treatments-xml/data/03/D5/87/03D587920C79B9621B420A19CE9720E4.xml

<document id="77009C134906F4414CF02E6DAB711AFE" ID-CLB-Dataset="231400" ID-DOI="10.1655/HERPETOLOGICA-D-15-00059" ID-GBIF-Dataset="c3c8c82e-cdab-46c5-9f80-f1ccd8fa0626" ID-ISSN="1938-5099" ID-Zenodo-Dep="7712201" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678362431454" checkinUser="felipe" docAuthor="Costa, Henrique C., Santana, Diego J., Leal, Fernando, Koroiva, Ricardo &amp; Garcia, Paulo C. A." docDate="2016" docId="03D587920C79B9621B420A19CE9720E4" docLanguage="en" docName="Herpetologica.72.2.157-166.pdf" docOrigin="Herpetologica 72 (2)" docSource="http://dx.doi.org/10.1655/herpetologica-d-15-00059" docStyle="DocumentStyle:7157CB4A8C4DC19627D49B44D928F812.2:Herpetologica.2015-.journal_article.1cover" docStyleId="7157CB4A8C4DC19627D49B44D928F812" docStyleName="Herpetologica.2015-.journal_article.1cover" docStyleVersion="2" docTitle="Helicops nentur Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="166" masterDocId="FFECFFEA0C7BB9651A7F0C21CD272634" masterDocTitle="A New Species of Helicops (Serpentes: Dipsadidae: Hydropsini) from Southeastern Brazil" masterLastPageNumber="166" masterPageNumber="157" pageNumber="157" updateTime="1702001217434" updateUser="plazi">
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<mods:title id="3BE03CCF33145AA72B6FE79A7B6A6665">A New Species of Helicops (Serpentes: Dipsadidae: Hydropsini) from Southeastern Brazil</mods:title>
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<mods:namePart id="B5ABCE12BE18D1FFC2F66830E656D25C">Costa, Henrique C.</mods:namePart>
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<mods:namePart id="E6DFFC6BB36A6B257F2CEF5A2FF5AD8B">Santana, Diego J.</mods:namePart>
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<mods:namePart id="E8B03C4A8C4D30BEB0834DE639485C55">Leal, Fernando</mods:namePart>
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<mods:namePart id="B7E24EC11C488CAEA5FA40515D972F3C">Koroiva, Ricardo</mods:namePart>
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<mods:namePart id="CA01E73A09E732151BFA7D946DD3AB03">Garcia, Paulo C. A.</mods:namePart>
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<treatment id="03D587920C79B9621B420A19CE9720E4" ID-DOI="http://doi.org/10.5281/zenodo.7716218" ID-GBIF-Taxon="209434938" ID-Zenodo-Dep="7716218" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4" httpUri="http://treatment.plazi.org/id/03D587920C79B9621B420A19CE9720E4" lastPageId="7" pageId="2">
<subSubSection id="C366650F0C79B9671B420A19CF722064" box="[317,597,1592,1616]" pageId="2" type="nomenclature">
<paragraph id="8BC336840C79B9671B420A19CF722064" blockId="2.[317,597,1592,1616]" box="[317,597,1592,1616]" pageId="2">
<heading id="D08B81E80C79B9671B420A19CF722064" box="[317,597,1592,1616]" centered="true" fontSize="10" level="2" pageId="2" reason="2">
<taxonomicName id="4C7C4D070C79B9671B420A19CCD2207B" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[317,501,1592,1616]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C79B9671B420A19CCD2207B" box="[317,501,1592,1616]" italics="true" pageId="2">
Helicops 
<emphasis id="B908EA960C79B9671BDA0A19CCD2207B" bold="true" box="[421,501,1592,1615]" italics="true" pageId="2">nentur</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C79B9671B810A18CF722064" box="[510,597,1593,1616]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C366650F0C79B9671B240A77CF10205A" box="[347,567,1622,1646]" pageId="2" type="description">
<paragraph id="8BC336840C79B9671B240A77CF10205A" blockId="2.[347,567,1622,1646]" box="[347,567,1622,1646]" pageId="2">
(
<figureCitation id="13472A010C79B9671B1C0A77CCEB205A" box="[355,460,1622,1646]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="2.[845,859,1933,1952]" captionStartId-1="3.[143,157,1112,1131]" captionStartId-2="3.[165,179,1909,1928]" captionTargetBox-0="[824,1496,1411,1915]" captionTargetBox-1="[143,1522,189,1095]" captionTargetBox-2="[143,814,1386,1891]" captionTargetId-0="figure-779@2.[824,1496,1411,1915]" captionTargetId-1="figure-335@3.[143,1522,189,1095]" captionTargetId-2="figure-374@3.[143,815,1386,1891]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIG. 1.—Holotype of Helicops nentur sp. nov. (UFMG 2486): dorsal (A) and ventral (B) view. Scale bar = 20 mm. A color version of this figure is available online." captionText-1="FIG. 2.—Holotype of Helicops nentur sp. nov. (UFMG 2486): head in dorsal (A), ventral (B), left (C), and right (D) view. Scale bar = 5 mm. A color version of this figure is available online." captionText-2="FIG. 3.—Right hemipenis of a paratype of Helicops nentur sp. nov. (MZUFV 960) fully everted and almost maximally expanded: sulcate (A) and asulcate (B) side. Scale bar = 10 mm. A color version of this figure is available online." figureDoi-0="http://doi.org/10.5281/zenodo.7712203" figureDoi-1="http://doi.org/10.5281/zenodo.7712205" figureDoi-2="http://doi.org/10.5281/zenodo.7712207" httpUri-0="https://zenodo.org/record/7712203/files/figure.png" httpUri-1="https://zenodo.org/record/7712205/files/figure.png" httpUri-2="https://zenodo.org/record/7712207/files/figure.png" pageId="2">Figs. 1–3</figureCitation>
; 
<tableCitation id="C6FE033F0C79B9671BA60A77CF08205A" box="[473,559,1622,1646]" captionStart="TABLE" captionStartId="4.[143,156,194,213]" captionTargetPageId="4" captionText="TABLE 2.—Meristic and morphometric characters in Helicops nentur sp. nov. PV = preventrals, VE = ventrals, C = cloacal shield, SC = subcaudals, SL = supralabials, SL + E = supralabials in contact with eye, IL = infralabials, IL + G1 = infralabials in contact with first pair of genials, IL + G2 = infralabials in contact with second pair of genials, NA = nasal (entire = E or semidivided = S), Pe = preocular, Po = postocular, AT = anterior temporals, PT = posterior temporals, MXT = maxillary teeth, SVL = snout–vent length (mm), TL = tail length (mm)." httpUri="http://table.plazi.org/id/DF03660C0C7FB9611AF00CE3CE8F2728" pageId="2" tableUuid="DF03660C0C7FB9611AF00CE3CE8F2728">Table 2</tableCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB6C11AA0C79B9671A050AA5CCDB20A8" box="[122,508,1668,1693]" pageId="2">
<taxonomicName id="4C7C4D070C79B9671A050AA5CC2620A8" box="[122,257,1668,1692]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B908EA960C79B9671A050AA5CDFE20A8" box="[122,217,1668,1692]" italics="true" pageId="2">Helicops</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="0ADD10950C79B9671B750AA4CCDF20A8" author="Costa, H. C. &amp; D. L. Pantoja &amp; J. L. Pontes &amp; R. N. Feio" box="[266,504,1668,1693]" page="359" pageId="2" year="2010">
<bibRefCitation id="EFED4B750C79B9671B750AA4CCDF20A8" author="Costa, H. C. &amp; D. L. Pantoja &amp; J. L. Pontes &amp; R. N. Feio" box="[266,504,1668,1693]" pageId="2" pagination="352 - 378" refId="ref9596" refString="Costa, H. C., D. L. Pantoja, J. L. Pontes, and R. N. Feio. 2010. Serpentes do municipio de Vicosa, Mata Atlantica do sudeste do Brasil. Biota Neotropica 10: 352 - 378. [In Portuguese.]" type="journal article" year="2010">Costa et al. 2010:359</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<treatmentCitationGroup id="AB6C11AA0C79B9671A050A82CCDB208F" box="[122,508,1699,1723]" pageId="2">
<taxonomicName id="4C7C4D070C79B9671A050A82CC26208E" box="[122,257,1699,1723]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B908EA960C79B9671A050A82CDFE208F" box="[122,217,1699,1723]" italics="true" pageId="2">Helicops</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="0ADD10950C79B9671B750A82CCDE208F" author="Costa, H. C. &amp; A. C. Rodrigues &amp; C. A. Coelho &amp; R. N. Feio" box="[266,505,1699,1723]" page="118" pageId="2" year="2014">
<bibRefCitation id="EFED4B750C79B9671B750A82CCDE208F" author="Costa, H. C. &amp; A. C. Rodrigues &amp; C. A. Coelho &amp; R. N. Feio" box="[266,505,1699,1723]" pageId="2" refId="ref9650" refString="Costa, H. C., A. C. Rodrigues, C. A. Coelho, and R. N. Feio. 2014. Repteis da Mata do Paraiso. Pp. 113 - 131 in Ecologia de Mata Atlantica, Estudos Ecologicos na Mata do Paraiso (G. S. Lima, G. A. Ribeiro, W. Goncalves, S. V. Martins, and M. P. Almeida, eds.). Supremo, Brazil. [In Portuguese.]" type="book" year="2014">Costa et al. 2014:118</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C366650F0C79B9671AEA0AEFC90F250F" pageId="2" type="materials_examined">
<paragraph id="8BC336840C79B9671AEA0AEFCFB7216F" blockId="2.[122,792,1742,2001]" pageId="2">
<materialsCitation id="3B143CD90C79B9671AEA0AEFCFB7216F" ID-GBIF-Occurrence="4052787303" collectingDate="2013-11-08" collectionCode="UFMG, WGS" country="Brazil" elevation="260" latitude="-18.329397" location="Sao Jose da Safira" longLatPrecision="1" longitude="-42.09202" pageId="2" specimenCode="UFMG 2486, WGS84" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="B908EA960C79B9671AEA0AEFCC2B20D2" bold="true" box="[149,268,1742,1766]" pageId="2">
<typeStatus id="54C788260C79B9671AEA0AEFCC2F20D2" box="[149,264,1742,1766]" pageId="2" type="holotype">Holotype</typeStatus>
.
</emphasis>
— 
<specimenCode id="DBDA9EFF0C79B9671B560AEFCC9D20D2" box="[297,442,1742,1766]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2">UFMG 2486</specimenCode>
, an adult female collected on 
<date id="FFC210440C79B96719750AEFCC082137" pageId="2" value="2013-11-08">
<collectingDate id="EF86E9AC0C79B96719750AEFCC082137" pageId="2" value="2013-11-08">8 November 2013</collectingDate>
</date>
by Fernando Leal at a swamp in Fazenda Papa Capim (
<geoCoordinate id="EE4850430C79B9671B690B28CC842115" box="[278,419,1801,1825]" degrees="18.329398" direction="south" orientation="latitude" pageId="2" precision="1" value="-18.329397">
18.329398 
<emphasis id="B908EA960C79B9671BF60B28CCB42115" bold="true" box="[393,403,1801,1825]" pageId="2">°</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="EE4850430C79B9671BCD0B28CF6D2114" box="[434,586,1801,1825]" degrees="42.092017" direction="west" orientation="longitude" pageId="2" precision="1" value="-42.09202">
42.092017 
<emphasis id="B908EA960C79B967185B0B28CF092115" bold="true" box="[548,558,1801,1825]" pageId="2">°</emphasis>
W
</geoCoordinate>
, 
<elevation id="0051D1B70C79B96718270B28CD8A210A" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" unit="m" value="260.0">
<quantity id="4C849B610C79B96718270B28CF872114" box="[600,672,1801,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" unit="m" value="260.0">260 m</quantity>
above sea level
</elevation>
[a.s.l.]; in all cases, datum = 
<specimenCode id="DBDA9EFF0C79B96718640B07CF51210A" box="[539,630,1830,1854]" collectionCode="WGS" pageId="2">WGS84</specimenCode>
), 
<location id="8EA3605F0C79B96718F10B07CD9B216F" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B96718F10B07CD9B216F" country="Brazil" latitude="-18.329397" longLatPrecision="1" longitude="-42.09202" name="Sao Jose da Safira" pageId="2" stateProvince="Minas Gerais">São José da Safira</location>
, state of 
<collectingRegion id="49B8F8660C79B9671B5B0B62CC9C216F" box="[292,443,1859,1883]" country="Brazil" name="Minas Gerais" pageId="2">Minas Gerais</collectingRegion>
, 
<collectingCountry id="F36B76140C79B9671BB70B62CF2E216F" box="[456,521,1859,1883]" name="Brazil" pageId="2">Brazil</collectingCountry>
(
<figureCitation id="13472A010C79B96718650B62CFA4216F" box="[538,643,1859,1883]" captionStart-0="FIG" captionStart-1="FIG" captionStartId-0="2.[845,859,1933,1952]" captionStartId-1="3.[143,157,1112,1131]" captionTargetBox-0="[824,1496,1411,1915]" captionTargetBox-1="[143,1522,189,1095]" captionTargetId-0="figure-779@2.[824,1496,1411,1915]" captionTargetId-1="figure-335@3.[143,1522,189,1095]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIG. 1.—Holotype of Helicops nentur sp. nov. (UFMG 2486): dorsal (A) and ventral (B) view. Scale bar = 20 mm. A color version of this figure is available online." captionText-1="FIG. 2.—Holotype of Helicops nentur sp. nov. (UFMG 2486): head in dorsal (A), ventral (B), left (C), and right (D) view. Scale bar = 5 mm. A color version of this figure is available online." figureDoi-0="http://doi.org/10.5281/zenodo.7712203" figureDoi-1="http://doi.org/10.5281/zenodo.7712205" httpUri-0="https://zenodo.org/record/7712203/files/figure.png" httpUri-1="https://zenodo.org/record/7712205/files/figure.png" pageId="2">Figs. 1–2</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC336840C79B9671AEA0B40C90F250F" blockId="2.[122,792,1742,2001]" lastBlockId="2.[824,1493,569,1381]" pageId="2">
<materialsCitation id="3B143CD90C79B9671AEA0B40CC9221A3" ID-GBIF-Occurrence="4052787309" collectorName="Four" location="Brazilian" pageId="2" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="B908EA960C79B9671AEA0B40CC33214D" bold="true" box="[149,276,1889,1913]" pageId="2">
<typeStatus id="54C788260C79B9671AEA0B40CC37214D" box="[149,272,1889,1913]" pageId="2" type="paratype">Paratypes</typeStatus>
.
</emphasis>
— 
<collectorName id="268953520C79B9671B4E0B43CC41214D" box="[305,358,1890,1913]" pageId="2">Four</collectorName>
specimens, all from the 
<location id="8EA3605F0C79B96718050B40CFFB214D" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B96718050B40CFFB214D" box="[634,732,1889,1913]" name="Brazilian" pageId="2" stateProvince="Minas Gerais">Brazilian</location>
state of 
<collectingRegion id="49B8F8660C79B9671AE40B5ECC1321A3" box="[155,308,1919,1943]" country="Brazil" name="Minas Gerais" pageId="2">Minas Gerais</collectingRegion>
, as follows
</materialsCitation>
: 
<materialsCitation id="3B143CD90C79B9671BBD0B5ECF5421E5" ID-GBIF-Occurrence="4052787310" collectionCode="MZUFV" collectorName="Licio W. Assad" elevation="480" latitude="-15.34027" location="Jaiba" longLatPrecision="1" longitude="-43.67686" pageId="2" specimenCode="MZUFV 742" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBDA9EFF0C79B9671BBD0B5ECF7121A3" box="[450,598,1919,1943]" collectionCode="MZUFV" pageId="2">MZUFV 742</specimenCode>
, an adult female collected by 
<collectorName id="268953520C79B9671B610BBDCCF82180" box="[286,479,1948,1972]" pageId="2">Lício W. Assad</collectorName>
, date unknown, at 
<location id="8EA3605F0C79B967189E0BBDCE3F2180" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B967189E0BBDCE3F2180" box="[737,792,1948,1972]" latitude="-15.34027" longLatPrecision="1" longitude="-43.67686" name="Jaiba" pageId="2" stateProvince="Minas Gerais">Jaíba</location>
(
<geoCoordinate id="EE4850430C79B9671AFE0B98CC2921E5" box="[129,270,1977,2001]" degrees="15.340270" direction="south" orientation="latitude" pageId="2" precision="1" value="-15.34027">
15.340270 
<emphasis id="B908EA960C79B9671A8B0B98CDD921E5" bold="true" box="[244,254,1977,2001]" pageId="2">°</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="EE4850430C79B9671B620B98CC9321E4" box="[285,436,1977,2001]" degrees="43.676862" direction="west" orientation="longitude" pageId="2" precision="1" value="-43.67686">
43.676862 
<emphasis id="B908EA960C79B9671BF10B98CCBF21E5" bold="true" box="[398,408,1977,2001]" pageId="2">°</emphasis>
W
</geoCoordinate>
, ca. 
<elevation id="0051D1B70C79B9671B970B98CF4E21E5" box="[488,617,1977,2001]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="2" unit="m" value="480.0">
<quantity id="4C849B610C79B9671B970B98CF0821E4" box="[488,559,1977,2001]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="2" unit="m" value="480.0">480 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
; 
<materialsCitation id="3B143CD90C79B96718FE0B98C8AD2465" ID-GBIF-Occurrence="4052787307" box="[641,1418,569,2001]" collectingDate="1998-02" collectionCode="MZUFV" pageId="2" specimenCode="MZUFV 960" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBDA9EFF0C79B96718FE0B98CE3421E5" box="[641,787,1977,2001]" collectionCode="MZUFV" pageId="2">MZUFV 960</specimenCode>
, a young male, collector unknown, 
<date id="FFC210440C79B9671EBB0E18C84C2466" box="[1220,1387,569,594]" pageId="2" value="1998-02">
<collectingDate id="EF86E9AC0C79B9671EBB0E18C84C2466" box="[1220,1387,569,594]" pageId="2" value="1998-02">February 1998</collectingDate>
</date>
at
</materialsCitation>
<materialsCitation id="3B143CD90C79B9671FEA0E1BC817245B" ID-GBIF-Occurrence="4052787304" collectionCode="CBIA" elevation="640" latitude="-20.71365" longLatPrecision="1" longitude="-42.864918" pageId="2" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectionCode id="ED6DAE410C79B9671FEA0E1BC8F22466" box="[1429,1493,570,594]" pageId="2">CBIA</collectionCode>
(
<geoCoordinate id="EE4850430C79B96719400E76CEEB245B" box="[831,972,599,623]" degrees="20.713650" direction="south" orientation="latitude" pageId="2" precision="1" value="-20.71365">
20.713650 
<emphasis id="B908EA960C79B96719CD0E76CE9B245B" bold="true" box="[946,956,599,623]" pageId="2">°</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="EE4850430C79B96719A50E76C956245A" box="[986,1137,599,623]" degrees="42.864917" direction="west" orientation="longitude" pageId="2" precision="1" value="-42.864918">
42.864917 
<emphasis id="B908EA960C79B9671E340E76C972245B" bold="true" box="[1099,1109,599,623]" pageId="2">°</emphasis>
W
</geoCoordinate>
, ca. 
<elevation id="0051D1B70C79B9671EDA0E76C802245B" box="[1189,1317,599,623]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="2" unit="m" value="640.0">
<quantity id="4C849B610C79B9671EDA0E76C9CB245A" box="[1189,1260,599,623]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="2" unit="m" value="640.0">640 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
, 
<materialsCitation id="3B143CD90C79B9671F410E76CEA624B8" ID-GBIF-Occurrence="4052787306" location="Vicosa" pageId="2" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
urban area of 
<location id="8EA3605F0C79B96719470E55CEA624B8" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B96719470E55CEA624B8" box="[824,897,628,652]" name="Vicosa" pageId="2" stateProvince="Minas Gerais">Viçosa</location>
</materialsCitation>
; 
<materialsCitation id="3B143CD90C79B96719F10E55C84724B8" ID-GBIF-Occurrence="4052787301" box="[910,1376,628,652]" collectionCode="UFMG" pageId="2" specimenCode="UFMG 175" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBDA9EFF0C79B96719F10E55C93324B8" box="[910,1044,628,652]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="2">UFMG 175</specimenCode>
, an adult female collected by
</materialsCitation>
<materialsCitation id="3B143CD90C79B9671F160E55C8F724F2" ID-GBIF-Occurrence="4052787302" collectingDate="1999-01-06" collectionCode="V" collectorName="X. Silva" elevation="820" latitude="-19.869394" location="Universidade Federal" longLatPrecision="1" longitude="-43.96648" pageId="2" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectionCode id="ED6DAE410C79B9671F160E55C85A24BF" box="[1385,1405,628,651]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" type="Museum">V</collectionCode>
. 
<collectorName id="268953520C79B9671FFF0E55C8F224B8" box="[1408,1493,628,652]" pageId="2">X. Silva</collectorName>
on 
<date id="FFC210440C79B96719250EB0CED8249D" box="[858,1023,657,682]" pageId="2" value="1999-01-06">
<collectingDate id="EF86E9AC0C79B96719250EB0CED8249D" box="[858,1023,657,682]" pageId="2" value="1999-01-06">6 January 1999</collectingDate>
</date>
at the campus of 
<location id="8EA3605F0C79B9671EBC0EB0C894249D" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B9671EBC0EB0C894249D" box="[1219,1459,657,681]" latitude="-19.869394" longLatPrecision="1" longitude="-43.96648" name="Universidade Federal" pageId="2" stateProvince="Minas Gerais">Universidade Federal</location>
de 
<collectingRegion id="49B8F8660C79B96719470E8FCEEA24F2" box="[824,973,686,710]" country="Brazil" name="Minas Gerais" pageId="2">Minas Gerais</collectingRegion>
(
<geoCoordinate id="EE4850430C79B96719A20E8FC94D24F3" box="[989,1130,686,711]" degrees="19.869395" direction="south" orientation="latitude" pageId="2" precision="1" value="-19.869394">
19.869395 
<emphasis id="B908EA960C79B9671E2F0E8FC97D24F2" bold="true" box="[1104,1114,686,710]" pageId="2">°</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="EE4850430C79B9671E070E8FC83724F2" box="[1144,1296,686,710]" degrees="43.966480" direction="west" orientation="longitude" pageId="2" precision="1" value="-43.96648">
43.966480 
<emphasis id="B908EA960C79B9671E950E8FC9D324F2" bold="true" box="[1258,1268,686,710]" pageId="2">°</emphasis>
W
</geoCoordinate>
, ca. 
<elevation id="0051D1B70C79B9671F3A0E8EC8E124F2" box="[1349,1478,686,711]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" unit="m" value="820.0">
<quantity id="4C849B610C79B9671F3A0E8EC8AB24F2" box="[1349,1420,687,711]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" unit="m" value="820.0">820 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
, 
<materialsCitation id="3B143CD90C79B96719470EEDCED324D0" ID-GBIF-Occurrence="4052787308" box="[824,1012,716,740]" location="Belo Horizonte" pageId="2" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<location id="8EA3605F0C79B96719470EEDCED324D0" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B96719470EEDCED324D0" box="[824,1012,716,740]" name="Belo Horizonte" pageId="2" stateProvince="Minas Gerais">Belo Horizonte</location>
</materialsCitation>
; 
<materialsCitation id="3B143CD90C79B9671E720EEDCEE5250F" ID-GBIF-Occurrence="4052787305" collectingDate="2013-03-04" collectionCode="ZUFMS-REP" collectorName="Carmen Cortes Santana Silva" elevation="224" latitude="-21.145254" location="District of Macuco" longLatPrecision="1" longitude="-42.493443" pageId="2" specimenCode="ZUFMS-REP 2291" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBDA9EFF0C79B9671E720EEDC9D124D0" box="[1037,1270,716,740]" collectionCode="ZUFMS-REP" pageId="2">ZUFMS-REP 2291</specimenCode>
, a young female collected by 
<collectorName id="268953520C79B96719AA0EC8C8142535" box="[981,1331,745,769]" pageId="2">Carmen Côrtes Santana Silva</collectorName>
on 
<date id="FFC210440C79B9671F100EC8CE4A252A" pageId="2" value="2013-03-04">
<collectingDate id="EF86E9AC0C79B9671F100EC8CE4A252A" pageId="2" value="2013-03-04">4 March 2013</collectingDate>
</date>
at 
<location id="8EA3605F0C79B96719EC0F27C940252A" LSID="urn:lsid:plazi:treatment:03D587920C79B9621B420A19CE9720E4:8EA3605F0C79B96719EC0F27C940252A" box="[915,1127,774,798]" latitude="-21.145254" longLatPrecision="1" longitude="-42.493443" name="District of Macuco" pageId="2" stateProvince="Minas Gerais">District of Macuco</location>
(
<geoCoordinate id="EE4850430C79B9671E080F27C823252A" box="[1143,1284,774,798]" degrees="21.145255" direction="south" orientation="latitude" pageId="2" precision="1" value="-21.145254">
21.145255 
<emphasis id="B908EA960C79B9671E950F27C9D3252A" bold="true" box="[1258,1268,774,798]" pageId="2">°</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="EE4850430C79B9671F6D0F27C88D252A" box="[1298,1450,774,798]" degrees="42.493443" direction="west" orientation="longitude" pageId="2" precision="1" value="-42.493443">
42.493443 
<emphasis id="B908EA960C79B9671FFC0F27C8AA252A" bold="true" box="[1411,1421,774,798]" pageId="2">°</emphasis>
W
</geoCoordinate>
, ca. 
<elevation id="0051D1B70C79B96719470F02CE9F250F" box="[824,952,803,827]" metricMagnitude="2" metricUnit="m" metricValue="2.2399999999999998" pageId="2" unit="m" value="224.0">
<quantity id="4C849B610C79B96719470F02CE59250F" box="[824,894,803,827]" metricMagnitude="2" metricUnit="m" metricValue="2.2399999999999998" pageId="2" unit="m" value="224.0">224 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
, Muriaé.
</paragraph>
</subSubSection>
<subSubSection id="C366650F0C79B966192D0F1EC848237E" lastPageId="3" pageId="2" type="diagnosis">
<paragraph id="8BC336840C79B967192D0F1EC81A2351" blockId="2.[824,1493,569,1381]" pageId="2">
<emphasis id="B908EA960C79B967192D0F1ECEF62563" bold="true" box="[850,977,831,855]" pageId="2">Diagnosis.</emphasis>
— 
<taxonomicName id="4C7C4D070C79B96719940F1EC9B82563" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1003,1183,831,855]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C79B96719940F1EC9B82563" box="[1003,1183,831,855]" italics="true" pageId="2">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C79B9671ED50F61C8242563" box="[1194,1283,832,855]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
can be diagnosed by the following combination of characters: (1) dorsal scale rows 17–17–15, smooth on the anterior third of body, becoming weakly keeled posteriorly; (2) ventral plates in the male 115, in females 111–117; (3) subcaudals in the male 56, in females 41–52; (4) subcaudal keels absent, dorsal caudal scales keeled; (5) supralabials 8, third and fourth contacting orbit; (6) infralabials 9–10, the first to fifth contacting the first pair of chinshields, the fifth and sixth contacting the second pair of chinshields; (7) intergenials absent; (8) nasal entire; (9) preocular one; (10) postoculars two; (11) temporals 2 + 2 or 2 + 3; (12) maxillary teeth 18–21 + 2; (13) hemipenis bilobed, semicalyculate, semicapitate, with lobes covered by encircling rows of spinulate flounces, body covered by numerous spinules and a few spines arranged in more or less oblique rows; (14) dorsum uniformly medium brown or dark brown; (15) venter cream, yellow, or light brown (sometimes mottled with gray) with two medial rows of black markings, usually semilunar in shape.
</paragraph>
<caption id="DF03660C0C79B96719320BACCEEF21FB" ID-DOI="http://doi.org/10.5281/zenodo.7712203" ID-Zenodo-Dep="7712203" httpUri="https://zenodo.org/record/7712203/files/figure.png" pageId="2" startId="2.[845,859,1933,1952]" targetBox="[824,1496,1411,1915]" targetPageId="2">
<paragraph id="8BC336840C79B96719320BACCEEF21FB" blockId="2.[824,1493,1932,1999]" pageId="2">
FIG. 1.—Holotype of 
<taxonomicName id="4C7C4D070C79B9671E700BADC9BB21AB" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1039,1180,1932,1951]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C79B9671E700BADC9BB21AB" box="[1039,1180,1932,1951]" italics="true" pageId="2">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C79B9671EDC0BACC9CE2194" box="[1187,1257,1933,1952]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
(UFMG 2486): dorsal (A) and ventral (B) view. Scale bar = 20 mm. A color version of this figure is available online.
</paragraph>
</caption>
<caption id="DF03660C0C78B9661AF00879CC8822B7" ID-DOI="http://doi.org/10.5281/zenodo.7712205" ID-Zenodo-Dep="7712205" httpUri="https://zenodo.org/record/7712205/files/figure.png" pageId="3" startId="3.[143,157,1112,1131]" targetBox="[143,1522,189,1095]" targetPageId="3">
<paragraph id="8BC336840C78B9661AF00879CC8822B7" blockId="3.[143,1514,1112,1155]" pageId="3">
FIG. 2.—Holotype of 
<taxonomicName id="4C7C4D070C78B9661B330879CCFF225F" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[332,472,1112,1131]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B9661B330879CCFF225F" box="[332,472,1112,1131]" italics="true" pageId="3">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B9661BA10879CF05225F" box="[478,546,1112,1131]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
(UFMG 2486): head in dorsal (A), ventral (B), left (C), and right (D) view. Scale bar = 5 mm. A color version of this figure is available online.
</paragraph>
</caption>
<paragraph id="8BC336840C78B9661AD50881C848237E" blockId="3.[143,813,1184,1357]" lastBlockId="3.[845,1515,1184,2001]" pageId="3">
<emphasis id="B908EA960C78B9661AD50881CCD3228C" bold="true" box="[170,500,1184,1208]" pageId="3">
Comparisons (
<tableCitation id="C6FE033F0C78B9661B210881CCC0228C" box="[350,487,1184,1208]" captionStart-0="TABLE" captionStart-1="TABLE" captionStartId-0="5.[118,131,410,430]" captionStartId-1="5.[692,705,1061,1081]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="TABLE 3.—Summary of morphological characters of Helicops species. AD = anterior dorsal scale rows, MD = midbody dorsal scale rows, PD = posterior dorsal scale rows, DK = dorsal keels on posterior third of body (medium = M, strong = S, or weak = W), V = ventral plates, SC = subcaudal plates, SCK = subcaudal keels (present = Yes or absent = No), SL = supralabials, SL + E = supralabials contacting the eye, IL = infralabials, IL + G1 = infralabials contacting the first pair of genials, IG = intergenials (present = Yes or absent = No), NA = nasal (entire = E or semidivided = S), Pe = preoculars, Po = postoculars, AT = anterior temporals, PT = posterior temporals, MXT = maxillary teeth." captionText-1="TABLE 4.—Descriptions of the color patterns seen in Helicops species." httpUri-0="http://table.plazi.org/id/DF03660C0C7EB9601A090DBBC97B27C2" httpUri-1="http://table.plazi.org/id/DF03660C0C7EB96018CB0804C80E220D" pageId="3" tableUuid-0="DF03660C0C7EB9601A090DBBC97B27C2" tableUuid-1="DF03660C0C7EB96018CB0804C80E220D">Tables 3–4</tableCitation>
).
</emphasis>
— 
<taxonomicName id="4C7C4D070C78B96618700881CFE2228C" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[527,709,1184,1208]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B96618700881CFE2228C" box="[527,709,1184,1208]" italics="true" pageId="3">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B96618AD0880CE0A228C" box="[722,813,1185,1208]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
differs from all congeners except 
<taxonomicName id="4C7C4D070C78B966187E089ECF9B22E2" box="[513,700,1214,1238]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="infrataeniatus">
<emphasis id="B908EA960C78B966187E089ECF9B22E2" box="[513,700,1214,1238]" italics="true" pageId="3">H. infrataeniatus</emphasis>
</taxonomicName>
by having 17–17–15 dorsal scale rows. It also differs from all other 
<taxonomicName id="4C7C4D070C78B9661AF008D8CDC92325" authorityName="Wagler" authorityYear="1828" box="[143,238,1273,1297]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="genus">
<emphasis id="B908EA960C78B9661AF008D8CDC92325" box="[143,238,1273,1297]" italics="true" pageId="3">Helicops</emphasis>
</taxonomicName>
except 
<emphasis id="B908EA960C78B9661B3708DBCE0A2325" box="[328,813,1273,1297]" italics="true" pageId="3">
<taxonomicName id="4C7C4D070C78B9661B3708DBCCF22325" authorityName="Gunther" authorityYear="1861" box="[328,469,1273,1297]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="modestus">H. modestus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661B9C08DBCFA02325" authorityName="Frota" authorityYear="2005" box="[483,647,1274,1297]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tapajonicus">H. tapajonicus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B96618EA08DBCE0F2325" box="[661,808,1274,1297]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="trivittatus">H. triƲittatus</taxonomicName>
,
</emphasis>
and some 
<taxonomicName id="4C7C4D070C78B9661B640939CCFF231B" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[283,472,1303,1327]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="carinicaudus">
<emphasis id="B908EA960C78B9661B640939CCFF231B" box="[283,472,1303,1327]" italics="true" pageId="3">H. carinicaudus</emphasis>
</taxonomicName>
by the presence of weak dorsal keels on the posterior third of body (character unknown in 
<taxonomicName id="4C7C4D070C78B96619950881C96D2283" box="[1002,1098,1184,1207]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C78B96619950881C96D2283" box="[1002,1098,1184,1207]" italics="true" pageId="3">H. ŋacu</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C7C4D070C78B9661E150881C8032283" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1130,1316,1184,1208]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B9661E150881C8032283" box="[1130,1316,1184,1208]" italics="true" pageId="3">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B9661F4B0881C8B42283" box="[1332,1427,1184,1207]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<emphasis id="B908EA960C78B96619F1089FC9EF22E1" box="[910,1224,1213,1237]" italics="true" pageId="3">
<taxonomicName id="4C7C4D070C78B96619F1089FC96222E1" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[910,1093,1213,1237]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="carinicaudus">H. carinicaudus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661E29089FC9EF22E1" box="[1110,1224,1213,1237]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="danieli">H. danieli</taxonomicName>
</emphasis>
, 
<emphasis id="B908EA960C78B9661EA5089FCEE322C6" italics="true" pageId="3">
<taxonomicName id="4C7C4D070C78B9661EA5089FC86822E1" box="[1242,1359,1214,1237]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="gomesi">H. gomesi</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661F20089FC8C122E1" box="[1375,1510,1214,1237]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="pastazae">H. pastazae</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B966193208FACE9822C6" box="[845,959,1243,1266]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="petersi">H. petersi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C78B9661E7B08FAC97922C6" box="[1028,1118,1243,1266]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C78B9661E7B08FAC97922C6" box="[1028,1118,1243,1266]" italics="true" pageId="3">H. ŋacu</emphasis>
</taxonomicName>
by having fewer than 119 ventral scales in both sexes. It further differs from 
<emphasis id="B908EA960C78B9661F0F08D9CEC22319" italics="true" pageId="3">
<taxonomicName id="4C7C4D070C78B9661F0F08D9C8C0233B" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[1392,1511,1271,1295]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="apiaka">H. apiaka</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B96619320934CEC62319" box="[845,993,1301,1325]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="hagmanni">H. hagmanni</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C78B9661E5E0934C9E42318" authorityName="Frota" authorityYear="2005" box="[1057,1219,1301,1324]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="B908EA960C78B9661E5E0934C9E42318" box="[1057,1219,1301,1324]" italics="true" pageId="3">H. tapajonicus</emphasis>
</taxonomicName>
by having fewer than 117 ventrals in males and fewer than 
<quantity id="4C849B610C78B9661EBA0913C82D237E" box="[1221,1290,1330,1354]" metricMagnitude="0" metricUnit="m" metricValue="3.0734" pageId="3" unit="in" value="121.0">121 in</quantity>
females.
</paragraph>
</subSubSection>
<subSubSection id="C366650F0C78B96319180973C8B921A2" lastPageId="6" pageId="3" type="description">
<paragraph id="8BC336840C78B96619180973C99520BB" blockId="3.[845,1515,1184,2001]" pageId="3">
<taxonomicName id="4C7C4D070C78B96619180973C904235E" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[871,1059,1362,1386]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B96619180973C904235E" box="[871,1059,1362,1386]" italics="true" pageId="3">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B9661E490972C9B0235E" box="[1078,1175,1363,1386]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C7C4D070C78B9661F340972C8C3235E" box="[1355,1508,1362,1386]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="angulatus">
<emphasis id="B908EA960C78B9661F340972C8C3235E" box="[1355,1508,1362,1386]" italics="true" pageId="3">H. angulatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B96619320951CE9A23BC" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[845,957,1392,1416]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="apiaka">
<emphasis id="B908EA960C78B96619320951CE9A23BC" box="[845,957,1392,1416]" italics="true" pageId="3">H. apiaka</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B96619AF0951C96523BC" box="[976,1090,1392,1416]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="danieli">
<emphasis id="B908EA960C78B96619AF0951C96523BC" box="[976,1090,1392,1416]" italics="true" pageId="3">H. danieli</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661E2A0951C9EF23B3" box="[1109,1224,1392,1415]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="gomesi">
<emphasis id="B908EA960C78B9661E2A0951C9EF23B3" box="[1109,1224,1392,1415]" italics="true" pageId="3">H. gomesi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661EA40951C84723B3" box="[1243,1376,1392,1415]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="pastazae">
<emphasis id="B908EA960C78B9661EA40951C84723B3" box="[1243,1376,1392,1415]" italics="true" pageId="3">H. pastazae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661F0C0951C8C323B3" box="[1395,1508,1392,1415]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B908EA960C78B9661F0C0951C8C323B3" box="[1395,1508,1392,1415]" italics="true" pageId="3">H. petersi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B966193209AFCEE92391" box="[845,974,1421,1445]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="polnlepis">
<emphasis id="B908EA960C78B966193209AFCEE92391" box="[845,974,1421,1445]" italics="true" pageId="3">H. polŋlepis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B96619A609AFC96C2391" box="[985,1099,1421,1445]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="scalaris">
<emphasis id="B908EA960C78B96619A609AFC96C2391" box="[985,1099,1421,1445]" italics="true" pageId="3">H. scalaris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661E2909AFC9D42390" authorityName="Frota" authorityYear="2005" box="[1110,1267,1421,1445]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="B908EA960C78B9661E2909AFC9D42390" box="[1110,1267,1421,1445]" italics="true" pageId="3">H. tapajonicus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7C4D070C78B9661F5409AFC8582391" box="[1323,1407,1422,1445]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C78B9661F5409AFC8582391" box="[1323,1407,1422,1445]" italics="true" pageId="3">H. ŋacu</emphasis>
</taxonomicName>
by having fewer than 60 subcaudals in both sexes. The presence of 56 subcaudals in the male and 
<quantity id="4C849B610C78B9661EB309E6C81523EB" box="[1228,1330,1479,1503]" metricMagnitude="0" metricUnit="m" metricValue="1.1811" metricValueMax="1.3208" metricValueMin="1.0413999999999999" pageId="3" unit="in" value="46.5" valueMax="52.0" valueMin="41.0">41–52 in</quantity>
the females of 
<taxonomicName id="4C7C4D070C78B966193209C4CE9A23C8" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[845,957,1509,1532]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B966193209C4CE9A23C8" box="[845,957,1509,1532]" italics="true" pageId="3">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B96619B709C4C90523C8" box="[968,1058,1509,1532]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
also distinguishes it from 
<taxonomicName id="4C7C4D070C78B9661F2809C4C8CD23C8" box="[1367,1514,1509,1532]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C78B9661F2809C4C8CD23C8" box="[1367,1514,1509,1532]" italics="true" pageId="3">H. triƲittatus</emphasis>
</taxonomicName>
(subcaudals 
<quantity id="4C849B610C78B96619A10A23C962202E" box="[990,1093,1538,1562]" metricMagnitude="0" metricUnit="m" metricValue="1.8669" metricValueMax="2.032" metricValueMin="1.7018" pageId="3" unit="in" value="73.5" valueMax="80.0" valueMin="67.0">67–80 in</quantity>
males and 
<quantity id="4C849B610C78B9661EA20A20C863202E" box="[1245,1348,1537,1562]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" metricValueMax="1.6764" metricValueMin="1.4224" pageId="3" unit="in" value="61.0" valueMax="66.0" valueMin="56.0">56–66 in</quantity>
females) and 
<taxonomicName id="4C7C4D070C78B96619320A01CEDE2003" box="[845,1017,1567,1591]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="leopardinus">
<emphasis id="B908EA960C78B96619320A01CEDE2003" box="[845,1017,1567,1591]" italics="true" pageId="3">H. leopardinus</emphasis>
</taxonomicName>
(
<quantity id="4C849B610C78B9661E6F0A3EC95E2003" box="[1040,1145,1567,1591]" metricMagnitude="0" metricUnit="m" metricValue="1.9431" metricValueMax="2.2606" metricValueMin="1.6256" pageId="3" unit="in" value="76.5" valueMax="89.0" valueMin="64.0">64–89 in</quantity>
males and 
<quantity id="4C849B610C78B9661F700A3EC8502003" box="[1295,1399,1567,1591]" metricMagnitude="0" metricUnit="m" metricValue="1.6383" metricValueMax="1.9304" metricValueMin="1.3462" pageId="3" unit="in" value="64.5" valueMax="76.0" valueMin="53.0">53–76 in</quantity>
females). The absence of subcaudal keels distinguish 
<taxonomicName id="4C7C4D070C78B9661F580A1CC8B62060" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1319,1425,1597,1620]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B9661F580A1CC8B62060" box="[1319,1425,1597,1620]" italics="true" pageId="3">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B9661FE80A1CC8CC2060" box="[1431,1515,1597,1620]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="4C7C4D070C78B96619F10A7BC9062045" box="[910,1057,1625,1649]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="angulatus">
<emphasis id="B908EA960C78B96619F10A7BC9062045" box="[910,1057,1625,1649]" italics="true" pageId="3">H. angulatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661E4C0A7BC9832045" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[1075,1188,1625,1649]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="apiaka">
<emphasis id="B908EA960C78B9661E4C0A7BC9832045" box="[1075,1188,1625,1649]" italics="true" pageId="3">H. apiaka</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661EC80A7BC80E2045" box="[1207,1321,1626,1649]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="gomesi">
<emphasis id="B908EA960C78B9661EC80A7BC80E2045" box="[1207,1321,1626,1649]" italics="true" pageId="3">H. gomesi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7C4D070C78B9661F0E0A7BC8CD2045" box="[1393,1514,1625,1649]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="scalaris">
<emphasis id="B908EA960C78B9661F0E0A7BC8CD2045" box="[1393,1514,1625,1649]" italics="true" pageId="3">H. scalaris</emphasis>
</taxonomicName>
(character unknown in 
<taxonomicName id="4C7C4D070C78B9661E320A56C98420BA" box="[1101,1187,1655,1678]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C78B9661E320A56C98420BA" box="[1101,1187,1655,1678]" italics="true" pageId="3">H. ŋacu</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC336840C78B96619180AB4C8BE21A2" blockId="3.[845,1515,1184,2001]" pageId="3">
In 
<taxonomicName id="4C7C4D070C78B96619FB0AB4CECD2098" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[900,1002,1685,1708]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B96619FB0AB4CECD2098" box="[900,1002,1685,1708]" italics="true" pageId="3">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B96619910AB4C9672098" box="[1006,1088,1685,1708]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
, the third and fourth supralabials contact the eye, whereas in 
<emphasis id="B908EA960C78B9661E450A93CEE420D3" italics="true" pageId="3">
<taxonomicName id="4C7C4D070C78B9661E450A93C98B20FD" box="[1082,1196,1713,1737]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="danieli">H. danieli</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661EC20A93C87520FD" box="[1213,1362,1713,1737]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="hagmanni">H. hagmanni</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B9661F1D0A93C8C020FD" box="[1378,1511,1714,1737]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="pastazae">H. pastazae</taxonomicName>
, 
<taxonomicName id="4C7C4D070C78B96619320AF1CEE720D3" box="[845,960,1744,1767]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="petersi">H. petersi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C78B9661E770AF1C98D20D3" authorityName="Frota" authorityYear="2005" box="[1032,1194,1744,1767]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="B908EA960C78B9661E770AF1C98D20D3" box="[1032,1194,1744,1767]" italics="true" pageId="3">H. tapajonicus</emphasis>
</taxonomicName>
only the fourth supralabial contacts the eye. This character also distinguishes 
<taxonomicName id="4C7C4D070C78B9661F200ACCC8E02130" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1375,1479,1773,1796]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B9661F200ACCC8E02130" box="[1375,1479,1773,1796]" italics="true" pageId="3">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B9661FB30ACDCE5C2115" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="4C7C4D070C78B96619AC0B2BC94F2115" box="[979,1128,1801,1825]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="angulatus">
<emphasis id="B908EA960C78B96619AC0B2BC94F2115" box="[979,1128,1801,1825]" italics="true" pageId="3">H. angulatus</emphasis>
</taxonomicName>
(fourth, fourth + fifth or fifth supralabial[s] contacting the eye), 
<taxonomicName id="4C7C4D070C78B9661EC10B06C80E210A" box="[1214,1321,1831,1854]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="gomesi">
<emphasis id="B908EA960C78B9661EC10B06C80E210A" box="[1214,1321,1831,1854]" italics="true" pageId="3">H. gomesi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7C4D070C78B9661F200B06C8CD210A" box="[1375,1514,1831,1854]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C78B9661F200B06C8CD210A" box="[1375,1514,1831,1854]" italics="true" pageId="3">H. triƲittatus</emphasis>
</taxonomicName>
(fourth or fifth supralabial contacting the eye), 
<taxonomicName id="4C7C4D070C78B9661F0C0B65C8CC2168" box="[1395,1515,1860,1884]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="scalaris">
<emphasis id="B908EA960C78B9661F0C0B65C8CC2168" box="[1395,1515,1860,1884]" italics="true" pageId="3">H. scalaris</emphasis>
</taxonomicName>
(fourth or fourth + fifth supralabial[s] contacting the eye), and 
<taxonomicName id="4C7C4D070C78B96619320B5ECE8621A2" box="[845,929,1919,1942]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C78B96619320B5ECE8621A2" box="[845,929,1919,1942]" italics="true" pageId="3">H. ŋacu</emphasis>
</taxonomicName>
(fourth + fifth supralabials contacting the eye).
</paragraph>
<caption id="DF03660C0C78B9661ADA0B54CC0721FB" ID-DOI="http://doi.org/10.5281/zenodo.7712207" ID-Zenodo-Dep="7712207" httpUri="https://zenodo.org/record/7712207/files/figure.png" pageId="3" startId="3.[165,179,1909,1928]" targetBox="[143,814,1386,1891]" targetPageId="3">
<paragraph id="8BC336840C78B9661ADA0B54CC0721FB" blockId="3.[143,814,1908,1999]" pageId="3">
FIG. 3.—Right hemipenis of a paratype of 
<taxonomicName id="4C7C4D070C78B966183A0B55CFFF21B3" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[581,728,1908,1927]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B966183A0B55CFFF21B3" box="[581,728,1908,1927]" italics="true" pageId="3">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B966189C0B54CE0921BC" box="[739,814,1909,1928]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
(MZUFV 960) fully everted and almost maximally expanded: sulcate (A) and asulcate (B) side. Scale bar = 10 mm. A color version of this figure is available online.
</paragraph>
</caption>
<paragraph id="8BC336840C78B96119180BBDCFF32445" blockId="3.[845,1515,1184,2001]" lastBlockId="4.[122,792,513,2001]" lastPageId="4" pageId="3">
The presence of 9–10 infralabials distinguishes 
<taxonomicName id="4C7C4D070C78B9661F020BBCC8CC2180" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1405,1515,1949,1972]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C78B9661F020BBCC8CC2180" box="[1405,1515,1949,1972]" italics="true" pageId="3">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C78B96619320B98CE8621E4" box="[845,929,1977,2000]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="4C7C4D070C78B966199D0B9BC94221E5" box="[994,1125,1977,2001]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="polnlepis">
<emphasis id="B908EA960C78B966199D0B9BC94221E5" box="[994,1125,1977,2001]" italics="true" pageId="3">H. polŋlepis</emphasis>
</taxonomicName>
(11–13), 
<taxonomicName id="4C7C4D070C78B9661EB30B9BC87D21E4" box="[1228,1370,1977,2001]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C78B9661EB30B9BC87D21E4" box="[1228,1370,1977,2001]" italics="true" pageId="3">H. triƲittatus</emphasis>
</taxonomicName>
(11–14), and 
<taxonomicName id="4C7C4D070C7FB9611A050E23CDF2242D" box="[122,213,514,537]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C7FB9611A050E23CDF2242D" box="[122,213,514,537]" italics="true" pageId="4">H. ŋacu</emphasis>
</taxonomicName>
(11–12). 
<taxonomicName id="4C7C4D070C7FB9611B360E20CCD9242D" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[329,510,513,537]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611B360E20CCD9242D" box="[329,510,513,537]" italics="true" pageId="4">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118750E23CF43242D" box="[522,612,514,537]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
can be further distinguished from 
<emphasis id="B908EA960C7FB9611B2D0E3ECCA52467" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB9611B2D0E3ECCC22403" box="[338,485,543,567]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="hagmanni">H. hagmanni</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611B8D0E3ECF522402" box="[498,629,543,566]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="pastazae">H. pastazae</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118FE0E3ECFD72402" box="[641,752,543,566]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="petersi">H. petersi</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118820E3ECDE82460" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="scalaris">H. scalaris</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611A9C0E1CCC5A2467" box="[227,381,572,596]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">H. triƲittatus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB9611BB30E1CCF0C2460" box="[460,555,573,596]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C7FB9611BB30E1CCF0C2460" box="[460,555,573,596]" italics="true" pageId="4">H. ŋacu</emphasis>
</taxonomicName>
by the absence of intergenials separating the second pair of chinshields.
</paragraph>
<caption id="DF03660C0C7FB9611AF00CE3CE8F2728" ID-Table-UUID="DF03660C0C7FB9611AF00CE3CE8F2728" httpUri="http://table.plazi.org/id/DF03660C0C7FB9611AF00CE3CE8F2728" pageId="4" startId="4.[143,156,194,213]" targetBox="[122,1493,315,465]" targetIsTable="true" targetPageId="4">
<paragraph id="8BC336840C7FB9611AF00CE3CE8F2728" blockId="4.[122,1494,193,284]" pageId="4">
TABLE 2.—Meristic and morphometric characters in 
<taxonomicName id="4C7C4D070C7FB96118160CE0CFD126E0" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[617,758,193,212]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB96118160CE0CFD126E0" box="[617,758,193,212]" italics="true" pageId="4">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118810CE3CE6326E1" box="[766,836,194,213]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
PV = preventrals, VE = ventrals, C = cloacal shield, SC = subcaudals, SL = supralabials, SL + E = supralabials in contact with eye, IL = infralabials, IL + G1 = infralabials in contact with first pair of genials, IL + G2 = infralabials in contact with second pair of genials, NA = nasal (entire = E or semidivided = S), Pe = preocular, Po = postocular, AT = anterior temporals, PT = posterior temporals, MXT = maxillary teeth, SVL = snout–vent length (mm), TL = tail length (mm).
</paragraph>
</caption>
<paragraph id="8BC336840C7FB9611A040D1AC89A27E5" pageId="4">
<table id="F97CC4240C7F469A1A050D1AC8F227E5" box="[122,1493,315,465]" gridcols="20" gridrows="6" pageId="4">
<tr id="354C34C60C7F469A1A050D1AC8F2277D" box="[122,1493,315,329]" gridrow="0" pageId="4">
<th id="769D5DBA0C7F469A1A050D1ACC68277D" box="[122,335,315,329]" gridcol="0" gridrow="0" pageId="4">Specimen</th>
<th id="769D5DBA0C7F469A1B1B0D1ACC5C277D" box="[356,379,315,329]" gridcol="1" gridrow="0" pageId="4">Sex</th>
<th id="769D5DBA0C7F469A1BEE0D1ACCC1277D" box="[401,486,315,329]" gridcol="2" gridrow="0" pageId="4">Dorsals</th>
<th id="769D5DBA0C7F469A1B840D1ACF29277D" box="[507,526,315,329]" gridcol="3" gridrow="0" pageId="4">PV</th>
<th id="769D5DBA0C7F469A185A0D1ACF62277D" box="[549,581,315,329]" gridcol="4" gridrow="0" pageId="4">VE</th>
<th id="769D5DBA0C7F469A18250D1ACFBC277D" box="[602,667,315,329]" gridcol="5" gridrow="0" pageId="4">C</th>
<th id="769D5DBA0C7F469A18CE0D1ACFCC277D" box="[689,747,315,329]" gridcol="6" gridrow="0" pageId="4">SC</th>
<th id="769D5DBA0C7F469A197D0D1ACE32277D" box="[770,789,315,329]" gridcol="7" gridrow="0" pageId="4">SL</th>
<th id="769D5DBA0C7F469A19570D1ACE7E277D" box="[808,857,315,329]" gridcol="8" gridrow="0" pageId="4">SL + E</th>
<th id="769D5DBA0C7F469A19100D1AC930277D" box="[879,1047,315,329]" colspan="3" colspanRight="2" gridcol="9" gridrow="0" pageId="4">IL IL + G1 IL + G2</th>
<th id="769D5DBA0C7F469A1E520D1AC96A277D" box="[1069,1101,315,329]" gridcol="12" gridrow="0" pageId="4">NA</th>
<th id="769D5DBA0C7F469A1E1D0D1AC954277D" box="[1122,1139,315,329]" gridcol="13" gridrow="0" pageId="4">Pe</th>
<th id="769D5DBA0C7F469A1EF60D1AC9BD277D" box="[1161,1178,315,329]" gridcol="14" gridrow="0" pageId="4">Po</th>
<th id="769D5DBA0C7F469A1ECE0D1AC9E2277D" box="[1201,1221,315,329]" gridcol="15" gridrow="0" pageId="4">AT</th>
<th id="769D5DBA0C7F469A1EA40D1AC9D1277D" box="[1243,1270,315,329]" gridcol="16" gridrow="0" pageId="4">PT</th>
<th id="769D5DBA0C7F469A1F740D1AC862277D" box="[1291,1349,315,329]" gridcol="17" gridrow="0" pageId="4">MXT</th>
<th id="769D5DBA0C7F469A1F250D1AC85D277D" box="[1370,1402,315,329]" gridcol="18" gridrow="0" pageId="4">SVL</th>
<th id="769D5DBA0C7F469A1FEF0D1AC8F2277D" box="[1424,1493,315,329]" gridcol="19" gridrow="0" pageId="4">TL</th>
</tr>
<tr id="354C34C60C7F469A1A050D7FC8F22745" box="[122,1493,350,369]" gridrow="1" pageId="4">
<th id="769D5DBA0C7F469A1A050D7FCC682745" box="[122,335,350,369]" gridcol="0" gridrow="1" pageId="4">UFMG 2486 (holotype)</th>
<td id="769D5DBA0C7F469A1B1B0D7FCC5C2745" box="[356,379,350,369]" gridcol="1" gridrow="1" pageId="4">F</td>
<td id="769D5DBA0C7F469A1BEE0D7FCCC12745" box="[401,486,350,369]" gridcol="2" gridrow="1" pageId="4">17–17–15</td>
<td id="769D5DBA0C7F469A1B840D7FCF292745" box="[507,526,350,369]" gridcol="3" gridrow="1" pageId="4">2</td>
<td id="769D5DBA0C7F469A185A0D7FCF622745" box="[549,581,350,369]" gridcol="4" gridrow="1" pageId="4">117</td>
<td id="769D5DBA0C7F469A18250D7FCFBC2745" box="[602,667,350,369]" gridcol="5" gridrow="1" pageId="4">divided</td>
<td id="769D5DBA0C7F469A18CE0D7FCFCC2745" box="[689,747,350,369]" gridcol="6" gridrow="1" pageId="4">41</td>
<td id="769D5DBA0C7F469A197D0D7FCE322745" box="[770,789,350,369]" gridcol="7" gridrow="1" pageId="4">8</td>
<td id="769D5DBA0C7F469A19570D7FCE7E2745" box="[808,857,350,369]" gridcol="8" gridrow="1" pageId="4">iii–iv</td>
<td id="769D5DBA0C7F469A19100D7FCEB22745" box="[879,917,350,369]" gridcol="9" gridrow="1" pageId="4">10</td>
<td id="769D5DBA0C7F469A19D40D7FCEE32745" box="[939,964,350,369]" gridcol="10" gridrow="1" pageId="4">i–v</td>
<td id="769D5DBA0C7F469A19950D7FC9302745" box="[1002,1047,350,369]" gridcol="11" gridrow="1" pageId="4">v–vi</td>
<td id="769D5DBA0C7F469A1E520D7FC96A2745" box="[1069,1101,350,369]" gridcol="12" gridrow="1" pageId="4">E:S</td>
<td id="769D5DBA0C7F469A1E1D0D7FC9542745" box="[1122,1139,350,369]" gridcol="13" gridrow="1" pageId="4">1</td>
<td id="769D5DBA0C7F469A1EF60D7FC9BD2745" box="[1161,1178,350,369]" gridcol="14" gridrow="1" pageId="4">2</td>
<td id="769D5DBA0C7F469A1ECE0D7FC9E22745" box="[1201,1221,350,369]" gridcol="15" gridrow="1" pageId="4">2</td>
<td id="769D5DBA0C7F469A1EA40D7FC9D12745" box="[1243,1270,350,369]" gridcol="16" gridrow="1" pageId="4">2</td>
<td id="769D5DBA0C7F469A1F740D7FC8622745" box="[1291,1349,350,369]" gridcol="17" gridrow="1" pageId="4">19 + 2</td>
<td id="769D5DBA0C7F469A1F250D7FC85D2745" box="[1370,1402,350,369]" gridcol="18" gridrow="1" pageId="4">334</td>
<td id="769D5DBA0C7F469A1FEF0D7FC8F22745" box="[1424,1493,350,369]" gridcol="19" gridrow="1" pageId="4">77</td>
</tr>
<tr id="354C34C60C7F469A1A050D57C8F227BD" box="[122,1493,374,393]" gridrow="2" pageId="4">
<th id="769D5DBA0C7F469A1A050D57CC6827BD" box="[122,335,374,393]" gridcol="0" gridrow="2" pageId="4">MZUFV 960</th>
<td id="769D5DBA0C7F469A1B1B0D57CC5C27BD" box="[356,379,374,393]" gridcol="1" gridrow="2" pageId="4">M</td>
<td id="769D5DBA0C7F469A1BEE0D57CCC127BD" box="[401,486,374,393]" gridcol="2" gridrow="2" pageId="4">17–17–15</td>
<td id="769D5DBA0C7F469A1B840D57CF2927BD" box="[507,526,374,393]" gridcol="3" gridrow="2" pageId="4">2</td>
<td id="769D5DBA0C7F469A185A0D57CF6227BD" box="[549,581,374,393]" gridcol="4" gridrow="2" pageId="4">115</td>
<td id="769D5DBA0C7F469A18250D57CFBC27BD" box="[602,667,374,393]" gridcol="5" gridrow="2" pageId="4">divided</td>
<td id="769D5DBA0C7F469A18CE0D57CFCC27BD" box="[689,747,374,393]" gridcol="6" gridrow="2" pageId="4">56</td>
<td id="769D5DBA0C7F469A197D0D57CE3227BD" box="[770,789,374,393]" gridcol="7" gridrow="2" pageId="4">8</td>
<td id="769D5DBA0C7F469A19570D57CE7E27BD" box="[808,857,374,393]" gridcol="8" gridrow="2" pageId="4">iii–iv</td>
<td id="769D5DBA0C7F469A19100D57CEB227BD" box="[879,917,374,393]" gridcol="9" gridrow="2" pageId="4">10</td>
<td id="769D5DBA0C7F469A19D40D57CEE327BD" box="[939,964,374,393]" gridcol="10" gridrow="2" pageId="4">i–v</td>
<td id="769D5DBA0C7F469A19950D57C93027BD" box="[1002,1047,374,393]" gridcol="11" gridrow="2" pageId="4">v–vi</td>
<td id="769D5DBA0C7F469A1E520D57C96A27BD" box="[1069,1101,374,393]" gridcol="12" gridrow="2" pageId="4">E</td>
<td id="769D5DBA0C7F469A1E1D0D57C95427BD" box="[1122,1139,374,393]" gridcol="13" gridrow="2" pageId="4">1</td>
<td id="769D5DBA0C7F469A1EF60D57C9BD27BD" box="[1161,1178,374,393]" gridcol="14" gridrow="2" pageId="4">2</td>
<td id="769D5DBA0C7F469A1ECE0D57C9E227BD" box="[1201,1221,374,393]" gridcol="15" gridrow="2" pageId="4">2</td>
<td id="769D5DBA0C7F469A1EA40D57C9D127BD" box="[1243,1270,374,393]" gridcol="16" gridrow="2" pageId="4">3:2</td>
<td id="769D5DBA0C7F469A1F740D57C86227BD" box="[1291,1349,374,393]" gridcol="17" gridrow="2" pageId="4">21 + 2</td>
<td id="769D5DBA0C7F469A1F250D57C85D27BD" box="[1370,1402,374,393]" gridcol="18" gridrow="2" pageId="4">318</td>
<td id="769D5DBA0C7F469A1FEF0D57C8F227BD" box="[1424,1493,374,393]" gridcol="19" gridrow="2" pageId="4">108</td>
</tr>
<tr id="354C34C60C7F469A1A050DAFC8F22795" box="[122,1493,398,417]" gridrow="3" pageId="4">
<th id="769D5DBA0C7F469A1A050DAFCC682795" box="[122,335,398,417]" gridcol="0" gridrow="3" pageId="4">MZUFV 742</th>
<td id="769D5DBA0C7F469A1B1B0DAFCC5C2795" box="[356,379,398,417]" gridcol="1" gridrow="3" pageId="4">F</td>
<td id="769D5DBA0C7F469A1BEE0DAFCCC12795" box="[401,486,398,417]" gridcol="2" gridrow="3" pageId="4">17–17–15</td>
<td id="769D5DBA0C7F469A1B840DAFCF292795" box="[507,526,398,417]" gridcol="3" gridrow="3" pageId="4">1</td>
<td id="769D5DBA0C7F469A185A0DAFCF622795" box="[549,581,398,417]" gridcol="4" gridrow="3" pageId="4">116</td>
<td id="769D5DBA0C7F469A18250DAFCFBC2795" box="[602,667,398,417]" gridcol="5" gridrow="3" pageId="4">divided</td>
<td id="769D5DBA0C7F469A18CE0DAFCFCC2795" box="[689,747,398,417]" gridcol="6" gridrow="3" pageId="4">44</td>
<td id="769D5DBA0C7F469A197D0DAFCE322795" box="[770,789,398,417]" gridcol="7" gridrow="3" pageId="4">8</td>
<td id="769D5DBA0C7F469A19570DAFCE7E2795" box="[808,857,398,417]" gridcol="8" gridrow="3" pageId="4">iii–iv</td>
<td id="769D5DBA0C7F469A19100DAFCEB22795" box="[879,917,398,417]" gridcol="9" gridrow="3" pageId="4">10</td>
<td id="769D5DBA0C7F469A19D40DAFCEE32795" box="[939,964,398,417]" gridcol="10" gridrow="3" pageId="4">i–v</td>
<td id="769D5DBA0C7F469A19950DAFC9302795" box="[1002,1047,398,417]" gridcol="11" gridrow="3" pageId="4">v–vi</td>
<td id="769D5DBA0C7F469A1E520DAFC96A2795" box="[1069,1101,398,417]" gridcol="12" gridrow="3" pageId="4">E</td>
<td id="769D5DBA0C7F469A1E1D0DAFC9542795" box="[1122,1139,398,417]" gridcol="13" gridrow="3" pageId="4">1</td>
<td id="769D5DBA0C7F469A1EF60DAFC9BD2795" box="[1161,1178,398,417]" gridcol="14" gridrow="3" pageId="4">2</td>
<td id="769D5DBA0C7F469A1ECE0DAFC9E22795" box="[1201,1221,398,417]" gridcol="15" gridrow="3" pageId="4">2</td>
<td id="769D5DBA0C7F469A1EA40DAFC9D12795" box="[1243,1270,398,417]" gridcol="16" gridrow="3" pageId="4">2:3</td>
<td id="769D5DBA0C7F469A1F740DAFC8622795" box="[1291,1349,398,417]" gridcol="17" gridrow="3" pageId="4">18 + 2</td>
<td id="769D5DBA0C7F469A1F250DAFC85D2795" box="[1370,1402,398,417]" gridcol="18" gridrow="3" pageId="4">432</td>
<td id="769D5DBA0C7F469A1FEF0DAFC8F22795" box="[1424,1493,398,417]" gridcol="19" gridrow="3" pageId="4">106</td>
</tr>
<tr id="354C34C60C7F469A1A050D84C8F2278D" box="[122,1493,421,441]" gridrow="4" pageId="4">
<th id="769D5DBA0C7F469A1A050D84CC68278D" box="[122,335,421,441]" gridcol="0" gridrow="4" pageId="4">UFMG 175</th>
<td id="769D5DBA0C7F469A1B1B0D84CC5C278D" box="[356,379,421,441]" gridcol="1" gridrow="4" pageId="4">F</td>
<td id="769D5DBA0C7F469A1BEE0D84CCC1278D" box="[401,486,421,441]" gridcol="2" gridrow="4" pageId="4">17–17–15</td>
<td id="769D5DBA0C7F469A1B840D84CF29278D" box="[507,526,421,441]" gridcol="3" gridrow="4" pageId="4">1</td>
<td id="769D5DBA0C7F469A185A0D84CF62278D" box="[549,581,421,441]" gridcol="4" gridrow="4" pageId="4">117</td>
<td id="769D5DBA0C7F469A18250D84CFBC278D" box="[602,667,421,441]" gridcol="5" gridrow="4" pageId="4">divided</td>
<td id="769D5DBA0C7F469A18CE0D84CFCC278D" box="[689,747,421,441]" gridcol="6" gridrow="4" pageId="4">40 + n</td>
<td id="769D5DBA0C7F469A197D0D84CE32278D" box="[770,789,421,441]" gridcol="7" gridrow="4" pageId="4">8</td>
<td id="769D5DBA0C7F469A19570D84CE7E278D" box="[808,857,421,441]" gridcol="8" gridrow="4" pageId="4">iii–iv</td>
<td id="769D5DBA0C7F469A19100D84CEB2278D" box="[879,917,421,441]" gridcol="9" gridrow="4" pageId="4">10</td>
<td id="769D5DBA0C7F469A19D40D84CEE3278D" box="[939,964,421,441]" gridcol="10" gridrow="4" pageId="4">i–v</td>
<td id="769D5DBA0C7F469A19950D84C930278D" box="[1002,1047,421,441]" gridcol="11" gridrow="4" pageId="4">v–vi</td>
<td id="769D5DBA0C7F469A1E520D84C96A278D" box="[1069,1101,421,441]" gridcol="12" gridrow="4" pageId="4">E</td>
<td id="769D5DBA0C7F469A1E1D0D84C954278D" box="[1122,1139,421,441]" gridcol="13" gridrow="4" pageId="4">1</td>
<td id="769D5DBA0C7F469A1EF60D84C9BD278D" box="[1161,1178,421,441]" gridcol="14" gridrow="4" pageId="4">2</td>
<td id="769D5DBA0C7F469A1ECE0D84C9E2278D" box="[1201,1221,421,441]" gridcol="15" gridrow="4" pageId="4">2</td>
<td id="769D5DBA0C7F469A1EA40D84C9D1278D" box="[1243,1270,421,441]" gridcol="16" gridrow="4" pageId="4">2</td>
<td id="769D5DBA0C7F469A1F740D84C862278D" box="[1291,1349,421,441]" gridcol="17" gridrow="4" pageId="4">20 + 2</td>
<td id="769D5DBA0C7F469A1F250D84C85D278D" box="[1370,1402,421,441]" gridcol="18" gridrow="4" pageId="4">650</td>
<td id="769D5DBA0C7F469A1FEF0D84C8F2278D" box="[1424,1493,421,441]" gridcol="19" gridrow="4" pageId="4">140 + n</td>
</tr>
<tr id="354C34C60C7F469A1A050D9FC8F227E5" box="[122,1493,446,465]" gridrow="5" pageId="4">
<th id="769D5DBA0C7F469A1A050D9FCC6827E5" box="[122,335,446,465]" gridcol="0" gridrow="5" pageId="4">ZUFMS-REP 2291</th>
<td id="769D5DBA0C7F469A1B1B0D9FCC5C27E5" box="[356,379,446,465]" gridcol="1" gridrow="5" pageId="4">F</td>
<td id="769D5DBA0C7F469A1BEE0D9FCCC127E5" box="[401,486,446,465]" gridcol="2" gridrow="5" pageId="4">17–17–15</td>
<td id="769D5DBA0C7F469A1B840D9FCF2927E5" box="[507,526,446,465]" gridcol="3" gridrow="5" pageId="4">1</td>
<td id="769D5DBA0C7F469A185A0D9FCF6227E5" box="[549,581,446,465]" gridcol="4" gridrow="5" pageId="4">111</td>
<td id="769D5DBA0C7F469A18250D9FCFBC27E5" box="[602,667,446,465]" gridcol="5" gridrow="5" pageId="4">divided</td>
<td id="769D5DBA0C7F469A18CE0D9FCFCC27E5" box="[689,747,446,465]" gridcol="6" gridrow="5" pageId="4">52</td>
<td id="769D5DBA0C7F469A197D0D9FCE3227E5" box="[770,789,446,465]" gridcol="7" gridrow="5" pageId="4">8</td>
<td id="769D5DBA0C7F469A19570D9FCE7E27E5" box="[808,857,446,465]" gridcol="8" gridrow="5" pageId="4">iii–iv</td>
<td id="769D5DBA0C7F469A19100D9FCEB227E5" box="[879,917,446,465]" gridcol="9" gridrow="5" pageId="4">9/10</td>
<td id="769D5DBA0C7F469A19D40D9FCEE327E5" box="[939,964,446,465]" gridcol="10" gridrow="5" pageId="4">i–v</td>
<td id="769D5DBA0C7F469A19950D9FC93027E5" box="[1002,1047,446,465]" gridcol="11" gridrow="5" pageId="4">v–vi</td>
<td id="769D5DBA0C7F469A1E520D9FC96A27E5" box="[1069,1101,446,465]" gridcol="12" gridrow="5" pageId="4">E</td>
<td id="769D5DBA0C7F469A1E1D0D9FC95427E5" box="[1122,1139,446,465]" gridcol="13" gridrow="5" pageId="4">1</td>
<td id="769D5DBA0C7F469A1EF60D9FC9BD27E5" box="[1161,1178,446,465]" gridcol="14" gridrow="5" pageId="4">2</td>
<td id="769D5DBA0C7F469A1ECE0D9FC9E227E5" box="[1201,1221,446,465]" gridcol="15" gridrow="5" pageId="4">2</td>
<td id="769D5DBA0C7F469A1EA40D9FC9D127E5" box="[1243,1270,446,465]" gridcol="16" gridrow="5" pageId="4">2</td>
<td id="769D5DBA0C7F469A1F740D9FC86227E5" box="[1291,1349,446,465]" gridcol="17" gridrow="5" pageId="4">19 + 2</td>
<td id="769D5DBA0C7F469A1F250D9FC85D27E5" box="[1370,1402,446,465]" gridcol="18" gridrow="5" pageId="4">153</td>
<td id="769D5DBA0C7F469A1FEF0D9FC8F227E5" box="[1424,1493,446,465]" gridcol="19" gridrow="5" pageId="4">45</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC336840C7FB9611AEA0E56CE2C25A2" blockId="4.[122,792,513,2001]" pageId="4">
An undivided nasal distinguishes 
<taxonomicName id="4C7C4D070C7FB96118760E59CF5224BB" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[521,629,632,655]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB96118760E59CF5224BB" box="[521,629,632,655]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118FD0E59CFFE24BB" box="[642,729,632,655]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
from all other 
<taxonomicName id="4C7C4D070C7FB9611A990EB5CC672498" authorityName="Wagler" authorityYear="1828" box="[230,320,660,684]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="genus">
<emphasis id="B908EA960C7FB9611A990EB5CC672498" box="[230,320,660,684]" italics="true" pageId="4">Helicops</emphasis>
</taxonomicName>
with the exception of 
<taxonomicName id="4C7C4D070C7FB96118300EB4CF8F2498" box="[591,680,661,684]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C7FB96118300EB4CF8F2498" box="[591,680,661,684]" italics="true" pageId="4">H. ŋacu</emphasis>
</taxonomicName>
. A single preocular distinguishes 
<taxonomicName id="4C7C4D070C7FB9611B050E93CCC224FD" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[378,485,690,713]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611B050E93CCC224FD" box="[378,485,690,713]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB9611B8F0E93CF6224FD" box="[496,581,690,713]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="4C7C4D070C7FB96118F20E93CE3024FD" box="[653,791,690,713]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C7FB96118F20E93CE3024FD" box="[653,791,690,713]" italics="true" pageId="4">H. triƲittatus</emphasis>
</taxonomicName>
(two preoculars), and the presence of two anterior temporals distinguishes 
<taxonomicName id="4C7C4D070C7FB9611B730ECCCC5E2530" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[268,377,749,772]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611B730ECCCC5E2530" box="[268,377,749,772]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB9611BFA0ECDCCFB2537" box="[389,476,748,771]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
from 
<emphasis id="B908EA960C7FB96118590ECCCCEB2515" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB96118590ECCCFB12530" box="[550,662,749,772]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="gomesi">H. gomesi</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118D90ECCCE332530" box="[678,788,749,772]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="petersi">H. petersi</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611A050F2BCC382515" authorityName="Frota" authorityYear="2005" box="[122,287,778,801]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="tapajonicus">H. tapajonicus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611B4B0F2BCCEF2515" box="[308,456,778,801]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">H. triƲittatus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB96118690F2BCF542515" box="[534,627,778,801]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C7FB96118690F2BCF542515" box="[534,627,778,801]" italics="true" pageId="4">H. ŋacu</emphasis>
</taxonomicName>
(one anterior temporal). The presence of ±18 (up to 21) prediastemal maxillary teeth distinguishes 
<taxonomicName id="4C7C4D070C7FB9611BE00F65CF24256F" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[415,515,836,859]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611BE00F65CF24256F" box="[415,515,836,859]" italics="true" pageId="4">H.nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118780F65CF72256F" box="[519,597,836,859]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
from 
<emphasis id="B908EA960C7FB96118F10F65CDC925A2" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB96118F10F65CE332568" box="[654,788,836,860]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="angulatus">H. angulatus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611A050F40CDC2254D" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[122,229,865,889]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="apiaka">H. apiaka</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611A8D0F40CCA7254D" box="[242,384,865,889]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="hagmanni">H. hagmanni</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611BF10F40CF2B254C" box="[398,524,865,888]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="pastazae">H. pastazae</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118650F40CFA2254C" box="[538,645,865,888]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="petersi">H. petersi</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118ED0F40CE33254D" box="[658,788,865,889]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="polnlepis">H. polŋlepis</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611A050F5ECDCC25A2" box="[122,235,894,918]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="scalaris">H. scalaris</taxonomicName>
,
</emphasis>
and 
<emphasis id="B908EA960C7FB9611B5D0F5ECC5C25A2" box="[290,379,895,918]" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB9611B5D0F5ECC5125A2" box="[290,374,895,918]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nacu">H. ŋacu</taxonomicName>
,
</emphasis>
all of which have fewer than 18 teeth.
</paragraph>
<paragraph id="8BC336840C7FB9611AEA0FBDCFA623B3" blockId="4.[122,792,513,2001]" pageId="4">
The uniformly colored dorsum of 
<taxonomicName id="4C7C4D070C7FB96118750FBCCF522580" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[522,629,925,948]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB96118750FBCCF522580" box="[522,629,925,948]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118040FBDCFF72587" box="[635,720,924,947]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
is also present in some specimens of 
<emphasis id="B908EA960C7FB9611BB50F9BCE3F25E5" box="[458,792,953,977]" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB9611BB50F9BCF5A25E5" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[458,637,953,977]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="carinicaudus">H. carinicaudus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118F60F9BCE3325E5" authorityName="Gunther" authorityYear="1861" box="[649,788,953,977]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="modestus">H. modestus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB9611AD30FF6CC6925DA" authorityName="Frota" authorityYear="2005" box="[172,334,983,1006]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="B908EA960C7FB9611AD30FF6CC6925DA" box="[172,334,983,1006]" italics="true" pageId="4">H. tapajonicus</emphasis>
</taxonomicName>
. A cream venter with two rows of black semilunar markings is present only in 
<taxonomicName id="4C7C4D070C7FB961185B0FD5CFB6223F" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[548,657,1012,1035]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB961185B0FD5CFB6223F" box="[548,657,1012,1035]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118E60FD5CFD6223F" box="[665,753,1012,1035]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
, 
<emphasis id="B908EA960C7FB96118820FD5CCA0221D" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB96118820FD5CC2D221D" baseAuthorityName="Wied" baseAuthorityYear="1824" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="carinicaudus">H. carinicaudus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611B6B0830CCA5221D" box="[276,386,1041,1065]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="danieli">H. danieli</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB9611BC40830CF6E221C" box="[443,585,1041,1064]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C7FB9611BC40830CF6E221C" box="[443,585,1041,1064]" italics="true" pageId="4">H. triƲittatus</emphasis>
</taxonomicName>
. In the latter three species the black ventral markings are strong and welldefined, whereas in 
<taxonomicName id="4C7C4D070C7FB9611B1F086DCCE92257" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[352,462,1100,1123]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611B1F086DCCE92257" box="[352,462,1100,1123]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB9611BA9086DCF092257" box="[470,558,1100,1123]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
they are usually less well-defined. In preserved specimens of 
<taxonomicName id="4C7C4D070C7FB961183A0848CF9522B4" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[581,690,1129,1152]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB961183A0848CF9522B4" box="[581,690,1129,1152]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118C40848CE3322B4" box="[699,788,1129,1152]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
, the first dorsal scale row (DSR1) is cream colored, as is the adjacent venter, on the anterior third of the body; posteriorly, the scales of DSR1 become gray dorsally, and occasionally totally gray. In 
<taxonomicName id="4C7C4D070C7FB9611B4908FFCF1122C1" authority="DSR" authorityName="DSR" box="[310,566,1245,1269]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="carinicaudus">
<emphasis id="B908EA960C7FB9611B4908FFCCCA22C1" box="[310,493,1245,1269]" italics="true" pageId="4">H. carinicaudus</emphasis>
DSR
</taxonomicName>
1 is cream colored, whereas DSR2 is gray anteriodorsally, usually becoming totally gray posteriorly; in 
<taxonomicName id="4C7C4D070C7FB9611BD60938CF472304" authority="DSR" authorityName="DSR" box="[425,608,1304,1328]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="danieli">
<emphasis id="B908EA960C7FB9611BD60938CF3D2304" box="[425,538,1304,1328]" italics="true" pageId="4">H. danieli</emphasis>
DSR
</taxonomicName>
1 and DSR2 are white or cream colored, similar to the venter (
<bibRefCitation id="EFED4B750C7FB96118CC0914CD98235E" author="Rossman, D. A." pageId="4" pagination="589 - 594" refId="ref10975" refString="Rossman, D. A. 2002 a. Morphological variation in the endemic Colombian water snake, Helicops danieli Amaral, 1937 (Serpentes: Xenodontidae). Revista da Academia Colombiana Ciencias 26: 589 - 594." type="journal article" year="2002">Rossman 2002a</bibRefCitation>
); in 
<taxonomicName id="4C7C4D070C7FB9611A930972CC5E235E" box="[236,377,1363,1386]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C7FB9611A930972CC5E235E" box="[236,377,1363,1386]" italics="true" pageId="4">H. triƲittatus</emphasis>
</taxonomicName>
the scales of DSR1 are cream colored dorsally along the entire body (
<bibRefCitation id="EFED4B750C7FB9611BAF094ECF5223B3" author="Rossman, D. A." box="[464,629,1391,1415]" pageId="4" pagination="271 - 278" refId="ref11061" refString="Rossman, D. A. 2010. Morphological variation in the striped water snake Helicops triVittatus (Gray, 1849) (Reptilia: Serpentes: Xenodontidae) of eastern Amazonia. Boletim do Museu Paraense Emilio Goeldi, Ciencias Naturais 5: 271 - 278." type="journal article" year="2010">Rossman 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BC336840C7FB9611AEA09AFCF9020BB" blockId="4.[122,792,513,2001]" pageId="4">
In 
<taxonomicName id="4C7C4D070C7FB9611AC709AFCC4C2391" box="[184,363,1421,1445]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="carinicaudus">
<emphasis id="B908EA960C7FB9611AC709AFCC4C2391" box="[184,363,1421,1445]" italics="true" pageId="4">H. carinicaudus</emphasis>
</taxonomicName>
there is no gray pigmentation on the outer edges of the ventrals, and sometimes there is a midventral row of poorly defined black semilunar markings (
<bibRefCitation id="EFED4B750C7FB9611AFE09C4CC5723C9" author="Yuki, R. N. &amp; T. Lema" box="[129,368,1509,1533]" pageId="4" pagination="85 - 128" refId="ref11677" refString="Yuki, R. N., and T. Lema. 2005. Analise comparativa entre as cobras d ̕ agua meridionais (Helicops carinicaudus (Wied, 1825) e Helicops infrataeniatus Jan, 1865), com a descricao do cranio e hemipenis (Serpentes: Colubridae: Xenodontinae). Comunicacoes do Museu de Ciencia e Tecnologa da PUCRS, Serie Zoologia 18: 85 - 128. [In Portuguese.]" type="journal article" year="2005">Yuki and Lema 2005</bibRefCitation>
; present study); in 
<taxonomicName id="4C7C4D070C7FB961183409C7CF9D23C9" box="[587,698,1509,1533]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="danieli">
<emphasis id="B908EA960C7FB961183409C7CF9D23C9" box="[587,698,1509,1533]" italics="true" pageId="4">H. danieli</emphasis>
</taxonomicName>
there is no gray pigmentation on the ventrals, and the black semilunar markings sometimes fuse midventrally (
<bibRefCitation id="EFED4B750C7FB96118CC0A3ECD982061" author="Rossman, D. A." pageId="4" pagination="589 - 594" refId="ref10975" refString="Rossman, D. A. 2002 a. Morphological variation in the endemic Colombian water snake, Helicops danieli Amaral, 1937 (Serpentes: Xenodontidae). Revista da Academia Colombiana Ciencias 26: 589 - 594." type="journal article" year="2002">Rossman 2002a</bibRefCitation>
); in 
<taxonomicName id="4C7C4D070C7FB9611A8C0A1CCCA22060" box="[243,389,1597,1620]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C7FB9611A8C0A1CCCA22060" box="[243,389,1597,1620]" italics="true" pageId="4">H. triƲittatus</emphasis>
</taxonomicName>
the black markings are proportionally larger than those of 
<taxonomicName id="4C7C4D070C7FB9611BF00A7ACCDA2046" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[399,509,1627,1650]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611BF00A7ACCDA2046" box="[399,509,1627,1650]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118790A7BCF472045" box="[518,608,1626,1649]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
, and there is no gray pigmentation on the ventrals (
<bibRefCitation id="EFED4B750C7FB961187B0A56CF8B20BB" author="Rossman, D. A." box="[516,684,1655,1679]" pageId="4" pagination="271 - 278" refId="ref11061" refString="Rossman, D. A. 2010. Morphological variation in the striped water snake Helicops triVittatus (Gray, 1849) (Reptilia: Serpentes: Xenodontidae) of eastern Amazonia. Boletim do Museu Paraense Emilio Goeldi, Ciencias Naturais 5: 271 - 278." type="journal article" year="2010">Rossman 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BC336840C7FB9611AEA0AB4C89724FD" blockId="4.[122,792,513,2001]" lastBlockId="4.[824,1494,514,2001]" pageId="4">
Among the species of 
<taxonomicName id="4C7C4D070C7FB9611BDC0AB4CF252099" authorityName="Wagler" authorityYear="1828" box="[419,514,1685,1709]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="genus">
<emphasis id="B908EA960C7FB9611BDC0AB4CF252099" box="[419,514,1685,1709]" italics="true" pageId="4">Helicops</emphasis>
</taxonomicName>
whose hemipenis morphology is described (all but 
<taxonomicName id="4C7C4D070C7FB9611BB00A92CF1820FE" box="[463,575,1715,1738]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B908EA960C7FB9611BB00A92CF1820FE" box="[463,575,1715,1738]" italics="true" pageId="4">H. petersi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7C4D070C7FB96118000A92CFFE20FE" box="[639,729,1715,1738]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C7FB96118000A92CFFE20FE" box="[639,729,1715,1738]" italics="true" pageId="4">H. ŋacu</emphasis>
</taxonomicName>
), the organ of 
<taxonomicName id="4C7C4D070C7FB9611A9B0AF1CC7420D3" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[228,339,1744,1767]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611A9B0AF1CC7420D3" box="[228,339,1744,1767]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB9611B210AF1CC9020D3" box="[350,439,1744,1767]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
is readily distinguishable from that of 
<emphasis id="B908EA960C7FB9611AB00ACCCFC22130" box="[207,741,1772,1797]" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB9611AB00ACCCCA82131" box="[207,399,1773,1797]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="infrataeniatus">H. infrataeniatus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611BE20ACCCF622130" box="[413,581,1772,1796]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="leopardinus">H. leopardinus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB961182B0ACCCFC72130" authorityName="Gunther" authorityYear="1861" box="[596,736,1772,1796]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="modestus">H. modestus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB9611A050B2BCC3B2115" authorityName="Frota" authorityYear="2005" box="[122,284,1802,1825]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="B908EA960C7FB9611A050B2BCC3B2115" box="[122,284,1802,1825]" italics="true" pageId="4">H. tapajonicus</emphasis>
</taxonomicName>
in having well-developed lobes. The asulcate side having lateral spines larger than central spines distinguishes the hemipenis of 
<taxonomicName id="4C7C4D070C7FB9611B8C0B65CF8C2168" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[499,683,1860,1884]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611B8C0B65CF8C2168" box="[499,683,1860,1884]" italics="true" pageId="4">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118C40B64CE3F2168" box="[699,792,1861,1884]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
from that of 
<emphasis id="B908EA960C7FB9611B790B43CC3721A3" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB9611B790B43CCBF214D" box="[262,408,1889,1913]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="hagmanni">H. hagmanni</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611BDB0B43CF45214D" box="[420,610,1889,1913]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="infrataeniatus">H. infrataeniatus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB96118120B43CE34214D" box="[621,787,1889,1913]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="leopardinus">H. leopardinus</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7FB9611A050B5ECC2B21A3" authorityName="Gunther" authorityYear="1861" box="[122,268,1919,1943]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="modestus">H. modestus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB9611B2B0B5ECCCD21A2" box="[340,490,1919,1942]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C7FB9611B2B0B5ECCCD21A2" box="[340,490,1919,1942]" italics="true" pageId="4">H. triƲittatus</emphasis>
</taxonomicName>
(small spines of uniform size). In 
<emphasis id="B908EA960C7FB9611AA20BBCCC432180" box="[221,356,1949,1972]" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB9611AA20BBCCC472180" box="[221,352,1949,1972]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="pastazae">H. pastazae</taxonomicName>
,
</emphasis>
the central spines on the asulcate side are considerably longer than in 
<taxonomicName id="4C7C4D070C7FB9611B8E0B9BCF4521E5" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[497,610,1978,2001]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611B8E0B9BCF4521E5" box="[497,610,1978,2001]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96118110B98CFEE21E4" box="[622,713,1977,2000]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
Lobes covered with spinulate flounces distinguish the organ of 
<taxonomicName id="4C7C4D070C7FB96119470E01CE832403" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[824,932,544,567]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB96119470E01CE832403" box="[824,932,544,567]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB96119D40E01C9272403" box="[939,1024,544,567]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
from that of 
<taxonomicName id="4C7C4D070C7FB9611EED0E01C9DA2403" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[1170,1277,543,567]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="apiaka">
<emphasis id="B908EA960C7FB9611EED0E01C9DA2403" box="[1170,1277,543,567]" italics="true" pageId="4">H. apiaka</emphasis>
</taxonomicName>
(lobes with plicae), 
<emphasis id="B908EA960C7FB96119470E1CCE942460" box="[824,947,572,596]" italics="true" pageId="4">
<taxonomicName id="4C7C4D070C7FB96119470E1CCE882460" box="[824,943,572,596]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="danieli">H. danieli</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7FB96119840E1CC9A02460" box="[1019,1159,572,596]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="polnlepis">
<emphasis id="B908EA960C7FB96119840E1CC9A02460" box="[1019,1159,572,596]" italics="true" pageId="4">H. polŋlepis</emphasis>
</taxonomicName>
(lobes with small papillae). The spines bordering the sulcus spermaticus in the hemipenis of 
<taxonomicName id="4C7C4D070C7FB96119A90E56C96124BA" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[982,1094,631,654]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB96119A90E56C96124BA" box="[982,1094,631,654]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB9611E2E0E56C98D24BA" box="[1105,1194,631,654]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
are smaller than those of 
<taxonomicName id="4C7C4D070C7FB96119470EB4CE832498" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[824,932,660,684]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="apiaka">
<emphasis id="B908EA960C7FB96119470EB4CE832498" box="[824,932,660,684]" italics="true" pageId="4">H. apiaka</emphasis>
</taxonomicName>
. The less developed basal pocket in 
<taxonomicName id="4C7C4D070C7FB9611F3E0EB4C8892498" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1345,1454,661,684]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7FB9611F3E0EB4C8892498" box="[1345,1454,661,684]" italics="true" pageId="4">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7FB9611FC90EB5CE4124FD" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
distinguishes its hemipenis from that of 
<taxonomicName id="4C7C4D070C7FB9611F4C0E93C88E24FD" box="[1331,1449,689,713]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="scalaris">
<emphasis id="B908EA960C7FB9611F4C0E93C88E24FD" box="[1331,1449,689,713]" italics="true" pageId="4">H. scalaris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC336840C7FB963192D0EEECFE924D0" blockId="4.[824,1494,514,2001]" lastBlockId="6.[122,792,190,2001]" lastPageId="6" pageId="4">
<emphasis id="B908EA960C7FB961192D0EEEC9A724DC" bold="true" box="[850,1152,719,744]" pageId="4">
Description of 
<typeStatus id="54C788260C7FB9611E700EF1C95C24DC" box="[1039,1147,720,744]" pageId="4" type="holotype">holotype</typeStatus>
.
</emphasis>
—Adult female, SVL 
<quantity id="4C849B610C7FB9611F0C0EF1C8F624D3" box="[1395,1489,720,744]" metricMagnitude="-1" metricUnit="m" metricValue="3.34" pageId="4" unit="mm" value="334.0">334 mm</quantity>
, tail length 
<quantity id="4C849B610C7FB96119CF0ECCCED92530" box="[944,1022,749,772]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="4" unit="mm" value="77.0">77 mm</quantity>
(23% SVL); head rounded in dorsal view, flattened in lateral view, head length 
<quantity id="4C849B610C7FB9611EA90F2BC8622515" box="[1238,1349,778,802]" metricMagnitude="-2" metricUnit="m" metricValue="1.9129999999999998" pageId="4" unit="mm" value="19.13">19.13 mm</quantity>
(5.8% SVL), head width 
<quantity id="4C849B610C7FB96119CA0F06C907250B" box="[949,1056,807,831]" metricMagnitude="-2" metricUnit="m" metricValue="1.192" pageId="4" unit="mm" value="11.92">11.92 mm</quantity>
(61.8% head length); no evident cervical constriction; snout rounded in dorsal and lateral views; rostrum–orbit distance 
<quantity id="4C849B610C7FB9611E410F43C9B3254D" box="[1086,1172,866,890]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="4" unit="mm" value="4.1">4.1 mm</quantity>
(21.2% head length); nasal– orbit distance 
<quantity id="4C849B610C7FB961199D0F5EC96D25A2" box="[994,1098,895,919]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="4" unit="mm" value="2.37">2.37 mm</quantity>
(12.3% head length); interorbital distance 
<quantity id="4C849B610C7FB96119E80FBDCED12580" box="[919,1014,924,948]" metricMagnitude="-3" metricUnit="m" metricValue="4.64" pageId="4" unit="mm" value="4.64">4.64 mm</quantity>
(38.9% head width); eye diameter 
<quantity id="4C849B610C7FB9611F100FBDC8F62580" box="[1391,1489,924,948]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="4" unit="mm" value="2.65">2.65 mm</quantity>
, pupil round; rostral subtriangular in frontal view, wider (
<quantity id="4C849B610C7FB9611FDA0F9BCE4225DB" metricMagnitude="-3" metricUnit="m" metricValue="3.22" pageId="4" unit="mm" value="3.22">3.22 mm</quantity>
) than high (
<quantity id="4C849B610C7FB96119830FF6C94F25DB" box="[1020,1128,983,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="4" unit="mm" value="1.78">1.78 mm</quantity>
), visible from above; internasal single, diamond-shaped, longer (
<quantity id="4C849B610C7FB9611ED00FD5C82B2238" box="[1199,1292,1012,1036]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" unit="mm" value="2.2">2.2 mm</quantity>
) than wide (
<quantity id="4C849B610C7FB9611FDA0FD5CE40221D" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="4" unit="mm" value="1.98">1.98 mm</quantity>
); prefrontals two, hexagonal, longer (2.34: 
<quantity id="4C849B610C7FB9611F4D0830C8B1221D" box="[1330,1430,1041,1065]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="4" unit="mm" value="2.35">2.35 mm</quantity>
) than wide (1.46: 
<quantity id="4C849B610C7FB96119D1080FC9332272" box="[942,1044,1070,1094]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="4" unit="mm" value="1.49">1.49 mm</quantity>
), in medial contact (prefontal suture 
<quantity id="4C849B610C7FB9611FD9080FCE402250" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="4" unit="mm" value="0.81">0.81 mm</quantity>
); frontal pentagonal, longer (
<quantity id="4C849B610C7FB9611ED1086DC8322250" box="[1198,1301,1100,1124]" metricMagnitude="-3" metricUnit="m" metricValue="4.84" pageId="4" unit="mm" value="4.84">4.84 mm</quantity>
) than wide (
<quantity id="4C849B610C7FB9611FDA086DCE4022B5" metricMagnitude="-3" metricUnit="m" metricValue="2.56" pageId="4" unit="mm" value="2.56">2.56 mm</quantity>
); supraoculars rectangular, longer (3.89: 
<quantity id="4C849B610C7FB9611F540848C8B422B5" box="[1323,1427,1129,1153]" metricMagnitude="-3" metricUnit="m" metricValue="3.98" pageId="4" unit="mm" value="3.98">3.98 mm</quantity>
) than wide (1.61: 
<quantity id="4C849B610C7FB96119CD08A7C93A22AA" box="[946,1053,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="4" unit="mm" value="1.63">1.63 mm</quantity>
); parietals longer (6.18: 
<quantity id="4C849B610C7FB9611F5708A7C8A922AA" box="[1320,1422,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="6.23" pageId="4" unit="mm" value="6.23">6.23 mm</quantity>
maximum length) than wide (3.47: 
<quantity id="4C849B610C7FB9611EE90882C9D8228F" box="[1174,1279,1187,1211]" metricMagnitude="-3" metricUnit="m" metricValue="3.52" pageId="4" unit="mm" value="3.52">3.52 mm</quantity>
maximum width); parietal suture 
<quantity id="4C849B610C7FB961199B08E0C96D22EC" box="[996,1098,1217,1241]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" pageId="4" unit="mm" value="3.95">3.95 mm</quantity>
; nasals quadrangular, separated by prefrontal, well visible from above; left nasal semidivided, right nasal entire; loreal quadrangular, contacting nasal anteriorly, prefrontal above, second and third supralabials below, and preocular posteriorly; preocular single, higher (1.90: 
<quantity id="4C849B610C7FB961190B0972CEFD235F" box="[884,986,1363,1387]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" pageId="4" unit="mm" value="1.97">1.97 mm</quantity>
) than wide (0.65: 
<quantity id="4C849B610C7FB9611EE20972C820235F" box="[1181,1287,1363,1387]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" unit="mm" value="0.6">0.60 mm</quantity>
), contacting loreal and prefrontal anteriorly, supraocular above, third supralabial below, and the eye posteriorly; postoculars two; upper postocular (1.33: 
<quantity id="4C849B610C7FB9611991098BC97623F6" box="[1006,1105,1450,1474]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="4" unit="mm" value="1.35">1.35 mm</quantity>
high; 1.48: 
<quantity id="4C849B610C7FB9611EB4098BC80823F6" box="[1227,1327,1450,1474]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" unit="mm" value="1.5">1.50 mm</quantity>
wide) contacting the eye anteriorly, supraocular and parietal above, lower postocular below, and upper anterior temporal posteriorly; lower postocular (2.17: 
<quantity id="4C849B610C7FB9611E440A22C985202E" box="[1083,1186,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="4" unit="mm" value="2.17">2.17 mm</quantity>
high; 1.14: 
<quantity id="4C849B610C7FB9611F5A0A22C8AB202E" box="[1317,1420,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="4" unit="mm" value="1.14">1.14 mm</quantity>
wide) contacting the eye anteriorly, upper postocular above, fourth to sixth supralabials below, and upper anterior temporal posteriorly; temporals 2 + 2, longer than wide; upper anterior temporal (1.97: 
<quantity id="4C849B610C7FB96119A40A56C91B20BB" box="[987,1084,1655,1679]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" pageId="4" unit="mm" value="1.96">1.96 mm</quantity>
high; 3.56: 
<quantity id="4C849B610C7FB9611ECE0A56C83620BB" box="[1201,1297,1655,1679]" metricMagnitude="-3" metricUnit="m" metricValue="3.58" pageId="4" unit="mm" value="3.58">3.58 mm</quantity>
long) contacts the postoculars anteriorly, parietal above, sixth supralabial below, lower anterior temporal and upper posterior temporal posteriorly; lower anterior temporal (2.06: 
<quantity id="4C849B610C7FB9611F590AEEC8A820D3" box="[1318,1423,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="4" unit="mm" value="1.65">1.65 mm</quantity>
high; 3.67: 
<quantity id="4C849B610C7FB96119140ACDCEE82130" box="[875,975,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="4" unit="mm" value="3.15">3.15 mm</quantity>
long) contacts upper anterior temporal above and anteriorly, sixth and seventh supralabials below, upper posterior temporal above, and lower posterior temporal posteriorly; upper posterior temporal (2.56: 
<quantity id="4C849B610C7FB9611F560B65C8B6216F" box="[1321,1425,1860,1884]" metricMagnitude="-3" metricUnit="m" metricValue="2.59" pageId="4" unit="mm" value="2.59">2.59 mm</quantity>
high; 2.92: 
<quantity id="4C849B610C7FB96119140B40CEFE214D" box="[875,985,1889,1913]" metricMagnitude="-3" metricUnit="m" metricValue="2.92" pageId="4" unit="mm" value="2.92">2.92 mm</quantity>
long) contacting both anterior temporals anteriorly, parietal above, lower anterior and posterior temporals below, and two small head scales posteriorly; lower posterior temporal (
<quantity id="4C849B610C7FB9611E540B98C9AC21E5" box="[1067,1163,1977,2001]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" unit="mm" value="2.2">2.2 mm</quantity>
high; 2.45: 
<quantity id="4C849B610C7FB9611F640B98C8AD21E5" box="[1307,1418,1977,2001]" metricMagnitude="-3" metricUnit="m" metricValue="2.52" pageId="4" unit="mm" value="2.52">2.52 mm</quantity>
long) contacting lower anterior temporal anteriorly, upper posterior temporal above, seventh and eighth supralabials below, and two small head scales posteriorly; supralabials eight, the third and fourth contacting the eye; sixth supralabial higher (3.05: 
<quantity id="4C849B610C7DB9631ACA0D13CC39277F" box="[181,286,306,331]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" unit="mm" value="3.2">3.20 mm</quantity>
) and wider (3.25: 
<quantity id="4C849B610C7DB9631B970D13CF76277F" box="[488,593,306,331]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="6" unit="mm" value="3.26">3.26 mm</quantity>
) than the others; symphyseal triangular, wider (
<quantity id="4C849B610C7DB9631BBC0D71CF02275C" box="[451,549,336,360]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="6" unit="mm" value="1.71">1.71 mm</quantity>
) than long (
<quantity id="4C849B610C7DB96318D80D71CE2B275C" box="[679,780,336,360]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="6" unit="mm" value="1.38">1.38 mm</quantity>
); infralabials 10, the first pair in contact behind symphyseal, preventing symphyseal–chinshield contact; first to fifth pair of infralabials contacting first pair of chinshields (genials); fifth and sixth pair of infralabials in contact with second pair of chinshields; intergenial scale absent; chinshields longer (first pair 4.62: 
<quantity id="4C849B610C7DB9631AA10DDECC642423" box="[222,323,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" unit="mm" value="4.5">4.50 mm</quantity>
; second pair 5.75: 
<quantity id="4C849B610C7DB963187D0DDECF412423" box="[514,614,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="6" unit="mm" value="4.7">4.70 mm</quantity>
) than wide (first pair 1.60: 
<quantity id="4C849B610C7DB9631AA10E3DCC652400" box="[222,322,540,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="6" unit="mm" value="1.62">1.62 mm</quantity>
; second pair 1.70: 
<quantity id="4C849B610C7DB963187D0E3CCF422401" box="[514,613,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="6" unit="mm" value="1.71">1.71 mm</quantity>
); dorsal scales in 17–17–15 rows, lacking apical pits, smooth behind the head, becoming weakly keeled posteriorly; keels are more visible on posterior-most region of body and on dorsal surface of tail; gular scales 4:5, between the last infralabial and first preventral; preventrals two, ventral scales 117; cloacal plate divided; 41 divided subcaudals; maxillary teeth 19 + 2.
</paragraph>
<caption id="DF03660C0C7EB9601A090DBBC97B27C2" ID-Table-UUID="DF03660C0C7EB9601A090DBBC97B27C2" httpUri="http://table.plazi.org/id/DF03660C0C7EB9601A090DBBC97B27C2" pageId="5" startId="5.[118,131,410,430]" targetBox="[96,1915,532,971]" targetIsTable="true" targetPageId="5">
<paragraph id="8BC336840C7EB9601A090DBBC97B27C2" blockId="5.[97,1914,410,502]" pageId="5">
TABLE 3.—Summary of morphological characters of 
<taxonomicName id="4C7C4D070C7EB960183E0DBBCFFF279A" box="[577,728,410,431]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B908EA960C7EB960183E0DBBCFAB279B" box="[577,652,410,431]" italics="true" pageId="5">Helicops</emphasis>
species.
</taxonomicName>
AD = anterior dorsal scale rows, MD = midbody dorsal scale rows, PD = posterior dorsal scale rows, DK = dorsal keels on posterior third of body (medium = M, strong = S, or weak = W), V = ventral plates, SC = subcaudal plates, SCK = subcaudal keels (present = Yes or absent = No), SL = supralabials, SL + E = supralabials contacting the eye, IL = infralabials, IL + G1 = infralabials contacting the first pair of genials, IG = intergenials (present = Yes or absent = No), NA = nasal (entire = E or semidivided = S), Pe = preoculars, Po = postoculars, AT = anterior temporals, PT = posterior temporals, MXT = maxillary teeth.
</paragraph>
</caption>
<paragraph id="8BC336840C7EB9601A1F0E35CA5C25FE" pageId="5">
<table id="F97CC4240C7E469A1A1F0E35CA5C25FF" box="[96,1915,532,971]" gridcols="19" gridrows="18" pageId="5">
<tr id="354C34C60C7E469A1A1F0E35CA5C2417" box="[96,1915,532,547]" gridrow="0" pageId="5">
<th id="769D5DBA0C7E469A1A1F0E35CC222417" box="[96,261,532,547]" gridcol="0" gridrow="0" pageId="5">Species</th>
<th id="769D5DBA0C7E469A1B5F0E35CC712417" box="[288,342,532,547]" gridcol="1" gridrow="0" pageId="5">AD</th>
<th id="769D5DBA0C7E469A1B0E0E35CC802417" box="[369,423,532,547]" gridcol="2" gridrow="0" pageId="5">MD</th>
<th id="769D5DBA0C7E469A1BBC0E35CCDF2417" box="[451,504,532,547]" gridcol="3" gridrow="0" pageId="5">PD</th>
<th id="769D5DBA0C7E469A186B0E35CF612417" box="[532,582,532,547]" gridcol="4" gridrow="0" pageId="5">DK</th>
<th id="769D5DBA0C7E469A181D0E35CFDA2417" box="[610,765,532,547]" gridcol="5" gridrow="0" pageId="5">V (male/female)</th>
<th id="769D5DBA0C7E469A19660E35CEB32417" box="[793,916,532,547]" gridcol="6" gridrow="0" pageId="5">SC (male/female)</th>
<th id="769D5DBA0C7E469A19CF0E35CEE82417" box="[944,975,532,547]" gridcol="7" gridrow="0" pageId="5">SCK</th>
<th id="769D5DBA0C7E469A19890E35C9072417" box="[1014,1056,532,547]" gridcol="8" gridrow="0" pageId="5">SL</th>
<th id="769D5DBA0C7E469A1E430E35C9942417" box="[1084,1203,532,547]" gridcol="9" gridrow="0" pageId="5">SL + E</th>
<th id="769D5DBA0C7E469A1EAE0E35C8202417" box="[1233,1287,532,547]" gridcol="10" gridrow="0" pageId="5">IL</th>
<th id="769D5DBA0C7E469A1F5C0E35C8B42417" box="[1315,1427,532,547]" gridcol="11" gridrow="0" pageId="5">IL + G1</th>
<th id="769D5DBA0C7E469A1FCF0E35C8E92417" box="[1456,1486,532,547]" gridcol="12" gridrow="0" pageId="5">IG</th>
<th id="769D5DBA0C7E469A1F950E35CB322417" box="[1514,1557,532,547]" gridcol="13" gridrow="0" pageId="5">NA</th>
<th id="769D5DBA0C7E469A1C4E0E35CB762417" box="[1585,1617,532,547]" gridcol="14" gridrow="0" pageId="5">Pe</th>
<th id="769D5DBA0C7E469A1C120E35CBAA2417" box="[1645,1677,532,547]" gridcol="15" gridrow="0" pageId="5">Po</th>
<th id="769D5DBA0C7E469A1CD60E35CBEE2417" box="[1705,1737,532,547]" gridcol="16" gridrow="0" pageId="5">AT</th>
<th id="769D5DBA0C7E469A1C9A0E35CA222417" box="[1765,1797,532,547]" gridcol="17" gridrow="0" pageId="5">PT</th>
<th id="769D5DBA0C7E469A1D5E0E35CA5C2417" box="[1825,1915,532,547]" gridcol="18" gridrow="0" pageId="5">MXT</th>
</tr>
<tr id="354C34C60C7E469A1A1F0E16CA5C2478" box="[96,1915,567,588]" gridrow="1" pageId="5">
<th id="769D5DBA0C7E469A1A1F0E16CC222478" box="[96,261,567,588]" gridcol="0" gridrow="1" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0E16CDF42478" box="[97,211,567,588]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="angulatus">
<emphasis id="B908EA960C7EB9601A1E0E16CDF42478" box="[97,211,567,588]" italics="true" pageId="5">H. angulatus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0E16CC712478" box="[288,342,567,588]" gridcol="1" gridrow="1" pageId="5">19–21</td>
<td id="769D5DBA0C7E469A1B0E0E16CC802478" box="[369,423,567,588]" gridcol="2" gridrow="1" pageId="5">19–20</td>
<td id="769D5DBA0C7E469A1BBC0E16CCDF2478" box="[451,504,567,588]" gridcol="3" gridrow="1" pageId="5">17–19</td>
<td id="769D5DBA0C7E469A186B0E16CF612478" box="[532,582,567,588]" gridcol="4" gridrow="1" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0E16CFDA2478" box="[610,765,567,588]" gridcol="5" gridrow="1" pageId="5">105–123/109–123</td>
<td id="769D5DBA0C7E469A19660E16CEB32478" box="[793,916,567,588]" gridcol="6" gridrow="1" pageId="5">74–96/66–84</td>
<td id="769D5DBA0C7E469A19CF0E16CEE82478" box="[944,975,567,588]" gridcol="7" gridrow="1" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A19890E16C9072478" box="[1014,1056,567,588]" gridcol="8" gridrow="1" pageId="5">7–9</td>
<td id="769D5DBA0C7E469A1E430E16C9942478" box="[1084,1203,567,588]" gridcol="9" gridrow="1" pageId="5">iv; iv–v; v</td>
<td id="769D5DBA0C7E469A1EAE0E16C8202478" box="[1233,1287,567,588]" gridcol="10" gridrow="1" pageId="5">9–11</td>
<td id="769D5DBA0C7E469A1F5C0E16C8B42478" box="[1315,1427,567,588]" gridcol="11" gridrow="1" pageId="5">i–iv; i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0E16C8E92478" box="[1456,1486,567,588]" gridcol="12" gridrow="1" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950E16CB322478" box="[1514,1557,567,588]" gridcol="13" gridrow="1" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0E16CB762478" box="[1585,1617,567,588]" gridcol="14" gridrow="1" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C120E16CBAA2478" box="[1645,1677,567,588]" gridcol="15" gridrow="1" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60E16CBEE2478" box="[1705,1737,567,588]" gridcol="16" gridrow="1" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1C9A0E16CA222478" box="[1765,1797,567,588]" gridcol="17" gridrow="1" pageId="5">3–4</td>
<td id="769D5DBA0C7E469A1D5E0E16CA5C2478" box="[1825,1915,567,588]" gridcol="18" gridrow="1" pageId="5">15 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0E6ECA5C2450" box="[96,1915,591,612]" gridrow="2" pageId="5">
<th id="769D5DBA0C7E469A1A1F0E6ECC222450" box="[96,261,591,612]" gridcol="0" gridrow="2" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0E6ECD9F2450" authorityName="Kawashita-Ribeiro, Avila and Morais" authorityYear="2013" box="[97,184,591,612]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="apiaka">
<emphasis id="B908EA960C7EB9601A1E0E6ECD9F2450" box="[97,184,591,612]" italics="true" pageId="5">H. apiaka</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0E6ECC712450" box="[288,342,591,612]" gridcol="1" gridrow="2" pageId="5">21–24</td>
<td id="769D5DBA0C7E469A1B0E0E6ECC802450" box="[369,423,591,612]" gridcol="2" gridrow="2" pageId="5">21–22</td>
<td id="769D5DBA0C7E469A1BBC0E6ECCDF2450" box="[451,504,591,612]" gridcol="3" gridrow="2" pageId="5">17–19</td>
<td id="769D5DBA0C7E469A186B0E6ECF612450" box="[532,582,591,612]" gridcol="4" gridrow="2" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0E6ECFDA2450" box="[610,765,591,612]" gridcol="5" gridrow="2" pageId="5">118–127/124–132</td>
<td id="769D5DBA0C7E469A19660E6ECEB32450" box="[793,916,591,612]" gridcol="6" gridrow="2" pageId="5">79–103/80–84</td>
<td id="769D5DBA0C7E469A19CF0E6ECEE82450" box="[944,975,591,612]" gridcol="7" gridrow="2" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A19890E6EC9072450" box="[1014,1056,591,612]" gridcol="8" gridrow="2" pageId="5">7–9</td>
<td id="769D5DBA0C7E469A1E430E6EC9942450" box="[1084,1203,591,612]" gridcol="9" gridrow="2" pageId="5">iii–iv</td>
<td id="769D5DBA0C7E469A1EAE0E6EC8202450" box="[1233,1287,591,612]" gridcol="10" gridrow="2" pageId="5">9–12</td>
<td id="769D5DBA0C7E469A1F5C0E6EC8B42450" box="[1315,1427,591,612]" gridcol="11" gridrow="2" pageId="5">i–iv; i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0E6EC8E92450" box="[1456,1486,591,612]" gridcol="12" gridrow="2" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950E6ECB322450" box="[1514,1557,591,612]" gridcol="13" gridrow="2" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0E6ECB762450" box="[1585,1617,591,612]" gridcol="14" gridrow="2" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120E6ECBAA2450" box="[1645,1677,591,612]" gridcol="15" gridrow="2" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60E6ECBEE2450" box="[1705,1737,591,612]" gridcol="16" gridrow="2" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1C9A0E6ECA222450" box="[1765,1797,591,612]" gridcol="17" gridrow="2" pageId="5">3</td>
<td id="769D5DBA0C7E469A1D5E0E6ECA5C2450" box="[1825,1915,591,612]" gridcol="18" gridrow="2" pageId="5">15–16 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0E46CA5C2448" box="[96,1915,615,636]" gridrow="3" pageId="5">
<th id="769D5DBA0C7E469A1A1F0E46CC222448" box="[96,261,615,636]" gridcol="0" gridrow="3" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0E46CDC82448" box="[97,239,615,636]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="carinicaudus">
<emphasis id="B908EA960C7EB9601A1E0E46CDC82448" box="[97,239,615,636]" italics="true" pageId="5">H. carinicaudus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0E46CC712448" box="[288,342,615,636]" gridcol="1" gridrow="3" pageId="5">19</td>
<td id="769D5DBA0C7E469A1B0E0E46CC802448" box="[369,423,615,636]" gridcol="2" gridrow="3" pageId="5">19</td>
<td id="769D5DBA0C7E469A1BBC0E46CCDF2448" box="[451,504,615,636]" gridcol="3" gridrow="3" pageId="5">17</td>
<td id="769D5DBA0C7E469A186B0E46CF612448" box="[532,582,615,636]" gridcol="4" gridrow="3" pageId="5">W–S</td>
<td id="769D5DBA0C7E469A181D0E46CFDA2448" box="[610,765,615,636]" gridcol="5" gridrow="3" pageId="5">130–141/135–148</td>
<td id="769D5DBA0C7E469A19660E46CEB32448" box="[793,916,615,636]" gridcol="6" gridrow="3" pageId="5">48–69/48–73</td>
<td id="769D5DBA0C7E469A19CF0E46CEE82448" box="[944,975,615,636]" gridcol="7" gridrow="3" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890E46C9072448" box="[1014,1056,615,636]" gridcol="8" gridrow="3" pageId="5">7–8</td>
<td id="769D5DBA0C7E469A1E430E46C9942448" box="[1084,1203,615,636]" gridcol="9" gridrow="3" pageId="5">iii–iv; iv; iv–v</td>
<td id="769D5DBA0C7E469A1EAE0E46C8202448" box="[1233,1287,615,636]" gridcol="10" gridrow="3" pageId="5">9–10</td>
<td id="769D5DBA0C7E469A1F5C0E46C8B42448" box="[1315,1427,615,636]" gridcol="11" gridrow="3" pageId="5">i–v</td>
<td id="769D5DBA0C7E469A1FCF0E46C8E92448" box="[1456,1486,615,636]" gridcol="12" gridrow="3" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950E46CB322448" box="[1514,1557,615,636]" gridcol="13" gridrow="3" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0E46CB762448" box="[1585,1617,615,636]" gridcol="14" gridrow="3" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120E46CBAA2448" box="[1645,1677,615,636]" gridcol="15" gridrow="3" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60E46CBEE2448" box="[1705,1737,615,636]" gridcol="16" gridrow="3" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C9A0E46CA222448" box="[1765,1797,615,636]" gridcol="17" gridrow="3" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1D5E0E46CA5C2448" box="[1825,1915,615,636]" gridcol="18" gridrow="3" pageId="5">16–19 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0E5ECA5C24A0" box="[96,1915,639,660]" gridrow="4" pageId="5">
<th id="769D5DBA0C7E469A1A1F0E5ECC2224A0" box="[96,261,639,660]" gridcol="0" gridrow="4" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0E5ECD9D24A0" box="[97,186,639,660]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="danieli">
<emphasis id="B908EA960C7EB9601A1E0E5ECD9D24A0" box="[97,186,639,660]" italics="true" pageId="5">H. danieli</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0E5ECC7124A0" box="[288,342,639,660]" gridcol="1" gridrow="4" pageId="5">19–21</td>
<td id="769D5DBA0C7E469A1B0E0E5ECC8024A0" box="[369,423,639,660]" gridcol="2" gridrow="4" pageId="5">19–20</td>
<td id="769D5DBA0C7E469A1BBC0E5ECCDF24A0" box="[451,504,639,660]" gridcol="3" gridrow="4" pageId="5">16–19</td>
<td id="769D5DBA0C7E469A186B0E5ECF6124A0" box="[532,582,639,660]" gridcol="4" gridrow="4" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0E5ECFDA24A0" box="[610,765,639,660]" gridcol="5" gridrow="4" pageId="5">125–135/130–141</td>
<td id="769D5DBA0C7E469A19660E5ECEB324A0" box="[793,916,639,660]" gridcol="6" gridrow="4" pageId="5">76–86/61–70</td>
<td id="769D5DBA0C7E469A19CF0E5ECEE824A0" box="[944,975,639,660]" gridcol="7" gridrow="4" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890E5EC90724A0" box="[1014,1056,639,660]" gridcol="8" gridrow="4" pageId="5">8–9</td>
<td id="769D5DBA0C7E469A1E430E5EC99424A0" box="[1084,1203,639,660]" gridcol="9" gridrow="4" pageId="5">iv</td>
<td id="769D5DBA0C7E469A1EAE0E5EC82024A0" box="[1233,1287,639,660]" gridcol="10" gridrow="4" pageId="5">10–11</td>
<td id="769D5DBA0C7E469A1F5C0E5EC8B424A0" box="[1315,1427,639,660]" gridcol="11" gridrow="4" pageId="5">i–v</td>
<td id="769D5DBA0C7E469A1FCF0E5EC8E924A0" box="[1456,1486,639,660]" gridcol="12" gridrow="4" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950E5ECB3224A0" box="[1514,1557,639,660]" gridcol="13" gridrow="4" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0E5ECB7624A0" box="[1585,1617,639,660]" gridcol="14" gridrow="4" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120E5ECBAA24A0" box="[1645,1677,639,660]" gridcol="15" gridrow="4" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60E5ECBEE24A0" box="[1705,1737,639,660]" gridcol="16" gridrow="4" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C9A0E5ECA2224A0" box="[1765,1797,639,660]" gridcol="17" gridrow="4" pageId="5">2</td>
<td id="769D5DBA0C7E469A1D5E0E5ECA5C24A0" box="[1825,1915,639,660]" gridcol="18" gridrow="4" pageId="5">16–20</td>
</tr>
<tr id="354C34C60C7E469A1A1F0EB6CA5C2498" box="[96,1915,663,684]" gridrow="5" pageId="5">
<th id="769D5DBA0C7E469A1A1F0EB6CC222498" box="[96,261,663,684]" gridcol="0" gridrow="5" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0EB6CD9D2498" box="[97,186,663,684]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gomesi">
<emphasis id="B908EA960C7EB9601A1E0EB6CD9D2498" box="[97,186,663,684]" italics="true" pageId="5">H. gomesi</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0EB6CC712498" box="[288,342,663,684]" gridcol="1" gridrow="5" pageId="5">19</td>
<td id="769D5DBA0C7E469A1B0E0EB6CC802498" box="[369,423,663,684]" gridcol="2" gridrow="5" pageId="5">19</td>
<td id="769D5DBA0C7E469A1BBC0EB6CCDF2498" box="[451,504,663,684]" gridcol="3" gridrow="5" pageId="5">19</td>
<td id="769D5DBA0C7E469A186B0EB6CF612498" box="[532,582,663,684]" gridcol="4" gridrow="5" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0EB6CFDA2498" box="[610,765,663,684]" gridcol="5" gridrow="5" pageId="5">125–132/128–132</td>
<td id="769D5DBA0C7E469A19660EB6CEB32498" box="[793,916,663,684]" gridcol="6" gridrow="5" pageId="5">71–86/67–73</td>
<td id="769D5DBA0C7E469A19CF0EB6CEE82498" box="[944,975,663,684]" gridcol="7" gridrow="5" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A19890EB6C9072498" box="[1014,1056,663,684]" gridcol="8" gridrow="5" pageId="5">8–9</td>
<td id="769D5DBA0C7E469A1E430EB6C9942498" box="[1084,1203,663,684]" gridcol="9" gridrow="5" pageId="5">iv; v</td>
<td id="769D5DBA0C7E469A1EAE0EB6C8202498" box="[1233,1287,663,684]" gridcol="10" gridrow="5" pageId="5">10</td>
<td id="769D5DBA0C7E469A1F5C0EB6C8B42498" box="[1315,1427,663,684]" gridcol="11" gridrow="5" pageId="5">i–iv; i–v</td>
<td id="769D5DBA0C7E469A1FCF0EB6C8E92498" box="[1456,1486,663,684]" gridcol="12" gridrow="5" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950EB6CB322498" box="[1514,1557,663,684]" gridcol="13" gridrow="5" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0EB6CB762498" box="[1585,1617,663,684]" gridcol="14" gridrow="5" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120EB6CBAA2498" box="[1645,1677,663,684]" gridcol="15" gridrow="5" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60EB6CBEE2498" box="[1705,1737,663,684]" gridcol="16" gridrow="5" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C9A0EB6CA222498" box="[1765,1797,663,684]" gridcol="17" gridrow="5" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1D5E0EB6CA5C2498" box="[1825,1915,663,684]" gridcol="18" gridrow="5" pageId="5">18</td>
</tr>
<tr id="354C34C60C7E469A1A1F0E8ECA5C24F0" box="[96,1915,687,708]" gridrow="6" pageId="5">
<th id="769D5DBA0C7E469A1A1F0E8ECC2224F0" box="[96,261,687,708]" gridcol="0" gridrow="6" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0E8ECDF224F0" box="[97,213,687,708]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="hagmanni">
<emphasis id="B908EA960C7EB9601A1E0E8ECDF224F0" box="[97,213,687,708]" italics="true" pageId="5">H. hagmanni</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0E8ECC7124F0" box="[288,342,687,708]" gridcol="1" gridrow="6" pageId="5">23–27</td>
<td id="769D5DBA0C7E469A1B0E0E8ECC8024F0" box="[369,423,687,708]" gridcol="2" gridrow="6" pageId="5">21–29</td>
<td id="769D5DBA0C7E469A1BBC0E8ECCDF24F0" box="[451,504,687,708]" gridcol="3" gridrow="6" pageId="5">19–23</td>
<td id="769D5DBA0C7E469A186B0E8ECF6124F0" box="[532,582,687,708]" gridcol="4" gridrow="6" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0E8ECFDA24F0" box="[610,765,687,708]" gridcol="5" gridrow="6" pageId="5">117–127/130–134</td>
<td id="769D5DBA0C7E469A19660E8ECEB324F0" box="[793,916,687,708]" gridcol="6" gridrow="6" pageId="5">55–67/51–53</td>
<td id="769D5DBA0C7E469A19CF0E8ECEE824F0" box="[944,975,687,708]" gridcol="7" gridrow="6" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890E8EC90724F0" box="[1014,1056,687,708]" gridcol="8" gridrow="6" pageId="5">8</td>
<td id="769D5DBA0C7E469A1E430E8EC99424F0" box="[1084,1203,687,708]" gridcol="9" gridrow="6" pageId="5">iv</td>
<td id="769D5DBA0C7E469A1EAE0E8EC82024F0" box="[1233,1287,687,708]" gridcol="10" gridrow="6" pageId="5">10–12</td>
<td id="769D5DBA0C7E469A1F5C0E8EC8B424F0" box="[1315,1427,687,708]" gridcol="11" gridrow="6" pageId="5">i–v</td>
<td id="769D5DBA0C7E469A1FCF0E8EC8E924F0" box="[1456,1486,687,708]" gridcol="12" gridrow="6" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A1F950E8ECB3224F0" box="[1514,1557,687,708]" gridcol="13" gridrow="6" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0E8ECB7624F0" box="[1585,1617,687,708]" gridcol="14" gridrow="6" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C120E8ECBAA24F0" box="[1645,1677,687,708]" gridcol="15" gridrow="6" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1CD60E8ECBEE24F0" box="[1705,1737,687,708]" gridcol="16" gridrow="6" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C9A0E8ECA2224F0" box="[1765,1797,687,708]" gridcol="17" gridrow="6" pageId="5">3</td>
<td id="769D5DBA0C7E469A1D5E0E8ECA5C24F0" box="[1825,1915,687,708]" gridcol="18" gridrow="6" pageId="5">15–17 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0EE6CA5C24E8" box="[96,1915,711,732]" gridrow="7" pageId="5">
<th id="769D5DBA0C7E469A1A1F0EE6CC2224E8" box="[96,261,711,732]" gridcol="0" gridrow="7" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0EE6CDDF24E8" box="[97,248,711,732]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="infrataeniatus">
<emphasis id="B908EA960C7EB9601A1E0EE6CDDF24E8" box="[97,248,711,732]" italics="true" pageId="5">H. infrataeniatus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0EE6CC7124E8" box="[288,342,711,732]" gridcol="1" gridrow="7" pageId="5">17–19</td>
<td id="769D5DBA0C7E469A1B0E0EE6CC8024E8" box="[369,423,711,732]" gridcol="2" gridrow="7" pageId="5">17–19</td>
<td id="769D5DBA0C7E469A1BBC0EE6CCDF24E8" box="[451,504,711,732]" gridcol="3" gridrow="7" pageId="5">15–19</td>
<td id="769D5DBA0C7E469A186B0EE6CF6124E8" box="[532,582,711,732]" gridcol="4" gridrow="7" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0EE6CFDA24E8" box="[610,765,711,732]" gridcol="5" gridrow="7" pageId="5">115–138/117–138</td>
<td id="769D5DBA0C7E469A19660EE6CEB324E8" box="[793,916,711,732]" gridcol="6" gridrow="7" pageId="5">52–88/50–83</td>
<td id="769D5DBA0C7E469A19CF0EE6CEE824E8" box="[944,975,711,732]" gridcol="7" gridrow="7" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890EE6C90724E8" box="[1014,1056,711,732]" gridcol="8" gridrow="7" pageId="5">7–9</td>
<td id="769D5DBA0C7E469A1E430EE6C99424E8" box="[1084,1203,711,732]" gridcol="9" gridrow="7" pageId="5">iii–iv; iv</td>
<td id="769D5DBA0C7E469A1EAE0EE6C82024E8" box="[1233,1287,711,732]" gridcol="10" gridrow="7" pageId="5">10–12</td>
<td id="769D5DBA0C7E469A1F5C0EE6C8B424E8" box="[1315,1427,711,732]" gridcol="11" gridrow="7" pageId="5">i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0EE6C8E924E8" box="[1456,1486,711,732]" gridcol="12" gridrow="7" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950EE6CB3224E8" box="[1514,1557,711,732]" gridcol="13" gridrow="7" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0EE6CB7624E8" box="[1585,1617,711,732]" gridcol="14" gridrow="7" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120EE6CBAA24E8" box="[1645,1677,711,732]" gridcol="15" gridrow="7" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60EE6CBEE24E8" box="[1705,1737,711,732]" gridcol="16" gridrow="7" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C9A0EE6CA2224E8" box="[1765,1797,711,732]" gridcol="17" gridrow="7" pageId="5">2</td>
<td id="769D5DBA0C7E469A1D5E0EE6CA5C24E8" box="[1825,1915,711,732]" gridcol="18" gridrow="7" pageId="5">17–20 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0EFFCA5C24C7" box="[96,1915,734,755]" gridrow="8" pageId="5">
<th id="769D5DBA0C7E469A1A1F0EFFCC2224C7" box="[96,261,734,755]" gridcol="0" gridrow="8" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0EFFCDC224C7" box="[97,229,734,755]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="leopardinus">
<emphasis id="B908EA960C7EB9601A1E0EFFCDC224C7" box="[97,229,734,755]" italics="true" pageId="5">H. leopardinus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0EFFCC7124C7" box="[288,342,734,755]" gridcol="1" gridrow="8" pageId="5">15–22</td>
<td id="769D5DBA0C7E469A1B0E0EFFCC8024C7" box="[369,423,734,755]" gridcol="2" gridrow="8" pageId="5">19–22</td>
<td id="769D5DBA0C7E469A1BBC0EFFCCDF24C7" box="[451,504,734,755]" gridcol="3" gridrow="8" pageId="5">17–19</td>
<td id="769D5DBA0C7E469A186B0EFFCF6124C7" box="[532,582,734,755]" gridcol="4" gridrow="8" pageId="5">M–S</td>
<td id="769D5DBA0C7E469A181D0EFFCFDA24C7" box="[610,765,734,755]" gridcol="5" gridrow="8" pageId="5">108–126/108–130</td>
<td id="769D5DBA0C7E469A19660EFFCEB324C7" box="[793,916,734,755]" gridcol="6" gridrow="8" pageId="5">64–89/53–76</td>
<td id="769D5DBA0C7E469A19CF0EFFCEE824C7" box="[944,975,734,755]" gridcol="7" gridrow="8" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890EFFC90724C7" box="[1014,1056,734,755]" gridcol="8" gridrow="8" pageId="5">8–10</td>
<td id="769D5DBA0C7E469A1E430EFFC99424C7" box="[1084,1203,734,755]" gridcol="9" gridrow="8" pageId="5">iii–iv; iv; iii–v</td>
<td id="769D5DBA0C7E469A1EAE0EFFC82024C7" box="[1233,1287,734,755]" gridcol="10" gridrow="8" pageId="5">8–11</td>
<td id="769D5DBA0C7E469A1F5C0EFFC8B424C7" box="[1315,1427,734,755]" gridcol="11" gridrow="8" pageId="5">i–iv; i–v</td>
<td id="769D5DBA0C7E469A1FCF0EFFC8E924C7" box="[1456,1486,734,755]" gridcol="12" gridrow="8" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950EFFCB3224C7" box="[1514,1557,734,755]" gridcol="13" gridrow="8" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0EFFCB7624C7" box="[1585,1617,734,755]" gridcol="14" gridrow="8" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C120EFFCBAA24C7" box="[1645,1677,734,755]" gridcol="15" gridrow="8" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60EFFCBEE24C7" box="[1705,1737,734,755]" gridcol="16" gridrow="8" pageId="5">2</td>
<td id="769D5DBA0C7E469A1C9A0EFFCA2224C7" box="[1765,1797,734,755]" gridcol="17" gridrow="8" pageId="5">2–4</td>
<td id="769D5DBA0C7E469A1D5E0EFFCA5C24C7" box="[1825,1915,734,755]" gridcol="18" gridrow="8" pageId="5">18 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0ED7CA5C253F" box="[96,1915,758,779]" gridrow="9" pageId="5">
<th id="769D5DBA0C7E469A1A1F0ED7CC22253F" box="[96,261,758,779]" gridcol="0" gridrow="9" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0ED7CDE8253F" authorityName="Gunther" authorityYear="1861" box="[97,207,758,779]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="modestus">
<emphasis id="B908EA960C7EB9601A1E0ED7CDE8253F" box="[97,207,758,779]" italics="true" pageId="5">H. modestus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0ED7CC71253F" box="[288,342,758,779]" gridcol="1" gridrow="9" pageId="5">19</td>
<td id="769D5DBA0C7E469A1B0E0ED7CC80253F" box="[369,423,758,779]" gridcol="2" gridrow="9" pageId="5">19</td>
<td id="769D5DBA0C7E469A1BBC0ED7CCDF253F" box="[451,504,758,779]" gridcol="3" gridrow="9" pageId="5">17–19</td>
<td id="769D5DBA0C7E469A186B0ED7CF61253F" box="[532,582,758,779]" gridcol="4" gridrow="9" pageId="5">W–M</td>
<td id="769D5DBA0C7E469A181D0ED7CFDA253F" box="[610,765,758,779]" gridcol="5" gridrow="9" pageId="5">112–125/116–122</td>
<td id="769D5DBA0C7E469A19660ED7CEB3253F" box="[793,916,758,779]" gridcol="6" gridrow="9" pageId="5">54–70/53–64</td>
<td id="769D5DBA0C7E469A19CF0ED7CEE8253F" box="[944,975,758,779]" gridcol="7" gridrow="9" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890ED7C907253F" box="[1014,1056,758,779]" gridcol="8" gridrow="9" pageId="5">8</td>
<td id="769D5DBA0C7E469A1E430ED7C994253F" box="[1084,1203,758,779]" gridcol="9" gridrow="9" pageId="5">iii–iv; iv</td>
<td id="769D5DBA0C7E469A1EAE0ED7C820253F" box="[1233,1287,758,779]" gridcol="10" gridrow="9" pageId="5">9–11</td>
<td id="769D5DBA0C7E469A1F5C0ED7C8B4253F" box="[1315,1427,758,779]" gridcol="11" gridrow="9" pageId="5">i–iv; i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0ED7C8E9253F" box="[1456,1486,758,779]" gridcol="12" gridrow="9" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950ED7CB32253F" box="[1514,1557,758,779]" gridcol="13" gridrow="9" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0ED7CB76253F" box="[1585,1617,758,779]" gridcol="14" gridrow="9" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C120ED7CBAA253F" box="[1645,1677,758,779]" gridcol="15" gridrow="9" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1CD60ED7CBEE253F" box="[1705,1737,758,779]" gridcol="16" gridrow="9" pageId="5">2</td>
<td id="769D5DBA0C7E469A1C9A0ED7CA22253F" box="[1765,1797,758,779]" gridcol="17" gridrow="9" pageId="5">2–4</td>
<td id="769D5DBA0C7E469A1D5E0ED7CA5C253F" box="[1825,1915,758,779]" gridcol="18" gridrow="9" pageId="5">19 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0F2FCA5C2517" box="[96,1915,782,803]" gridrow="10" pageId="5">
<th id="769D5DBA0C7E469A1A1F0F2FCC222517" box="[96,261,782,803]" gridcol="0" gridrow="10" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0F2FCDEE2517" box="[97,201,782,803]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="pastazae">
<emphasis id="B908EA960C7EB9601A1E0F2FCDEE2517" box="[97,201,782,803]" italics="true" pageId="5">H. pastazae</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0F2FCC712517" box="[288,342,782,803]" gridcol="1" gridrow="10" pageId="5">23</td>
<td id="769D5DBA0C7E469A1B0E0F2FCC802517" box="[369,423,782,803]" gridcol="2" gridrow="10" pageId="5">23–25</td>
<td id="769D5DBA0C7E469A1BBC0F2FCCDF2517" box="[451,504,782,803]" gridcol="3" gridrow="10" pageId="5">19</td>
<td id="769D5DBA0C7E469A186B0F2FCF612517" box="[532,582,782,803]" gridcol="4" gridrow="10" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0F2FCFDA2517" box="[610,765,782,803]" gridcol="5" gridrow="10" pageId="5">121–134/130–145</td>
<td id="769D5DBA0C7E469A19660F2FCEB32517" box="[793,916,782,803]" gridcol="6" gridrow="10" pageId="5">93–117/72–97</td>
<td id="769D5DBA0C7E469A19CF0F2FCEE82517" box="[944,975,782,803]" gridcol="7" gridrow="10" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890F2FC9072517" box="[1014,1056,782,803]" gridcol="8" gridrow="10" pageId="5">8–10</td>
<td id="769D5DBA0C7E469A1E430F2FC9942517" box="[1084,1203,782,803]" gridcol="9" gridrow="10" pageId="5">iv</td>
<td id="769D5DBA0C7E469A1EAE0F2FC8202517" box="[1233,1287,782,803]" gridcol="10" gridrow="10" pageId="5">9–13</td>
<td id="769D5DBA0C7E469A1F5C0F2FC8B42517" box="[1315,1427,782,803]" gridcol="11" gridrow="10" pageId="5">i–iv; i–v</td>
<td id="769D5DBA0C7E469A1FCF0F2FC8E92517" box="[1456,1486,782,803]" gridcol="12" gridrow="10" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A1F950F2FCB322517" box="[1514,1557,782,803]" gridcol="13" gridrow="10" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0F2FCB762517" box="[1585,1617,782,803]" gridcol="14" gridrow="10" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120F2FCBAA2517" box="[1645,1677,782,803]" gridcol="15" gridrow="10" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60F2FCBEE2517" box="[1705,1737,782,803]" gridcol="16" gridrow="10" pageId="5">2</td>
<td id="769D5DBA0C7E469A1C9A0F2FCA222517" box="[1765,1797,782,803]" gridcol="17" gridrow="10" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1D5E0F2FCA5C2517" box="[1825,1915,782,803]" gridcol="18" gridrow="10" pageId="5">14–17 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0F07CA5C250F" box="[96,1915,806,827]" gridrow="11" pageId="5">
<th id="769D5DBA0C7E469A1A1F0F07CC22250F" box="[96,261,806,827]" gridcol="0" gridrow="11" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0F07CD9E250F" box="[97,185,806,827]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B908EA960C7EB9601A1E0F07CD9E250F" box="[97,185,806,827]" italics="true" pageId="5">H. petersi</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0F07CC71250F" box="[288,342,806,827]" gridcol="1" gridrow="11" pageId="5">21</td>
<td id="769D5DBA0C7E469A1B0E0F07CC80250F" box="[369,423,806,827]" gridcol="2" gridrow="11" pageId="5">21–23</td>
<td id="769D5DBA0C7E469A1BBC0F07CCDF250F" box="[451,504,806,827]" gridcol="3" gridrow="11" pageId="5">16</td>
<td id="769D5DBA0C7E469A186B0F07CF61250F" box="[532,582,806,827]" gridcol="4" gridrow="11" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0F07CFDA250F" box="[610,765,806,827]" gridcol="5" gridrow="11" pageId="5">135–142/137–150</td>
<td id="769D5DBA0C7E469A19660F07CEB3250F" box="[793,916,806,827]" gridcol="6" gridrow="11" pageId="5">85–91/67–73</td>
<td id="769D5DBA0C7E469A19CF0F07CEE8250F" box="[944,975,806,827]" gridcol="7" gridrow="11" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890F07C907250F" box="[1014,1056,806,827]" gridcol="8" gridrow="11" pageId="5">8</td>
<td id="769D5DBA0C7E469A1E430F07C994250F" box="[1084,1203,806,827]" gridcol="9" gridrow="11" pageId="5">iv</td>
<td id="769D5DBA0C7E469A1EAE0F07C820250F" box="[1233,1287,806,827]" gridcol="10" gridrow="11" pageId="5">9–12</td>
<td id="769D5DBA0C7E469A1F5C0F07C8B4250F" box="[1315,1427,806,827]" gridcol="11" gridrow="11" pageId="5">i–v</td>
<td id="769D5DBA0C7E469A1FCF0F07C8E9250F" box="[1456,1486,806,827]" gridcol="12" gridrow="11" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A1F950F07CB32250F" box="[1514,1557,806,827]" gridcol="13" gridrow="11" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0F07CB76250F" box="[1585,1617,806,827]" gridcol="14" gridrow="11" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120F07CBAA250F" box="[1645,1677,806,827]" gridcol="15" gridrow="11" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60F07CBEE250F" box="[1705,1737,806,827]" gridcol="16" gridrow="11" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C9A0F07CA22250F" box="[1765,1797,806,827]" gridcol="17" gridrow="11" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1D5E0F07CA5C250F" box="[1825,1915,806,827]" gridcol="18" gridrow="11" pageId="5">16–17 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0F1FCA5C2567" box="[96,1915,830,851]" gridrow="12" pageId="5">
<th id="769D5DBA0C7E469A1A1F0F1FCC222567" box="[96,261,830,851]" gridcol="0" gridrow="12" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0F1FCDEB2567" box="[97,204,830,851]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="polnlepis">
<emphasis id="B908EA960C7EB9601A1E0F1FCDEB2567" box="[97,204,830,851]" italics="true" pageId="5">H. polŋlepis</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0F1FCC712567" box="[288,342,830,851]" gridcol="1" gridrow="12" pageId="5">23–25</td>
<td id="769D5DBA0C7E469A1B0E0F1FCC802567" box="[369,423,830,851]" gridcol="2" gridrow="12" pageId="5">23–25</td>
<td id="769D5DBA0C7E469A1BBC0F1FCCDF2567" box="[451,504,830,851]" gridcol="3" gridrow="12" pageId="5">19</td>
<td id="769D5DBA0C7E469A186B0F1FCF612567" box="[532,582,830,851]" gridcol="4" gridrow="12" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0F1FCFDA2567" box="[610,765,830,851]" gridcol="5" gridrow="12" pageId="5">112–131/121–133</td>
<td id="769D5DBA0C7E469A19660F1FCEB32567" box="[793,916,830,851]" gridcol="6" gridrow="12" pageId="5">70–102/71–81</td>
<td id="769D5DBA0C7E469A19CF0F1FCEE82567" box="[944,975,830,851]" gridcol="7" gridrow="12" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890F1FC9072567" box="[1014,1056,830,851]" gridcol="8" gridrow="12" pageId="5">8–9</td>
<td id="769D5DBA0C7E469A1E430F1FC9942567" box="[1084,1203,830,851]" gridcol="9" gridrow="12" pageId="5">iii–iv; iv</td>
<td id="769D5DBA0C7E469A1EAE0F1FC8202567" box="[1233,1287,830,851]" gridcol="10" gridrow="12" pageId="5">11–13</td>
<td id="769D5DBA0C7E469A1F5C0F1FC8B42567" box="[1315,1427,830,851]" gridcol="11" gridrow="12" pageId="5">i–iv; i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0F1FC8E92567" box="[1456,1486,830,851]" gridcol="12" gridrow="12" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950F1FCB322567" box="[1514,1557,830,851]" gridcol="13" gridrow="12" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0F1FCB762567" box="[1585,1617,830,851]" gridcol="14" gridrow="12" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120F1FCBAA2567" box="[1645,1677,830,851]" gridcol="15" gridrow="12" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60F1FCBEE2567" box="[1705,1737,830,851]" gridcol="16" gridrow="12" pageId="5">1–2</td>
<td id="769D5DBA0C7E469A1C9A0F1FCA222567" box="[1765,1797,830,851]" gridcol="17" gridrow="12" pageId="5">2–4</td>
<td id="769D5DBA0C7E469A1D5E0F1FCA5C2567" box="[1825,1915,830,851]" gridcol="18" gridrow="12" pageId="5">16 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0F77CA5C255F" box="[96,1915,854,875]" gridrow="13" pageId="5">
<th id="769D5DBA0C7E469A1A1F0F77CC22255F" box="[96,261,854,875]" gridcol="0" gridrow="13" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0F77CD98255F" box="[97,191,854,875]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="scalaris">
<emphasis id="B908EA960C7EB9601A1E0F77CD98255F" box="[97,191,854,875]" italics="true" pageId="5">H. scalaris</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0F77CC71255F" box="[288,342,854,875]" gridcol="1" gridrow="13" pageId="5">21–25</td>
<td id="769D5DBA0C7E469A1B0E0F77CC80255F" box="[369,423,854,875]" gridcol="2" gridrow="13" pageId="5">19–21</td>
<td id="769D5DBA0C7E469A1BBC0F77CCDF255F" box="[451,504,854,875]" gridcol="3" gridrow="13" pageId="5">16–19</td>
<td id="769D5DBA0C7E469A186B0F77CF61255F" box="[532,582,854,875]" gridcol="4" gridrow="13" pageId="5">S</td>
<td id="769D5DBA0C7E469A181D0F77CFDA255F" box="[610,765,854,875]" gridcol="5" gridrow="13" pageId="5">110–119/113–125</td>
<td id="769D5DBA0C7E469A19660F77CEB3255F" box="[793,916,854,875]" gridcol="6" gridrow="13" pageId="5">83–95/67–81</td>
<td id="769D5DBA0C7E469A19CF0F77CEE8255F" box="[944,975,854,875]" gridcol="7" gridrow="13" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A19890F77C907255F" box="[1014,1056,854,875]" gridcol="8" gridrow="13" pageId="5">8–9</td>
<td id="769D5DBA0C7E469A1E430F77C994255F" box="[1084,1203,854,875]" gridcol="9" gridrow="13" pageId="5">iv; iv–v</td>
<td id="769D5DBA0C7E469A1EAE0F77C820255F" box="[1233,1287,854,875]" gridcol="10" gridrow="13" pageId="5">10–14</td>
<td id="769D5DBA0C7E469A1F5C0F77C8B4255F" box="[1315,1427,854,875]" gridcol="11" gridrow="13" pageId="5">i–v</td>
<td id="769D5DBA0C7E469A1FCF0F77C8E9255F" box="[1456,1486,854,875]" gridcol="12" gridrow="13" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A1F950F77CB32255F" box="[1514,1557,854,875]" gridcol="13" gridrow="13" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0F77CB76255F" box="[1585,1617,854,875]" gridcol="14" gridrow="13" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120F77CBAA255F" box="[1645,1677,854,875]" gridcol="15" gridrow="13" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60F77CBEE255F" box="[1705,1737,854,875]" gridcol="16" gridrow="13" pageId="5">1–3</td>
<td id="769D5DBA0C7E469A1C9A0F77CA22255F" box="[1765,1797,854,875]" gridcol="17" gridrow="13" pageId="5">3</td>
<td id="769D5DBA0C7E469A1D5E0F77CA5C255F" box="[1825,1915,854,875]" gridcol="18" gridrow="13" pageId="5">15–17 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0F4FCA5C25B7" box="[96,1915,878,899]" gridrow="14" pageId="5">
<th id="769D5DBA0C7E469A1A1F0F4FCC2225B7" box="[96,261,878,899]" gridcol="0" gridrow="14" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0F4FCDC525B7" authorityName="Frota" authorityYear="2005" box="[97,226,878,899]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="B908EA960C7EB9601A1E0F4FCDC525B7" box="[97,226,878,899]" italics="true" pageId="5">H. tapajonicus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0F4FCC7125B7" box="[288,342,878,899]" gridcol="1" gridrow="14" pageId="5">19</td>
<td id="769D5DBA0C7E469A1B0E0F4FCC8025B7" box="[369,423,878,899]" gridcol="2" gridrow="14" pageId="5">19</td>
<td id="769D5DBA0C7E469A1BBC0F4FCCDF25B7" box="[451,504,878,899]" gridcol="3" gridrow="14" pageId="5">17</td>
<td id="769D5DBA0C7E469A186B0F4FCF6125B7" box="[532,582,878,899]" gridcol="4" gridrow="14" pageId="5">W</td>
<td id="769D5DBA0C7E469A181D0F4FCFDA25B7" box="[610,765,878,899]" gridcol="5" gridrow="14" pageId="5">118/121–123</td>
<td id="769D5DBA0C7E469A19660F4FCEB325B7" box="[793,916,878,899]" gridcol="6" gridrow="14" pageId="5">79/67–76</td>
<td id="769D5DBA0C7E469A19CF0F4FCEE825B7" box="[944,975,878,899]" gridcol="7" gridrow="14" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890F4FC90725B7" box="[1014,1056,878,899]" gridcol="8" gridrow="14" pageId="5">8</td>
<td id="769D5DBA0C7E469A1E430F4FC99425B7" box="[1084,1203,878,899]" gridcol="9" gridrow="14" pageId="5">iv</td>
<td id="769D5DBA0C7E469A1EAE0F4FC82025B7" box="[1233,1287,878,899]" gridcol="10" gridrow="14" pageId="5">9–11</td>
<td id="769D5DBA0C7E469A1F5C0F4FC8B425B7" box="[1315,1427,878,899]" gridcol="11" gridrow="14" pageId="5">i–iv; i–v</td>
<td id="769D5DBA0C7E469A1FCF0F4FC8E925B7" box="[1456,1486,878,899]" gridcol="12" gridrow="14" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950F4FCB3225B7" box="[1514,1557,878,899]" gridcol="13" gridrow="14" pageId="5">S (?)</td>
<td id="769D5DBA0C7E469A1C4E0F4FCB7625B7" box="[1585,1617,878,899]" gridcol="14" gridrow="14" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120F4FCBAA25B7" box="[1645,1677,878,899]" gridcol="15" gridrow="14" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60F4FCBEE25B7" box="[1705,1737,878,899]" gridcol="16" gridrow="14" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C9A0F4FCA2225B7" box="[1765,1797,878,899]" gridcol="17" gridrow="14" pageId="5">2</td>
<td id="769D5DBA0C7E469A1D5E0F4FCA5C25B7" box="[1825,1915,878,899]" gridcol="18" gridrow="14" pageId="5">18 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0FA7CA5C25AF" box="[96,1915,902,923]" gridrow="15" pageId="5">
<th id="769D5DBA0C7E469A1A1F0FA7CC2225AF" box="[96,261,902,923]" gridcol="0" gridrow="15" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0FA7CDF225AF" box="[97,213,902,923]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="trivittatus">
<emphasis id="B908EA960C7EB9601A1E0FA7CDF225AF" box="[97,213,902,923]" italics="true" pageId="5">H. triƲittatus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0FA7CC7125AF" box="[288,342,902,923]" gridcol="1" gridrow="15" pageId="5">21–25</td>
<td id="769D5DBA0C7E469A1B0E0FA7CC8025AF" box="[369,423,902,923]" gridcol="2" gridrow="15" pageId="5">20–23</td>
<td id="769D5DBA0C7E469A1BBC0FA7CCDF25AF" box="[451,504,902,923]" gridcol="3" gridrow="15" pageId="5">16–19</td>
<td id="769D5DBA0C7E469A186B0FA7CF6125AF" box="[532,582,902,923]" gridcol="4" gridrow="15" pageId="5">W</td>
<td id="769D5DBA0C7E469A181D0FA7CFDA25AF" box="[610,765,902,923]" gridcol="5" gridrow="15" pageId="5">114–128/115–129</td>
<td id="769D5DBA0C7E469A19660FA7CEB325AF" box="[793,916,902,923]" gridcol="6" gridrow="15" pageId="5">67–80/56–66</td>
<td id="769D5DBA0C7E469A19CF0FA7CEE825AF" box="[944,975,902,923]" gridcol="7" gridrow="15" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890FA7C90725AF" box="[1014,1056,902,923]" gridcol="8" gridrow="15" pageId="5">8–10</td>
<td id="769D5DBA0C7E469A1E430FA7C99425AF" box="[1084,1203,902,923]" gridcol="9" gridrow="15" pageId="5">iv; v</td>
<td id="769D5DBA0C7E469A1EAE0FA7C82025AF" box="[1233,1287,902,923]" gridcol="10" gridrow="15" pageId="5">11–14</td>
<td id="769D5DBA0C7E469A1F5C0FA7C8B425AF" box="[1315,1427,902,923]" gridcol="11" gridrow="15" pageId="5">i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0FA7C8E925AF" box="[1456,1486,902,923]" gridcol="12" gridrow="15" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A1F950FA7CB3225AF" box="[1514,1557,902,923]" gridcol="13" gridrow="15" pageId="5">S</td>
<td id="769D5DBA0C7E469A1C4E0FA7CB7625AF" box="[1585,1617,902,923]" gridcol="14" gridrow="15" pageId="5">2</td>
<td id="769D5DBA0C7E469A1C120FA7CBAA25AF" box="[1645,1677,902,923]" gridcol="15" gridrow="15" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60FA7CBEE25AF" box="[1705,1737,902,923]" gridcol="16" gridrow="15" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C9A0FA7CA2225AF" box="[1765,1797,902,923]" gridcol="17" gridrow="15" pageId="5">2</td>
<td id="769D5DBA0C7E469A1D5E0FA7CA5C25AF" box="[1825,1915,902,923]" gridcol="18" gridrow="15" pageId="5">19–22 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0FBFCA5C2587" box="[96,1915,926,947]" gridrow="16" pageId="5">
<th id="769D5DBA0C7E469A1A1F0FBFCC222587" box="[96,261,926,947]" gridcol="0" gridrow="16" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0FBFCD802587" box="[97,167,926,947]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nacu">
<emphasis id="B908EA960C7EB9601A1E0FBFCD802587" box="[97,167,926,947]" italics="true" pageId="5">H. ŋacu</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B5F0FBFCC712587" box="[288,342,926,947]" gridcol="1" gridrow="16" pageId="5">25–29</td>
<td id="769D5DBA0C7E469A1B0E0FBFCC802587" box="[369,423,926,947]" gridcol="2" gridrow="16" pageId="5">25–28</td>
<td id="769D5DBA0C7E469A1BBC0FBFCCDF2587" box="[451,504,926,947]" gridcol="3" gridrow="16" pageId="5">18–20</td>
<td id="769D5DBA0C7E469A186B0FBFCF612587" box="[532,582,926,947]" gridcol="4" gridrow="16" pageId="5">?</td>
<td id="769D5DBA0C7E469A181D0FBFCFDA2587" box="[610,765,926,947]" gridcol="5" gridrow="16" pageId="5">124/129–136</td>
<td id="769D5DBA0C7E469A19660FBFCEB32587" box="[793,916,926,947]" gridcol="6" gridrow="16" pageId="5">?/85–96</td>
<td id="769D5DBA0C7E469A19CF0FBFCEE82587" box="[944,975,926,947]" gridcol="7" gridrow="16" pageId="5">?</td>
<td id="769D5DBA0C7E469A19890FBFC9072587" box="[1014,1056,926,947]" gridcol="8" gridrow="16" pageId="5">8–9</td>
<td id="769D5DBA0C7E469A1E430FBFC9942587" box="[1084,1203,926,947]" gridcol="9" gridrow="16" pageId="5">iv–v</td>
<td id="769D5DBA0C7E469A1EAE0FBFC8202587" box="[1233,1287,926,947]" gridcol="10" gridrow="16" pageId="5">11–12</td>
<td id="769D5DBA0C7E469A1F5C0FBFC8B42587" box="[1315,1427,926,947]" gridcol="11" gridrow="16" pageId="5">i–v; i–vi</td>
<td id="769D5DBA0C7E469A1FCF0FBFC8E92587" box="[1456,1486,926,947]" gridcol="12" gridrow="16" pageId="5">Yes</td>
<td id="769D5DBA0C7E469A1F950FBFCB322587" box="[1514,1557,926,947]" gridcol="13" gridrow="16" pageId="5">E</td>
<td id="769D5DBA0C7E469A1C4E0FBFCB762587" box="[1585,1617,926,947]" gridcol="14" gridrow="16" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120FBFCBAA2587" box="[1645,1677,926,947]" gridcol="15" gridrow="16" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60FBFCBEE2587" box="[1705,1737,926,947]" gridcol="16" gridrow="16" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C9A0FBFCA222587" box="[1765,1797,926,947]" gridcol="17" gridrow="16" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1D5E0FBFCA5C2587" box="[1825,1915,926,947]" gridcol="18" gridrow="16" pageId="5">14–15 + 2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0F97CA5C25FF" box="[96,1915,950,971]" gridrow="17" pageId="5">
<th id="769D5DBA0C7E469A1A1F0F97CC2225FF" box="[96,261,950,971]" gridcol="0" gridrow="17" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1E0F97CD9E25FF" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[97,185,950,971]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7EB9601A1E0F97CD9E25FF" box="[97,185,950,971]" italics="true" pageId="5">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7EB9601AC00F97CC2225FE" box="[191,261,950,970]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="769D5DBA0C7E469A1B5F0F97CC7125FF" box="[288,342,950,971]" gridcol="1" gridrow="17" pageId="5">17</td>
<td id="769D5DBA0C7E469A1B0E0F97CC8025FF" box="[369,423,950,971]" gridcol="2" gridrow="17" pageId="5">17</td>
<td id="769D5DBA0C7E469A1BBC0F97CCDF25FF" box="[451,504,950,971]" gridcol="3" gridrow="17" pageId="5">15</td>
<td id="769D5DBA0C7E469A186B0F97CF6125FF" box="[532,582,950,971]" gridcol="4" gridrow="17" pageId="5">W</td>
<td id="769D5DBA0C7E469A181D0F97CFDA25FF" box="[610,765,950,971]" gridcol="5" gridrow="17" pageId="5">115/111–117</td>
<td id="769D5DBA0C7E469A19660F97CEB325FF" box="[793,916,950,971]" gridcol="6" gridrow="17" pageId="5">56/41–52</td>
<td id="769D5DBA0C7E469A19CF0F97CEE825FF" box="[944,975,950,971]" gridcol="7" gridrow="17" pageId="5">No</td>
<td id="769D5DBA0C7E469A19890F97C90725FF" box="[1014,1056,950,971]" gridcol="8" gridrow="17" pageId="5">8</td>
<td id="769D5DBA0C7E469A1E430F97C99425FF" box="[1084,1203,950,971]" gridcol="9" gridrow="17" pageId="5">iii–iv</td>
<td id="769D5DBA0C7E469A1EAE0F97C82025FF" box="[1233,1287,950,971]" gridcol="10" gridrow="17" pageId="5">9–10</td>
<td id="769D5DBA0C7E469A1F5C0F97C8B425FF" box="[1315,1427,950,971]" gridcol="11" gridrow="17" pageId="5">i–v</td>
<td id="769D5DBA0C7E469A1FCF0F97C8E925FF" box="[1456,1486,950,971]" gridcol="12" gridrow="17" pageId="5">No</td>
<td id="769D5DBA0C7E469A1F950F97CB3225FF" box="[1514,1557,950,971]" gridcol="13" gridrow="17" pageId="5">E</td>
<td id="769D5DBA0C7E469A1C4E0F97CB7625FF" box="[1585,1617,950,971]" gridcol="14" gridrow="17" pageId="5">1</td>
<td id="769D5DBA0C7E469A1C120F97CBAA25FF" box="[1645,1677,950,971]" gridcol="15" gridrow="17" pageId="5">2</td>
<td id="769D5DBA0C7E469A1CD60F97CBEE25FF" box="[1705,1737,950,971]" gridcol="16" gridrow="17" pageId="5">2</td>
<td id="769D5DBA0C7E469A1C9A0F97CA2225FF" box="[1765,1797,950,971]" gridcol="17" gridrow="17" pageId="5">2–3</td>
<td id="769D5DBA0C7E469A1D5E0F97CA5C25FF" box="[1825,1915,950,971]" gridcol="18" gridrow="17" pageId="5">18–21 + 2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC336840C7EB9601A090FF9CDE625CE" blockId="5.[97,1914,984,1018]" pageId="5">
<tableNote id="769A370A0C7EB9601A090FF9CDE625CE" pageId="5" targetBox="[96,1915,532,971]" targetPageId="5">
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<bibRefCitation id="EFED4B750C7EB9601EF80FF9C81925D3" author="Rossman, D. A. &amp; J. R. Dixon" box="[1159,1342,984,999]" pageId="5" pagination="412 - 414" refId="ref11152" refString="Rossman, D. A., and J. R. Dixon. 1975. A new colubrid snake of the genus Helicops from Peru. Herpetologica 31: 412 - 414." type="journal article" year="1975">Rossman and Dixon (1975)</bibRefCitation>
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, 
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, and the present study.
</tableNote>
</paragraph>
<caption id="DF03660C0C7EB96018CB0804C80E220D" ID-Table-UUID="DF03660C0C7EB96018CB0804C80E220D" box="[692,1321,1061,1082]" httpUri="http://table.plazi.org/id/DF03660C0C7EB96018CB0804C80E220D" pageId="5" startId="5.[692,705,1061,1081]" targetBox="[96,1908,1111,1716]" targetIsTable="true" targetPageId="5">
<paragraph id="8BC336840C7EB96018CB0804C80E220D" blockId="5.[692,1321,1061,1082]" box="[692,1321,1061,1082]" pageId="5">
TABLE 4.—Descriptions of the color patterns seen in 
<taxonomicName id="4C7C4D070C7EB9601EEE0804C80E220D" box="[1169,1321,1061,1082]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B908EA960C7EB9601EEE0804C9FB220E" box="[1169,1244,1061,1082]" italics="true" pageId="5">Helicops</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BC336840C7EB9601A1F0876C87A2080" pageId="5">
<table id="F97CC4240C7E469A1A1F0876CA532080" box="[96,1908,1111,1716]" gridcols="4" gridrows="25" pageId="5">
<tr id="354C34C60C7E469A1A1F0876CA532252" box="[96,1908,1111,1126]" gridrow="0" pageId="5">
<th id="769D5DBA0C7E469A1A1F0876CDDF2252" box="[96,248,1111,1126]" gridcol="0" gridrow="0" pageId="5">Species</th>
<th id="769D5DBA0C7E469A1B730876C9272252" box="[268,1024,1111,1126]" gridcol="1" gridrow="0" pageId="5">Dorsal color pattern</th>
<th id="769D5DBA0C7E469A1E6E0876CA282252" box="[1041,1807,1111,1126]" gridcol="2" gridrow="0" pageId="5">Ventral color pattern</th>
<th id="769D5DBA0C7E469A1D5A0876CA532252" box="[1829,1908,1111,1126]" gridcol="3" gridrow="0" pageId="5">
Reference 
<emphasis id="B908EA960C7EB9601D140876CA542251" bold="true" box="[1899,1907,1111,1125]" pageId="5">*</emphasis>
</th>
</tr>
<tr id="354C34C60C7E469A1A1F085BCA5322BB" box="[96,1908,1146,1167]" gridrow="1" pageId="5">
<th id="769D5DBA0C7E469A1A1F085BCDDF22BB" box="[96,248,1146,1167]" gridcol="0" gridrow="1" pageId="5">
<emphasis id="B908EA960C7EB9601A1D085BCDF422BB" box="[98,211,1146,1167]" italics="true" pageId="5">H. angulatus</emphasis>
</th>
<td id="769D5DBA0C7E469A1B73085BC92722BB" box="[268,1024,1146,1167]" gridcol="1" gridrow="1" pageId="5">Olive gray, tan or olive brown, with reddish brown to dark brown crossbands.</td>
<td id="769D5DBA0C7E469A1E6E085BCA2822BB" box="[1041,1807,1146,1167]" gridcol="2" gridrow="1" pageId="5">Cream to light tan, with or without extensions of dorsal crossbands, making a checkered</td>
<td id="769D5DBA0C7E469A1D5A085BCA5322BB" box="[1829,1908,1146,1167]" gridcol="3" gridrow="1" pageId="5">11, 16</td>
</tr>
<tr id="354C34C60C7E469A1A1F08B3CA532292" box="[96,1908,1170,1190]" gridrow="2" pageId="5">
<th id="769D5DBA0C7E469A1A1F08B3CA532292" box="[96,1908,1170,1190]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="5">pattern.</th>
</tr>
<tr id="354C34C60C7E469A1A1F088BCA53228B" box="[96,1908,1194,1215]" gridrow="3" pageId="5">
<th id="769D5DBA0C7E469A1A1F088BCDDF228B" box="[96,248,1194,1215]" gridcol="0" gridrow="3" pageId="5">
<emphasis id="B908EA960C7EB9601A1D088BCD9F228B" box="[98,184,1194,1215]" italics="true" pageId="5">H. apiaka</emphasis>
</th>
<td id="769D5DBA0C7E469A1B73088BC927228B" box="[268,1024,1194,1215]" gridcol="1" gridrow="3" pageId="5">Approximately 15 dark crossbands.</td>
<td id="769D5DBA0C7E469A1E6E088BCA28228B" box="[1041,1807,1194,1215]" gridcol="2" gridrow="3" pageId="5">Creamy white with the extensions of large black dorsal crossbands.</td>
<td id="769D5DBA0C7E469A1D5A088BCA53228B" box="[1829,1908,1194,1215]" gridcol="3" gridrow="3" pageId="5">15</td>
</tr>
<tr id="354C34C60C7E469A1A1F08E3CA5322E3" box="[96,1908,1218,1239]" gridrow="4" pageId="5">
<th id="769D5DBA0C7E469A1A1F08E3CDDF22E3" box="[96,248,1218,1239]" gridcol="0" gridrow="4" pageId="5">
<emphasis id="B908EA960C7EB9601A1D08E3CDC822E3" box="[98,239,1218,1239]" italics="true" pageId="5">H. carinicaudus</emphasis>
</th>
<td id="769D5DBA0C7E469A1B7308E3C92722E3" box="[268,1024,1218,1239]" gridcol="1" gridrow="4" pageId="5">Uniformly dark-brown or black, sometimes with broad longitudinal stripes lighter</td>
<td id="769D5DBA0C7E469A1E6E08E3CA2822E3" box="[1041,1807,1218,1239]" gridcol="2" gridrow="4" pageId="5">Cream with two (sometimes three) medial rows of black semilunar marks.</td>
<td id="769D5DBA0C7E469A1D5A08E3CA5322E3" box="[1829,1908,1218,1239]" gridcol="3" gridrow="4" pageId="5">13, 16</td>
</tr>
<tr id="354C34C60C7E469A1A1F08FBCA5322DA" box="[96,1908,1242,1262]" gridrow="5" pageId="5">
<th id="769D5DBA0C7E469A1A1F08FBCA5322DA" box="[96,1908,1242,1262]" colspan="4" colspanRight="3" gridcol="0" gridrow="5" pageId="5">than ground color.</th>
</tr>
<tr id="354C34C60C7E469A1A1F08D3CA532333" box="[96,1908,1266,1287]" gridrow="6" pageId="5">
<th id="769D5DBA0C7E469A1A1F08D3CDDF2333" box="[96,248,1266,1287]" gridcol="0" gridrow="6" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1D08D3CD9D2333" box="[98,186,1266,1287]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="danieli">
<emphasis id="B908EA960C7EB9601A1D08D3CD9D2333" box="[98,186,1266,1287]" italics="true" pageId="5">H. danieli</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B7308D3C9272333" box="[268,1024,1266,1287]" gridcol="1" gridrow="6" pageId="5">Reddish-brown or olive-brown with 4–5 rows of alternating dark spots.</td>
<td id="769D5DBA0C7E469A1E6E08D3CA282333" box="[1041,1807,1266,1287]" gridcol="2" gridrow="6" pageId="5">Cream with two medial rows of black semilunar marks, sometimes fused midventrally.</td>
<td id="769D5DBA0C7E469A1D5A08D3CA532333" box="[1829,1908,1266,1287]" gridcol="3" gridrow="6" pageId="5">3, 9</td>
</tr>
<tr id="354C34C60C7E469A1A1F092BCA53232B" box="[96,1908,1290,1311]" gridrow="7" pageId="5">
<th id="769D5DBA0C7E469A1A1F092BCDDF232B" box="[96,248,1290,1311]" gridcol="0" gridrow="7" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1D092BCD9D232B" box="[98,186,1290,1311]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="gomesi">
<emphasis id="B908EA960C7EB9601A1D092BCD9D232B" box="[98,186,1290,1311]" italics="true" pageId="5">H. gomesi</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B73092BC927232B" box="[268,1024,1290,1311]" gridcol="1" gridrow="7" pageId="5">Olive-brown with irregular dark blotches.</td>
<td id="769D5DBA0C7E469A1E6E092BCA28232B" box="[1041,1807,1290,1311]" gridcol="2" gridrow="7" pageId="5">Yellowish-brown or olive, with the extensions of dark dorsal blotches laterally.</td>
<td id="769D5DBA0C7E469A1D5A092BCA53232B" box="[1829,1908,1290,1311]" gridcol="3" gridrow="7" pageId="5">2</td>
</tr>
<tr id="354C34C60C7E469A1A1F0903CA532303" box="[96,1908,1314,1335]" gridrow="8" pageId="5">
<th id="769D5DBA0C7E469A1A1F0903CDDF2303" box="[96,248,1314,1335]" gridcol="0" gridrow="8" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1D0903CDF22303" box="[98,213,1314,1335]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="hagmanni">
<emphasis id="B908EA960C7EB9601A1D0903CDF22303" box="[98,213,1314,1335]" italics="true" pageId="5">H. hagmanni</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B730903C9272303" box="[268,1024,1314,1335]" gridcol="1" gridrow="8" pageId="5">Gray-brown with alternating light and dark blotches.</td>
<td id="769D5DBA0C7E469A1E6E0903CA282303" box="[1041,1807,1314,1335]" gridcol="2" gridrow="8" pageId="5">Irregularly checkered light/red and dark color.</td>
<td id="769D5DBA0C7E469A1D5A0903CA532303" box="[1829,1908,1314,1335]" gridcol="3" gridrow="8" pageId="5">5</td>
</tr>
<tr id="354C34C60C7E469A1A1F0918CA53237A" box="[96,1908,1337,1358]" gridrow="9" pageId="5">
<th id="769D5DBA0C7E469A1A1F0918CDDF237A" box="[96,248,1337,1358]" gridcol="0" gridrow="9" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1D0918CDDF237A" box="[98,248,1337,1358]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="infrataeniatus">
<emphasis id="B908EA960C7EB9601A1D0918CDDF237A" box="[98,248,1337,1358]" italics="true" pageId="5">H. infrataeniatus</emphasis>
</taxonomicName>
</th>
<td id="769D5DBA0C7E469A1B730918C927237A" box="[268,1024,1337,1358]" gridcol="1" gridrow="9" pageId="5">Dark-brown with light-brown stripes.</td>
<td id="769D5DBA0C7E469A1E6E0918CA28237A" box="[1041,1807,1337,1358]" gridcol="2" gridrow="9" pageId="5">Cream or red with 1–3 dark stripes or darkly checkered.</td>
<td id="769D5DBA0C7E469A1D5A0918CA53237A" box="[1829,1908,1337,1358]" gridcol="3" gridrow="9" pageId="5">13, 16</td>
</tr>
<tr id="354C34C60C7E469A1A1F0970CA532352" box="[96,1908,1361,1382]" gridrow="10" pageId="5">
<th id="769D5DBA0C7E469A1A1F0970CDDF2352" box="[96,248,1361,1382]" gridcol="0" gridrow="10" pageId="5">
<emphasis id="B908EA960C7EB9601A1D0970CDC22352" box="[98,229,1361,1382]" italics="true" pageId="5">H. leopardinus</emphasis>
</th>
<td id="769D5DBA0C7E469A1B730970C9272352" box="[268,1024,1361,1382]" gridcol="1" gridrow="10" pageId="5">Medium brown with dark brown or black blotches.</td>
<td id="769D5DBA0C7E469A1E6E0970CA282352" box="[1041,1807,1361,1382]" gridcol="2" gridrow="10" pageId="5">Checkerboard of black and red, banded black and red, or a combination of both.</td>
<td id="769D5DBA0C7E469A1D5A0970CA532352" box="[1829,1908,1361,1382]" gridcol="3" gridrow="10" pageId="5">11, 16</td>
</tr>
<tr id="354C34C60C7E469A1A1F0948CA53234A" box="[96,1908,1385,1406]" gridrow="11" pageId="5">
<th id="769D5DBA0C7E469A1A1F0948CDDF234A" box="[96,248,1385,1406]" gridcol="0" gridrow="11" pageId="5">
<emphasis id="B908EA960C7EB9601A1D0948CDE8234A" box="[98,207,1385,1406]" italics="true" pageId="5">H. modestus</emphasis>
</th>
<td id="769D5DBA0C7E469A1B730948C927234A" box="[268,1024,1385,1406]" gridcol="1" gridrow="11" pageId="5">Dark brown with 2–3 narrow light brown stripes, or uniformly dark olive.</td>
<td id="769D5DBA0C7E469A1E6E0948CA28234A" box="[1041,1807,1385,1406]" gridcol="2" gridrow="11" pageId="5">Cream, with or without faint brown flecks.</td>
<td id="769D5DBA0C7E469A1D5A0948CA53234A" box="[1829,1908,1385,1406]" gridcol="3" gridrow="11" pageId="5">4, 15, 16</td>
</tr>
<tr id="354C34C60C7E469A1A1F09A0CA5323A2" box="[96,1908,1409,1430]" gridrow="12" pageId="5">
<th id="769D5DBA0C7E469A1A1F09A0CDDF23A2" box="[96,248,1409,1430]" gridcol="0" gridrow="12" pageId="5">
<emphasis id="B908EA960C7EB9601A1D09A0CDEE23A2" box="[98,201,1409,1430]" italics="true" pageId="5">H. pastazae</emphasis>
</th>
<td id="769D5DBA0C7E469A1B7309A0C92723A2" box="[268,1024,1409,1430]" gridcol="1" gridrow="12" pageId="5">Tan to gray brown with four or five rows of alternating dark spots, the paravertebral</td>
<td id="769D5DBA0C7E469A1E6E09A0CA2823A2" box="[1041,1807,1409,1430]" gridcol="2" gridrow="12" pageId="5">Cream with a series of dark crossbands or alternating checks, the light ventral color</td>
<td id="769D5DBA0C7E469A1D5A09A0CA5323A2" box="[1829,1908,1409,1430]" gridcol="3" gridrow="12" pageId="5">7</td>
</tr>
<tr id="354C34C60C7E469A1A1F09B8CA532399" box="[96,1908,1433,1453]" gridrow="13" pageId="5" rowspan-0="1" rowspan-3="1">
<td id="769D5DBA0C7E469A1B7309B8C9272399" box="[268,1024,1433,1453]" gridcol="1" gridrow="13" pageId="5">spots large and often fused transversely.</td>
<td id="769D5DBA0C7E469A1E6E09B8CA282399" box="[1041,1807,1433,1453]" gridcol="2" gridrow="13" pageId="5">extending onto several dorsal scale rows.</td>
</tr>
<tr id="354C34C60C7E469A1A1F0990CA5323F2" box="[96,1908,1457,1478]" gridrow="14" pageId="5">
<th id="769D5DBA0C7E469A1A1F0990CDDF23F2" box="[96,248,1457,1478]" gridcol="0" gridrow="14" pageId="5">
<emphasis id="B908EA960C7EB9601A1D0990CD9E23F2" box="[98,185,1457,1478]" italics="true" pageId="5">H. petersi</emphasis>
</th>
<td id="769D5DBA0C7E469A1B730990C92723F2" box="[268,1024,1457,1478]" gridcol="1" gridrow="14" pageId="5">Dark olive brown with four or five rows of alternating dark spots.</td>
<td id="769D5DBA0C7E469A1E6E0990CA2823F2" box="[1041,1807,1457,1478]" gridcol="2" gridrow="14" pageId="5">Cream with a lateral series of dark checks.</td>
<td id="769D5DBA0C7E469A1D5A0990CA5323F2" box="[1829,1908,1457,1478]" gridcol="3" gridrow="14" pageId="5">7</td>
</tr>
<tr id="354C34C60C7E469A1A1F09E8CA5323EA" box="[96,1908,1481,1502]" gridrow="15" pageId="5">
<th id="769D5DBA0C7E469A1A1F09E8CDDF23EA" box="[96,248,1481,1502]" gridcol="0" gridrow="15" pageId="5">
<emphasis id="B908EA960C7EB9601A1D09E8CDEB23EA" box="[98,204,1481,1502]" italics="true" pageId="5">H. polŋlepis</emphasis>
</th>
<td id="769D5DBA0C7E469A1B7309E8C92723EA" box="[268,1024,1481,1502]" gridcol="1" gridrow="15" pageId="5">Brownish or olive gray, with blackish spots in five alternating rows.</td>
<td id="769D5DBA0C7E469A1E6E09E8CA2823EA" box="[1041,1807,1481,1502]" gridcol="2" gridrow="15" pageId="5">Dark with pale spots.</td>
<td id="769D5DBA0C7E469A1D5A09E8CA5323EA" box="[1829,1908,1481,1502]" gridcol="3" gridrow="15" pageId="5">1, 7</td>
</tr>
<tr id="354C34C60C7E469A1A1F09C0CA5323C2" box="[96,1908,1505,1526]" gridrow="16" pageId="5">
<th id="769D5DBA0C7E469A1A1F09C0CDDF23C2" box="[96,248,1505,1526]" gridcol="0" gridrow="16" pageId="5">
<emphasis id="B908EA960C7EB9601A1D09C0CD9823C2" box="[98,191,1505,1526]" italics="true" pageId="5">H. scalaris</emphasis>
</th>
<td id="769D5DBA0C7E469A1B7309C0C92723C2" box="[268,1024,1505,1526]" gridcol="1" gridrow="16" pageId="5">Grayish tan with 3–5 rows of irregular dark blotches, the vertebral blotches larger</td>
<td id="769D5DBA0C7E469A1E6E09C0CA2823C2" box="[1041,1807,1505,1526]" gridcol="2" gridrow="16" pageId="5">Cream medially with two rows of brown pigment forming either an irregular checkered</td>
<td id="769D5DBA0C7E469A1D5A09C0CA5323C2" box="[1829,1908,1505,1526]" gridcol="3" gridrow="16" pageId="5">10</td>
</tr>
<tr id="354C34C60C7E469A1A1F09D8CA532039" box="[96,1908,1529,1549]" gridrow="17" pageId="5" rowspan-0="1" rowspan-3="1">
<td id="769D5DBA0C7E469A1B7309D8C9272039" box="[268,1024,1529,1549]" gridcol="1" gridrow="17" pageId="5">than the laterals, and all 3 usually fused longitudinally on the neck to form short,</td>
<td id="769D5DBA0C7E469A1E6E09D8CA282039" box="[1041,1807,1529,1549]" gridcol="2" gridrow="17" pageId="5">pattern or broad ventrolateral stripes, at least anteriorly.</td>
</tr>
<tr id="354C34C60C7E469A1A1F0A30CA532011" box="[96,1908,1553,1573]" gridrow="18" pageId="5">
<th id="769D5DBA0C7E469A1A1F0A30CA532011" box="[96,1908,1553,1573]" colspan="4" colspanRight="3" gridcol="0" gridrow="18" pageId="5">broad stripes.</th>
</tr>
<tr id="354C34C60C7E469A1A1F0A08CA53200A" box="[96,1908,1577,1598]" gridrow="19" pageId="5">
<th id="769D5DBA0C7E469A1A1F0A08CDDF200A" box="[96,248,1577,1598]" gridcol="0" gridrow="19" pageId="5">
<emphasis id="B908EA960C7EB9601A1D0A08CDC5200A" box="[98,226,1577,1598]" italics="true" pageId="5">H. tapajonicus</emphasis>
</th>
<td id="769D5DBA0C7E469A1B730A08C927200A" box="[268,1024,1577,1598]" gridcol="1" gridrow="19" pageId="5">Uniformly moss-green.</td>
<td id="769D5DBA0C7E469A1E6E0A08CA28200A" box="[1041,1807,1577,1598]" gridcol="2" gridrow="19" pageId="5">Checkered light and dark brown.</td>
<td id="769D5DBA0C7E469A1D5A0A08CA53200A" box="[1829,1908,1577,1598]" gridcol="3" gridrow="19" pageId="5">12</td>
</tr>
<tr id="354C34C60C7E469A1A1F0A60CA532062" box="[96,1908,1601,1622]" gridrow="20" pageId="5">
<th id="769D5DBA0C7E469A1A1F0A60CDDF2062" box="[96,248,1601,1622]" gridcol="0" gridrow="20" pageId="5">
<emphasis id="B908EA960C7EB9601A1D0A60CDF22062" box="[98,213,1601,1622]" italics="true" pageId="5">H. triƲittatus</emphasis>
</th>
<td id="769D5DBA0C7E469A1B730A60C9272062" box="[268,1024,1601,1622]" gridcol="1" gridrow="20" pageId="5">Medium brown with five narrow longitudinal light stripes.</td>
<td id="769D5DBA0C7E469A1E6E0A60CA282062" box="[1041,1807,1601,1622]" gridcol="2" gridrow="20" pageId="5">Cream with two medial rows of black semilunar markings.</td>
<td id="769D5DBA0C7E469A1D5A0A60CA532062" box="[1829,1908,1601,1622]" gridcol="3" gridrow="20" pageId="5">14</td>
</tr>
<tr id="354C34C60C7E469A1A1F0A79CA532059" box="[96,1908,1624,1645]" gridrow="21" pageId="5">
<th id="769D5DBA0C7E469A1A1F0A79CDDF2059" box="[96,248,1624,1645]" gridcol="0" gridrow="21" pageId="5">
<emphasis id="B908EA960C7EB9601A1D0A79CD802059" box="[98,167,1624,1645]" italics="true" pageId="5">H. ŋacu</emphasis>
</th>
<td id="769D5DBA0C7E469A1B730A79C9272059" box="[268,1024,1624,1645]" gridcol="1" gridrow="21" pageId="5">Light to medium gray-brown, with four alternating rows of relatively small dark</td>
<td id="769D5DBA0C7E469A1E6E0A79CA282059" box="[1041,1807,1624,1645]" gridcol="2" gridrow="21" pageId="5">Cream with an irregular series of large black checks or a median row of black crescents.</td>
<td id="769D5DBA0C7E469A1D5A0A79CA532059" box="[1829,1908,1624,1645]" gridcol="3" gridrow="21" pageId="5">6, 8</td>
</tr>
<tr id="354C34C60C7E469A1A1F0A50CA5320B1" box="[96,1908,1649,1669]" gridrow="22" pageId="5">
<th id="769D5DBA0C7E469A1A1F0A50CA5320B1" box="[96,1908,1649,1669]" colspan="4" colspanRight="3" gridcol="0" gridrow="22" pageId="5">spots.</th>
</tr>
<tr id="354C34C60C7E469A1A1F0AA9CA5320A9" box="[96,1908,1672,1693]" gridrow="23" pageId="5">
<th id="769D5DBA0C7E469A1A1F0AA9C92720A9" box="[96,1024,1672,1693]" colspan="2" colspanRight="1" gridcol="0" gridrow="23" pageId="5">
<taxonomicName id="4C7C4D070C7EB9601A1D0AA9CD9E20A9" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[98,185,1672,1693]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7EB9601A1D0AA9CD9E20A9" box="[98,185,1672,1693]" italics="true" pageId="5">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7EB9601ABF0AA9CC2120A8" box="[192,262,1672,1692]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
Uniformly dark-olive green, dark brown or dark gray.
</th>
<td id="769D5DBA0C7E469A1E6E0AA9CA2820A9" box="[1041,1807,1672,1693]" gridcol="2" gridrow="23" pageId="5">Yellow (in life), cream or light brown (sometimes mottled with gray) with two medial</td>
<td id="769D5DBA0C7E469A1D5A0AA9CA5320A9" box="[1829,1908,1672,1693]" gridcol="3" gridrow="23" pageId="5">16</td>
</tr>
<tr id="354C34C60C7E469A1A1F0A81CA532080" box="[96,1908,1696,1716]" gridrow="24" pageId="5" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="769D5DBA0C7E469A1E6E0A81CA282080" box="[1041,1807,1696,1716]" gridcol="2" gridrow="24" pageId="5">rows of black semilunar markings.</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC336840C7EB9601A080AE3CFEB20D0" blockId="5.[98,1915,1730,1764]" pageId="5">
<tableNote id="769A370A0C7EB9601A080AE3CFEB20D0" pageId="5" targetBox="[96,1908,1111,1716]" targetPageId="5">
<emphasis id="B908EA960C7EB9601A080AE3CD5820E4" bold="true" box="[119,127,1730,1744]" pageId="5">*</emphasis>
(1) 
<bibRefCitation id="EFED4B750C7EB9601AE40AE2CC2220E6" author="Boulenger, G. A." box="[155,261,1731,1746]" pageId="5" refId="ref9505" refString="Boulenger, G. A. 1893. Catalogue of the Snakes in the British Museum (Natural History). Volume I. Containing the Families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae Aglyphae, part. Order of the Trustees, British Museum, UK." type="book" year="1893">Boulenger 1893</bibRefCitation>
; (2) 
<bibRefCitation id="EFED4B750C7EB9601B5D0AE2CC5020E6" author="Amaral, A." box="[290,375,1731,1746]" pageId="5" pagination="1 - 37" refId="ref9327" refString="Amaral, A. 1922 [dated 1921]. Contribuicao para o conhecimento dos ofidios do Brasil. Parte I. Quatro novas especies de serpentes brasileiras. Memorias do Instituto Butantan, 1: 1 - 37. [In Portuguese.]" type="journal article" year="1922">Amaral 1922</bibRefCitation>
; (3) 
<bibRefCitation id="EFED4B750C7EB9601BEB0AE2CCCE20E6" author="Amaral, A." box="[404,489,1731,1746]" pageId="5" pagination="232 - 240" refId="ref9372" refString="Amaral, A. 1938. Estudo sobre ophidios neotropicos 34. Novas notas sobre a fauna da Colombia e descripcao de uma especie nova de Colubrideo aglypho. Memorias do Instituto Butantan 11: 232 - 240. [In Portuguese.]" type="journal article" year="1938">Amaral 1938</bibRefCitation>
; (4) 
<bibRefCitation id="EFED4B750C7EB96018780AE2CF4020E6" author="Rossman, D. A." box="[519,615,1731,1746]" pageId="5" pagination="189 - 191" refId="ref10855" refString="Rossman, D. A. 1974. Miscellaneous notes on the South American water snakes genus Helicops. HISS News-Journal 1: 189 - 191." type="journal article" year="1974">Rossman 1974</bibRefCitation>
; (5) Rossman 1975; (6) 
<bibRefCitation id="EFED4B750C7EB960197C0AE2CE8A20E6" author="Rossman, D. A. &amp; J. R. Dixon" box="[771,941,1731,1746]" pageId="5" pagination="412 - 414" refId="ref11152" refString="Rossman, D. A., and J. R. Dixon. 1975. A new colubrid snake of the genus Helicops from Peru. Herpetologica 31: 412 - 414." type="journal article" year="1975">Rossman and Dixon 1975</bibRefCitation>
; (7) 
<bibRefCitation id="EFED4B750C7EB96019B50AE2C90C20E6" author="Rossman, D. A." box="[970,1067,1731,1746]" pageId="5" pagination="1 - 13" refId="ref10908" refString="Rossman, D. A. 1976. Revision of the South American colubrid snakes of the Helicops pastazae complex. Occasional Papers of the Museum of Zoology, Louisiana State University 50: 1 - 13." type="journal article" year="1976">Rossman 1976</bibRefCitation>
; (8) 
<bibRefCitation id="EFED4B750C7EB9601E370AE2C9C320E6" author="Rossman, D. A. &amp; A. S. Abe" box="[1096,1252,1731,1746]" pageId="5" pagination="7 - 9" refId="ref11109" refString="Rossman, D. A., and A. S. Abe. 1979. Comments on the taxonomic status of Helicops nacu (Serpentes: Colubridae). Proceedings of the Louisiana Academy of Science 42: 7 - 9." type="journal article" year="1979">Rossman and Abe 1979</bibRefCitation>
; (9) 
<bibRefCitation id="EFED4B750C7EB9601F7D0AE2C84D20E6" author="Rossman, D. A." box="[1282,1386,1731,1746]" pageId="5" pagination="589 - 594" refId="ref10975" refString="Rossman, D. A. 2002 a. Morphological variation in the endemic Colombian water snake, Helicops danieli Amaral, 1937 (Serpentes: Xenodontidae). Revista da Academia Colombiana Ciencias 26: 589 - 594." type="journal article" year="2002">Rossman 2002a</bibRefCitation>
; (10) 
<bibRefCitation id="EFED4B750C7EB9601FF00AE2C8DE20E6" author="Rossman, D. A." box="[1423,1529,1731,1746]" pageId="5" pagination="1 - 18" refId="ref11015" refString="Rossman, D. A. 2002 b. Variation in the xenodontid water snake Helicops scalaris Jan, and the status of H. hogei Lancini. Occasional Papers of the Museum of Natural Science, Louisiana State University 78: 1 - 18." type="journal article" year="2002">Rossman 2002b</bibRefCitation>
; (11) 
<bibRefCitation id="EFED4B750C7EB9601C610AE2CBEF20E6" author="Bartlett, R. D. &amp; P. Bartlett" box="[1566,1736,1731,1746]" pageId="5" refId="ref9416" refString="Bartlett, R. D., and P. Bartlett. 2003. Reptiles and Amphibians of the Amazon: An Ecotourist ̕ s Guide. University Press of Florida, USA." type="book" year="2003">Bartlett and Bartlett 2003</bibRefCitation>
; (12) 
<bibRefCitation id="EFED4B750C7EB9601C910AE2CA1020E6" author="Frota, J. G." box="[1774,1847,1731,1746]" pageId="5" pagination="61 - 67" refId="ref9998" refString="Frota, J. G. 2005. Nova especie de Helicops Wagler, 1830 (Serpentes, Colubridae) do rio Tapajos, Amazonia, Brasil. Phyllomedusa 4: 61 - 67. [In Portuguese.]" type="journal article" year="2005">Frota 2005</bibRefCitation>
; (13) 
<bibRefCitation id="EFED4B750C7EB9601D220AE2CDEC20D0" author="Yuki, R. N. &amp; T. Lema" pageId="5" pagination="85 - 128" refId="ref11677" refString="Yuki, R. N., and T. Lema. 2005. Analise comparativa entre as cobras d ̕ agua meridionais (Helicops carinicaudus (Wied, 1825) e Helicops infrataeniatus Jan, 1865), com a descricao do cranio e hemipenis (Serpentes: Colubridae: Xenodontinae). Comunicacoes do Museu de Ciencia e Tecnologa da PUCRS, Serie Zoologia 18: 85 - 128. [In Portuguese.]" type="journal article" year="2005">Yuki and Lema 2005</bibRefCitation>
; (14) 
<bibRefCitation id="EFED4B750C7EB9601A8C0AF4CC7220D0" author="Rossman, D. A." box="[243,341,1749,1764]" pageId="5" pagination="271 - 278" refId="ref11061" refString="Rossman, D. A. 2010. Morphological variation in the striped water snake Helicops triVittatus (Gray, 1849) (Reptilia: Serpentes: Xenodontidae) of eastern Amazonia. Boletim do Museu Paraense Emilio Goeldi, Ciencias Naturais 5: 271 - 278." type="journal article" year="2010">Rossman 2010</bibRefCitation>
; (15) 
<bibRefCitation id="EFED4B750C7EB9601B020AF4CF6320D0" author="Kawashita-Ribeiro, R. A. &amp; R. W. Avila &amp; D. H. Morais" box="[381,580,1749,1764]" pageId="5" pagination="80 - 90" refId="ref10310" refString="Kawashita-Ribeiro, R. A., R. W. Avila, and D. H. Morais. 2013. A new snake of the genus Helicops Wagler, 1830 (Dipsadidae, Xenodontinae) from Brazil. Herpetologica 69: 80 - 90." type="journal article" year="2013">Kawashita-Ribeiro et al. 2013</bibRefCitation>
; (16) present study.
</tableNote>
</paragraph>
<paragraph id="8BC336840C7DB9631AEA0ECBCF302212" blockId="6.[122,792,190,2001]" pageId="6">
<emphasis id="B908EA960C7DB9631AEA0ECBCC9B2536" bold="true" box="[149,444,746,770]" pageId="6">
Color in life of 
<typeStatus id="54C788260C7DB9631B340ECBCC902536" box="[331,439,746,770]" pageId="6" type="holotype">holotype</typeStatus>
.
</emphasis>
—Dorsal and lateral surfaces of head dark olive-green; supralabials dark olive-green above, becoming yellowish-green below; infralabials and gular region yellow with scattered olive-green pigment. Ventrals yellow, with two medial rows of black round or semilunar marks (two marks on each ventral); gray pigment between the black marks on each ventral anteriorly; gray pigment more intense and covering entire ventral surface posteriorly. Subcaudals gray, with black medial markings on subcaudals 1–6. Dorsal ground color dark olive-green; DSR1 yellow with olive-green pigments anteriorly.
</paragraph>
<paragraph id="8BC336840C7DB9631AEA080DCF9223B2" blockId="6.[122,792,190,2001]" pageId="6">
<emphasis id="B908EA960C7DB9631AEA080DCF942270" bold="true" box="[149,691,1068,1093]" pageId="6">
Color in preservative of 
<typeStatus id="54C788260C7DB9631BA5080CCF632271" box="[474,580,1069,1093]" pageId="6" type="holotype">holotype</typeStatus>
(
<figureCitation id="13472A010C7DB9631824080DCF852270" box="[603,674,1068,1092]" captionStart="FIG" captionStartId="2.[845,859,1933,1952]" captionTargetBox="[824,1496,1411,1915]" captionTargetId="figure-779@2.[824,1496,1411,1915]" captionTargetPageId="2" captionText="FIG. 1.—Holotype of Helicops nentur sp. nov. (UFMG 2486): dorsal (A) and ventral (B) view. Scale bar = 20 mm. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712203" httpUri="https://zenodo.org/record/7712203/files/figure.png" pageId="6">Fig. 1</figureCitation>
).
</emphasis>
—Dorsal and lateral surfaces of head dark gray; supralabials 1–5 light gray; supralabials 6–7 cream with gray pigmentation, more intense above; supralabial 8 dark gray above and cream below; infralabials and gular region cream with scattered gray pigment. Ventrals cream, with two medial rows of black round or semilunar marks (two marks on each ventral); gray pigment between the black marks on each ventral anteriorly; gray pigment becoming more intense and covering entire ventral surface posteriorly. Subcaudals grayish, with black medial markings on subcaudals 1–6. Dorsal ground color dark gray; DSR1 cream with gray flecks anteriorly.
</paragraph>
<caption id="DF03660C0C7DB9631932084FC9BA2284" ID-DOI="http://doi.org/10.5281/zenodo.7712209" ID-Zenodo-Dep="7712209" httpUri="https://zenodo.org/record/7712209/files/figure.png" pageId="6" startId="6.[845,859,1134,1153]" targetBox="[824,1495,189,1116]" targetPageId="6">
<paragraph id="8BC336840C7DB9631932084FC9BA2284" blockId="6.[824,1494,1133,1200]" pageId="6">
FIG. 4.—Paratype (ZUFMS-REP 2291) of 
<taxonomicName id="4C7C4D070C7DB9631E95084CC85A22B4" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1258,1405,1133,1152]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7DB9631E95084CC85A22B4" box="[1258,1405,1133,1152]" italics="true" pageId="6">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7DB9631FF5084FC8F122B5" box="[1418,1494,1134,1153]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
immediately postmortem in (A) dorsal view and (B) ventral view. A color version of this figure is available online.
</paragraph>
</caption>
<paragraph id="8BC336840C7DB9631AEA09ADCF2D21A1" blockId="6.[122,792,190,2001]" pageId="6">
<emphasis id="B908EA960C7DB9631AEA09ADCCFC2390" bold="true" box="[149,475,1420,1444]" pageId="6">Description of hemipenis.</emphasis>
—Based on the right hemipenis of the male 
<typeStatus id="54C788260C7DB9631B33098BCC8C23F6" box="[332,427,1450,1474]" pageId="6" type="paratype">paratype</typeStatus>
(MZUFV 960), which was fully everted and almost maximally expanded (
<figureCitation id="13472A010C7DB963183B09E6CFAE23EB" box="[580,649,1479,1503]" captionStart="FIG" captionStartId="3.[165,179,1909,1928]" captionTargetBox="[143,814,1386,1891]" captionTargetId="figure-374@3.[143,815,1386,1891]" captionTargetPageId="3" captionText="FIG. 3.—Right hemipenis of a paratype of Helicops nentur sp. nov. (MZUFV 960) fully everted and almost maximally expanded: sulcate (A) and asulcate (B) side. Scale bar = 10 mm. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712207" httpUri="https://zenodo.org/record/7712207/files/figure.png" pageId="6">Fig. 3</figureCitation>
). In situ, the everted organ extends to the 14th subcaudal. Hemipenis bilobed, semicalyculate, and semicapitate; body 
<quantity id="4C849B610C7DB96318CF0A23CE30202D" box="[688,791,1538,1562]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="6" unit="mm" value="12.9">12.9 mm</quantity>
long, with well-developed lobes (
<quantity id="4C849B610C7DB9631B9A0A3ECF1A2002" box="[485,573,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" unit="mm" value="7.5">7.5 mm</quantity>
; 58% of hemipenial body length), distinct from the body by a constriction from lateral to asulcate side; lobular crotch with a line of small spines; lobes covered by encircling rows of horizontally oriented spinulate flounces, except at the apices, which are nude; sulcate side covered by several spinules and some small spines arranged in more or less oblique rows; sulcus spermaticus bifurcating in the proximal region, with branches having a S-shaped condition; the asulcate side is covered with spinules and bordered by lateral spines increasing in size from the base to the apex; a small oblique nude area is present in the left and right portions of the asulcate side, close to the edge of capitulum; a basal pocket is present.
</paragraph>
<paragraph id="8BC336840C7DB9631AEA0BBDCEF3238B" blockId="6.[122,792,190,2001]" lastBlockId="6.[824,1493,1242,2001]" pageId="6">
<emphasis id="B908EA960C7DB9631AEA0BBDCC2A2180" bold="true" box="[149,269,1948,1972]" pageId="6">Variation.</emphasis>
—The single male 
<quantity id="4C849B610C7DB9631B960BBDCF602187" box="[489,583,1948,1972]" metricMagnitude="-1" metricUnit="m" metricValue="1.54" pageId="6" unit="mm" value="154.0">154 mm</quantity>
SVL, 
<quantity id="4C849B610C7DB96318EB0BBDCFC32187" box="[660,740,1948,1972]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="6" unit="mm" value="45.0">45 mm</quantity>
TL; largest female UFMG 
<quantity id="4C849B610C7DB9631BF60B98CF0121E5" box="[393,550,1977,2001]" metricMagnitude="2" metricUnit="m" metricValue="1.7565" pageId="6" unit="mm" value="175650.0">175, 650 mm</quantity>
SVL, 140+ mm TL (broken tail); tail 33.9% SVL in the male, 23–29.2% SVL (mean = 25.56 ± 3.23%, 
<emphasis id="B908EA960C7DB9631E2908D9C943233B" box="[1110,1124,1272,1295]" italics="true" pageId="6">n</emphasis>
= 3) in females (UFMG 175 not included); ventrals in the male 115, in females 111–117 (mean = 115.2 ± 2.87, 
<emphasis id="B908EA960C7DB9631E410913C96B237D" box="[1086,1100,1330,1353]" italics="true" pageId="6">n</emphasis>
= 4); subcaudals 
<quantity id="4C849B610C7DB9631F6B0910C86C237D" box="[1300,1355,1329,1353]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="6" unit="in" value="56.0">56 in</quantity>
the male, in females 41–52 (mean = 45.6 ± 5.68, 
<emphasis id="B908EA960C7DB9631EAA0971C9C42353" box="[1237,1251,1360,1383]" italics="true" pageId="6">n</emphasis>
= 3); preventrals 1–2 (mean = 1.40 ± 0.54, 
<emphasis id="B908EA960C7DB9631E43094CC96D23B0" box="[1084,1098,1389,1412]" italics="true" pageId="6">n</emphasis>
= 5); prediastemal maxillary teeth 18–21 (mean = 19.40 ± 1.14, 
<emphasis id="B908EA960C7DB9631EF909ABC9B32395" box="[1158,1172,1418,1441]" italics="true" pageId="6">n</emphasis>
= 5); infralabials on left side in the male 9.
</paragraph>
<paragraph id="8BC336840C7DB963192D09E4C97820F3" blockId="6.[824,1493,1242,2001]" pageId="6">The nasals are undivided in all specimens, with two exceptions: The right nasal of UFMG 2486 is semidivided and the right nasal of MZUFV 742 shows a small sulcus that does not reach the lower part of the nostril. All specimens have two anterior temporals, the upper longer than the lower. Usually there are two posterior temporals unequal in size, but MZUFV 742 has three posterior temporals on the right side, whereas MZUFV 960 has three posterior temporals on the left side.</paragraph>
<paragraph id="8BC336840C7DB963192D0AEFC8B921A2" blockId="6.[824,1493,1242,2001]" pageId="6">
Color pattern is similar among all examined specimens of 
<taxonomicName id="4C7C4D070C7DB96319470ACDCE882137" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[824,943,1772,1795]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7DB96319470ACDCE882137" box="[824,943,1772,1795]" italics="true" pageId="6">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7DB96319BD0ACAC9032136" box="[962,1060,1771,1794]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
, with few differences, probably of ontogenetic origin. The two smallest specimens—MZUFV 960 and ZUFMS-REP 2291—have less gray pigment in the medial region of the ventrals. In life, the venter was cream in the gular region, yellow on the anterior half of body, and light orange posteriorly in ZUFMS-REP 2291 (
<figureCitation id="13472A010C7DB9631F330B5CC8B621A2" box="[1356,1425,1917,1942]" captionStart="FIG" captionStartId="6.[845,859,1134,1153]" captionTargetBox="[824,1495,189,1116]" captionTargetId="figure-796@6.[824,1496,189,1116]" captionTargetPageId="6" captionText="FIG. 4.—Paratype (ZUFMS-REP 2291) of Helicops nentur sp. nov. immediately postmortem in (A) dorsal view and (B) ventral view. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712209" httpUri="https://zenodo.org/record/7712209/files/figure.png" pageId="6">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366650F0C7DB962192D0BBDCDCF2392" lastPageId="7" pageId="6" type="biology_ecology">
<paragraph id="8BC336840C7DB962192D0BBDCDCF2392" blockId="6.[824,1493,1242,2001]" lastBlockId="7.[143,813,1217,2001]" lastPageId="7" pageId="6">
<emphasis id="B908EA960C7DB963192D0BBDC9E42180" bold="true" box="[850,1219,1948,1972]" pageId="6">Natural history and habitat.</emphasis>
—Available information (from the 
<typeStatus id="54C788260C7DB96319D90B98C92221E5" box="[934,1029,1977,2001]" pageId="6" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D7AFD0D0C7DB9631E450B98C9ED21E5" box="[1082,1226,1977,2001]" pageId="6" type="generic" typeStatus="paratype">one paratype</specimenCount>
) suggests that 
<taxonomicName id="4C7C4D070C7DB9631F150B9BC8F221E5" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1386,1493,1978,2001]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7DB9631F150B9BC8F221E5" box="[1386,1493,1978,2001]" italics="true" pageId="6">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB9621AF008E3CDCE22ED" box="[143,233,1218,1241]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
inhabits lentic water bodies such as swamps. The 
<typeStatus id="54C788260C7CB9621AF008FFCDC922C2" box="[143,238,1246,1270]" pageId="7" type="holotype">holotype</typeStatus>
was collected while active at 0300 h, partially submerged. After collection, it gave birth to two stillborn specimens not measured nor preserved. 
<specimenCode id="DBDA9EFF0C7CB962182A0938CE0A2305" box="[597,813,1305,1329]" collectionCode="ZUFMS-REP" pageId="7">ZUFMS-REP 2291</specimenCode>
has an umbilical scar on ventrals 91–95. Date of collection of these 
<specimenCount id="9D7AFD0D0C7CB9621AAD0975CC52235F" box="[210,373,1363,1387]" pageId="7" type="generic">two specimens</specimenCount>
suggest that reproduction in 
<taxonomicName id="4C7C4D070C7CB96218BF0975CE0A235F" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[704,813,1364,1387]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7CB96218BF0975CE0A235F" box="[704,813,1364,1387]" italics="true" pageId="7">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB9621AF00950CDCB23BC" box="[143,236,1393,1416]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
might occur during the rainy season (October– March).
</paragraph>
</subSubSection>
<caption id="DF03660C0C7CB9621ACE08AAC8ED22A9" ID-DOI="http://doi.org/10.5281/zenodo.7712211" ID-Zenodo-Dep="7712211" box="[177,1482,1162,1182]" httpUri="https://zenodo.org/record/7712211/files/figure.png" pageId="7" startId="7.[177,191,1163,1182]" targetBox="[145,1520,190,1145]" targetPageId="7">
<paragraph id="8BC336840C7CB9621ACE08AAC8ED22A9" blockId="7.[177,1482,1162,1182]" box="[177,1482,1162,1182]" pageId="7">
FIG. 5.—Locality records of 
<taxonomicName id="4C7C4D070C7CB9621BCF08ABCF1A22A9" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[432,573,1162,1181]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7CB9621BCF08ABCF1A22A9" box="[432,573,1162,1181]" italics="true" pageId="7">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB962183B08AACFAD22AA" box="[580,650,1163,1182]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
in the state of Minas Gerais, Brazil (insert). A color version of this figure is available online.
</paragraph>
</caption>
<subSubSection id="C366650F0C7CB9621AD5098ACC12214D" pageId="7" type="distribution">
<paragraph id="8BC336840C7CB9621AD5098ACC12214D" blockId="7.[143,813,1217,2001]" pageId="7">
<emphasis id="B908EA960C7CB9621AD5098ACCFC23F7" bold="true" box="[170,475,1451,1475]" pageId="7">Geographic distribution.</emphasis>
— 
<taxonomicName id="4C7C4D070C7CB9621B89098ACF8D23F7" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[502,682,1451,1475]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7CB9621B89098ACF8D23F7" box="[502,682,1451,1475]" italics="true" pageId="7">Helicops nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB96218C9098ACE3723F6" box="[694,784,1451,1474]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
is known only from five localities in the Brazilian state of 
<collectingRegion id="49B8F8660C7CB9621AF009C4CC0B23C9" box="[143,300,1509,1533]" country="Brazil" name="Minas Gerais" pageId="7">Minas Gerais</collectingRegion>
: Belo Horizonte, Jaíba, Muriaé, São José da Safira, and Viçosa (
<figureCitation id="13472A010C7CB9621B130A22CC94202E" box="[364,435,1538,1563]" captionStart="FIG" captionStartId="7.[177,191,1163,1182]" captionTargetBox="[145,1520,190,1145]" captionTargetId="figure-456@7.[143,1522,189,1145]" captionTargetPageId="7" captionText="FIG. 5.—Locality records of Helicops nentur sp. nov. in the state of Minas Gerais, Brazil (insert). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712211" httpUri="https://zenodo.org/record/7712211/files/figure.png" pageId="7">Fig. 5</figureCitation>
). The distribution of this species (sensu 
<bibRefCitation id="EFED4B750C7CB9621A9B0A01CC96200C" author="Olson, D. M. &amp; E. Dinerstein &amp; E. D. Wikramanayake &amp; K. R. Kassem" box="[228,433,1568,1592]" pageId="7" pagination="933 - 938" refId="ref10728" refString="Olson, D. M., E. Dinerstein, E. D. Wikramanayake, ..., K. R. Kassem. 2001. Terrestrial ecoregions of the world: A new map of life on Earth. Bioscience 51: 933 - 938." type="journal article" year="2001">Olson et al. 2001</bibRefCitation>
) is in the 
<collectingRegion id="49B8F8660C7CB962184C0A01CF54200C" box="[563,627,1568,1592]" country="Brazil" name="Bahia" pageId="7">Bahia</collectingRegion>
Interior Forest ecoregion (Tropical and Subtropical Moist Broadleaf Forests biome), its border with the Cerrado ecoregion (Tropical and Subtropical Grasslands, Savannas, and Shrublands biome), and the ecotone between the Atlantic Dry Forest and the Caatinga ecoregions (Tropical and Subtropical Dry Broadleaf Forests biome and Deserts and Xeric Shrublands biome). This indicates that 
<taxonomicName id="4C7C4D070C7CB96218E90ACCCE222130" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[662,773,1773,1796]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7CB96218E90ACCCE222130" box="[662,773,1773,1796]" italics="true" pageId="7">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB96219720ACCCDE72115" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
might occupy different environments and climatic conditions. The distance between collection localities also indicates that there are many distribution gaps in the range of the species.
</paragraph>
</subSubSection>
<subSubSection id="C366650F0C7CB9621AD50B5ECEE12359" pageId="7" type="etymology">
<paragraph id="8BC336840C7CB9621AD50B5ECEE12359" blockId="7.[143,813,1217,2001]" lastBlockId="7.[845,1515,1218,1744]" pageId="7">
<emphasis id="B908EA960C7CB9621AD50B5ECC1321A3" bold="true" box="[170,308,1919,1943]" pageId="7">Etymology.</emphasis>
—The specific name 
<taxonomicName id="4C7C4D070C7CB96218570B5ECF5521A2" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[552,626,1919,1942]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur">
<emphasis id="B908EA960C7CB96218570B5ECF5521A2" box="[552,626,1919,1942]" italics="true" pageId="7">nentur</emphasis>
</taxonomicName>
has its origin in Quenya, a fictional language created by the late writer and philologist J.R.R. Tolkien. In Tolkien̕s mythology, Quenya is spoken by the Eldar, the High Elves of Valinor, which is a land in the blessed continent of Aman. The name 
<taxonomicName id="4C7C4D070C7CB9621FF608C1C8F422C3" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1417,1491,1248,1271]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur">
<emphasis id="B908EA960C7CB9621FF608C1C8F422C3" box="[1417,1491,1248,1271]" italics="true" pageId="7">nentur</emphasis>
</taxonomicName>
is formed by the words 
<emphasis id="B908EA960C7CB9621E3208DCC9522320" box="[1101,1141,1277,1300]" italics="true" pageId="7">nen</emphasis>
(water) and 
<emphasis id="B908EA960C7CB9621F6E08DCC8142320" box="[1297,1331,1277,1300]" italics="true" pageId="7">tur</emphasis>
(ruler, master), meaning “master of waters,” which is a reference to the aquatic habits of 
<taxonomicName id="4C7C4D070C7CB9621E720919C94C2364" authorityName="Wagler" authorityYear="1828" box="[1037,1131,1336,1360]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="genus">
<emphasis id="B908EA960C7CB9621E720919C94C2364" box="[1037,1131,1336,1360]" italics="true" pageId="7">Helicops</emphasis>
</taxonomicName>
. The name is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C366650F0C7CB96219180955CE9720E4" pageId="7" type="discussion">
<paragraph id="8BC336840C7CB96219180955CE9720E4" blockId="7.[845,1515,1218,1744]" pageId="7">
<emphasis id="B908EA960C7CB96219180955CECD239D" bold="true" pageId="7">Molecular phylogeny and genetic divergence of the new species.</emphasis>
—Sequence divergences between 
<taxonomicName id="4C7C4D070C7CB9621F0609B3C8CC239D" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1401,1515,1426,1449]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7CB9621F0609B3C8CC239D" box="[1401,1515,1426,1449]" italics="true" pageId="7">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB9621932098ECE8423F2" box="[845,931,1455,1478]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
and other congeners range between 4.0% and 6.0% (mean 4.8%) for 16S rRNA and between 0.0% and 0.4% (mean 0.2%) for C-mos (
<tableCitation id="C6FE033F0C7CB9621E1809C8C9982035" box="[1127,1215,1513,1537]" captionStart="TABLE" captionStartId="8.[143,156,194,213]" captionTargetPageId="8" captionText="TABLE 5.—Uncorrected pairwise distances of nucleotide substitution between 16S rRNA (below the diagonal) and C-mos (above the diagonal) sequences of some Helicops species." httpUri="http://table.plazi.org/id/DF03660C0C73B96D1AF00CE3CC7826D8" pageId="7" tableUuid="DF03660C0C73B96D1AF00CE3CC7826D8">Table 5</tableCitation>
). Among those species for which 16S genetic data are available, 
<taxonomicName id="4C7C4D070C7CB9621E810A29C84A202B" authority="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia, 2016" authorityName="Costa &amp; Santana &amp; Leal &amp; Koroiva &amp; Garcia" authorityYear="2016" box="[1278,1389,1544,1567]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="nentur" status="sp. nov.">
<emphasis id="B908EA960C7CB9621E810A29C84A202B" box="[1278,1389,1544,1567]" italics="true" pageId="7">H. nentur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57ED0C7CB9621F080A26C8F7202A" box="[1399,1488,1543,1566]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
is sister to a clade formed by 
<emphasis id="B908EA960C7CB9621EFD0A04C89F2008" box="[1154,1464,1572,1596]" italics="true" pageId="7">
<taxonomicName id="4C7C4D070C7CB9621EFD0A04C8312008" box="[1154,1302,1572,1596]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="hagmanni">H. hagmanni</taxonomicName>
, 
<taxonomicName id="4C7C4D070C7CB9621F5C0A04C8932008" box="[1315,1460,1572,1596]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="angulatus">H. angulatus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7C4D070C7CB96219320A62CE99206E" box="[845,958,1603,1626]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="gomesi">
<emphasis id="B908EA960C7CB96219320A62CE99206E" box="[845,958,1603,1626]" italics="true" pageId="7">H. gomesi</emphasis>
</taxonomicName>
(
<figureCitation id="13472A010C7CB96219AD0A63C90F206E" box="[978,1064,1602,1626]" captionStart="FIG" captionStartId="8.[143,157,1934,1953]" captionTargetBox="[128,1499,1186,1914]" captionTargetId="figure-515@8.[122,1500,1182,1916]" captionTargetPageId="8" captionText="FIG. 6.—Phylogenetic relationships of sampled species of Helicops based on (A) 16s rRNA and (B) C-mos sequence data. Numbers at nodes indicate the bootstrap values/posterior probabilities; the bars below each cladogram indicate base substitution rate for each gene (# of substitutions per site). Hŋdrops triangularis and Pseudoerŋx plicatilis were used as outgroups." figureDoi="http://doi.org/10.5281/zenodo.7712213" httpUri="https://zenodo.org/record/7712213/files/figure.png" pageId="7">Fig. 6A</figureCitation>
) with high support (bootstrap value = 81%, and posterior probability = 0.99). A partial C-mos tree shows the new species as sister to 
<taxonomicName id="4C7C4D070C7CB9621EAE0A5FC8A420A1" box="[1233,1411,1661,1685]" class="Reptilia" family="Dipsadidae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" phylum="Chordata" rank="species" species="carinicaudus">
<emphasis id="B908EA960C7CB9621EAE0A5FC8A420A1" box="[1233,1411,1661,1685]" italics="true" pageId="7">H. carinicaudus</emphasis>
</taxonomicName>
with low bootstrap value (64%) but high posterior probability (0.99; 
<figureCitation id="13472A010C7CB96219320A99CE8420E4" box="[845,931,1720,1744]" captionStart="FIG" captionStartId="8.[143,157,1934,1953]" captionTargetBox="[128,1499,1186,1914]" captionTargetId="figure-515@8.[122,1500,1182,1916]" captionTargetPageId="8" captionText="FIG. 6.—Phylogenetic relationships of sampled species of Helicops based on (A) 16s rRNA and (B) C-mos sequence data. Numbers at nodes indicate the bootstrap values/posterior probabilities; the bars below each cladogram indicate base substitution rate for each gene (# of substitutions per site). Hŋdrops triangularis and Pseudoerŋx plicatilis were used as outgroups." figureDoi="http://doi.org/10.5281/zenodo.7712213" httpUri="https://zenodo.org/record/7712213/files/figure.png" pageId="7">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>