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<document id="3EF3932E66CFE465BAA339E3224CE22D" ID-DOI="10.11646/zootaxa.5284.2.8" ID-GBIF-Dataset="b4bc4bdf-927c-43d9-ac2b-c39d33137048" ID-ISSN="1175-5326" ID-Zenodo-Dep="7923530" ID-ZooBank="770F88D2-FA38-499B-AC81-61B8BE61992F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683787118133" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Tri, Ngo Van" docDate="2023" docId="039387F1FFBD3A0CFF2E596BFCB9FA03" docLanguage="en" docName="zootaxa.5284.2.8.pdf" docOrigin="Zootaxa 5284 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tortomon Huang, Wang &amp; Shih 2020" docType="treatment" docVersion="2" lastPageNumber="365" masterDocId="FFAAFF89FFBC3A0DFFB95804FFC4FFBF" masterDocTitle="Two new species of semiaquatic freshwater crabs of the genera Tortomon and Aiyunamon from karst forests in northern Vietnam (Crustacea: Brachyura: Potamidae)" masterLastPageNumber="376" masterPageNumber="364" pageNumber="365" updateTime="1683787784201" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FA392EC90F0DA6E54ADCAAB24ABB4F80">Two new species of semiaquatic freshwater crabs of the genera Tortomon and Aiyunamon from karst forests in northern Vietnam (Crustacea: Brachyura: Potamidae)</mods:title>
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<mods:namePart id="9B9AF4CE474F629F09029F96E615A738">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="67591F9F33544C8C4D590280A71EC7F6">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:namePart id="E04A7E574E35D162363C2246272E322B">Tri, Ngo Van</mods:namePart>
<mods:affiliation id="4D7330522FE229477C89D78384FD0DF2">National Key Laboratory, Institute of Tropical Biology, Vietnamese Academy of Sciences and Technology, 9 / 621 Hanoi Highway, Thu Duc City, Ho Chi Minh City, Vietnam.</mods:affiliation>
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<subSubSection id="C320656CFFBD3A0CFF2E596BFD06FE36" box="[151,706,367,393]" pageId="1" pageNumber="365" type="nomenclature">
<paragraph id="8B8536E7FFBD3A0CFF2E596BFD06FE36" blockId="1.[151,706,367,393]" box="[151,706,367,393]" pageId="1" pageNumber="365">
<heading id="D0CD818BFFBD3A0CFF2E596BFD06FE36" bold="true" box="[151,706,367,393]" fontSize="11" level="1" pageId="1" pageNumber="365" reason="1">
<emphasis id="B94EEAF5FFBD3A0CFF2E596BFD06FE36" bold="true" box="[151,706,367,393]" pageId="1" pageNumber="365">
Genus
<taxonomicName id="4C3A4D64FFBD3A0CFF57596BFD06FE36" ID-CoL="84GL5" ID-ENA="2768179" authority="Huang, Wang &amp; Shih, 2020" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[238,706,367,393]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFF57596BFEA0FE36" bold="true" box="[238,356,367,393]" italics="true" pageId="1" pageNumber="365">Tortomon</emphasis>
<bibRefCitation id="EFAB4B16FFBD3A0CFED5596BFD06FE36" author="Huang, C. &amp; Wang, J. &amp; Shih, H. - T." box="[364,706,367,393]" pageId="1" pageNumber="365" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. &amp; Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">Huang, Wang &amp; Shih, 2020</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C320656CFFBD3A0CFF2E59B0FC19FE73" box="[151,989,435,460]" pageId="1" pageNumber="365" type="type_taxon">
<paragraph id="8B8536E7FFBD3A0CFF2E59B0FC19FE73" blockId="1.[151,989,435,460]" box="[151,989,435,460]" pageId="1" pageNumber="365">
<emphasis id="B94EEAF5FFBD3A0CFF2E59B0FEE7FE74" bold="true" box="[151,291,435,460]" pageId="1" pageNumber="365">
<typeStatus id="54818845FFBD3A0CFF2E59B0FF08FE73" box="[151,204,436,460]" pageId="1" pageNumber="365">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3A4D64FFBD3A0CFE9359B0FD25FE73" authority="Huang, Wang &amp; Shih, 2020" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[298,737,436,460]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="puer">
<emphasis id="B94EEAF5FFBD3A0CFE9359B0FE04FE74" box="[298,448,436,459]" italics="true" pageId="1" pageNumber="365">Tortomon puer</emphasis>
<bibRefCitation id="EFAB4B16FFBD3A0CFE7E59B0FD25FE73" author="Huang, C. &amp; Wang, J. &amp; Shih, H. - T." box="[455,737,436,460]" pageId="1" pageNumber="365" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. &amp; Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">Huang, Wang &amp; Shih, 2020</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C320656CFFBD3A0CFF2E59FDFC25FC56" pageId="1" pageNumber="365" type="diagnosis">
<paragraph id="8B8536E7FFBD3A0CFF2E59FDFC25FC56" blockId="1.[151,1437,505,1468]" pageId="1" pageNumber="365">
<emphasis id="B94EEAF5FFBD3A0CFF2E59FDFED6FDAC" bold="true" box="[151,274,505,531]" pageId="1" pageNumber="365">Diagnosis.</emphasis>
(Emended from
<bibRefCitation id="EFAB4B16FFBD3A0CFE6359FEFD0BFDAB" author="Huang, C. &amp; Wang, J. &amp; Shih, H. - T." box="[474,719,506,532]" pageId="1" pageNumber="365" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. &amp; Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
Huang
<emphasis id="B94EEAF5FFBD3A0CFD9659FFFDAFFDAB" box="[559,619,506,532]" italics="true" pageId="1" pageNumber="365">et al.</emphasis>
2020: 2
</bibRefCitation>
) Carapace width less than
<quantity id="4CC29B02FFBD3A0CFBB759FEFBA7FDAB" box="[1038,1123,506,533]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="365" unit="mm" value="25.0">25 mm</quantity>
; carapace wider than long, dorsal surface smooth, evenly convex, regions indistinct; epigastric cristae and postfrontal cristae very low to inconspicuous, smooth; external orbital tooth not demarcated from rest of anterolateral margin, no trace of epibranchial tooth, margin smooth to weakly cristate; median lobe of epistome triangular. Third maxilliped with quadrate to rectangular ischium, exopod strongly tapering, reaching beyond anterior edge of ischium, no trace of flagellum. Cheliped merus margins smooth, surface of chela smooth. Ambulatory legs slender, unarmed. Male anterior thoracic sternum relatively wide, sternites 3 and 4 fused but demarcated by shallow but visible oblique grooves, distal edge of sternopleonal cavity with low ridge; sternal tubercle of male pleonal locking mechanism relatively small or large, on submedian part or distal third of sternite 5. Male pleon variously triangular; telson with gently to strongly concave lateral margins. G1 relatively short, almost straight to gently curved, relatively stout, terminal segment short, stout, conical to more sharply tapering. G2 subequal or longer than G1, flagellum half to two-thirds length of basal segment, with proximal part tubular and gently twisted, distalmost part very slender, prominently coiled. Vulvae with relatively wide space between them, ovate, convex, medium-sized, positioned on proximal half or one-third of sternite 6, pushing onto suture with sternite 5.
</paragraph>
</subSubSection>
<subSubSection id="C320656CFFBD3A0CFF7E5BF6FCB9FA03" pageId="1" pageNumber="365" type="discussion">
<paragraph id="8B8536E7FFBD3A0CFF7E5BF6FB52FB5B" blockId="1.[151,1437,505,1468]" pageId="1" pageNumber="365">
<emphasis id="B94EEAF5FFBD3A0CFF7E5BF6FEF0FBB3" bold="true" box="[199,308,1010,1036]" pageId="1" pageNumber="365">Remarks</emphasis>
.
<bibRefCitation id="EFAB4B16FFBD3A0CFEFC5BF6FDEEFBB3" author="Huang, C. &amp; Wang, J. &amp; Shih, H. - T." box="[325,554,1010,1036]" pageId="1" pageNumber="365" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. &amp; Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">
Huang
<emphasis id="B94EEAF5FFBD3A0CFE235BF7FE11FBB3" box="[410,469,1010,1036]" italics="true" pageId="1" pageNumber="365">et al.</emphasis>
(2020)
</bibRefCitation>
described
<taxonomicName id="4C3A4D64FFBD3A0CFD105BF7FCD1FBB3" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[681,789,1011,1036]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFD105BF7FCD1FBB3" box="[681,789,1011,1036]" italics="true" pageId="1" pageNumber="365">Tortomon</emphasis>
</taxonomicName>
on the basis of two new species from southern
<collectingRegion id="49FEF805FFBD3A0CFAFE5BF6FA59FBB3" box="[1351,1437,1010,1036]" country="China" name="Yunnan" pageId="1" pageNumber="365">Yunnan</collectingRegion>
in
<collectingCountry id="F32D7677FFBD3A0CFF0C5C12FF3FFB8F" box="[181,251,1046,1072]" name="China" pageId="1" pageNumber="365">China</collectingCountry>
,
<taxonomicName id="4C3A4D64FFBD3A0CFEB15C13FD5BFB8F" authority="Huang, Wang &amp; Shih, 2020" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[264,671,1046,1072]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="puer">
<emphasis id="B94EEAF5FFBD3A0CFEB15C13FE9CFB8F" box="[264,344,1047,1072]" italics="true" pageId="1" pageNumber="365">T. puer</emphasis>
<bibRefCitation id="EFAB4B16FFBD3A0CFED95C12FD5BFB8F" author="Huang, C. &amp; Wang, J. &amp; Shih, H. - T." box="[352,671,1046,1072]" pageId="1" pageNumber="365" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. &amp; Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">Huang, Wang &amp; Shih, 2020</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C3A4D64FFBD3A0CFD655C13FBBDFB8F" authority="Huang, Wang &amp; Shih, 2020" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[732,1145,1046,1072]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="gejiu">
<emphasis id="B94EEAF5FFBD3A0CFD655C13FCF5FB8F" box="[732,817,1047,1072]" italics="true" pageId="1" pageNumber="365">T. gejiu</emphasis>
<bibRefCitation id="EFAB4B16FFBD3A0CFC805C12FBBDFB8F" author="Huang, C. &amp; Wang, J. &amp; Shih, H. - T." box="[825,1145,1046,1072]" pageId="1" pageNumber="365" pagination="1 - 14" refId="ref5493" refString="Huang, C., Wang, J. &amp; Shih, H. - T. (2020) A new genus and two new species of freshwater crab (Crustacea: Brachyura: Potamidae) with unusual coiled tip of male second gonopods from Yunnan, southwestern China. Zoological Studies, 59 (24), 1 - 14. https: // doi. org / 10.6620 / ZS. 2020.59 - 24" type="journal article" year="2020">Huang, Wang &amp; Shih, 2020</bibRefCitation>
</taxonomicName>
. Both are relatively small species (carapace width less than
<quantity id="4CC29B02FFBD3A0CFDAB5C3EFDA6FBEB" box="[530,610,1082,1109]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="365" unit="mm" value="25.0">25 mm</quantity>
) with a smooth carapace having entire anterolateral margins. The exopod of their third maxilliped lacks a flagellum, a feature typically associated with the genus
<taxonomicName id="4C3A4D64FFBD3A0CFBCF5C5BFA5CFBC7" authority="Alcock, 1909" authorityName="Alcock" authorityYear="1909" box="[1142,1432,1118,1144]" class="Malacostraca" family="Potamidae" genus="Potamiscus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFBCF5C5BFB33FBC7" box="[1142,1271,1119,1144]" italics="true" pageId="1" pageNumber="365">Potamiscus</emphasis>
<bibRefCitation id="EFAB4B16FFBD3A0CFB475C5AFA5CFBC7" author="Alcock, A." box="[1278,1432,1118,1144]" pageId="1" pageNumber="365" pagination="375 - 381" refId="ref4990" refString="Alcock, A. (1909) Diagnoses of new species and varieties of freshwater crabs. No. 4. Records of the Indian Museum, 3, 375 - 381. https: // doi. org / 10.26515 / rzsi / v 3 / i 4 / 1909 / 163297" type="journal article" year="1909">Alcock, 1909</bibRefCitation>
</taxonomicName>
. Unlike
<taxonomicName id="4C3A4D64FFBD3A0CFF555C87FEA9FB23" authorityName="Alcock" authorityYear="1909" box="[236,365,1155,1180]" class="Malacostraca" family="Potamidae" genus="Potamiscus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFF555C87FEA9FB23" box="[236,365,1155,1180]" italics="true" pageId="1" pageNumber="365">Potamiscus</emphasis>
</taxonomicName>
s. lato, however (cf.
<bibRefCitation id="EFAB4B16FFBD3A0CFDE65C86FD0FFB23" author="Dai, A. Y." box="[607,715,1154,1180]" pageId="1" pageNumber="365" refId="ref5227" refString="Dai, A. Y. (1999) Fauna Sinica: Arthropoda Crustacea Malacostraca Decapod Parathelphusidae Potamidae. Science Press, Beijing, 501 pp., 30 pls. [in Chinese with English summary]" type="book" year="1999">Dai 1999</bibRefCitation>
;
<bibRefCitation id="EFAB4B16FFBD3A0CFD6E5C86FC4BFB23" author="Yeo, D. C. J. &amp; Ng, P. K. L." box="[727,911,1154,1180]" pageId="1" pageNumber="365" pagination="273 - 308" refId="ref6318" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (2007) On the genus &quot; Potamon &quot; and allies in Indochina (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, Supplement 16, 273 - 308." type="journal article" year="2007">Yeo &amp; Ng 2007</bibRefCitation>
;
<bibRefCitation id="EFAB4B16FFBD3A0CFC255C86FB87FB23" author="Ng, P. K. L. &amp; Htoo, H. &amp; Mar, W." box="[924,1091,1154,1180]" pageId="1" pageNumber="365" pagination="129 - 136" refId="ref5743" refString="Ng, P. K. L., Htoo, H. &amp; Mar, W. (2019) Potamiscus whitteni, a new freshwater crab from Chin State, Myanmar (Crustacea: Brachyura: Potamidae). Raffles Bulletin of Zoology, Supplement 35, 129 - 136." type="journal article" year="2019">
Ng
<emphasis id="B94EEAF5FFBD3A0CFC7F5C87FBC4FB23" box="[966,1024,1154,1180]" italics="true" pageId="1" pageNumber="365">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="EFAB4B16FFBD3A0CFBE95C86FAC5FB23" author="Pati, S. K. &amp; Mitra, S. &amp; Ng, P. K. L." box="[1104,1281,1154,1180]" pageId="1" pageNumber="365" pagination="703 - 725" refId="ref5991" refString="Pati, S. K., Mitra, S. &amp; Ng, P. K. L. (2020) The identity of the freshwater crab Indochinamon manipurense (Alcock, 1909), with description of a new species of Potamiscus Alcock, 1909, from Manipur State, India (Decapoda, Brachyura, Potamidae). Crustaceana, 93 (7), 703 - 725. https: // doi. org / 10.1163 / 15685403 - bja 10009" type="journal article" year="2020">
Pati
<emphasis id="B94EEAF5FFBD3A0CFB3A5C87FB79FB23" box="[1155,1213,1154,1180]" italics="true" pageId="1" pageNumber="365">et al.</emphasis>
2020
</bibRefCitation>
),
<taxonomicName id="4C3A4D64FFBD3A0CFAAC5C87FA45FB23" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[1301,1409,1155,1180]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFAAC5C87FA45FB23" box="[1301,1409,1155,1180]" italics="true" pageId="1" pageNumber="365">Tortomon</emphasis>
</taxonomicName>
is unique in possessing an unusual male G
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which the flagellum is twisted, especially at the distalmost part, which is very slender and twisted. Such a male G2 form is not known in any other potamid genus.
</paragraph>
<paragraph id="8B8536E7FFBD3A0CFF7E5CEAFCB9FA03" blockId="1.[151,1437,505,1468]" pageId="1" pageNumber="365">
A note on the G1 is useful. The G1 of
<taxonomicName id="4C3A4D64FFBD3A0CFD2F5CEBFCC6FAB7" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[662,770,1263,1288]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFD2F5CEBFCC6FAB7" box="[662,770,1263,1288]" italics="true" pageId="1" pageNumber="365">Tortomon</emphasis>
</taxonomicName>
is relatively short and stout, and the terminal segment is not demarcated from the subterminal segment by deep sutures. In most potamids, the suture separating the two segments is deep and prominent, visible from both ventral and dorsal views. In the case of
<taxonomicName id="4C3A4D64FFBD3A0CFB205D33FAC1FAEF" authorityName="Huang, Wang &amp; Shih" authorityYear="2020" box="[1177,1285,1335,1360]" class="Malacostraca" family="Potamidae" genus="Tortomon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF5FFBD3A0CFB205D33FAC1FAEF" box="[1177,1285,1335,1360]" italics="true" pageId="1" pageNumber="365">Tortomon</emphasis>
</taxonomicName>
, the suture is shallow and more obvious in dorsal view (
<figureCitation id="13012A62FFBD3A0CFDC55D5EFD11FACB" box="[636,725,1370,1396]" captionStart="FIGURE 3" captionStartId="4.[151,250,1900,1924]" captionTargetBox="[167,1421,181,1876]" captionTargetId="figure-19@4.[167,1421,181,1876]" captionTargetPageId="4" captionText="FIGURE 3. Tortomon distinctum n. sp.A–D, G–I, holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam; E, paratype male (18.3 × 12.4 mm) (ZRC 2022.798), Vietnam; F, paratype male (19.5 × 12.8 mm) (ZRC 2022.798), Vietnam. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity; C, right fourth ambulatory leg; D–F, pleon; G, left G1 (ventral view); H, left G1 (dorsal view); I, left G2, ventral view). G–I, same scale." figureDoi="http://doi.org/10.5281/zenodo.7923536" httpUri="https://zenodo.org/record/7923536/files/figure.png" pageId="1" pageNumber="365">Fig. 3H</figureCitation>
,
<figureCitation id="13012A62FFBD3A0CFD5A5D5EFCCDFACB" box="[739,777,1370,1396]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[153,1434,183,1044]" captionTargetId="figure-32@5.[151,1436,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Tortomon distinctum n. sp., holotype male (19.2 × 13.0 mm) (ZRC 2022.797), Vietnam. A, left G1 (ventral view); B, distal part of left G1 (ventral view); C, distal part of left G1 (dorsomesial view); D, distal part of left G1 (dorsal view); E, left G2 (ventral view); F, distal part of left G2 (ventral view); G, distal part of left G2 (dorsal view). Scales: A, E = 0.5 mm; B–D, F, G = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7923538" httpUri="https://zenodo.org/record/7923538/files/figure.png" pageId="1" pageNumber="365">4D</figureCitation>
). The G2 flagellum is unusual in that proximal part of the structure is normal in thickness and only slightly twisted medially; but the distalmost part tapers suddenly to form a flagellum-like structure, which is strongly coiled like a corkscrew.
</paragraph>
</subSubSection>
</treatment>
</document>
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