treatments-xml/data/03/D5/87/03D587CD4725FFCD228CFAC9609BAE77.xml

<document id="B7458CF7A485312C3B6313C1F142E949" ID-DOI="10.5281/zenodo.5342032" ID-ISSN="2345-7600" ID-Zenodo-Dep="5342032" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630192200825" checkinUser="diego" docAuthor="Zettel, Herbert &amp; Tran, A. D." docDate="2009" docId="03D587CD4725FFCD228CFAC9609BAE77" docLanguage="en" docName="57rbz279-295.pdf" docOrigin="Raffles Bulletin of Zoology 57 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Hydrotrephes langkawicus Zettel &amp; Tran 2009, new species" docType="treatment" docVersion="3" lastPageNumber="284" masterDocId="FFECFFB54726FFC82107FFF2637DAC02" masterDocTitle="First Inventory Of The Water Bugs (Heteroptera: Nepomorpha: Gerromorpha) Of Langkawi Island, Kedah, Malaysia" masterLastPageNumber="295" masterPageNumber="279" pageNumber="282" updateTime="1699181942476" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9A7A2DB9B7E3D23EFED07767A3ED843D">First Inventory Of The Water Bugs (Heteroptera: Nepomorpha: Gerromorpha) Of Langkawi Island, Kedah, Malaysia</mods:title>
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<mods:namePart id="6AAE1EB83107CE7594C7FA98083B2C26">Zettel, Herbert</mods:namePart>
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<mods:namePart id="DA7D571A54DAECFA7B211ADDD6B8399E">Tran, A. D.</mods:namePart>
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<paragraph id="8BC336DB4725FFCB228CFAC96635A950" blockId="3.[907,1352,1338,1394]" box="[907,1352,1338,1363]" pageId="3" pageNumber="282">
<heading id="D08B81B74725FFCB228CFAC96635A950" bold="true" box="[907,1352,1338,1363]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="282" reason="2">
<emphasis id="B908EAC94725FFCB228CFAC96635A950" bold="true" box="[907,1352,1338,1363]" pageId="3" pageNumber="282">
<taxonomicName id="4C7C4D584725FFCB228CFAC967CEA951" ID-CoL="6ML28" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" box="[907,1203,1339,1363]" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="langkawicus" status="sp. nov.">
<emphasis id="B908EAC94725FFCB228CFAC967CEA951" bold="true" box="[907,1203,1339,1363]" italics="true" pageId="3" pageNumber="282">Hydrotrephes langkawicus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23B57B24725FFCB25C4FAC86635A950" box="[1219,1352,1338,1362]" pageId="3" pageNumber="282" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C36665504725FFCB251CFAA967CAA970" box="[1051,1207,1371,1394]" pageId="3" pageNumber="282" type="description">
<paragraph id="8BC336DB4725FFCB251CFAA967CAA970" blockId="3.[907,1352,1338,1394]" box="[1051,1207,1371,1394]" pageId="3" pageNumber="282">
(
<figureCitation id="13472A5E4725FFCB2524FAA967CDA970" box="[1059,1200,1371,1394]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="4.[134,178,2004,2025]" captionStartId-1="5.[111,155,2032,2053]" captionTargetBox-0="[135,1471,616,1978]" captionTargetBox-1="[116,1447,1264,2003]" captionTargetId-0="figure-214@4.[135,1471,616,1978]" captionTargetId-1="figure-423@5.[97,1466,1237,2017]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs. 2–6. Habitus of (2) Hydrotrephes langkawicus, new species (macropterous male, paratype). (3) Strongylovelia narusei, new species (apterous female, paratype). (4) Microvelia (Picaultia) minutissima, new species (apterous female, paratype). (5) Microvelia (Picaultia) minutissima, new species (macropterous male, holotype, right forewing broken off). (6) Frontal view of cephalonotum of Hydrotrephes langkawicus, new species (macropterous male, paratype)." captionText-1="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi-0="http://doi.org/10.5281/zenodo.5342036" figureDoi-1="http://doi.org/10.5281/zenodo.5342038" httpUri-0="https://zenodo.org/record/5342036/files/figure.png" httpUri-1="https://zenodo.org/record/5342038/files/figure.png" pageId="3" pageNumber="282">Figs. 2, 6-12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36665504725FFCB222DFA6B6706AA3C" pageId="3" pageNumber="282" type="materials_examined">
<paragraph id="8BC336DB4725FFCB222DFA6B6706AA3C" blockId="3.[810,1449,1433,1598]" pageId="3" pageNumber="282">
<emphasis id="B908EAC94725FFCB222DFA6B6761A9AC" bold="true" box="[810,1052,1433,1454]" italics="true" pageId="3" pageNumber="282">Material examined. –</emphasis>
<materialsCitation id="3B143C864725FFCB252FFA6B6706AA3C" collectingDate="2006-11-22" collectionCode="NHMW, ZRC" collectorName="Gn. Raya &amp; H. Zettel" country="Malaysia" location="Kedah" pageId="3" pageNumber="282" specimenCode="HZL9" specimenCount="1" specimenCount-male="1" stateProvince="Kedah" typeStatus="holotype">
<typeStatus id="54C788794725FFCB252FFA6B67F6A9AC" box="[1064,1163,1433,1454]" pageId="3" pageNumber="282" type="holotype">Holotype</typeStatus>
(macropterous male) and 
<typeStatus id="54C788794725FFCB222DFA4460F1A9C9" box="[810,908,1462,1483]" pageId="3" pageNumber="282" type="paratype">paratypes</typeStatus>
(1 hindwing-micropterous female, 3 macropterous males, 2 macropterous females), 
<specimenCode id="DBDA9EA04725FFCB257BFA2167C4A9EA" box="[1148,1209,1491,1512]" collectionCode="HZL" pageId="3" pageNumber="282">HZL9</specimenCode>
, labeled “ 
<collectingCountry id="F36B764B4725FFCB2427FA2166D8A9EA" box="[1312,1445,1491,1512]" name="Malaysia" pageId="3" pageNumber="282">MALAYSIA</collectingCountry>
, 
<collectingRegion id="49B8F8394725FFCB222DFA026017AA07" box="[810,874,1520,1541]" country="Malaysia" name="Kedah" pageId="3" pageNumber="282">Kedah</collectingRegion>
\ C-Langkawi, S slopes\ of 
<collectorName id="2689530D4725FFCB258AFA02678CAA07" box="[1165,1265,1520,1541]" pageId="3" pageNumber="282">Gn. Raya</collectorName>
, lowland\ stream, 
<date id="FFC2101B4725FFCB222DF9FF60EAAA20" box="[810,919,1549,1570]" pageId="3" pageNumber="282" value="2006-11-22">
<collectingDate id="EF86E9F34725FFCB222DF9FF60EAAA20" box="[810,919,1549,1570]" pageId="3" pageNumber="282" value="2006-11-22">22.11.2006</collectingDate>
</date>
\ leg. 
<collectorName id="2689530D4725FFCB22D3F9FF6753AA20" box="[980,1070,1549,1570]" pageId="3" pageNumber="282">H. Zettel</collectorName>
(
<specimenCode id="DBDA9EA04725FFCB2539F9FF6703AA20" box="[1086,1150,1549,1570]" collectionCode="HZL" pageId="3" pageNumber="282">HZL9</specimenCode>
)”, 
<typeStatus id="54C788794725FFCB259BF9FF6787AA20" box="[1180,1274,1549,1570]" pageId="3" pageNumber="282" type="holotype">holotypes</typeStatus>
and 
<typeStatus id="54C788794725FFCB2437F9FF66F0AA20" box="[1328,1421,1549,1570]" pageId="3" pageNumber="282" type="paratype">paratypes</typeStatus>
in 
<collectionCode id="ED6DAE1E4725FFCB222DF9DB6006AA3C" box="[810,891,1577,1598]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="282" type="Museum">NHMW</collectionCode>
, 
<specimenCount id="9D7AFD524725FFCB2281F9DB6759AA3C" box="[902,1060,1577,1598]" pageId="3" pageNumber="282" type="male" typeStatus="paratype">1 paratype male</specimenCount>
in 
<collectionCode id="ED6DAE1E4725FFCB2541F9DB6705AA3C" box="[1094,1144,1577,1598]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="282">ZRC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36665504725FFCD222DF995614DAF97" lastPageId="5" lastPageNumber="284" pageId="3" pageNumber="282" type="description">
<paragraph id="8BC336DB4725FFCB222DF9956611AA9D" blockId="3.[810,1449,1639,1695]" pageId="3" pageNumber="282">
<emphasis id="B908EAC94725FFCB222DF99567C4AA7D" bold="true" box="[810,1209,1639,1663]" italics="true" pageId="3" pageNumber="282">Description of macropterous male. –</emphasis>
Habitus: see 
<figureCitation id="13472A5E4725FFCB244EF99A66D8AA7D" box="[1353,1445,1640,1663]" captionStart="Figs" captionStartId="4.[134,178,2004,2025]" captionTargetBox="[135,1471,616,1978]" captionTargetId="figure-214@4.[135,1471,616,1978]" captionTargetPageId="4" captionText="Figs. 2–6. Habitus of (2) Hydrotrephes langkawicus, new species (macropterous male, paratype). (3) Strongylovelia narusei, new species (apterous female, paratype). (4) Microvelia (Picaultia) minutissima, new species (apterous female, paratype). (5) Microvelia (Picaultia) minutissima, new species (macropterous male, holotype, right forewing broken off). (6) Frontal view of cephalonotum of Hydrotrephes langkawicus, new species (macropterous male, paratype)." figureDoi="http://doi.org/10.5281/zenodo.5342036" httpUri="https://zenodo.org/record/5342036/files/figure.png" pageId="3" pageNumber="282">Figure 2</figureCitation>
. Body size: length 1.9–2.0 mm, width 
<quantity id="4C849B3E4725FFCB25CFF97A6614AA9D" box="[1224,1385,1672,1695]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="3" pageNumber="282" unit="mm" value="1.375" valueMax="1.4" valueMin="1.35">1.35–1.40 mm</quantity>
.
</paragraph>
<paragraph id="8BC336DB4725FFCB222DF93A66D4AB1D" blockId="3.[810,1449,1736,1823]" pageId="3" pageNumber="282">
Colour: Yellow with extended dark brown markings on dorsum, especially on head (
<figureCitation id="13472A5E4725FFCB257EF91A6790AAFD" box="[1145,1261,1768,1791]" captionStart="Figs" captionStartId="4.[134,178,2004,2025]" captionTargetBox="[135,1471,616,1978]" captionTargetId="figure-214@4.[135,1471,616,1978]" captionTargetPageId="4" captionText="Figs. 2–6. Habitus of (2) Hydrotrephes langkawicus, new species (macropterous male, paratype). (3) Strongylovelia narusei, new species (apterous female, paratype). (4) Microvelia (Picaultia) minutissima, new species (apterous female, paratype). (5) Microvelia (Picaultia) minutissima, new species (macropterous male, holotype, right forewing broken off). (6) Frontal view of cephalonotum of Hydrotrephes langkawicus, new species (macropterous male, paratype)." figureDoi="http://doi.org/10.5281/zenodo.5342036" httpUri="https://zenodo.org/record/5342036/files/figure.png" pageId="3" pageNumber="282">Figs. 2, 6</figureCitation>
). Venter mostly brownish. Legs and antennae yellow; rostrum light brown.
</paragraph>
<paragraph id="8BC336DB4725FFCC222DF8BA6149ADD7" blockId="3.[810,1449,1864,2079]" lastBlockId="4.[133,773,190,469]" lastPageId="4" lastPageNumber="283" pageId="3" pageNumber="282">
Cephalonotum with lateral margins behind eyes slightly concave in dorsal view; sharp margin of pronotal part ending in short distance from posterolateral corner and there forming a small, rather blunt angulation; head with densely set small punctures, with very narrow interspaces, almost dull; disk of pronotum set with numerous very fine micropunctures, and with scattered, unequally distributed and somewhat larger punctures becoming denser towards sides. Mesoscutellum 0.9 times as long as wide, with puncturation of similar size, but slightly denser than on disk of pronotum, micropunctures medially obliterated. Hemelytron with slightly stronger puncturation, interspaces shagreened and dull. Genal plate stout; pronotal plate (
<figureCitation id="13472A5E4722FFCC232FFEAC6117AD77" box="[552,618,350,373]" captionStart="Figs" captionStartId="5.[111,155,2032,2053]" captionTargetBox="[116,1447,1264,2003]" captionTargetId="figure-423@5.[97,1466,1237,2017]" captionTargetPageId="5" captionText="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi="http://doi.org/10.5281/zenodo.5342038" httpUri="https://zenodo.org/record/5342038/files/figure.png" pageId="4" pageNumber="283">Fig. 7</figureCitation>
) with shallow incision, anteriorly relatively wide. Inner corner of propleural plate broadly convex. Eye index: 2.1–2.2. Fourth rostral segment 2.3 times as long as segment 3.
</paragraph>
<paragraph id="8BC336DB4722FFCC2246FF4C60A5AD97" blockId="4.[833,1472,190,405]" pageId="4" pageNumber="283">
Ventral carinae (
<figureCitation id="13472A5E4722FFCC22F9FF4C6738ACD7" box="[1022,1093,190,213]" captionStart="Figs" captionStartId="5.[111,155,2032,2053]" captionTargetBox="[116,1447,1264,2003]" captionTargetId="figure-423@5.[97,1466,1237,2017]" captionTargetPageId="5" captionText="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi="http://doi.org/10.5281/zenodo.5342038" httpUri="https://zenodo.org/record/5342038/files/figure.png" pageId="4" pageNumber="283">Fig. 8</figureCitation>
): Prosternal carina with posterior corner approximately rectangular, with posterior edge slightly concave; apices of meso- and metasternal carina thin-laminate, that on mesosternum strongly reduced; carina of sternite 3 with few small denticles, the proximal one variably enlarged. Abdominal segments strongly asymmetrical.
</paragraph>
<paragraph id="8BC336DB4722FFCD2246FE4C62B4AD57" blockId="4.[833,1471,446,501]" lastBlockId="5.[111,750,190,341]" lastPageId="5" lastPageNumber="284" pageId="4" pageNumber="283">
Aedeagus (
<figureCitation id="13472A5E4722FFCC22C2FE4C6773ADD7" box="[965,1038,446,469]" captionStart="Figs" captionStartId="5.[111,155,2032,2053]" captionTargetBox="[116,1447,1264,2003]" captionTargetId="figure-423@5.[97,1466,1237,2017]" captionTargetPageId="5" captionText="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi="http://doi.org/10.5281/zenodo.5342038" httpUri="https://zenodo.org/record/5342038/files/figure.png" pageId="4" pageNumber="283">Fig. 9</figureCitation>
) moderately slender, apically rather stout and slightly bent to right side; with narrow apical plate posteriorly extended into short spine. Right paramere (
<figureCitation id="13472A5E4723FFCD2170FF2C63B6ACF7" box="[119,203,222,245]" captionStart="Figs" captionStartId="5.[111,155,2032,2053]" captionTargetBox="[116,1447,1264,2003]" captionTargetId="figure-423@5.[97,1466,1237,2017]" captionTargetPageId="5" captionText="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi="http://doi.org/10.5281/zenodo.5342038" httpUri="https://zenodo.org/record/5342038/files/figure.png" pageId="5" pageNumber="284">Fig. 10</figureCitation>
) shorter than left paramere, slender-lanceolate, curved, apex minutely pointed. Left paramere (
<figureCitation id="13472A5E4723FFCD2396FF0C6199AD17" box="[657,740,254,277]" captionStart="Figs" captionStartId="5.[111,155,2032,2053]" captionTargetBox="[116,1447,1264,2003]" captionTargetId="figure-423@5.[97,1466,1237,2017]" captionTargetPageId="5" captionText="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi="http://doi.org/10.5281/zenodo.5342038" httpUri="https://zenodo.org/record/5342038/files/figure.png" pageId="5" pageNumber="284">Fig. 11</figureCitation>
) moderately slender, distally evenly narrowed and slightly curved, apex narrowly rounded.
</paragraph>
<caption id="DF0366534722FFCC2181F82661C4A442" ID-DOI="http://doi.org/10.5281/zenodo.5342036" ID-Zenodo-Dep="5342036" httpUri="https://zenodo.org/record/5342036/files/figure.png" pageId="4" pageNumber="283" startId="4.[134,178,2004,2025]" targetBox="[135,1471,616,1978]" targetPageId="4">
<paragraph id="8BC336DB4722FFCC2181F82661C4A442" blockId="4.[134,1471,2004,2112]" pageId="4" pageNumber="283">
Figs. 2–6. Habitus of (2) 
<taxonomicName id="4C7C4D584722FFCC2079F82661FDABEB" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" box="[382,640,2004,2025]" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="langkawicus" status="sp. nov.">
<emphasis id="B908EAC94722FFCC2079F82661FDABEB" box="[382,640,2004,2025]" italics="true" pageId="4" pageNumber="283">Hydrotrephes langkawicus</emphasis>
</taxonomicName>
, 
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(macropterous male, paratype). (3) 
<taxonomicName id="4C7C4D584722FFCC2564F8266640ABEB" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" box="[1123,1341,2004,2025]" class="Insecta" family="Veliidae" genus="Strongylovelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="narusei" status="sp. nov.">
<emphasis id="B908EAC94722FFCC2564F8266640ABEB" box="[1123,1341,2004,2025]" italics="true" pageId="4" pageNumber="283">Strongylovelia narusei</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23B57B24722FFCC244CF82666C2ABEB" box="[1355,1471,2004,2025]" pageId="4" pageNumber="283" rank="species">new species</taxonomicNameLabel>
(apterous female, paratype). (4) 
<taxonomicName id="4C7C4D584722FFCC20C2F8036067A404" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" box="[453,794,2033,2054]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="minutissima" status="sp. nov." subGenus="Picaultia">
<emphasis id="B908EAC94722FFCC20C2F8036150A404" box="[453,557,2033,2054]" italics="true" pageId="4" pageNumber="283">Microvelia</emphasis>
(
<emphasis id="B908EAC94722FFCC2339F80361EBA404" box="[574,662,2033,2054]" italics="true" pageId="4" pageNumber="283">Picaultia</emphasis>
) 
<emphasis id="B908EAC94722FFCC23A1F8036067A404" box="[678,794,2033,2054]" italics="true" pageId="4" pageNumber="283">minutissima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23B57B24722FFCC222EF80360E3A404" box="[809,926,2033,2054]" pageId="4" pageNumber="283" rank="species">new species</taxonomicNameLabel>
(apterous female, paratype). (5) 
<taxonomicName id="4C7C4D584722FFCC25E1F8036387A421" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="minutissima" status="sp. nov." subGenus="Picaultia">
<emphasis id="B908EAC94722FFCC25E1F8036633A404" box="[1254,1358,2033,2054]" italics="true" pageId="4" pageNumber="283">Microvelia</emphasis>
(
<emphasis id="B908EAC94722FFCC2458F80366CAA404" box="[1375,1463,2033,2054]" italics="true" pageId="4" pageNumber="283">Picaultia</emphasis>
) 
<emphasis id="B908EAC94722FFCC2181F7FC6387A421" box="[134,250,2062,2083]" italics="true" pageId="4" pageNumber="283">minutissima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23B57B24722FFCC2000F7FC6201A421" box="[263,380,2062,2083]" pageId="4" pageNumber="283" rank="species">new species</taxonomicNameLabel>
(macropterous male, holotype, right forewing broken off). (6) Frontal view of cephalonotum of 
<taxonomicName id="4C7C4D584722FFCC243AF7FC6383A442" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="langkawicus" status="sp. nov.">
<emphasis id="B908EAC94722FFCC243AF7FC6383A442" italics="true" pageId="4" pageNumber="283">Hydrotrephes langkawicus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23B57B24722FFCC200CF7D962FDA442" box="[267,384,2091,2112]" pageId="4" pageNumber="283" rank="species">new species</taxonomicNameLabel>
(macropterous male, paratype).
</paragraph>
</caption>
<paragraph id="8BC336DB4723FFCD2168FE8C63B8AE77" blockId="5.[111,750,382,629]" pageId="5" pageNumber="284">
<emphasis id="B908EAC94723FFCD2168FE8C615EAD94" bold="true" box="[111,547,382,406]" italics="true" pageId="5" pageNumber="284">Description of macropterous female. –</emphasis>
Body size: length 
<quantity id="4C849B3E4723FFCD2168FE6C6385ADB7" box="[111,248,414,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="5" pageNumber="284" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1–2.2 mm</quantity>
, width 
<quantity id="4C849B3E4723FFCD2049FE6C628FADB7" box="[334,498,414,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.49" metricValueMin="1.46" pageId="5" pageNumber="284" unit="mm" value="1.475" valueMax="1.49" valueMin="1.46">1.46–1.49 mm</quantity>
. Eye index: 2.1. Most characteristics similar to that of male, except the following. Abdomen symmetrical. Sternite 6 with concave hind margin. Subgenital plate (
<figureCitation id="13472A5E4723FFCD2089FE0C62A3AE17" box="[398,478,510,533]" captionStart="Figs" captionStartId="5.[111,155,2032,2053]" captionTargetBox="[116,1447,1264,2003]" captionTargetId="figure-423@5.[97,1466,1237,2017]" captionTargetPageId="5" captionText="Figs. 7–12. Hydrotrephes langkawicus, new species: 7, genal and pronotal plate; 8, ventromedian carina of thorax and abdomen, with variations of prosternal carina, mesosternal carina, and carinae of sternites 2 and 3; 9, aedeagus, right view; 10, right paramere, right view; 11, left paramere, right view; 12, sternite 7 of female, ventral view, pilosity omitted." figureDoi="http://doi.org/10.5281/zenodo.5342038" httpUri="https://zenodo.org/record/5342038/files/figure.png" pageId="5" pageNumber="284">Fig. 12</figureCitation>
) with broad mediodistal lobe with broadly convex hind margin; inner ridge strongly concave, almost semicircular. Ventral laterotergites 8 slender.
</paragraph>
<paragraph id="8BC336DB4723FFCD2168FD6C63BDAF57" blockId="5.[111,749,670,853]" pageId="5" pageNumber="284">
Hindwing-micropterous female: body size: length 
<quantity id="4C849B3E4723FFCD23CDFD6C63E0AED7" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="284" unit="mm" value="2.1">2.1 mm</quantity>
, width 
<quantity id="4C849B3E4723FFCD21FBFD4C6217AED7" box="[252,362,702,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="5" pageNumber="284" unit="mm" value="1.42">1.42 mm</quantity>
; eye index: 2.6; pronotum with posterolateral corners less elevated and with angulation of lateral margin more blunt; hemelytron without embolar and claval sutures; other characteristics as in brachypterous female.
</paragraph>
<paragraph id="8BC336DB4723FFCD2168FC8C614DAF97" blockId="5.[111,560,894,917]" box="[111,560,894,917]" pageId="5" pageNumber="284">Hindwing-micropterous male: unknown.</paragraph>
</subSubSection>
<subSubSection id="C36665504723FFCD2168FC4C6159AFF7" pageId="5" pageNumber="284" type="etymology">
<paragraph id="8BC336DB4723FFCD2168FC4C6159AFF7" blockId="5.[111,749,958,1013]" pageId="5" pageNumber="284">
<emphasis id="B908EAC94723FFCD2168FC4C627CAFD4" bold="true" box="[111,257,958,982]" italics="true" pageId="5" pageNumber="284">Etymology. –</emphasis>
Named after the island of 
<typeStatus id="54C788794723FFCD232CFC4C6127AFD7" box="[555,602,958,981]" pageId="5" pageNumber="284">type</typeStatus>
locality. The species epithet is a Latinized adjective.
</paragraph>
</subSubSection>
<subSubSection id="C36665504723FFCD2168FBEC6097AE17" pageId="5" pageNumber="284" type="discussion">
<paragraph id="8BC336DB4723FFCD2168FBEC6097AE17" blockId="5.[111,749,1054,1141]" lastBlockId="5.[808,1449,190,533]" pageId="5" pageNumber="284">
<emphasis id="B908EAC94723FFCD2168FBEC638BA834" bold="true" box="[111,246,1054,1078]" italics="true" pageId="5" pageNumber="284">Remarks. –</emphasis>
This species belongs to the 
<taxonomicName id="4C7C4D584723FFCD2354FBEC63C2A857" baseAuthorityName="Kirkaldy" baseAuthorityYear="1904" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="B908EAC94723FFCD2354FBEC63C2A857" italics="true" pageId="5" pageNumber="284">Hydrotrephes martini</emphasis>
</taxonomicName>
group as defined by 
<bibRefCitation id="EFED4B2A4723FFCD20A9FBCC6132A857" author="Zettel, H" box="[430,591,1086,1109]" pageId="5" pageNumber="284" pagination="1 - 22" refId="ref13667" refString="Zettel, H., 2000 a. The Helotrephidae (Heteroptera) of Borneo. Entomological Problems, 31 (1): 1 - 22." type="journal article" year="2000">Zettel (2000a)</bibRefCitation>
. Hitherto, this species group contained eight described species, distributed on the islands of Borneo (five species), 
<collectingRegion id="49B8F8394723FFCD25E0FF4C6639ACD7" box="[1255,1348,190,213]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="284">Sumatra</collectingRegion>
, Siberut, and Nias (one species each) (
<bibRefCitation id="EFED4B2A4723FFCD2585FF2C6663ACF7" author="Zettel, H" box="[1154,1310,222,245]" pageId="5" pageNumber="284" pagination="1359 - 1382" refId="ref13776" refString="Zettel, H., 2004 a. Weitere neue Helotrephini (Heteroptera: Helotrephidae) aus China, Indonesien und Malaysien sowie von den Philippinen. Linzer Biologische Beitrage, 36 (2): 1359 - 1382." type="journal article" year="2004">Zettel, 2004a</bibRefCitation>
; Papá ek &amp; 
<bibRefCitation id="EFED4B2A4723FFCD222DFF0C60CBAD17" author="Zettel, H" box="[810,950,254,277]" pageId="5" pageNumber="284" pagination="409 - 420" refId="ref13892" refString="Zettel, H., 2005. A new sea-skater from the southern Philippines, with a key to the Philippine species of Halobates (Heteroptera: Gerridae). Entomofauna, 26 (24): 409 - 420." type="journal article" year="2005">Zettel, 2005</bibRefCitation>
). 
<emphasis id="B908EAC94723FFCD22CAFF0D6786AD14" box="[973,1275,255,278]" italics="true" pageId="5" pageNumber="284">
<taxonomicName id="4C7C4D584723FFCD22CAFF0D678BAD14" authority="Zettel &amp; Tran, 2009" authorityName="Zettel &amp; Tran" authorityYear="2009" box="[973,1270,255,278]" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="langkawicus" status="sp. nov.">Hydrotrephes langkawicus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23B57B24723FFCD2402FF0C66F1AD17" box="[1285,1420,254,277]" pageId="5" pageNumber="284" rank="species">new species</taxonomicNameLabel>
is one of the smallest species of the genus; in the 
<taxonomicName id="4C7C4D584723FFCD2431FEED66D4AD34" baseAuthorityName="Kirkaldy" baseAuthorityYear="1904" box="[1334,1449,287,310]" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="B908EAC94723FFCD2431FEED66D4AD34" box="[1334,1449,287,310]" italics="true" pageId="5" pageNumber="284">H. martini</emphasis>
</taxonomicName>
group only 
<taxonomicName id="4C7C4D584723FFCD22A1FECD67E0AD57" authority="Zettel, 2004" authorityName="Zettel" authorityYear="2004" box="[934,1181,318,342]" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B908EAC94723FFCD22A1FECD676CAD54" box="[934,1041,319,342]" italics="true" pageId="5" pageNumber="284">H. nitidus</emphasis>
Zettel, 2004
</taxonomicName>
, a strongly shiny species with deeply emarginate prosternal carina, is of similar size. The genital structures of the male of 
<taxonomicName id="4C7C4D584723FFCD25F9FE8D66D4AD94" authorityName="Zettel &amp; Tran" authorityYear="2009" box="[1278,1449,383,406]" class="Insecta" family="Helotrephidae" genus="Hydrotrephes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="langkawicus">
<emphasis id="B908EAC94723FFCD25F9FE8D66D4AD94" box="[1278,1449,383,406]" italics="true" pageId="5" pageNumber="284">H. langkawicus</emphasis>
</taxonomicName>
are characteristic. Further, the combination of the weakly emarginate prosternal carina, the reduced lamina on the mesosternal carina, and the lobate subgenital plate of the female is unique.
</paragraph>
</subSubSection>
<subSubSection id="C36665504723FFCD222DFDCC609BAE77" pageId="5" pageNumber="284" type="distribution">
<paragraph id="8BC336DB4723FFCD222DFDCC609BAE77" blockId="5.[810,1448,574,629]" pageId="5" pageNumber="284">
<emphasis id="B908EAC94723FFCD222DFDCC60A4AE54" bold="true" box="[810,985,574,598]" italics="true" pageId="5" pageNumber="284">Distribution. –</emphasis>
Only known from the 
<typeStatus id="54C788794723FFCD25F7FDCC665DAE57" box="[1264,1312,574,597]" pageId="5" pageNumber="284">type</typeStatus>
locality on Langkawi Island.
</paragraph>
</subSubSection>
</treatment>
</document>