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<document id="65820666BE7DDA3E206F289CD41ABB67" ID-DOI="10.5281/zenodo.178539" ID-GBIF-Dataset="dda53384-9974-4b83-b780-a9cd51fffacf" ID-ISSN="1175-5326" ID-Zenodo-Dep="178539" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459899930198" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03939707FFAE2554FF2CFE42E10AFC3D" docLanguage="en" docName="zt01590p059.pdf" docOrigin="Zootaxa 1590" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ancyronyx helgeschneideri Freitag &amp; Jach, sp. n." docType="treatment" docVersion="10" lastPageNumber="58" masterDocId="FFAAEF7FFFBC2541FFBBFFBDE027FF97" masterDocTitle="The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species" masterLastPageNumber="59" masterPageNumber="37" pageNumber="55" updateTime="1698228715028" updateUser="plazi">
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<mods:title id="A191B1D2841526625B6FC946C1804445">The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species</mods:title>
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<mods:namePart id="7420736FD377AFB80BDEEC000EB9B0AF">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="25D47543455A3F2C89A46663D24F0A7E">Jäch, Manfred A.</mods:namePart>
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<mods:date id="9DC88B65827FD8C2D4399A0947192F4B">2007</mods:date>
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<treatment id="03939707FFAE2554FF2CFE42E10AFC3D" ID-DOI="http://doi.org/10.5281/zenodo.6243459" ID-GBIF-Taxon="119372908" ID-Zenodo-Dep="6243459" LSID="urn:lsid:zoobank.org:act:BECA2458-DBF9-42E7-A42C-EAE45784D8C8" httpUri="http://treatment.plazi.org/id/03939707FFAE2554FF2CFE42E10AFC3D" lastPageId="21" lastPageNumber="58" pageId="18" pageNumber="55">
<subSubSection id="C320759AFFAE2553FF2CFE42E179FDAB" pageId="18" pageNumber="55" type="nomenclature">
<paragraph id="8B852611FFAE2553FF2CFE42E2CAFD8E" blockId="18.[151,749,511,572]" box="[151,749,511,537]" pageId="18" pageNumber="55">
<heading id="D0CD917DFFAE2553FF2CFE42E2CAFD8E" bold="true" box="[151,749,511,537]" fontSize="11" level="1" pageId="18" pageNumber="55" reason="1">
<emphasis id="B94EFA03FFAE2553FF2CFE42E2CAFD8E" bold="true" box="[151,749,511,537]" pageId="18" pageNumber="55">
<taxonomicName id="4C3A5D92FFAE2553FF2CFE42E283FD8E" ID-CoL="DPSH" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,676,511,537]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="helgeschneideri" status="sp. nov.">
<emphasis id="B94EFA03FFAE2553FF2CFE42E1C7FD8E" bold="true" box="[151,480,511,537]" italics="true" pageId="18" pageNumber="55">Ancyronyx helgeschneideri</emphasis>
Freitag &amp; Jäch
</taxonomicName>
<taxonomicNameLabel id="A27D4778FFAE2553FD10FE42E2CAFD8E" box="[683,749,511,537]" pageId="18" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B852611FFAE2553FF2CFD9FE179FDAB" blockId="18.[151,749,511,572]" box="[151,350,546,572]" pageId="18" pageNumber="55">
(
<figureCitation id="13013A94FFAE2553FF24FD9FE125FDAB" box="[159,258,546,572]" captionStart="FIGURES 7 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="18" pageNumber="55">Figs. 10</figureCitation>
,
<figureCitation id="13013A94FFAE2553FEB5FD9FE10DFDAB" box="[270,298,546,572]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="18" pageNumber="55">16</figureCitation>
ah)
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFAE2553FF2CFDCFE3D4FC5B" pageId="18" pageNumber="55" type="materials_examined">
<paragraph id="8B852611FFAE2553FF2CFDCFE104FD23" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
<emphasis id="B94EFA03FFAE2553FF2CFDCFE119FD1B" bold="true" box="[151,318,626,652]" pageId="18" pageNumber="55">
<typeStatus id="548198B3FFAE2553FF2CFDCFE0F3FD1B" box="[151,212,626,652]" pageId="18" pageNumber="55">Type</typeStatus>
locality.
</emphasis>
Cabayugan River, St. Paul National Park, Cabayugan District, Puerto Princesa City, Palawan,
<collectingCountry id="F32D6681FFAE2553FF2CFD27E138FD23" box="[151,287,666,692]" name="Philippines" pageId="18" pageNumber="55">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8B852611FFAE2553FF7DFD7FE3D4FC5B" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
<emphasis id="B94EFA03FFAE2553FF7DFD7FE22EFD4B" bold="true" box="[198,521,706,732]" pageId="18" pageNumber="55">
<typeStatus id="548198B3FFAE2553FF7DFD7FE126FD4B" box="[198,257,706,732]" pageId="18" pageNumber="55">Type</typeStatus>
material.
<typeStatus id="548198B3FFAE2553FE2CFD7FE22EFD4B" box="[407,521,706,732]" pageId="18" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
ď (
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): “
<collectionCode id="ED2BBED4FFAE2553FD78FD7FE329FD4B" box="[707,782,706,732]" pageId="18" pageNumber="55">PHIL</collectionCode>
.: Palawan, P. Princesa S Manturon, Cabayugan R.
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118°52'30''
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</geoCoordinate>
<date id="FF8400D1FFAE2553FE7CFD57E218FC93" box="[455,575,746,772]" pageId="18" pageNumber="55" value="2000-06-17">17.6.2000</date>
, leg. Freitag (CR3)M”, dissected (terminal parts of abdomen, aedeagus and one tarsus glued separately), one distal tarsal segment lacking, right aedeagal paramere incomplete.
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<typeStatus id="548198B3FFAE2553FF2CFC87E133FCC3" box="[151,276,826,852]" pageId="18" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
: 1 ď (
<collectionCode id="ED2BBED4FFAE2553FEDAFC87E193FCC3" box="[353,436,826,852]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="55">NMW</collectionCode>
): “
<collectionCode id="ED2BBED4FFAE2553FE68FC87E239FCC3" box="[467,542,826,852]" pageId="18" pageNumber="55">PHIL</collectionCode>
.: Palawan, P. Princesa S Manturon, Cabayugan R.
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118°52'30''
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</geoCoordinate>
<date id="FF8400D1FFAE2553FF2CFCDFE128FCEB" box="[151,271,866,892]" pageId="18" pageNumber="55" value="2001-05-25">25.5.2001</date>
, leg. Freitag (CR3)D”; 2 ΨΨ (
<collectionCode id="ED2BBED4FFAE2553FDD1FCDFE290FCEB" box="[618,695,866,892]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="55">NMW</collectionCode>
,
<collectionCode id="ED2BBED4FFAE2553FD79FCDFE334FCEB" box="[706,787,866,892]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="18" pageNumber="55">UPLB</collectionCode>
): “
<collectionCode id="ED2BBED4FFAE2553FC8BFCDFE35CFCEB" box="[816,891,866,892]" pageId="18" pageNumber="55">PHIL</collectionCode>
.: Palawan, P. Princesa S Manturon, Cabayugan R.
<geoCoordinate id="EE0E40D6FFAE2553FF05FC37E161FC33" box="[190,326,906,932]" direction="north" orientation="latitude" pageId="18" pageNumber="55" precision="15" value="10.157778">10°09'28''N</geoCoordinate>
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118°53'26''
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<date id="FF8400D1FFAE2553FE49FC37E24CFC33" box="[498,619,906,932]" pageId="18" pageNumber="55" value="2001-05-25">25.5.2001</date>
, leg. Freitag (CR4)D”; 1 Ψ (
<collectionCode id="ED2BBED4FFAE2553FC6DFC37E439FC33" box="[982,1054,906,932]" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="18" pageNumber="55">MTD</collectionCode>
): “
<collectionCode id="ED2BBED4FFAE2553FBF9FC37E4AAFC33" box="[1090,1165,906,932]" pageId="18" pageNumber="55">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.
<geoCoordinate id="EE0E40D6FFAE2553FEF2FC0FE1F5FC5B" box="[329,466,946,972]" direction="north" orientation="latitude" pageId="18" pageNumber="55" precision="15" value="10.2525">10°15'09''N</geoCoordinate>
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118°58'03''
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</geoCoordinate>
<date id="FF8400D1FFAE2553FDC8FC0FE2CCFC5B" box="[627,747,946,972]" pageId="18" pageNumber="55" value="2001-05-17">17.5.2001</date>
, leg. Freitag (PR1)D”.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFAE2554FF7CFC67E2C7FD2D" lastPageId="21" lastPageNumber="58" pageId="18" pageNumber="55" type="description">
<paragraph id="8B852611FFAE2553FF7CFC67E38EFB8B" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
<emphasis id="B94EFA03FFAE2553FF7CFC67E178FC63" bold="true" box="[199,351,986,1012]" pageId="18" pageNumber="55">Description.</emphasis>
Body 1.72.0 mm long (CL),
<quantity id="4CC28BF4FFAE2553FD68FC67E3A7FC63" box="[723,896,986,1012]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.6" metricValueMin="7.6" pageId="18" pageNumber="55" unit="mm" value="0.81" valueMax="0.86" valueMin="0.76">0.760.86 mm</quantity>
broad (EW), 2.42.6 times as broad as wide (CL/EW). Body form elongately oval, moderately convex dorsally.
</paragraph>
<paragraph id="8B852611FFAE2553FF7DFB97E26CFAA3" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
Colouration (
<figureCitation id="13013A94FFAE2553FED9FB97E19DFBD3" box="[354,442,1066,1092]" captionStart="FIGURES 7 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="18" pageNumber="55">Fig. 10</figureCitation>
). Labrum brown laterally, slightly paler in middle; mouth parts and antennae yellowish; frons and clypeus dirty yellowish; small area around the eyes and remainder of head dark brown to black; pronotum yellowish, lateroposteriorly and anteriorly dark brown to black; at least margins of scutellum dark; elytra dark brown, medially with a subquadrate or X-shaped yellowish marking, apices yellowish; ventral side yellowish; coxae pale yellowish; femora yellow (one specimen with indistinct dark anterodorsal median spot on middle and hind femur); tibiae basally black, distally yellowish; tarsomeres basally dark brown, distally yellowish; claws pale reddish brown.
</paragraph>
<paragraph id="8B852611FFAE2553FF7DFAFFE43FFACB" blockId="18.[151,1437,626,2012]" box="[198,1048,1346,1372]" pageId="18" pageNumber="55">Plastron hardly discernible with stereoscopic microscope, not examined.</paragraph>
<paragraph id="8B852611FFAE2553FF7DFAD7E34CFA6B" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
Head
<quantity id="4CC28BF4FFAE2553FEB5FAD7E19CFA13" box="[270,443,1386,1412]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="18" pageNumber="55" unit="mm" value="0.43" valueMax="0.46" valueMin="0.4">0.400.46 mm</quantity>
broad (HW); ID
<quantity id="4CC28BF4FFAE2553FD34FAD7E319FA13" box="[655,830,1386,1412]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="18" pageNumber="55" unit="mm" value="0.24" valueMax="0.26" valueMin="0.22">0.220.26 mm</quantity>
; labrum and clypeus micropunctate; setae distinct; frons granulose, granules arranged in longitudinal lines; frontoclypeal suture slightly arcuate, distinctly impressed. Eyes protruding. Antennae 11-segmented, slender, rather long (longer than HW). Gula microgranulose; gular sutures conspicuous, mesally narrowly separated.
</paragraph>
<paragraph id="8B852611FFAE2553FF7DF9B7E366F883" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
Pronotum
<quantity id="4CC28BF4FFAE2553FEF9F9B7E1CAF9B3" box="[322,493,1546,1572]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8" metricValueMin="5.3" pageId="18" pageNumber="55" unit="mm" value="0.5549999999999999" valueMax="0.58" valueMin="0.53">0.530.58 mm</quantity>
long (PL),
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broad (MW), distinctly wider than long (PL/MW: ca. 0.8), widest at about posterior 0.4, anteriorly attenuate; anterior margin moderately arcuate; margin between pronotum and hypomeron inconspicuous; anterior part with inconspicuous mesal longitudinal carina; anterior transverse groove distinct and deeply impressed, seemingly dividing the pronotum; posteriorly of transverse groove with conspicuous gibbosity; posterolateral oblique grooves moderately deeply impressed; pronotal surface and hypomeron reticulate. Prosternum irregularly punctate; prosternal process distinctly transverse, medially slightly impressed, posterior margin subtruncate.
</paragraph>
<paragraph id="8B852611FFAE2553FF7DF89FE5BAF84B" blockId="18.[151,1437,626,2012]" pageId="18" pageNumber="55">
Scutellum subpentagonal, glabrous. Elytra elongate,
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long (EL), ca. 1.51.7 times as long as wide (EL/EW), very slightly convergent in anterior 0.10.7, posteriorly roundly convergent to apices; with ca. 10 longitudinal, quite regular rows of punctures (seven strial rows between suture and shoulder); punctures large and deeply impressed, except in the almost flat anterior area; interstices and intervals flat to convex, rugulous; lateral elytral gutter very narrow; humeri moderately prominent; elytral apices conjointly rounded.
</paragraph>
<paragraph id="8B852611FFAF2552FF2CFF25E166FEBD" blockId="19.[151,1437,152,298]" pageId="19" pageNumber="56">
Meso- and metaventrite moderately densely granulose and pubescent, lateral parts distinctly punctate. Mesoventrite very short, with one medial and a pair of sublateral impressions. Metaventrite prominent, with distinct discrimen; anepisternum 3 not visible in ventral view, with one row of deeply impressed punctures.
<collectingCountry id="F32D6681FFAF2552FADAFF55E5BBFE95" box="[1377,1436,232,258]" name="India" pageId="19" pageNumber="56">Hind</collectingCountry>
wings present.
</paragraph>
<caption id="DF457699FFAF2552FF2CF8FDE3C2F82F" httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="19" pageNumber="56" targetBox="[195,1367,349,1794]" targetPageId="19">
<paragraph id="8B852611FFAF2552FF2CF8FDE3C2F82F" blockId="19.[151,1436,1856,1976]" pageId="19" pageNumber="56">
<emphasis id="B94EFA03FFAF2552FF2CF8FDE100F8CF" bold="true" box="[151,295,1856,1880]" pageId="19" pageNumber="56">FIGURE 16.</emphasis>
<taxonomicName id="4C3A5D92FFAF2552FE94F8FCE276F8CF" box="[303,593,1857,1880]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="56" phylum="Arthropoda" rank="species" species="helgeschneideri" status="sp. nov.">
<emphasis id="B94EFA03FFAF2552FE94F8FCE276F8CF" box="[303,593,1857,1880]" italics="true" pageId="19" pageNumber="56">Ancyronyx helgeschneideri</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFAF2552FDE2F8FDE2B1F8CF" bold="true" box="[601,662,1856,1880]" pageId="19" pageNumber="56">
<taxonomicNameLabel id="A27D4778FFAF2552FDE2F8FDE2B1F8CF" box="[601,662,1856,1880]" pageId="19" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♂ paratype from CR3, Ψ paratype from PR1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted).
</paragraph>
</caption>
<paragraph id="8B852611FFA82555FF2CFB5CE35FFADF" blockId="20.[151,1436,1249,1352]" pageId="20" pageNumber="57">
<emphasis id="B94EFA03FFA82555FF2CFB5CE10FFB6E" bold="true" box="[151,296,1249,1273]" pageId="20" pageNumber="57">FIGURE 17.</emphasis>
Collecting sites and geographical distribution of
<taxonomicName id="4C3A5D92FFA82555FCF9FB5FE394FB6E" box="[834,947,1250,1273]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B94EFA03FFA82555FCF9FB5FE394FB6E" box="[834,947,1250,1273]" italics="true" pageId="20" pageNumber="57">Ancyronyx</emphasis>
</taxonomicName>
species in Palawan and Busuanga, including enlarged map of St. Paul National Park:
<taxonomicName id="4C3A5D92FFA82555FDE4FAB7E308FAB6" box="[607,815,1290,1313]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="patrolus" status="sp. nov.">
<emphasis id="B94EFA03FFA82555FDE4FAB7E308FAB6" box="[607,815,1290,1313]" italics="true" pageId="20" pageNumber="57">Ancyronyx patrolus</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFA82555FC8EFAB4E356FAB6" bold="true" box="[821,881,1289,1313]" pageId="20" pageNumber="57">
<taxonomicNameLabel id="A27D4778FFA82555FC8EFAB4E356FAB6" box="[821,881,1289,1313]" pageId="20" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C3A5D92FFA82555FC21FAB7E47CFAB6" box="[922,1115,1290,1313]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="pseudopatrolus" status="sp. nov.">
<emphasis id="B94EFA03FFA82555FC21FAB7E47CFAB6" box="[922,1115,1290,1313]" italics="true" pageId="20" pageNumber="57">A. pseudopatrolus</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFA82555FBD9FAB4E4BAFAB6" bold="true" box="[1122,1181,1289,1313]" pageId="20" pageNumber="57">
<taxonomicNameLabel id="A27D4778FFA82555FBD9FAB4E4BAFAB6" box="[1122,1181,1289,1313]" pageId="20" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♦
<taxonomicName id="4C3A5D92FFA82555FB6BFAB7E517FAB6" box="[1232,1328,1290,1313]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="punkti" status="sp. nov.">
<emphasis id="B94EFA03FFA82555FB6BFAB7E517FAB6" box="[1232,1328,1290,1313]" italics="true" pageId="20" pageNumber="57">A. punkti</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFA82555FA8DFAB4E555FAB6" bold="true" box="[1334,1394,1289,1313]" pageId="20" pageNumber="57">
<taxonomicNameLabel id="A27D4778FFA82555FA8DFAB4E555FAB6" box="[1334,1394,1289,1313]" pageId="20" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C3A5D92FFA82555FF2CFA8CE12CFADF" box="[151,267,1329,1352]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="minerva" status="sp. nov.">
<emphasis id="B94EFA03FFA82555FF2CFA8CE12CFADF" box="[151,267,1329,1352]" italics="true" pageId="20" pageNumber="57">A. minerva</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFA82555FEA9FA8DE169FADF" bold="true" box="[274,334,1328,1352]" pageId="20" pageNumber="57">
<taxonomicNameLabel id="A27D4778FFA82555FEA9FA8DE169FADF" box="[274,334,1328,1352]" pageId="20" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C3A5D92FFA82555FEC4FA8CE221FADF" box="[383,518,1329,1352]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="minutulus" status="sp. nov.">
<emphasis id="B94EFA03FFA82555FEC4FA8CE221FADF" box="[383,518,1329,1352]" italics="true" pageId="20" pageNumber="57">A. minutulus</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFA82555FDB7FA8DE26FFADF" bold="true" box="[524,584,1328,1352]" pageId="20" pageNumber="57">
<taxonomicNameLabel id="A27D4778FFA82555FDB7FA8DE26FFADF" box="[524,584,1328,1352]" pageId="20" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C3A5D92FFA82555FDCBFA8CE311FADF" box="[624,822,1329,1352]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="helgeschneideri" status="sp. nov.">
<emphasis id="B94EFA03FFA82555FDCBFA8CE311FADF" box="[624,822,1329,1352]" italics="true" pageId="20" pageNumber="57">A. helgeschneideri</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFA82555FC86FA8DE35FFADF" bold="true" box="[829,888,1328,1352]" pageId="20" pageNumber="57">
<taxonomicNameLabel id="A27D4778FFA82555FC86FA8DE35FFADF" box="[829,888,1328,1352]" pageId="20" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B852611FFA82555FF7DFAC1E213FA01" blockId="20.[151,1436,1404,2030]" box="[198,564,1404,1430]" pageId="20" pageNumber="57">Ventrites granulose, pubescent.</paragraph>
<paragraph id="8B852611FFA82555FF7DFA19E530F9A1" blockId="20.[151,1436,1404,2030]" pageId="20" pageNumber="57">Legs approximately 1/4 longer than body; pro- and mesocoxae large, bluntly drop-shaped; metacoxae distinctly smaller and only slightly protruding laterally; femora, tibiae, and tarsi covered with inconspicuous setae, tibiae with few additional longer setae; distal tarsal segments with several conspicuous setae near base of claws; claws well developed, base with three teeth, basal one very small and hardly discernible.</paragraph>
<paragraph id="8B852611FFA82555FF7DF9F9E4BAF969" blockId="20.[151,1436,1404,2030]" pageId="20" pageNumber="57">
Sternite IX (
<figureCitation id="13013A94FFA82555FEE3F9F9E18AF9C9" box="[344,429,1604,1630]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="20" pageNumber="57">Fig. 16</figureCitation>
c) ca. 450 µm long; apical margin distinctly emarginate, lateroapically finely striolate and covered with microsetae; paraprocts not or only slightly reaching beyond apical margin. Tergite VIII in both sexes without distinct reticulation, with conspicuous median transverse complete ridge separating posterior and anterior portion; basal half with microstrial pattern; apical half darker, with hyaline apical margin and subapical fringe of trichoid setae, with ca. six conspicuously strong and very long setae.
</paragraph>
<paragraph id="8B852611FFA82555FF7DF8B1E2C0F879" blockId="20.[151,1436,1404,2030]" pageId="20" pageNumber="57">
Aedeagus (
<figureCitation id="13013A94FFA82555FEF5F8B1E180F8B1" box="[334,423,1804,1830]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="20" pageNumber="57">Fig. 16</figureCitation>
a, b) ca. 440 µm long. Median lobe rather short and conical, subapically distinctly curved ventrad (lateral view), with numerous distinct microtube-like structures, most conspicuous apically; ventral sac apically ending in two separately tapered tips, mediobasally plicate; fibula inconspicuous; corona weekly developed. Phallobase asymmetrical, approximately half as long as parameres. Parameres long (almost reaching apex of aedeagus), dorsally widened at base (lateral view), ventral margin distinctly sinuous; basal margin emarginate, not contiguous ventrally.
</paragraph>
<paragraph id="8B852611FFA92554FF7DFF25E0F0FEED" blockId="21.[151,1437,152,938]" pageId="21" pageNumber="58">
Ovipositor (
<figureCitation id="13013A94FFA92554FEE8FF25E192FF25" box="[339,437,152,178]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="21" pageNumber="58">Figs. 16</figureCitation>
d). Total length ca. 480 µm. Stylus slender, very slightly bent outwards (dorsal view). Coxite short and stout, distal portion with several rather short and broad, peg-like setae, most densely set lateroapically; inner margin moderately pubescent; basal portion approximately as long as distal portion, with rather short, peg-like setae, most densely set at lateral margin, medially with striolate patches. Valvifer slightly longer than coxite, apical margin with one long spine, finely striolate near apex; fibula curved inwards caudally.
</paragraph>
<paragraph id="8B852611FFA92554FF7DFE35E55AFDD5" blockId="21.[151,1437,152,938]" pageId="21" pageNumber="58">
Secondary sexual characters. Sternite VIII (
<figureCitation id="13013A94FFA92554FD6DFE35E308FE35" box="[726,815,392,418]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="21" pageNumber="58">
Fig.
<quantity id="4CC28BF4FFA92554FCAAFE35E308FE35" box="[785,815,392,418]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="21" pageNumber="58" unit="g" value="16.0">16</quantity>
</figureCitation>
g) in male weekly sclerotized, without conspicuous setae; median strut short, apically roundly tapered; female sternite VIII not dissected. Tergite VIII in male (
<figureCitation id="13013A94FFA92554FF24FE65E0DFFE65" box="[159,248,472,498]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="21" pageNumber="58">Fig. 16</figureCitation>
e) distinctly broader than long, condyles straight and prominent. Tergite VIII in female (
<figureCitation id="13013A94FFA92554FA97FE65E5A4FE65" box="[1324,1411,472,498]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="21" pageNumber="58">Fig. 16</figureCitation>
h) slightly broader than long; condyles inconspicuous, more or less straight and not distinctly reaching beyond anterior margin. Ventrite
<quantity id="4CC28BF4FFA92554FE05FD95E1CCFDD5" box="[446,491,552,578]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="58" unit="in" value="5.0">5 in</quantity>
male (
<figureCitation id="13013A94FFA92554FD87FD95E2A9FDD5" box="[572,654,552,578]" captionStart="FIGURE 16" captionStartId="19.[151,255,1856,1880]" captionTargetBox="[195,1367,349,1794]" captionTargetId="figure@19.[195,1368,349,1794]" captionTargetPageId="19" captionText="FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178549/files/figure.png" pageId="21" pageNumber="58">Fig. 16</figureCitation>
f) suboval, in female similar, but longer and more subtriangular.
</paragraph>
<paragraph id="8B852611FFA92554FF7DFDF2E2C7FD2D" blockId="21.[151,1437,152,938]" pageId="21" pageNumber="58">
<emphasis id="B94EFA03FFA92554FF7DFDF2E1FFFDFE" bold="true" box="[198,472,591,617]" pageId="21" pageNumber="58">Differential diagnosis.</emphasis>
The general habitus and colour patterns of this species somewhat resemble
<taxonomicName id="4C3A5D92FFA92554FAF0FDEDE179FD05" authority="Jach, 1994" authorityName="Jach" authorityYear="1994" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="58" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B94EFA03FFA92554FAF0FDEDE0F0FD06" italics="true" pageId="21" pageNumber="58">A. procerus</emphasis>
<bibRefCitation id="EFAB5BE0FFA92554FF64FDC5E179FD05" author="Jach" box="[223,350,632,658]" pageId="21" pageNumber="59" refString="Jach, M. A. (1994) A taxonomic review of the Oriental species of the genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae). Revue suisse de Zoologie, 101, 601 - 622." type="journal article" year="1994">Jäch, 1994</bibRefCitation>
</taxonomicName>
(see Fig. 28,
<bibRefCitation id="EFAB5BE0FFA92554FDB9FDC5E25AFD05" author="Jach" box="[514,637,632,658]" pageId="21" pageNumber="59" refString="Jach, M. A. (1994) A taxonomic review of the Oriental species of the genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae). Revue suisse de Zoologie, 101, 601 - 622." type="journal article" year="1994">Jäch 1994</bibRefCitation>
), from which it can be distinguished by smaller size, long aedeagal parameres and presence of a spine on the valvifer.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFA92554FF7DFD7AE44FFCCD" pageId="21" pageNumber="58" type="distribution">
<paragraph id="8B852611FFA92554FF7DFD7AE547FD75" blockId="21.[151,1437,152,938]" box="[198,1376,711,738]" pageId="21" pageNumber="58">
<emphasis id="B94EFA03FFA92554FF7DFD7AE178FD76" bold="true" box="[198,351,711,737]" pageId="21" pageNumber="58">Distribution</emphasis>
(Fig. 17). This species is known only from the St. Paul National Park, central Palawan.
</paragraph>
<paragraph id="8B852611FFA92554FF7DFD52E44FFCCD" blockId="21.[151,1437,152,938]" pageId="21" pageNumber="58">
<emphasis id="B94EFA03FFA92554FF7DFD52E108FC9E" bold="true" box="[198,303,751,777]" pageId="21" pageNumber="58">Ecology.</emphasis>
The distribution of this species in river courses distinctly varying in their morphology and physicochemical character (CR3 vs. PR1; see
<bibRefCitation id="EFAB5BE0FFA92554FDC1FCA5E335FCA5" author="Freitag" box="[634,786,792,818]" pageId="21" pageNumber="59" refString="Freitag, H. (2005) Longitudinal zonation patterns and determinants in Decapoda (Crustacea) in rivers of Palawan Island, the Philippines. Archiv fur Hydrobiologie, Supplement, 151, 243 - 268." type="journal article" year="2005">Freitag 2005</bibRefCitation>
), including slightly disturbed sections suggests a higher ecological tolerance. This species seems to prefer stream sections of medium order.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFA92554FF7DFCDAE10AFC3D" pageId="21" pageNumber="58" type="etymology">
<paragraph id="8B852611FFA92554FF7DFCDAE10AFC3D" blockId="21.[151,1437,152,938]" pageId="21" pageNumber="58">
<emphasis id="B94EFA03FFA92554FF7DFCDAE174FC16" bold="true" box="[198,339,871,897]" pageId="21" pageNumber="58">Etymology.</emphasis>
Named for Helge Schneider, famous
<collectingCountry id="F32D6681FFA92554FCB2FCD5E340FC15" box="[777,871,872,898]" name="Germany" pageId="21" pageNumber="58">German</collectingCountry>
multi-talented artist, with special interest in animal wildlife.
</paragraph>
</subSubSection>
</treatment>
</document>