fish from Papua New Guinea. Revue francaise d'Aquariologie Herpetologie, 11, 47 - 50." type="journal article" year="1984">Allen (1984)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="89904D324C65E567E623423AFCFBFEE0" authority="Anderson, Parin &amp; Randall (1990)" authorityName="Anderson, Parin &amp; Randall" authorityYear="1990" box="[283,843,295,320]" class="Actinopterygii" family="Serranidae" genus="Anatolanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E623423AFE08FEE0" box="[283,440,295,320]" italics="true" pageId="8" pageNumber="75">Anatolanthias</emphasis>
<bibRefCitation id="2A014B404C65E567E6F8423AFCFBFEE0" box="[448,843,295,320]" pageId="8" pageNumber="75" refString="Anderson, W. D., Jr., Parin, N. V. &amp; Randall, J. E. (1990) A new genus and species of anthiine fish (Pisces: Serranidae) from the eastern South Pacific with comments on anthiine relationships. Proceedings of the Biological Society of Washington, 103, 922 - 930." type="journal article">Anderson, Parin &amp; Randall (1990)</bibRefCitation>
</taxonomicName>
. Classification within the genus is also problematic. Generally species have been assigned to either the nominate subgenus (
<taxonomicName id="89904D324C65E567E3304251FB2FFEC5" authorityName="Bleeker" authorityYear="1871" box="[1032,1183,332,357]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E3304251FB2FFEC5" box="[1032,1183,332,357]" italics="true" pageId="8" pageNumber="75">Pseudanthias</emphasis>
</taxonomicName>
) or to the subgenus
<taxonomicName id="89904D324C65E567E7AF4272FE5AFE28" authority="Herre (1927)" authorityName="Herre" authorityYear="1927" box="[151,490,367,392]" family="Serranidae" genus="Mirolabrichthys" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E7AF4272FEFCFE28" box="[151,332,367,392]" italics="true" pageId="8" pageNumber="75">Mirolabrichthys</emphasis>
Herre (1927)
</taxonomicName>
. Randall &amp; Lubbock (1981) recognised the latter on the basis of a single character, upper lip of males hypertrophied. However,
<bibRefCitation id="2A014B404C65E567E5A24289FC12FE0C" author="Randall" box="[666,930,404,429]" pageId="8" pageNumber="75" refString="Randall, J. E. &amp; Pyle, R. L. (2001) Four new serranid fishes of the anthiine genus Pseudanthias from the South Pacific. Raffles Bulletin of Zoology, 49, 19 - 34." type="journal article" year="2001">Randall &amp; Pyle (2001)</bibRefCitation>
noted that this character is expressed to an intermediate degree in
<taxonomicName id="89904D324C65E567E6A342A4FD9EFE70" authorityName="Randall &amp; Pyle" authorityYear="2001" box="[411,558,439,464]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="flavicauda">
<emphasis id="7CE4EAA34C65E567E6A342A4FD9EFE70" box="[411,558,439,464]" italics="true" pageId="8" pageNumber="75">P. flavicauda</emphasis>
</taxonomicName>
, and
<bibRefCitation id="2A014B404C65E567E55542A5FC07FE70" author="Heemstra" box="[621,951,439,464]" pageId="8" pageNumber="75" refString="Heemstra, P. C. &amp; Akhilesh, K. V. (2012) A review of the anthiine fish genus Pseudanthias (Perciformes: Serranidae) of the western Indian Ocean, with description of a new species and a key to the species. aqua, International Journal of Ichthyology, 18, 121 - 164." type="journal article" year="2012">Heemstra &amp; Akhilesh (2012)</bibRefCitation>
reported similar development of the upper lip in
<taxonomicName id="89904D324C65E567E7EF42C0FE37FE55" box="[215,391,476,501]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="7CE4EAA34C65E567E7EF42C0FE37FE55" box="[215,391,476,501]" italics="true" pageId="8" pageNumber="75">P. pulcherrimus</emphasis>
</taxonomicName>
. Various degrees of hypertrophy also occur in at least
<taxonomicName id="89904D324C65E567E4D942C0FBE6FE55" baseAuthorityName="Lubbock &amp; Allen" baseAuthorityYear="1978" box="[993,1110,476,501]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="randalli">
<emphasis id="7CE4EAA34C65E567E4D942C0FBE6FE55" box="[993,1110,476,501]" italics="true" pageId="8" pageNumber="75">P. randalli</emphasis>
</taxonomicName>
and
<taxonomicName id="89904D324C65E567E3B442C0FB44FE55" authorityName="Gill &amp; Tea &amp; Senou" authorityYear="2017" box="[1164,1268,476,501]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="tequila">
<emphasis id="7CE4EAA34C65E567E3B442C0FB44FE55" box="[1164,1268,476,501]" italics="true" pageId="8" pageNumber="75">P. tequila</emphasis>
</taxonomicName>
. This might be taken as evidence for placing all four species (and possibly
<taxonomicName id="89904D324C65E567E45A411CFC60FDB8" authorityName="Williams, Delrieu-Trottin &amp; Planes" authorityYear="2013" box="[866,976,513,536]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="oumati">
<emphasis id="7CE4EAA34C65E567E45A411CFC60FDB8" box="[866,976,513,536]" italics="true" pageId="8" pageNumber="75">P. oumati</emphasis>
</taxonomicName>
, for which males are unknown) in the subgenus
<taxonomicName id="89904D324C65E567E6324139FE0FFD9D" authorityName="Herre" authorityYear="1927" box="[266,447,548,573]" family="Serranidae" genus="Mirolabrichthys" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E6324139FE0FFD9D" box="[266,447,548,573]" italics="true" pageId="8" pageNumber="75">Mirolabrichthys</emphasis>
</taxonomicName>
. However, upper lip hypertrophy also occurs in
<taxonomicName id="89904D324C65E567E4C34139FB31FD9D" authorityName="Smith" authorityYear="1954" box="[1019,1153,548,573]" class="Actinopterygii" family="Serranidae" genus="Nemanthias" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E4C34139FB31FD9D" box="[1019,1153,548,573]" italics="true" pageId="8" pageNumber="75">Nemanthias</emphasis>
</taxonomicName>
. Moreover, taken in the context of other characters (including specialisations associated with orientation of the first dorsal pterygiophore, pectoral ray ornamentation, number of epineural bones, and predorsal formulae),
<taxonomicName id="89904D324C65E567E3144171FB51FD25" authorityName="Herre" authorityYear="1927" box="[1068,1249,620,645]" family="Serranidae" genus="Mirolabrichthys" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="7CE4EAA34C65E567E3144171FB51FD25" box="[1068,1249,620,645]" italics="true" pageId="8" pageNumber="75">Mirolabrichthys</emphasis>
</taxonomicName>
as diagnosed by Randall &amp; Lubbock (1981) does not appear to be monophyletic, and hypertrophy appears to have arisen multiple times within anthiadines. Preliminary molecular studies also support this conclusion (e.g.
<bibRefCitation id="2A014B404C65E567E3BC41A9FADBFD6C" author="Williams" box="[1156,1387,692,717]" pageId="8" pageNumber="75" refString="Williams, J. T., Delrieu-Trottin, E. &amp; Planes, S. (2013) Two new fish species of the subfamily Anthiinae (Perciformes, Serranidae) from the Marquesas. Zootaxa, 3647 (1), 167 - 180." type="journal article" year="2013">
Williams
<emphasis id="7CE4EAA34C65E567E3C841A8FA99FD6D" box="[1264,1321,692,717]" italics="true" pageId="8" pageNumber="75">et al.</emphasis>
2013
</bibRefCitation>
: fig. 13).
</paragraph>
</subSubSection>
</treatment>
</document>